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<mods:title id="10E2AF9DA81471592473A5FC4999600D">Revision of the genus Harmothoe Kinberg, 1856 (Polychaeta: Polynoidae) in the Northeast Atlantic</mods:title>
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<emphasis id="B97B747DC207FFFDFF18FDD9FEDB1087" bold="true" box="[151,383,151,177]" italics="true" pageId="36" pageNumber="37">Harmothoe fragilis</emphasis>
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(
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, 
<figureCitation id="1334B4EAC207FFFDFF70FDF4FEF210E2" box="[255,342,186,212]" captionStart="FIGURE 17" captionStartId="37.[151,255,1869,1893]" captionTargetBox="[273,1373,341,1841]" captionTargetId="figure-181@37.[151,1436,329,1852]" captionTargetPageId="37" captionText="FIGURE 17. Harmothoe fragilis (additional specimen, SMF 17309). (A) anterior end; styles of tentacular cirri and left dorsal cirrus of segment 3 missing; (B) right elytron from segment 5; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus missing; (E) long notochaeta; (F) tip of same; (G) short notochaeta; (H) middle neurochaeta; (I) tip of same; (J) tip of uppermost neurochaeta; (K) tip of lower neurochaeta. (Scale: A, B = 2 mm; C, E, G, H = 250 µm; D = 1 mm; F, I–K = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361475" httpUri="https://zenodo.org/record/5361475/files/figure.png" pageId="36" pageNumber="37">17A–K</figureCitation>
)
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<emphasis id="B97B747DC207FFFDFF18FC46FEB81117" box="[151,284,264,289]" italics="true" pageId="36" pageNumber="37">Harmothoe</emphasis>
(
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) 
<emphasis id="B97B747DC207FFFDFE1CFC46FE4C1117" box="[403,488,264,289]" italics="true" pageId="36" pageNumber="37">fragilis</emphasis>
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, pl. 29 figs. 29, 30, pl. 30 figs. 31, 32.
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<taxonomicName id="4C0FD3ECC207FFFDFF18FC61FD6F117E" ID-CoL="6LCG5" authority=": Hartmann-Schroder (1996)" authorityName=": Hartmann-Schroder" authorityYear="1996" box="[151,715,302,328]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="fragilis">
<emphasis id="B97B747DC207FFFDFF18FC61FEDD117E" box="[151,377,303,328]" italics="true" pageId="36" pageNumber="37">Harmothoe fragilis</emphasis>
: 
<bibRefCitation id="EF9ED59EC207FFFDFE07FC60FD6F117E" author="Hartmann-Schroder, G." box="[392,715,302,328]" pageId="36" pageNumber="37" pagination="1 - 648" refId="ref37937" refString="Hartmann-Schroder, G. (1996) Annelida, Borstenwurmer, Polychaeta. Die Tierwelt Deutschlands. 2. neubearbeitete Auflage, 58, 1 - 648." type="journal article" year="1996">Hartmann-Schröder (1996)</bibRefCitation>
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: 44
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; 
<treatmentCitation id="0AAE8E7EC207FFFDFC8CFC60FBCB117E" author="Chambers, S. J. &amp; Muir, A. I." box="[771,1135,302,328]" page="100" pageId="36" pageNumber="37" year="1997">
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: 100
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, fig. 24; 
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<bibRefCitation id="EF9ED59EC207FFFDFB57FC60FF4D1159" author="Barnich, R. &amp; Fiege, D." pageId="36" pageNumber="37" pagination="1889 - 1938" refId="ref36372" refString="Barnich, R. &amp; Fiege, D. (2000) Revision of the Mediterranean species of Harmothoe Kinberg, 1856 and Lagisca Malmgren, 1865 (Polychaeta, Polynoidae, Polynoinae) with descriptions of a new genus and a new species. Journal of Natural History, 34, 1889 - 1938." type="journal article" year="2000">Barnich &amp; Fiege (2000)</bibRefCitation>
: 1911
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, fig. 11A–D; 
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<bibRefCitation id="EF9ED59EC207FFFDFE5FFC1BFD491159" author="Barnich, R. &amp; Fiege, D." box="[464,749,341,367]" pageId="36" pageNumber="37" pagination="1 - 167" refId="ref36428" refString="Barnich, R. &amp; Fiege, D. (2003) The Aphroditoidea (Annelida, Polychaeta) of the Mediterranean Sea. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 559, 1 - 167." type="journal article" year="2003">Barnich &amp; Fiege (2003)</bibRefCitation>
: 41
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, fig. 17A–D.
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<treatmentCitation id="0AAE8E7EC207FFFDFF18FC32FDE311A0" author="Malmgren, A. J." box="[151,583,380,406]" page="71" pageId="36" pageNumber="37" year="1866">
<taxonomicName id="4C0FD3ECC207FFFDFF18FC32FDB811A0" ID-CoL="3DGYT" authority=": Malmgren (1866)" authorityName="Malmgren" authorityYear="1866" box="[151,540,380,406]" class="Polychaeta" family="Polynoidae" genus="Evarne" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="impar">
<emphasis id="B97B747DC207FFFDFF18FC32FE9D11A3" box="[151,313,380,405]" italics="true" pageId="36" pageNumber="37">Evarne impar</emphasis>
: 
<bibRefCitation id="EF9ED59EC207FFFDFEC7FC32FDB811A0" author="Malmgren, A. J." box="[328,540,380,406]" pageId="36" pageNumber="37" pagination="51 - 110" refId="ref38655" refString="Malmgren, A. J. (1866) Nordiska Hafs-Annulata. Ofversigt af Konglia Vetenskaps-Akademiens Forhhandlingar Stockholm, 22 (1), 51 - 110 [read 1865, printed 1866]." type="journal article" year="1866">Malmgren (1866)</bibRefCitation>
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: 71
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, pl. 9 fig. 7 [not 
<bibRefCitation id="EF9ED59EC207FFFDFC85FC32FC6511A0" author="Johnston, G." box="[778,961,380,406]" pageId="36" pageNumber="37" pagination="424 - 441" refId="ref38156" refString="Johnston, G. (1839) Miscellanea Zoologica. VI. On the British Aphroditaceae. Annals and Magazine of Natural History, 2 (12), 424 - 441." type="journal article" year="1839">Johnston, 1839</bibRefCitation>
].
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<emphasis id="B97B747DC207FFFDFF18FC87FEE211D4" box="[151,326,457,482]" italics="true" pageId="36" pageNumber="37">Type material.</emphasis>
<taxonomicName id="4C0FD3ECC207FFFDFEC0FC87FD0911D4" authorityName=": Hartmann-Schroder" authorityYear="1996" box="[335,685,457,483]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="36" pageNumber="37" phylum="Annelida" rank="species" species="fragilis" subGenus="Evarne">
<emphasis id="B97B747DC207FFFDFEC0FC87FE7C11D4" box="[335,472,457,482]" italics="true" pageId="36" pageNumber="37">Harmothoe</emphasis>
(
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) 
<emphasis id="B97B747DC207FFFDFDDAFC87FD0911D4" box="[597,685,457,482]" italics="true" pageId="36" pageNumber="37">fragilis</emphasis>
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: 
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, 
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, RV &quot;Albatross&quot; St. 4413, off 
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(
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, 152 fms
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.
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<emphasis id="B97B747DC207FFFDFF49FF59FE101206" box="[198,436,535,560]" italics="true" pageId="36" pageNumber="37">Additional material.</emphasis>
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: 1 spm., SMF 17324, &quot;Johan Ruud&quot; St. 734, 
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, N 
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, 
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i 
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</date>
, epibenthic sledge, 
<quantity id="4CF7058AC207FFFDFC2FFF73FC441261" box="[928,992,573,599]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="37" unit="m" value="60.0">
<elevation id="00224F5CC207FFFDFC2FFF73FC441261" box="[928,992,573,599]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="36" pageNumber="37" unit="m" value="60.0">60 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFBF3FF73FA211261" box="[1148,1413,573,599]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
1 spm. (af), SMF 17310, &quot;Johan Ruud&quot; St. 819, 
<geoCoordinate id="EE3BCEA8C207FFFDFD41FF2AFCFB1248" box="[718,863,612,638]" degrees="69" direction="north" minutes="32.66" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="69.544334">69°32.66’N</geoCoordinate>
, 
<geoCoordinate id="EE3BCEA8C207FFFDFCE3FF2AFC5E1248" box="[876,1018,612,638]" degrees="18" direction="east" minutes="06.69" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="18.1115">18°06.69’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFBA9FF2AFB2C1248" box="[1062,1160,612,638]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, Malangen Buvika, 
<date id="FFB18EAFC207FFFDFAF1FF2AFEE61292" pageId="36" pageNumber="37" value="2002-10-29">
<collectingDate id="EFF57747C207FFFDFAF1FF2AFEE61292" pageId="36" pageNumber="37" value="2002-10-29">29 October 2002</collectingDate>
</date>
, epibenthic sledge, 
<quantity id="4CF7058AC207FFFDFDB6FFC4FDDF1292" box="[569,635,650,676]" metricMagnitude="1" metricUnit="m" metricValue="3.9" pageId="36" pageNumber="37" unit="m" value="39.0">
<elevation id="00224F5CC207FFFDFDB6FFC4FDDF1292" box="[569,635,650,676]" metricMagnitude="1" metricUnit="m" metricValue="3.9" pageId="36" pageNumber="37" unit="m" value="39.0">39 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFC92FFC4FB891292" box="[797,1069,650,676]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
2 spms., SMF 17308–17309, &quot;Johan Ruud&quot; St. 737, 
<geoCoordinate id="EE3BCEA8C207FFFDFE32FFFFFDF012FD" box="[445,596,689,715]" degrees="69" direction="north" minutes="49.90" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="69.831665">69°49.90’N</geoCoordinate>
, 
<geoCoordinate id="EE3BCEA8C207FFFDFDECFFFFFD5312FD" box="[611,759,689,715]" degrees="17" direction="east" minutes="53.25" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="17.8875">17°53.25’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFCA9FFFFFC2812FD" box="[806,908,689,715]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, 
<locationDeviation id="9A02B14DC207FFFDFC15FFFFFB0B12FD" box="[922,1199,689,715]" location="Malangsgrunnen" pageId="36" pageNumber="37">S of Malangsgrunnen</locationDeviation>
, 
<date id="FFB18EAFC207FFFDFB31FFFFFA3312FD" box="[1214,1431,689,715]" pageId="36" pageNumber="37" value="2002-10-08">
<collectingDate id="EFF57747C207FFFDFB31FFFFFA3312FD" box="[1214,1431,689,715]" pageId="36" pageNumber="37" value="2002-10-08">08 October 2002</collectingDate>
</date>
, epibenthic sledge, 
<quantity id="4CF7058AC207FFFDFEF4FF96FE6F12C4" box="[379,459,728,754]" metricMagnitude="2" metricUnit="m" metricValue="1.39" pageId="36" pageNumber="37" unit="m" value="139.0">
<elevation id="00224F5CC207FFFDFEF4FF96FE6F12C4" box="[379,459,728,754]" metricMagnitude="2" metricUnit="m" metricValue="1.39" pageId="36" pageNumber="37" unit="m" value="139.0">139 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFDE5FF96FCD212C4" box="[618,886,728,754]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
3 spms., SMF 17312, &quot;Johan Ruud&quot; St. 297, 
<geoCoordinate id="EE3BCEA8C207FFFDFF18FFB0FF59132E" box="[151,253,766,792]" degrees="68" direction="north" minutes="22" orientation="latitude" pageId="36" pageNumber="37" precision="925" value="68.36667">68°22’N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C207FFFDFE8AFFB0FECF132E" box="[261,363,766,792]" degrees="15" direction="east" minutes="47" orientation="longitude" pageId="36" pageNumber="37" precision="925" value="15.783334">15°47’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFE19FFB0FE53132E" box="[406,503,766,792]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, Kanstadfjord Katnesholmen, 
<date id="FFB18EAFC207FFFDFCEDFFB0FBA0132E" box="[866,1028,766,792]" pageId="36" pageNumber="37" value="2004-05-06">
<collectingDate id="EFF57747C207FFFDFCEDFFB0FBA0132E" box="[866,1028,766,792]" pageId="36" pageNumber="37" value="2004-05-06">
06 May 
<quantity id="4CF7058AC207FFFDFC49FFB0FBA0132E" box="[966,1028,766,792]" metricMagnitude="3" metricUnit="m" metricValue="2.004132" pageId="36" pageNumber="37" unit="m" value="2004.132">2004</quantity>
</collectingDate>
</date>
, 132 m, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFB74FFB0FF511309" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
1 spm. (af, mf), SMF 17311, &quot;Johan Ruud&quot; St. 289, 
<geoCoordinate id="EE3BCEA8C207FFFDFCEEFE6BFC4F1309" box="[865,1003,805,831]" degrees="70" direction="north" minutes="13.87" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="70.23116">70°13.87’N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C207FFFDFC7DFE6BFBD81309" box="[1010,1148,805,831]" degrees="22" direction="east" minutes="31.93" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="22.532167">22°31.93’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFB23FE6BFAAF1309" box="[1196,1291,805,831]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, Stjernsund, outer Lokkarfjord, 
<date id="FFB18EAFC207FFFDFEF8FE02FDB91350" box="[375,541,844,870]" pageId="36" pageNumber="37" value="2005-04-12">
<collectingDate id="EFF57747C207FFFDFEF8FE02FDB91350" box="[375,541,844,870]" pageId="36" pageNumber="37" value="2005-04-12">12 April 2005</collectingDate>
</date>
, dive, 
<quantity id="4CF7058AC207FFFDFDE4FE02FD0C1350" box="[619,680,844,870]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">
<elevation id="00224F5CC207FFFDFDE4FE02FD0C1350" box="[619,680,844,870]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">20 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFCB0FE02FBE61350" box="[831,1090,844,870]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
3 spms., SMF 17314–17316, &quot;Johan Ruud&quot; St. 298, 
<geoCoordinate id="EE3BCEA8C207FFFDFE24FE3CFD9313BA" box="[427,567,882,908]" degrees="70" direction="north" minutes="58.73" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="70.978836">70°58.73’N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C207FFFDFDCFFE3CFD6F13BA" box="[576,715,882,908]" degrees="25" direction="east" minutes="17.78" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="25.296333">25°17.78’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFD79FE3CFCF313BA" box="[758,855,882,908]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, Magerøysundet Stikkelsvågnaeringen, 
<date id="FFB18EAFC207FFFDFAB8FE3CFF721385" pageId="36" pageNumber="37" value="2005-04-13">
<collectingDate id="EFF57747C207FFFDFAB8FE3CFF721385" pageId="36" pageNumber="37" value="2005-04-13">13 April 2005</collectingDate>
</date>
, dive, 
<quantity id="4CF7058AC207FFFDFEA6FED7FECE1385" box="[297,362,921,947]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">
<elevation id="00224F5CC207FFFDFEA6FED7FECE1385" box="[297,362,921,947]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">20 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFD85FED7FCB31385" box="[522,791,921,947]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
3 spms., SMF 17317–17319, &quot;Johan Ruud&quot; St. 299, 
<geoCoordinate id="EE3BCEA8C207FFFDFF18FE8EFE8413EC" box="[151,288,960,986]" degrees="70" direction="north" minutes="59.26" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="70.98766">70°59.26’N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C207FFFDFEA6FE8EFE1613EC" box="[297,434,960,986]" degrees="25" direction="east" minutes="19.99" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="25.333166">25°19.99’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFE53FE8EFD9813EC" box="[476,572,960,986]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, Magerøysundet Stikkelsvågnaeringen, 
<date id="FFB18EAFC207FFFDFB9FFE8EFB1C13EC" box="[1040,1208,960,986]" pageId="36" pageNumber="37" value="2005-04-13">
<collectingDate id="EFF57747C207FFFDFB9FFE8EFB1C13EC" box="[1040,1208,960,986]" pageId="36" pageNumber="37" value="2005-04-13">13 April 2005</collectingDate>
</date>
, dive, 
<quantity id="4CF7058AC207FFFDFA89FE8EFAE113EC" box="[1286,1349,960,986]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">
<elevation id="00224F5CC207FFFDFA89FE8EFAE113EC" box="[1286,1349,960,986]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">20 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFF5EFEA8FE7D1436" box="[209,473,998,1024]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
1 spm., SMF 17320, &quot;Johan Ruud&quot; St. 300, 
<geoCoordinate id="EE3BCEA8C207FFFDFC61FEA8FBDD1436" box="[1006,1145,998,1024]" degrees="71" direction="north" minutes="00.28" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="71.00467">71°00.28’N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C207FFFDFB0DFEA8FAA81436" box="[1154,1292,998,1024]" degrees="25" direction="east" minutes="12.49" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="25.208166">25°12.49’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFAB8FEA8FA331436" box="[1335,1431,998,1024]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, Vannfjordryggen, 
<date id="FFB18EAFC207FFFDFEE2F943FDB11411" box="[365,533,1037,1063]" pageId="36" pageNumber="37" value="2005-04-13">
<collectingDate id="EFF57747C207FFFDFEE2F943FDB11411" box="[365,533,1037,1063]" pageId="36" pageNumber="37" value="2005-04-13">13 April 2005</collectingDate>
</date>
, dive, 
<quantity id="4CF7058AC207FFFDFDEBF943FD061411" box="[612,674,1037,1063]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">
<elevation id="00224F5CC207FFFDFDEBF943FD061411" box="[612,674,1037,1063]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">20 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFCB4F943FB9B1411" box="[827,1087,1037,1063]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
2 spms., SMF 17321–17322, &quot;Johan Ruud&quot; St. 301, 
<geoCoordinate id="EE3BCEA8C207FFFDFE24F97AFD921478" box="[427,566,1076,1102]" degrees="71" direction="north" minutes="00.29" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="71.00484">71°00.29’N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C207FFFDFDB0F97AFD6F1478" box="[575,715,1076,1102]" degrees="25" direction="east" minutes="21.59" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="25.359833">25°21.59’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFD79F97AFCF31478" box="[758,855,1076,1102]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, Vannfjorden, 
<date id="FFB18EAFC207FFFDFB88F97AFB171478" box="[1031,1203,1076,1102]" pageId="36" pageNumber="37" value="2005-04-13">
<collectingDate id="EFF57747C207FFFDFB88F97AFB171478" box="[1031,1203,1076,1102]" pageId="36" pageNumber="37" value="2005-04-13">13 April 2005</collectingDate>
</date>
, dive, 
<quantity id="4CF7058AC207FFFDFA8CF97AFAE71478" box="[1283,1347,1076,1102]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">
<elevation id="00224F5CC207FFFDFA8CF97AFAE71478" box="[1283,1347,1076,1102]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">20 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFF5EF914FE7D1442" box="[209,473,1114,1140]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
1 spm., SMF 17323, &quot;Johan Ruud&quot; St. 309, 
<geoCoordinate id="EE3BCEA8C207FFFDFC61F914FBDE1442" box="[1006,1146,1114,1140]" degrees="70" direction="north" minutes="59.32" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="70.98867">70°59.32’N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C207FFFDFB0DF914FAA91442" box="[1154,1293,1114,1140]" degrees="25" direction="east" minutes="20.50" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="25.341667">25°20.50’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFAB8F914FA331442" box="[1335,1431,1114,1140]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, Stikkelsvågnaeringen, 
<date id="FFB18EAFC207FFFDFE23F9CFFDFE14AD" box="[428,602,1153,1179]" pageId="36" pageNumber="37" value="2005-04-14">
<collectingDate id="EFF57747C207FFFDFE23F9CFFDFE14AD" box="[428,602,1153,1179]" pageId="36" pageNumber="37" value="2005-04-14">14 April 2005</collectingDate>
</date>
, dive, 
<quantity id="4CF7058AC207FFFDFD22F9CFFD4914AD" box="[685,749,1153,1179]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">
<elevation id="00224F5CC207FFFDFD22F9CFFD4914AD" box="[685,749,1153,1179]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="36" pageNumber="37" unit="m" value="20.0">20 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFC03F9CFFB3D14AD" box="[908,1177,1153,1179]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz.</collectorName>
1 spm., SMF 17313, &quot;Johan Ruud&quot; St. 465, 
<geoCoordinate id="EE3BCEA8C207FFFDFE28F9E6FD9614F4" box="[423,562,1192,1218]" degrees="69" direction="north" minutes="34.92" orientation="latitude" pageId="36" pageNumber="37" precision="9" value="69.582">69°34.92’N</geoCoordinate>
<geoCoordinate id="EE3BCEA8C207FFFDFDB5F9E6FD6114F4" box="[570,709,1192,1218]" degrees="18" direction="east" minutes="55.92" orientation="longitude" pageId="36" pageNumber="37" precision="9" value="18.932">18°55.92’E</geoCoordinate>
, N 
<collectingCountry id="F318E8FFC207FFFDFD60F9E6FCEB14F4" box="[751,847,1192,1218]" name="Norway" pageId="36" pageNumber="37">Norway</collectingCountry>
, 
<collectingCounty id="62D1D0E3C207FFFDFCD3F9E6FBAA14F4" box="[860,1038,1192,1218]" pageId="36" pageNumber="37">Balsfjord Berg</collectingCounty>
, 
<date id="FFB18EAFC207FFFDFB94F9E6FB1814F4" box="[1051,1212,1192,1218]" pageId="36" pageNumber="37" value="2006-05-08">
<collectingDate id="EFF57747C207FFFDFB94F9E6FB1814F4" box="[1051,1212,1192,1218]" pageId="36" pageNumber="37" value="2006-05-08">08 May 2006</collectingDate>
</date>
, triangular dredge, 
<quantity id="4CF7058AC207FFFDFF18F980FF7014DE" box="[151,212,1230,1256]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="36" pageNumber="37" unit="m" value="29.0">
<elevation id="00224F5CC207FFFDFF18F980FF7014DE" box="[151,212,1230,1256]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="36" pageNumber="37" unit="m" value="29.0">29 m</elevation>
</quantity>
, leg. &amp; ded. 
<collectorName id="26FACDB9C207FFFDFEE5F980FDCD14DE" box="[362,617,1230,1256]" pageId="36" pageNumber="37">C. d’Udekem d’Acoz</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB0A86FC207FFFDFF4AF9BBFCD01539" blockId="36.[151,1436,457,2030]" box="[197,884,1269,1295]" pageId="36" pageNumber="37">
(For further material see 
<bibRefCitation id="EF9ED59EC207FFFDFE66F9BBFD4B1539" author="Barnich, R. &amp; Fiege, D." box="[489,751,1269,1295]" pageId="36" pageNumber="37" pagination="1889 - 1938" refId="ref36372" refString="Barnich, R. &amp; Fiege, D. (2000) Revision of the Mediterranean species of Harmothoe Kinberg, 1856 and Lagisca Malmgren, 1865 (Polychaeta, Polynoidae, Polynoinae) with descriptions of a new genus and a new species. Journal of Natural History, 34, 1889 - 1938." type="journal article" year="2000">Barnich &amp; Fiege 2000</bibRefCitation>
and 
<bibRefCitation id="EF9ED59EC207FFFDFCA7F9BBFCC31539" author="Barnich, R. &amp; Fiege, D." box="[808,871,1269,1295]" pageId="36" pageNumber="37" pagination="1 - 167" refId="ref36428" refString="Barnich, R. &amp; Fiege, D. (2003) The Aphroditoidea (Annelida, Polychaeta) of the Mediterranean Sea. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 559, 1 - 167." type="journal article" year="2003">2003</bibRefCitation>
).
</paragraph>
<paragraph id="8BB0A86FC207FFFDFF4AF852FB5815B5" blockId="36.[151,1436,457,2030]" pageId="36" pageNumber="37">
<emphasis id="B97B747DC207FFFDFF4AF852FE991503" box="[197,317,1308,1333]" italics="true" pageId="36" pageNumber="37">Diagnosis</emphasis>
. Anterior pair of eyes dorsolateral at widest part of prostomium. Elytral margin with fringing papillae; surface covered by conical microtubercles and few scattered papillae, with a row of rounded, soft and flattened macrotubercles near posterior margin, giving margin a scallop-shaped appearence.
</paragraph>
<paragraph id="8BB0A86FC207FFFDFF4AF8DEFA97159C" blockId="36.[151,1436,457,2030]" box="[197,1331,1424,1450]" pageId="36" pageNumber="37">
<emphasis id="B97B747DC207FFFDFF4AF8DEFEEB159F" box="[197,335,1424,1449]" italics="true" pageId="36" pageNumber="37">Description</emphasis>
(based on additional specimen SMF 17309, since 
<typeStatus id="54B416CDC207FFFDFC10F8DEFBA2159C" box="[927,1030,1424,1450]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
in rather poor condition).
</paragraph>
<paragraph id="8BB0A86FC207FFFDFF4AF8F8FBE91673" blockId="36.[151,1436,457,2030]" pageId="36" pageNumber="37">
Body with 39 segments. At anterior end (
<figureCitation id="1334B4EAC207FFFDFD35F8F8FC8E15E6" box="[698,810,1462,1488]" captionStart="FIGURE 17" captionStartId="37.[151,255,1869,1893]" captionTargetBox="[273,1373,341,1841]" captionTargetId="figure-181@37.[151,1436,329,1852]" captionTargetPageId="37" captionText="FIGURE 17. Harmothoe fragilis (additional specimen, SMF 17309). (A) anterior end; styles of tentacular cirri and left dorsal cirrus of segment 3 missing; (B) right elytron from segment 5; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus missing; (E) long notochaeta; (F) tip of same; (G) short notochaeta; (H) middle neurochaeta; (I) tip of same; (J) tip of uppermost neurochaeta; (K) tip of lower neurochaeta. (Scale: A, B = 2 mm; C, E, G, H = 250 µm; D = 1 mm; F, I–K = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361475" httpUri="https://zenodo.org/record/5361475/files/figure.png" pageId="36" pageNumber="37">Fig. 17A</figureCitation>
), prostomium bilobed, with distinct cephalic peaks; ceratophore of median antenna in anterior notch, lateral antennae inserted ventrally, styles of antennae papillate, tapering to filiform tip; anterior pair of eyes situated dorsolaterally at widest part of prostomium, posterior pair dorsally near hind margin of prostomium; palps papillate, tapering.
</paragraph>
<paragraph id="8BB0A86FC207FFFDFF4AFB1FFB5B168E" blockId="36.[151,1436,457,2030]" pageId="36" pageNumber="37">Tentaculophores inserted laterally to prostomium, each with two notochaetae and a dorsal and ventral tentacular cirrus, styles of cirri papillate, tapering to filiform tip. Second segment with first pair of elytra, biramous parapodia, and long buccal cirri. Following segments with tapering, short ventral cirri.</paragraph>
<paragraph id="8BB0A86FC207FFFDFF4AFB8BFB4B174C" blockId="36.[151,1436,457,2030]" pageId="36" pageNumber="37">
Fifteen pairs of elytra, covering dorsum, on segments 2, 4, 5, 7, then on every second segment to 23, 26, 29, 32; last seven segments cirrigerous; elytral margin with fringing papillae; surface covered by conical microtubercles and few scattered papillae, with a row of rounded, soft and flattened macrotubercles near posterior margin, giving margin a scallop-shaped appearence (
<figureCitation id="1334B4EAC207FFFDFCE0FA77FC511765" box="[879,1013,1849,1875]" captionStart="FIGURE 17" captionStartId="37.[151,255,1869,1893]" captionTargetBox="[273,1373,341,1841]" captionTargetId="figure-181@37.[151,1436,329,1852]" captionTargetPageId="37" captionText="FIGURE 17. Harmothoe fragilis (additional specimen, SMF 17309). (A) anterior end; styles of tentacular cirri and left dorsal cirrus of segment 3 missing; (B) right elytron from segment 5; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus missing; (E) long notochaeta; (F) tip of same; (G) short notochaeta; (H) middle neurochaeta; (I) tip of same; (J) tip of uppermost neurochaeta; (K) tip of lower neurochaeta. (Scale: A, B = 2 mm; C, E, G, H = 250 µm; D = 1 mm; F, I–K = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361475" httpUri="https://zenodo.org/record/5361475/files/figure.png" pageId="36" pageNumber="37">Fig. 17B,C</figureCitation>
). Cirrigerous segments with distinct dorsal tubercles; dorsal cirri with cylindrical cirrophore, style papillate, tapering to filiform tip.
</paragraph>
<paragraph id="8BB0A86FC207FFFCFF4AFAC8FA2F10EE" blockId="36.[151,1436,457,2030]" lastBlockId="37.[151,1435,152,294]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
Parapodia biramous; notopodia with elongate acicular lobe; neuropodia with elongate prechaetal acicular lobe with digitiform supra-acicular process; neuropodial postchaetal lobe shorter than prechaetal lobe, rounded; tips of noto- and neuroacicula penetrating epidermis (
<figureCitation id="1334B4EAC207FFFDFC44FA9AFBE717D8" box="[971,1091,2004,2030]" captionStart="FIGURE 17" captionStartId="37.[151,255,1869,1893]" captionTargetBox="[273,1373,341,1841]" captionTargetId="figure-181@37.[151,1436,329,1852]" captionTargetPageId="37" captionText="FIGURE 17. Harmothoe fragilis (additional specimen, SMF 17309). (A) anterior end; styles of tentacular cirri and left dorsal cirrus of segment 3 missing; (B) right elytron from segment 5; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus missing; (E) long notochaeta; (F) tip of same; (G) short notochaeta; (H) middle neurochaeta; (I) tip of same; (J) tip of uppermost neurochaeta; (K) tip of lower neurochaeta. (Scale: A, B = 2 mm; C, E, G, H = 250 µm; D = 1 mm; F, I–K = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361475" httpUri="https://zenodo.org/record/5361475/files/figure.png" pageId="36" pageNumber="37">Fig. 17D</figureCitation>
). Notochaetae stouter than neurochaetae, with distinct rows of spines and blunt tip (
<figureCitation id="1334B4EAC206FFFCFCC2FDD6FC441084" box="[845,992,152,178]" captionStart="FIGURE 17" captionStartId="37.[151,255,1869,1893]" captionTargetBox="[273,1373,341,1841]" captionTargetId="figure-181@37.[151,1436,329,1852]" captionTargetPageId="37" captionText="FIGURE 17. Harmothoe fragilis (additional specimen, SMF 17309). (A) anterior end; styles of tentacular cirri and left dorsal cirrus of segment 3 missing; (B) right elytron from segment 5; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus missing; (E) long notochaeta; (F) tip of same; (G) short notochaeta; (H) middle neurochaeta; (I) tip of same; (J) tip of uppermost neurochaeta; (K) tip of lower neurochaeta. (Scale: A, B = 2 mm; C, E, G, H = 250 µm; D = 1 mm; F, I–K = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361475" httpUri="https://zenodo.org/record/5361475/files/figure.png" pageId="37" pageNumber="38">Fig. 17E–G</figureCitation>
); neurochaetae with distinct rows of spines, mostly bidentate with slender secondary tooth (often abraded), some lower unidentate (
<figureCitation id="1334B4EAC206FFFCFB62FDF0FADA10EE" box="[1261,1406,190,216]" captionStart="FIGURE 17" captionStartId="37.[151,255,1869,1893]" captionTargetBox="[273,1373,341,1841]" captionTargetId="figure-181@37.[151,1436,329,1852]" captionTargetPageId="37" captionText="FIGURE 17. Harmothoe fragilis (additional specimen, SMF 17309). (A) anterior end; styles of tentacular cirri and left dorsal cirrus of segment 3 missing; (B) right elytron from segment 5; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus missing; (E) long notochaeta; (F) tip of same; (G) short notochaeta; (H) middle neurochaeta; (I) tip of same; (J) tip of uppermost neurochaeta; (K) tip of lower neurochaeta. (Scale: A, B = 2 mm; C, E, G, H = 250 µm; D = 1 mm; F, I–K = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361475" httpUri="https://zenodo.org/record/5361475/files/figure.png" pageId="37" pageNumber="38">Fig. 17H–K</figureCitation>
).
</paragraph>
<paragraph id="8BB0A86FC206FFFCFF4AFDABFC421110" blockId="37.[151,1435,152,294]" pageId="37" pageNumber="38">
<emphasis id="B97B747DC206FFFCFF4AFDABFED210C8" box="[197,374,229,254]" italics="true" pageId="37" pageNumber="38">Measurements</emphasis>
. 
<taxonomicName id="4C0FD3ECC206FFFCFE07FDABFDA310C8" authorityName=": Hartmann-Schroder" authorityYear="1996" box="[392,519,229,254]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="fragilis">
<emphasis id="B97B747DC206FFFCFE07FDABFDA310C8" box="[392,519,229,254]" italics="true" pageId="37" pageNumber="38">H. fragilis</emphasis>
</taxonomicName>
, 
<typeStatus id="54B416CDC206FFFCFD96FDABFD2310C9" box="[537,647,229,255]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
, 
<collectionCode id="ED1E30AAC206FFFCFD19FDABFD4910C9" box="[662,749,229,255]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="37" pageNumber="38" type="Museum">USNM</collectionCode>
17148: L 
<quantity id="4CF7058AC206FFFCFCFFFDABFC6D10C9" box="[880,969,229,255]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="37" pageNumber="38" unit="mm" value="19.0">19 mm</quantity>
, W 
<quantity id="4CF7058AC206FFFCFC73FDABFBE610C9" box="[1020,1090,229,255]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="37" pageNumber="38" unit="mm" value="6.0">6 mm</quantity>
for 38 segments. Specimen figured, SMF 17309 (
<figureCitation id="1334B4EAC206FFFCFE1AFC42FD831110" box="[405,551,268,294]" captionStart="FIGURE 17" captionStartId="37.[151,255,1869,1893]" captionTargetBox="[273,1373,341,1841]" captionTargetId="figure-181@37.[151,1436,329,1852]" captionTargetPageId="37" captionText="FIGURE 17. Harmothoe fragilis (additional specimen, SMF 17309). (A) anterior end; styles of tentacular cirri and left dorsal cirrus of segment 3 missing; (B) right elytron from segment 5; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus missing; (E) long notochaeta; (F) tip of same; (G) short notochaeta; (H) middle neurochaeta; (I) tip of same; (J) tip of uppermost neurochaeta; (K) tip of lower neurochaeta. (Scale: A, B = 2 mm; C, E, G, H = 250 µm; D = 1 mm; F, I–K = 50 µm)." figureDoi="http://doi.org/10.5281/zenodo.5361475" httpUri="https://zenodo.org/record/5361475/files/figure.png" pageId="37" pageNumber="38">Fig. 17A–K</figureCitation>
): L 
<quantity id="4CF7058AC206FFFCFDDAFC42FD0E1110" box="[597,682,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="38" unit="mm" value="20.0">20 mm</quantity>
, W 
<quantity id="4CF7058AC206FFFCFD56FC42FCB81110" box="[729,796,268,294]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="37" pageNumber="38" unit="mm" value="5.0">5 mm</quantity>
for 39 segments.
</paragraph>
<caption id="DF70F8E7C206FFFCFF18FA03FB8E17C6" ID-DOI="http://doi.org/10.5281/zenodo.5361475" ID-Zenodo-Dep="5361475" httpUri="https://zenodo.org/record/5361475/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,255,1869,1893]" targetBox="[273,1373,341,1841]" targetPageId="37">
<paragraph id="8BB0A86FC206FFFCFF18FA03FB8E17C6" blockId="37.[151,1436,1869,2032]" pageId="37" pageNumber="38">
<emphasis id="B97B747DC206FFFCFF18FA03FE831753" bold="true" box="[151,295,1869,1893]" pageId="37" pageNumber="38">FIGURE 17.</emphasis>
<taxonomicName id="4C0FD3ECC206FFFCFEA1FA00FE581753" authorityName=": Hartmann-Schroder" authorityYear="1996" box="[302,508,1870,1893]" class="Polychaeta" family="Polynoidae" genus="Harmothoe" kingdom="Animalia" order="Phyllodocida" pageId="37" pageNumber="38" phylum="Annelida" rank="species" species="fragilis">
<emphasis id="B97B747DC206FFFCFEA1FA00FE581753" box="[302,508,1870,1893]" italics="true" pageId="37" pageNumber="38">Harmothoe fragilis</emphasis>
</taxonomicName>
(additional specimen, SMF 17309). (A) anterior end; styles of tentacular cirri and left dorsal cirrus of segment 3 missing; (B) right elytron from segment 5; (C) detail of posterior margin of same; (D) right cirrigerous parapodium from segment 14, posterior view, style of dorsal cirrus missing; (E) long notochaeta; (F) tip of same; (G) short notochaeta; (H) middle neurochaeta; (I) tip of same; (J) tip of uppermost neurochaeta; (K) tip of lower neurochaeta. (Scale: A, B = 2 mm; C, E, G, H = 250 µm; D = 1 mm; F, I–K = 50 µm).
</paragraph>
</caption>
<paragraph id="8BB0A86FC205FFFFFF4AFDD6FCD010EE" blockId="38.[197,1422,152,216]" pageId="38" pageNumber="39">
<emphasis id="B97B747DC205FFFFFF4AFDD6FEF01087" box="[197,340,152,177]" italics="true" pageId="38" pageNumber="39">Distribution</emphasis>
. Arctic, North Pacific, North Atlantic, Skagerrak, Kattegat, and western Mediterranean Sea. 
<emphasis id="B97B747DC205FFFFFF49FDF1FE8410EE" box="[198,288,191,216]" italics="true" pageId="38" pageNumber="39">Habitat</emphasis>
. On muddy and sandy substrata from 
<quantity id="4CF7058AC205FFFFFD50FDF0FCCB10EE" box="[735,879,190,216]" metricMagnitude="2" metricUnit="m" metricValue="1.6" metricValueMax="3.0" metricValueMin="0.2" pageId="38" pageNumber="39" unit="m" value="160.0" valueMax="300.0" valueMin="20.0">
20 to 
<specimenCount id="9D0963E6C205FFFFFCADFDF0FCCB10EE" box="[802,879,190,216]" pageId="38" pageNumber="39" type="generic">300 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>