treatments-xml/data/03/FE/87/03FE879673062928FF35F976FC60FA39.xml

<document id="61905BF3BD684FA2DB9138DD09C91C3F" ID-DOI="10.11646/phytotaxa.619.4.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8430545" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1697019535391" checkinUser="plazi" docAuthor="Averyanov, Leonid V., Nguyen, Van Canh, Vuong, Truong Ba, Nguyen, Khang Sinh, Nuraliev, Maxim S., Nguyen, Cuong Huu, Ormerod, Paul A., Maisak, Tatiana V., Diep, Dinh Quang, Lyskov, Dmitry F. &amp; Nong, Duy Van" docDate="2023" docId="03FE879673062928FF35F976FC60FA39" docLanguage="en" docName="phytotaxa.619.4.1.pdf" docOrigin="Phytotaxa 619 (4)" docSource="http://dx.doi.org/10.11646/phytotaxa.619.4.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Goodyera umbonata Aver., V. C. Nguyen &amp; Ormerod 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="260" masterDocId="FFC7FFEE7304292DFFBDFFF0FF9BFFD5" masterDocTitle="New orchids in the flora of Vietnam VII (Orchidaceae: tribes Cypripedieae, Cranichideae, Orchideae, and Collabieae)" masterLastPageNumber="276" masterPageNumber="255" pageNumber="257" updateTime="1697020034893" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2948A31802BBDC67AA0047183DA72079">New orchids in the flora of Vietnam VII (Orchidaceae: tribes Cypripedieae, Cranichideae, Orchideae, and Collabieae)</mods:title>
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<mods:namePart id="5B6EDEA9645E0E10D62C283F11087C67">Averyanov, Leonid V.</mods:namePart>
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<mods:namePart id="70CE5C4DA6E89D6CDF8C3DD970240986">Nguyen, Van Canh</mods:namePart>
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<mods:affiliation id="BDDC2203EA97173FF9ABCD7837EB36C3">Department of Higher Plants, Biological Faculty, M. V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234 Moscow, Russia &amp; Joint Russian-Vietnamese Tropical Scientific and Technological Center, 63, Nguyen Van Huyen Street, Cau Giay, 122000 Hanoi, Vietnam &amp; max. nuraliev @ gmail. com; https: // orcid. org / 0000 - 0001 - 8291 - 2633</mods:affiliation>
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<mods:affiliation id="E2D7AED3057A4670568F03F1DBA1A935">Vietnam National University of Forestry, Xuan Mai, Chuong My, Hanoi, Vietnam &amp; nguyenhuucuong. tvr @ gmail. com; https: // orcid. org / 0000 - 0003 - 2281 - 741 X</mods:affiliation>
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<emphasis id="B923EA927306292FFF35F976FEEFF974" bold="true" box="[136,372,1670,1697]" italics="true" pageId="2" pageNumber="257">Goodyera umbonata</emphasis>
Aver., V.C.Nguyen &amp; Ormerod
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, 
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(
<figureCitation id="136C2A057306292FFF2DF95AFF4FF911" box="[144,212,1706,1732]" captionStart="FIGURE 2" captionStartId="4.[136,229,1866,1888]" captionTargetBox="[243,1344,192,1841]" captionTargetId="figure-24@4.[243,1344,190,1841]" captionTargetPageId="4" captionText="FIGURE 2. Goodyera umbonata. A. Flowering plant. B. Portion of inflorescence. C. Floral bract, adaxial side. D. Flattened median sepal and petals, adaxial side. E. Median sepal, adaxial side. F. Lateral sepals, adaxial side. G. Petals, adaxial side. H. Lip, view from above and from below. I. Lip, side view and sagittal section. J. Artificially flattened lip, adaxial view. K. Intact column, and column with anther removed. L. Column with anther, view from above. M. Column with anther removed, view from above. N. Column and stigma, view from below. O. Anther, side view. P. Anther and pollinarium, view from below. Drawn from the type specimen (V.C.Nguyen AL 1545) by L.Averyanov and T.Maisak." figureDoi="http://doi.org/10.5281/zenodo.8430555" httpUri="https://zenodo.org/record/8430555/files/figure.png" pageId="2" pageNumber="257">Fig. 2</figureCitation>
)
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<emphasis id="B923EA927306292FFF35F902FF60F8D9" bold="true" box="[136,251,1778,1804]" pageId="2" pageNumber="257">Diagnosis</emphasis>
:—The new species differs from the morphologically closest 
<taxonomicName id="4C574D037306292FFC0CF902FB52F8D9" authorityName="Averyanov" authorityYear="2007" box="[945,1225,1778,1804]" class="Liliopsida" family="Orchidaceae" genus="Goodyera" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="257" phylum="Tracheophyta" rank="species" species="rhombodoides">
<emphasis id="B923EA927306292FFC0CF902FB52F8D9" box="[945,1225,1778,1804]" italics="true" pageId="2" pageNumber="257">Goodyera rhombodoides</emphasis>
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, 
<taxonomicName id="4C574D037306292FFB65F903FAE0F8D9" authorityName="Seidenfaden" authorityYear="1969" box="[1240,1403,1778,1805]" class="Liliopsida" family="Orchidaceae" genus="Goodyera" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="257" phylum="Tracheophyta" rank="species" species="thailandica">
<emphasis id="B923EA927306292FFB65F903FAE0F8D9" box="[1240,1403,1778,1805]" italics="true" pageId="2" pageNumber="257">G. thailandica</emphasis>
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and 
<taxonomicName id="4C574D037306292FFF35F8E7FE92F8E5" authorityName="Fukuyama" authorityYear="1936" box="[136,265,1815,1841]" class="Liliopsida" family="Orchidaceae" genus="Goodyera" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="257" phylum="Tracheophyta" rank="species" species="yamiana">
<emphasis id="B923EA927306292FFF35F8E7FE92F8E5" box="[136,265,1815,1841]" italics="true" pageId="2" pageNumber="257">G. yamiana</emphasis>
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in smaller flowers, obliquely spatulate to narrowly obovate petals with truncate apices, cymbiform epichile with thin plicate margin, and prominent umbo at epichile center.
</paragraph>
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<emphasis id="B923EA927306292FFF00F8AEFF63F8AD" bold="true" box="[189,248,1886,1912]" pageId="2" pageNumber="257">Type</emphasis>
:— 
<materialsCitation id="3B3F3CDD7306292FFEA4F8AEFDA7F815" collectingDate="2022-02" collectionCode="AL, LE" collectorName="Nguyen Van Canh" country="Vietnam" location="Chu Mu Mountain" municipality="M'Drak District" pageId="2" pageNumber="257" specimenCode="AL 1545, LE01169141, LE01124095" specimenCount="1" stateProvince="Dak Lak" typeStatus="holotype">
<collectingCountry id="F34076107306292FFEA4F8AEFE3BF8AD" box="[281,416,1886,1912]" name="Vietnam" pageId="2" pageNumber="257">VIETNAM</collectingCountry>
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<collectingRegion id="4993F8627306292FFE0CF8AEFD15F8AD" box="[433,654,1886,1912]" country="Vietnam" name="Dak Lak" pageId="2" pageNumber="257">Dak Lak Province</collectingRegion>
: 
<collectingMunicipality id="6B8CACFA7306292FFD22F8AEFCC6F8AD" box="[671,861,1886,1912]" pageId="2" pageNumber="257">M’Drak District</collectingMunicipality>
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, 
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<collectorName id="26A253567306292FFAA3F8AFFF5EF849" pageId="2" pageNumber="257">Nguyen Van Canh</collectorName>
<specimenCode id="DBF19EFB7306292FFF72F873FEA8F849" box="[207,307,1922,1949]" collectionCode="AL" country="Algeria" lsid="urn:lsid:biocol.org:col:14432" name="Université d'Alger" pageId="2" pageNumber="257" type="Herbarium">AL 1545</specimenCode>
</emphasis>
(
<typeStatus id="54EC88227306292FFEFBF872FE31F849" box="[326,426,1922,1948]" pageId="2" pageNumber="257">holotype</typeStatus>
<specimenCode id="DBF19EFB7306292FFE08F872FDDDF849" box="[437,582,1922,1948]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="2" pageNumber="257">LE01169141</specimenCode>
https://en.herbariumle.ru/?t=occ&amp;id=141019, drawing 
<specimenCode id="DBF19EFB7306292FFB7CF872FAC9F848" box="[1217,1362,1922,1949]" collectionCode="LE" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="2" pageNumber="257">LE01124095</specimenCode>
https:// en.herbariumle.ru/?t=occ&amp;id=168135)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D650B7306292FFF00F83AFE40F7DC" pageId="2" pageNumber="257" type="etymology">
<paragraph id="8BE836807306292FFF00F83AFE40F7DC" blockId="2.[136,1452,1778,2057]" pageId="2" pageNumber="257">
<emphasis id="B923EA927306292FFF00F83AFEA5F831" bold="true" box="[189,318,1994,2020]" pageId="2" pageNumber="257">Etymology</emphasis>
:—The species name refers to the umbo-like callus on the lip epichile which distinguishes the new species from all its congeners.
</paragraph>
</subSubSection>
<caption id="DF2866087307292EFF35F8C8FA14F7D6" ID-DOI="http://doi.org/10.5281/zenodo.8430551" ID-Zenodo-Dep="8430551" httpUri="https://zenodo.org/record/8430551/files/figure.png" pageId="3" pageNumber="258" startId="3.[136,229,1848,1870]" targetBox="[249,1338,190,1824]" targetPageId="3" targetType="figure">
<paragraph id="8BE836807307292EFF35F8C8FA14F7D6" blockId="3.[136,1452,1848,2051]" pageId="3" pageNumber="258">
<emphasis id="B923EA927307292EFF35F8C8FF65F898" bold="true" box="[136,254,1848,1870]" pageId="3" pageNumber="258">FIGURE 1.</emphasis>
<taxonomicName id="4C574D037307292EFEB9F8C8FD20F89B" authority="var. barbigerum" box="[260,699,1848,1870]" class="Liliopsida" family="Orchidaceae" genus="Paphiopedilum" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="variety" species="barbigerum" variety="barbigerum">
<emphasis id="B923EA927307292EFEB9F8C8FD8EF89B" bold="true" box="[260,533,1848,1870]" italics="true" pageId="3" pageNumber="258">Paphiopedilum barbigerum</emphasis>
var. barbigerum
</taxonomicName>
<emphasis id="B923EA927307292EFD06F8C8FD46F898" bold="true" box="[699,733,1848,1870]" pageId="3" pageNumber="258">
<emphasis id="B923EA927307292EFD06F8C8FD24F89B" bold="true" box="[699,703,1848,1870]" italics="true" pageId="3" pageNumber="258">.</emphasis>
A.
</emphasis>
Flowering plant in natural habitat. 
<emphasis id="B923EA927307292EFB8CF8C9FBDAF89B" bold="true" box="[1073,1089,1849,1870]" pageId="3" pageNumber="258">B</emphasis>
&amp; 
<emphasis id="B923EA927307292EFBDCF8C8FBE3F89B" bold="true" box="[1121,1144,1848,1870]" pageId="3" pageNumber="258">C.</emphasis>
Flowers, front view. Specimen: 
<emphasis id="B923EA927307292EFF35F8ADFE32F8A7" box="[136,425,1885,1906]" italics="true" pageId="3" pageNumber="258">L.T.Van, K.S.Nguyen AL 1605</emphasis>
. 
<emphasis id="B923EA927307292EFE0BF8ACFD35F8A7" bold="true" box="[438,686,1884,1906]" pageId="3" pageNumber="258">
<taxonomicName id="4C574D037307292EFE0BF8ACFD12F8A7" box="[438,649,1884,1906]" class="Liliopsida" family="Orchidaceae" genus="Myrmechis" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="species" species="urceolata">
<emphasis id="B923EA927307292EFE0BF8ACFD12F8A7" bold="true" box="[438,649,1884,1906]" italics="true" pageId="3" pageNumber="258">Myrmechis urceolata</emphasis>
</taxonomicName>
. D.
</emphasis>
Flattened flowering and fruiting plants. 
<emphasis id="B923EA927307292EFB84F8ADFBD2F8A7" bold="true" box="[1081,1097,1885,1906]" pageId="3" pageNumber="258">E</emphasis>
&amp; 
<emphasis id="B923EA927307292EFBD7F8ADFBE6F8A7" bold="true" box="[1130,1149,1885,1906]" pageId="3" pageNumber="258">F.</emphasis>
Apical part of floriferous stem with flowers. 
<emphasis id="B923EA927307292EFEAFF870FEBEF843" bold="true" box="[274,293,1920,1942]" pageId="3" pageNumber="258">G</emphasis>
&amp; 
<emphasis id="B923EA927307292EFEF4F871FEF8F843" bold="true" box="[329,355,1921,1942]" pageId="3" pageNumber="258">H.</emphasis>
Flowers and flower buds, side and front view. Specimen: 
<emphasis id="B923EA927307292EFC10F871FB4EF843" box="[941,1237,1921,1942]" italics="true" pageId="3" pageNumber="258">T.B.Vuong, N.T.Luan AL 1237</emphasis>
. 
<emphasis id="B923EA927307292EFB5EF870FA30F843" bold="true" box="[1251,1451,1920,1942]" pageId="3" pageNumber="258">
<taxonomicName id="4C574D037307292EFB5EF870FA15F843" authorityName="Seidenfaden" authorityYear="1977" baseAuthorityName="D. Don" box="[1251,1422,1920,1942]" class="Liliopsida" family="Orchidaceae" genus="Peristylus" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="258" phylum="Tracheophyta" rank="species" species="affinis">
<emphasis id="B923EA927307292EFB5EF870FA15F843" bold="true" box="[1251,1422,1920,1942]" italics="true" pageId="3" pageNumber="258">Peristylus affinis</emphasis>
</taxonomicName>
. I.
</emphasis>
Flattened plant. 
<emphasis id="B923EA927307292EFE95F855FEA0F86F" bold="true" box="[296,315,1957,1978]" pageId="3" pageNumber="258">J.</emphasis>
Portion of inflorescence. 
<emphasis id="B923EA927307292EFD80F855FDCDF86F" bold="true" box="[573,598,1957,1978]" pageId="3" pageNumber="258">K.</emphasis>
Flower, side view. 
<emphasis id="B923EA927307292EFCA7F855FCAAF86F" bold="true" box="[794,817,1957,1978]" pageId="3" pageNumber="258">L.</emphasis>
Detached flower, side view, with sepals removed. 
<emphasis id="B923EA927307292EFA91F855FAD3F86F" bold="true" box="[1324,1352,1957,1978]" pageId="3" pageNumber="258">M.</emphasis>
Detached flower, sagittal section. 
<emphasis id="B923EA927307292EFED1F839FE18F80B" bold="true" box="[364,387,1993,2014]" pageId="3" pageNumber="258">N.</emphasis>
Detached flattened sepals, petals and lip. Specimen: 
<emphasis id="B923EA927307292EFCC3F839FADAF80B" box="[894,1345,1993,2014]" italics="true" pageId="3" pageNumber="258">S.K.Nguyen, T.H.Nguyen, M.H.Giang NSK 950</emphasis>
. Photos by L.T.Van &amp; K.S.Nguyen (A–C), T.B.Vuong (D–H) and K.S.Nguyen (I–N); photo correction and design by L.Averyanov and T.Maisak.
</paragraph>
</caption>
<caption id="DF28660873002929FF35F8BAFE0FF7C1" ID-DOI="http://doi.org/10.5281/zenodo.8430555" ID-Zenodo-Dep="8430555" httpUri="https://zenodo.org/record/8430555/files/figure.png" pageId="4" pageNumber="259" startId="4.[136,229,1866,1888]" targetBox="[243,1344,192,1841]" targetPageId="4" targetType="figure">
<paragraph id="8BE8368073002929FF35F8BAFE0FF7C1" blockId="4.[136,1452,1866,2068]" pageId="4" pageNumber="259">
<emphasis id="B923EA9273002929FF35F8BAFE75F88A" bold="true" box="[136,494,1866,1888]" pageId="4" pageNumber="259">
FIGURE 2. 
<taxonomicName id="4C574D0373002929FEBFF8BAFE50F8B5" box="[258,459,1866,1888]" class="Liliopsida" family="Orchidaceae" genus="Goodyera" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="259" phylum="Tracheophyta" rank="species" species="umbonata">
<emphasis id="B923EA9273002929FEBFF8BAFE50F8B5" bold="true" box="[258,459,1866,1888]" italics="true" pageId="4" pageNumber="259">Goodyera umbonata</emphasis>
</taxonomicName>
. A.
</emphasis>
Flowering plant. 
<emphasis id="B923EA9273002929FD2BF8BAFD37F88A" bold="true" box="[662,684,1866,1887]" pageId="4" pageNumber="259">B.</emphasis>
Portion of inflorescence. 
<emphasis id="B923EA9273002929FC1CF8BAFC23F8B5" bold="true" box="[929,952,1866,1888]" pageId="4" pageNumber="259">C.</emphasis>
Floral bract, adaxial side. 
<emphasis id="B923EA9273002929FB0EF8BAFB50F88A" bold="true" box="[1203,1227,1866,1887]" pageId="4" pageNumber="259">D.</emphasis>
Flattened median sepal and petals, adaxial side. 
<emphasis id="B923EA9273002929FECAF89EFE15F856" bold="true" box="[375,398,1902,1923]" pageId="4" pageNumber="259">E.</emphasis>
Median sepal, adaxial side. 
<emphasis id="B923EA9273002929FD19F89EFD2DF856" bold="true" box="[676,694,1902,1923]" pageId="4" pageNumber="259">F.</emphasis>
Lateral sepals, adaxial side. 
<emphasis id="B923EA9273002929FC6CF89EFC71F851" bold="true" box="[977,1002,1902,1924]" pageId="4" pageNumber="259">G.</emphasis>
Petals, adaxial side. 
<emphasis id="B923EA9273002929FB05F89EFB4AF856" bold="true" box="[1208,1233,1902,1923]" pageId="4" pageNumber="259">H.</emphasis>
Lip, view from above and from below. 
<emphasis id="B923EA9273002929FE97F862FEA2F872" bold="true" box="[298,313,1938,1959]" pageId="4" pageNumber="259">I.</emphasis>
Lip, side view and sagittal section. 
<emphasis id="B923EA9273002929FD2DF862FD38F872" bold="true" box="[656,675,1938,1959]" pageId="4" pageNumber="259">J.</emphasis>
Artificially flattened lip, adaxial view. 
<emphasis id="B923EA9273002929FBA7F862FBA8F872" bold="true" box="[1050,1075,1938,1959]" pageId="4" pageNumber="259">K.</emphasis>
Intact column, and column with anther removed. 
<emphasis id="B923EA9273002929FF57F846FE9BF81E" bold="true" box="[234,256,1974,1995]" pageId="4" pageNumber="259">L.</emphasis>
Column with anther, view from above. 
<emphasis id="B923EA9273002929FD36F846FD33F81E" bold="true" box="[651,680,1974,1995]" pageId="4" pageNumber="259">M.</emphasis>
Column with anther removed, view from above. 
<emphasis id="B923EA9273002929FB2DF846FB33F81E" bold="true" box="[1168,1192,1974,1995]" pageId="4" pageNumber="259">N.</emphasis>
Column and stigma, view from below. 
<emphasis id="B923EA9273002929FEBFF82AFE80F825" bold="true" box="[258,283,2010,2032]" pageId="4" pageNumber="259">O.</emphasis>
Anther, side view. 
<emphasis id="B923EA9273002929FE6EF82AFE7EF83A" bold="true" box="[467,485,2010,2031]" pageId="4" pageNumber="259">P.</emphasis>
Anther and pollinarium, view from below. Drawn from the type specimen (
<emphasis id="B923EA9273002929FB00F82AFA1EF83A" box="[1213,1413,2010,2031]" italics="true" pageId="4" pageNumber="259">V.C.Nguyen AL 1545</emphasis>
) by L.Averyanov and T.Maisak.
</paragraph>
</caption>
<subSubSection id="C34D650B73012928FF00FF6EFF49FBCD" pageId="5" pageNumber="260" type="description">
<paragraph id="8BE8368073012928FF00FF6EFF49FBCD" blockId="5.[136,1452,158,1517]" pageId="5" pageNumber="260">
<emphasis id="B923EA9273012928FF00FF6EFDB5FF6D" bold="true" box="[189,558,158,185]" pageId="5" pageNumber="260">Description:—Terrestrial herb</emphasis>
with creeping rhizome and ascending leafy floriferous stem. 
<emphasis id="B923EA9273012928FB4EFF6EFAC6FF6D" bold="true" box="[1267,1373,158,184]" pageId="5" pageNumber="260">Rhizome</emphasis>
green, fleshy, (4)5–6(8) cm long, rooting at nodes. 
<emphasis id="B923EA9273012928FDC2FF32FCDEFF09" bold="true" box="[639,837,194,220]" pageId="5" pageNumber="260">Floriferous stem</emphasis>
(8)10–12(14) cm tall, glabrous, with (3)4–5(6) leaves distant in lower half. 
<emphasis id="B923EA9273012928FEC1FF16FE55FED5" bold="true" box="[380,462,230,256]" pageId="5" pageNumber="260">Leaves</emphasis>
petiolate, glabrous; petiole and sheath (6)8–22(26) mm long, 
<quantity id="4CAF9B6573012928FB2DFF16FB75FED4" box="[1168,1262,230,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="260" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
wide; leaf blade uniformly grassy green adaxially, pale green abaxially, narrowly ovate, sometimes slightly oblique, gradually tapering to the apex, acute, (2.2)2.5–4.5(5) cm long, (1)1.2–1.5(1.7) cm wide. 
<emphasis id="B923EA9273012928FC7DFEDEFBB6FE9D" bold="true" box="[960,1069,302,328]" pageId="5" pageNumber="260">Peduncle</emphasis>
erect, (6)7–8(9) cm long, finely papillose to shortly hairy, with 2–3 sterile bracts; bracts greenish, cuneate, acuminate, (5)6–15(16) mm long, 
<quantity id="4CAF9B6573012928FAEAFEA2FF2FFE44" metricMagnitude="-3" metricUnit="m" metricValue="1.05" metricValueMax="1.6" metricValueMin="0.5" pageId="5" pageNumber="260" unit="mm" value="1.05" valueMax="1.6" valueMin="0.5">0.5–1.6 mm</quantity>
wide, shortly ciliate or fimbriate along margin, entirely glabrous; rachis finely hairy, usually with (10)12–16(18) laxly arranged flowers. 
<emphasis id="B923EA9273012928FE22FE6AFDA5FE61" bold="true" box="[415,574,410,436]" pageId="5" pageNumber="260">Floral bracts</emphasis>
light green, herbaceous, broadly ovate, concave, attenuate, finely serrulate or shortly fimbriate along margin, sparsely verruculose abaxially, (4)4.5–6(7) mm long, (0.8)1–1.5(1.6) mm wide, entirely glabrous. 
<emphasis id="B923EA9273012928FEE6FE12FDACFE29" bold="true" box="[347,567,482,508]" pageId="5" pageNumber="260">Pedicel and ovary</emphasis>
grassy green, cylindrical, twisted, straight to strongly curved, (3.5)4–7.5(8) mm long, 
<quantity id="4CAF9B6573012928FEBCFDF6FEECFDF4" box="[257,375,518,545]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="5" pageNumber="260" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
in diameter, shortly sparsely papillulate to subglabrous. 
<emphasis id="B923EA9273012928FBB6FDF6FBF1FDF5" bold="true" box="[1035,1130,518,544]" pageId="5" pageNumber="260">Flowers</emphasis>
resupinate, hardly opening, (2.8)3–3.5(3.8) mm long. 
<emphasis id="B923EA9273012928FE14FDDAFE6EFD91" bold="true" box="[425,501,554,580]" pageId="5" pageNumber="260">Sepals</emphasis>
subequal, forward directed, narrowly ovate, cymbiform, slightly cucullate, obtuse, 1- nerved, (2.6)2.8–3(3.2) mm long, (1.4)1.6–1.8(2) mm wide, adaxially glabrous, abaxially sparsely papillulate to almost glabrous; lateral sepals slightly oblique. 
<emphasis id="B923EA9273012928FDF1FD82FD0FFD59" bold="true" box="[588,660,626,652]" pageId="5" pageNumber="260">Petals</emphasis>
obliquely spatulate to narrowly obovate, truncate to almost rounded at apex, 1-nerved, as long as sepals, 
<quantity id="4CAF9B6573012928FE46FD66FD1AFD64" box="[507,641,662,689]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="5" pageNumber="260" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.2–1.4 mm</quantity>
wide, glabrous, slightly eroded or almost entire at apex, tightly connivent in apical half with the median sepal forming together forward directed hood. 
<emphasis id="B923EA9273012928FC5BFD4AFB8BFD01" bold="true" box="[998,1040,698,724]" pageId="5" pageNumber="260">Lip</emphasis>
(3.2)3.4–3.8(4) mm long, 
<quantity id="4CAF9B6573012928FA87FD4AFA37FD00" box="[1338,1452,698,725]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="5" pageNumber="260" unit="mm" value="2.2" valueMax="2.4" valueMin="2.0">2–2.4 mm</quantity>
wide (being flattened), distinctly divided into hypochile and epichile; hypochile rather fleshy, concave, hemispheric, cup-like, 
<quantity id="4CAF9B6573012928FF4EFCF2FEE0FCC8" box="[243,379,770,797]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="5" pageNumber="260" unit="mm" value="1.55" valueMax="1.6" valueMin="1.5">1.5–1.6 mm</quantity>
across, with numerous long thick multicellular hairs inside; epichile forward directed, broadly ovate, cymbiform, 
<quantity id="4CAF9B6573012928FEE0FCD6FE7FFC94" box="[349,484,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="5" pageNumber="260" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
long, 
<quantity id="4CAF9B6573012928FD96FCD6FD29FC94" box="[555,690,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="5" pageNumber="260" unit="mm" value="2.4" valueMax="2.6" valueMin="2.2">2.2–2.6 mm</quantity>
wide (being flattened), entire, with thin plicate margin, at base with 2 distinct papillulate callosities and in center with prominent umbo-like callus. 
<emphasis id="B923EA9273012928FC40FCBAFBC6FCB1" bold="true" box="[1021,1117,842,868]" pageId="5" pageNumber="260">Column</emphasis>
(1.5)1.6–1.8(2) mm tall, with stout, shortly cylindrical base and 2 finger-like forward directed incurved rostellar arms 
<quantity id="4CAF9B6573012928FB3FFC9EFA97FC5C" box="[1154,1292,878,905]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="5" pageNumber="260" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.7–0.8 mm</quantity>
long, bearing anther distally and elliptic verruculose stigma proximally. 
<emphasis id="B923EA9273012928FC92FC62FC1FFC79" bold="true" box="[815,900,914,940]" pageId="5" pageNumber="260">Anther</emphasis>
semi-globose, (1.2)1.4–1.6(1.7) mm long, with short triangular beak at front; pollinarium with very small, narrowly elliptic viscidium and numerous polyads arranged in two rows. 
<emphasis id="B923EA9273012928FEA4FC2AFEE2FC21" bold="true" box="[281,377,986,1012]" pageId="5" pageNumber="260">Capsule</emphasis>
suberect, fusiform, slightly attenuate at apex, 
<quantity id="4CAF9B6573012928FCC1FC2AFC7CFC20" box="[892,999,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="5" pageNumber="260" unit="mm" value="9.0" valueMax="10.0" valueMin="8.0">8–10 mm</quantity>
long, 
<quantity id="4CAF9B6573012928FB91FC2AFB05FC20" box="[1068,1182,986,1013]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="5" pageNumber="260" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
in diameter, opening by 6 slits.
</paragraph>
</subSubSection>
<subSubSection id="C34D650B73012928FF00FBD2FACAFBE9" box="[189,1361,1058,1085]" pageId="5" pageNumber="260" type="biology_ecology">
<paragraph id="8BE8368073012928FF00FBD2FACAFBE9" blockId="5.[136,1452,158,1517]" box="[189,1361,1058,1085]" pageId="5" pageNumber="260">
<emphasis id="B923EA9273012928FF00FBD2FE49FBE9" bold="true" box="[189,466,1058,1084]" pageId="5" pageNumber="260">Ecology and phenology</emphasis>
:—Evergreen broad-leaved submontane forests on granite. Flowers in February.
</paragraph>
</subSubSection>
<subSubSection id="C34D650B73012928FF00FBB6FC60FA39" pageId="5" pageNumber="260" type="distribution">
<paragraph id="8BE8368073012928FF00FBB6FB73FBB5" blockId="5.[136,1452,158,1517]" box="[189,1256,1094,1121]" pageId="5" pageNumber="260">
<emphasis id="B923EA9273012928FF00FBB6FED4FBB5" bold="true" box="[189,335,1094,1120]" pageId="5" pageNumber="260">Distribution</emphasis>
:—Endemic to 
<collectingCountry id="F340761073012928FE47FBB6FDC2FBB5" box="[506,601,1094,1120]" name="Vietnam" pageId="5" pageNumber="260">Vietnam</collectingCountry>
(
<collectingRegion id="4993F86273012928FDDAFBB6FCA2FBB5" box="[615,825,1094,1120]" country="Vietnam" name="Dak Lak" pageId="5" pageNumber="260">Dak Lak Province</collectingRegion>
: M’Drak District, Chu Mu Mountain).
</paragraph>
<paragraph id="8BE8368073012928FF00FB9AFF41FB25" blockId="5.[136,1452,158,1517]" pageId="5" pageNumber="260">
<emphasis id="B923EA9273012928FF00FB9AFE37FB51" bold="true" box="[189,428,1130,1156]" pageId="5" pageNumber="260">Conservation status</emphasis>
:—The species is known from a single location with several mature individuals observed. Pending comprehensive field studies in the area of its 
<emphasis id="B923EA9273012928FCBAFB7EFC2AFB7D" box="[775,945,1166,1192]" italics="true" pageId="5" pageNumber="260">locus classicus</emphasis>
, we estimate the conservation status of the species as Data Deficient (DD) according to the criteria proposed by the 
<bibRefCitation id="EFC64B7173012928FC73FB42FF4DFB25" author="IUCN Standards &amp; Petitions Committee" pageId="5" pageNumber="260" refId="ref16566" refString="IUCN Standards and Petitions Committee (2023) Guidelines for Using the IUCN Red List Categories and Criteria. Version 14. Prepared by the Standards and Petitions Committee. Available from: https: // www. iucnredlist. org / documents / RedListGuidelines. pdf (accessed on 16 June 2023)." type="book" year="2023">IUCN Standards and Petitions Committee (2023)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE8368073012928FF00FB0AFC60FA39" blockId="5.[136,1452,158,1517]" pageId="5" pageNumber="260">
<emphasis id="B923EA9273012928FF00FB0AFE9BFAC1" bold="true" box="[189,256,1274,1300]" pageId="5" pageNumber="260">Notes</emphasis>
:—In its habit and floral morphology, 
<taxonomicName id="4C574D0373012928FD1EFB0AFC12FAC1" box="[675,905,1274,1300]" class="Liliopsida" family="Orchidaceae" genus="Goodyera" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="260" phylum="Tracheophyta" rank="species" species="umbonata">
<emphasis id="B923EA9273012928FD1EFB0AFC12FAC1" box="[675,905,1274,1300]" italics="true" pageId="5" pageNumber="260">Goodyera umbonata</emphasis>
</taxonomicName>
is somewhat similar to its SE Asian congeners 
<taxonomicName id="4C574D0373012928FA2DFB0BFDA9FAED" authority="Averyanov (2007: 297)" authorityName="Averyanov" authorityPageNumber="297" authorityYear="2007" class="Liliopsida" family="Orchidaceae" genus="Goodyera" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="260" phylum="Tracheophyta" rank="species" species="rhombodoides">
<emphasis id="B923EA9273012928FA2DFB0BFEB3FAED" italics="true" pageId="5" pageNumber="260">G. rhombodoides</emphasis>
<bibRefCitation id="EFC64B7173012928FE91FAEEFDA9FAED" author="Averyanov, L. V." box="[300,562,1310,1336]" pageId="5" pageNumber="260" pagination="287 - 306" refId="ref14359" refString="Averyanov, L. V. (2007) New species of orchids from Vietnam. Taiwania 52 (4): 287 - 306. https: // doi. org / 10.6165 / tai. 2007.52 (4). 287" type="journal article" year="2007">Averyanov (2007: 297)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C574D0373012928FD80FAEFFC6CFAED" authority="Seidenfaden (1969: 109)" authorityName="Seidenfaden" authorityPageNumber="109" authorityYear="1969" box="[573,1015,1310,1337]" class="Liliopsida" family="Orchidaceae" genus="Goodyera" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="260" phylum="Tracheophyta" rank="species" species="thailandica">
<emphasis id="B923EA9273012928FD80FAEFFD45FAED" box="[573,734,1310,1337]" italics="true" pageId="5" pageNumber="260">G. thailandica</emphasis>
<bibRefCitation id="EFC64B7173012928FD59FAEEFC6CFAED" author="Seidenfaden, G." box="[740,1015,1310,1336]" pageId="5" pageNumber="260" pagination="100 - 162" refId="ref18013" refString="Seidenfaden, G. (1969) Contributions to the orchid flora of Thailand. Botanisk Tidsskrift 65 (1 - 2): 100 - 162." type="journal article" year="1969">Seidenfaden (1969: 109)</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C574D0373012928FB96FAEFFA3CFAED" authority="Fukuyama (1936: 18)" authorityName="Fukuyama" authorityPageNumber="18" authorityYear="1936" box="[1067,1447,1310,1337]" class="Liliopsida" family="Orchidaceae" genus="Goodyera" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="260" phylum="Tracheophyta" rank="species" species="yamiana">
<emphasis id="B923EA9273012928FB96FAEFFB36FAED" box="[1067,1197,1311,1337]" italics="true" pageId="5" pageNumber="260">G. yamiana</emphasis>
<bibRefCitation id="EFC64B7173012928FB0EFAEEFA3CFAED" author="Fukuyama, N." box="[1203,1447,1310,1336]" pageId="5" pageNumber="260" pagination="16 - 24" refId="ref16171" refString="Fukuyama, N. (1936) Studia orchidacearum Japonicarum. VIII. Orchidaceae Formosae et Insulae Koto-syo novae atque criticae. Botanical Magazine (Tokyo) 50 (589): 16 - 24. https: // doi. org / 10.15281 / jplantres 1887.50.16" type="journal article" year="1936">Fukuyama (1936: 18)</bibRefCitation>
</taxonomicName>
. The new species differs from them in smaller flowers, obliquely spatulate to narrowly obovate petals truncate to almost rounded at apex, cymbiform epichile with thin plicate margin, and prominent umbo at epichile center. 
<taxonomicName id="4C574D0373012928FA86FA96FF63FA71" class="Liliopsida" family="Orchidaceae" genus="Goodyera" kingdom="Plantae" order="Asparagales" pageId="5" pageNumber="260" phylum="Tracheophyta" rank="species" species="umbonata">
<emphasis id="B923EA9273012928FA86FA96FF63FA71" italics="true" pageId="5" pageNumber="260">Goodyera umbonata</emphasis>
</taxonomicName>
represents an example of strict endemics of the mountainous area surrounding Langbian Plateau (a part of the so-called Central Highlands of 
<collectingCountry id="F340761073012928FDAEFA5EFDEDFA1D" box="[531,630,1454,1480]" name="Vietnam" pageId="5" pageNumber="260">Vietnam</collectingCountry>
: 
<bibRefCitation id="EFC64B7173012928FD3CFA5EFCEAFA1D" author="Poyarkov, N. A. &amp; Nguyen, T. V. &amp; Popov, E. S. &amp; Geissler, P. &amp; Pawangkhanant, P. &amp; Neang, T. &amp; Suwannapoom, C. &amp; Orlov, N. L." box="[641,881,1454,1480]" pageId="5" pageNumber="260" pagination="1 - 110" refId="ref17693" refString="Poyarkov, N. A., Nguyen, T. V., Popov, E. S., Geissler, P., Pawangkhanant, P., Neang, T., Suwannapoom, C. &amp; Orlov, N. L. (2021) Recent progress in taxonomic studies, biogeographic analysis and revised checklist of amphibians in Indochina. Russian Journal of Herpetology 28 (3 A): 1 - 110. https: // doi. org / 10.30906 / 1026 - 2296 - 2021 - 28 - 3 A- 1 - 110" type="journal article" year="2021">
Poyarkov 
<emphasis id="B923EA9273012928FD49FA5FFCB6FA1D" box="[756,813,1454,1480]" italics="true" pageId="5" pageNumber="260">et al.</emphasis>
2021
</bibRefCitation>
), which is outlined in the biogeography of Eastern Indochina as the South Annamese Floristic Region (
<bibRefCitation id="EFC64B7173012928FD73FA22FC40FA39" author="Averyanov, L. &amp; Cribb, P. &amp; Loc, P. K. &amp; Nguyen, T. H." box="[718,987,1490,1516]" pageId="5" pageNumber="260" refId="ref14556" refString="Averyanov, L., Cribb, P., Loc, P. K. &amp; Nguyen, T. H. (2003 a) Slipper orchids of Vietnam. With an introduction to the flora of Vietnam. Compass Press Limited, Kew, 308 pp." type="book" year="2003">
Averyanov 
<emphasis id="B923EA9273012928FCECFA23FC11FA39" box="[849,906,1490,1516]" italics="true" pageId="5" pageNumber="260">et al.</emphasis>
2003a
</bibRefCitation>
,b).
</paragraph>
</subSubSection>
</treatment>
</document>