360 lines
47 KiB
XML
360 lines
47 KiB
XML
<document id="F9386D7C5E8A62A16C6F282B598D0915" ID-DOI="10.1111/j.1096-3642.2005.00189.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5435260" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630434122956" checkinUser="diego" docAuthor="Daugeron, Christophe & Grootaert, Patrick" docDate="2005" docId="039D7838FFC5FFD03C53D835FC71FB49" docLanguage="en" docName="j.1096-3642.2005.00189.x.pdf" docOrigin="Zoological Journal of the Linnean Society 145 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2005.00189.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Empis (Coptophlebia) jacobsoni De Meijere 1907" docType="treatment" docVersion="4" lastPageNumber="370" masterDocId="FFA40040FFD8FFCF3F28DF3EFFC4FFD9" masterDocTitle="Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae)" masterLastPageNumber="391" masterPageNumber="339" pageNumber="368" updateTime="1699269823662" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="390AC2700DC3A29EFC8DDF1527EC1E22">
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<mods:title id="A1B71AE064FF2D82613204BC3963988B">Phylogenetic systematics of the Empis (Coptophlebia) hyalea-group (Insecta: Diptera: Empididae)</mods:title>
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<mods:namePart id="0F6D4C81B33E3BB535B6886D3989760F">Daugeron, Christophe</mods:namePart>
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<mods:namePart id="7BE5086DFE7403BD5E44C7361BAABA23">Grootaert, Patrick</mods:namePart>
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<mods:typeOfResource id="CD2A1E650DB2D70CEFFA04F7C1324079">text</mods:typeOfResource>
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<mods:title id="72734F2F32222BF28FA6BB48886D6595">Zoological Journal of the Linnean Society</mods:title>
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<mods:part id="AEAC235B09B69E3A3D0F49DDB0BD8A41">
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<mods:date id="7101873A66D22EF848CCFC47B64C57A6">2005</mods:date>
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<mods:number id="7F1216037DEC1CE278CD957DD98BDD7F">2005-11-30</mods:number>
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<mods:number id="B9C790186169FF6319B6C792FB826A6A">145</mods:number>
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<mods:url id="3022CAF8816B1CDC9A280C2618E187CC">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2005.00189.x</mods:url>
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<mods:identifier id="F0B4D96EDE1991286A5E3FD8F733D803" type="DOI">10.1111/j.1096-3642.2005.00189.x</mods:identifier>
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<mods:identifier id="BD17CD85E3F11A54CB86791B193BEC4D" type="ISSN">0024-4082</mods:identifier>
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<treatment id="039D7838FFC5FFD03C53D835FC71FB49" ID-DOI="http://doi.org/10.5281/zenodo.5490724" ID-Zenodo-Dep="5490724" LSID="urn:lsid:plazi:treatment:039D7838FFC5FFD03C53D835FC71FB49" httpUri="http://treatment.plazi.org/id/039D7838FFC5FFD03C53D835FC71FB49" lastPageId="31" lastPageNumber="370" pageId="29" pageNumber="368">
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<subSubSection id="C32E9AA5FFC5FFD13C53D835FE43FF21" lastPageId="30" lastPageNumber="369" pageId="29" pageNumber="368" type="nomenclature">
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<paragraph id="8B8BC92EFFC5FFD23C53D835FAA5F8F9" blockId="29.[891,1377,1802,1826]" box="[891,1377,1802,1826]" pageId="29" pageNumber="368">
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<heading id="D0C37E42FFC5FFD23C53D835FAA5F8F9" box="[891,1377,1802,1826]" centered="true" fontSize="9" level="2" pageId="29" pageNumber="368" reason="2">
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<taxonomicName id="4C34B2ADFFC5FFD23C53D835FAD4F8F9" ID-CoL="8BZ5H" authorityName="De Meijere" authorityYear="1907" box="[891,1296,1802,1826]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="380" phylum="Arthropoda" rank="species" species="jacobsoni" subGenus="Coptophlebia">
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<emphasis id="B940153CFFC5FFD23C53D835FC00F8F9" box="[891,964,1803,1826]" italics="true" pageId="29" pageNumber="368">EMPIS</emphasis>
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(
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<emphasis id="B940153CFFC5FFD23CFDD834FB45F8F9" box="[981,1153,1802,1826]" italics="true" pageId="29" pageNumber="368">COPTOPHLEBIA</emphasis>
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)
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<emphasis id="B940153CFFC5FFD23BBAD833FAD4F8F9" box="[1170,1296,1805,1824]" italics="true" pageId="29" pageNumber="368">JACOBSONI</emphasis>
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</taxonomicName>
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CLADE
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</heading>
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</paragraph>
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<paragraph id="8B8BC92EFFC5FFD13C12D805FE43FF21" blockId="29.[826,1441,1851,1903]" lastBlockId="30.[142,756,195,248]" lastPageId="30" lastPageNumber="369" pageId="29" pageNumber="368">
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<emphasis id="B940153CFFC5FFD23C12D805FC7AF889" box="[826,958,1851,1872]" italics="true" pageId="29" pageNumber="368">Monophyly:</emphasis>
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The monophyly of this clade is supported by one character: male fore tibia with a cluster or row of strong to spine-like anterodorsal bristles on basal third [character 6(1)].
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</paragraph>
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</subSubSection>
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<subSubSection id="C32E9AA5FFC6FFD03FA6DE2EFB15FDF2" lastPageId="31" lastPageNumber="370" pageId="30" pageNumber="369" type="distribution">
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<paragraph id="8B8BC92EFFC6FFD13FA6DE2EFD73FE66" blockId="30.[142,758,272,447]" pageId="30" pageNumber="369">
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<emphasis id="B940153CFFC6FFD13FA6DE2EFE4CFEFC" box="[142,392,272,293]" italics="true" pageId="30" pageNumber="369">
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Distribution (
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<figureCitation id="130FD5ABFFC6FFD13E02DE2EFEBFFEFC" box="[298,379,272,293]" captionStart="Figure 35" captionStartId="41.[162,240,1766,1785]" captionTargetBox="[333,1270,1448,1733]" captionTargetId="figure-336@41.[323,1283,1435,1746]" captionTargetPageId="41" captionText="Figure 35. Distribution of the Empis (Coptophlebia) jacobsoni clade: E. (C.) jacobsoni De Meijere = 6 (Java); E. (C.) lachaisei sp. nov. = 1 (Ivory Coast); E. (C.) loici sp. nov. = 2 (Central African Republic); E. (C.) menglunensis sp. nov. = 3 (China); E. (C.) mengyangensis sp. nov. = 3 (China); E. (C.) cameronsensis sp. nov. = 4 (Malaysia); E. (C.) nimbaensis sp. nov. = 1 (Guinea); E. (C.) nondouensis Daugeron = 7 (New Caledonia); E. (C.) padangensis sp. nov. = 5 (Sumatra)." figureDoi="http://doi.org/10.5281/zenodo.5435336" httpUri="https://zenodo.org/record/5435336/files/figure.png" pageId="30" pageNumber="369">Fig. 35</figureCitation>
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):
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</emphasis>
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This group of species is widely distributed including tropical Africa (
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<collectingCountry id="F32389BEFFC6FFD13D14DE11FD56FE9C" box="[572,658,303,325]" name="Guinea-Bissau" pageId="30" pageNumber="369">Guinea</collectingCountry>
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,
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<collectingCountry id="F32389BEFFC6FFD13DB6DE11FE97FEBA" name="Central African Republic" pageId="30" pageNumber="369">Central African Republic</collectingCountry>
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and
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<collectingCountry id="F32389BEFFC6FFD13EBFDE70FDE7FEBA" box="[407,547,333,355]" name="Ivory Coast" pageId="30" pageNumber="369">Ivory Coast</collectingCountry>
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), South-east Asia (
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<collectingCountry id="F32389BEFFC6FFD13FBEDE52FF1AFE5B" box="[150,222,364,386]" name="China" pageId="30" pageNumber="369">China</collectingCountry>
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:
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<collectingRegion id="49F007CCFFC6FFD13FC1DE52FE80FE58" box="[233,324,364,385]" country="China" name="Yunnan" pageId="30" pageNumber="369">Yunnan</collectingRegion>
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,
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<collectingCountry id="F32389BEFFC6FFD13E78DE52FE7FFE5B" box="[336,443,364,386]" name="Malaysia" pageId="30" pageNumber="369">Malaysia</collectingCountry>
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,
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<collectingCountry id="F32389BEFFC6FFD13EEFDE52FDFEFE5B" box="[455,570,364,386]" name="Indonesia" pageId="30" pageNumber="369">Indonesia</collectingCountry>
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:
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<collectingRegion id="49F007CCFFC6FFD13D6DDE52FDB8FE58" box="[581,636,364,385]" country="Indonesia" name="Jawa Timur" pageId="30" pageNumber="369">Java</collectingRegion>
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,
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<collectingRegion id="49F007CCFFC6FFD13DA1DE52FD2AFE5B" box="[649,750,364,386]" country="Indonesia" name="Sumatera Utara" pageId="30" pageNumber="369">Sumatra</collectingRegion>
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) and
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<collectingCountry id="F32389BEFFC6FFD13FECDEB5FEBDFE78" box="[196,377,395,417]" name="New Caledonia" pageId="30" pageNumber="369">New Caledonia</collectingCountry>
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. This distribution suggests that this group should also be found in New
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<collectingCountry id="F32389BEFFC6FFD13D74DE97FD77FE66" box="[604,691,425,447]" name="Guinea-Bissau" pageId="30" pageNumber="369">Guinea</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B8BC92EFFC6FFD13FB5DD3FFD22FDC0" blockId="30.[157,742,513,569]" box="[157,742,513,537]" pageId="30" pageNumber="369">
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<taxonomicName id="4C34B2ADFFC6FFD13FB5DD3FFE7DFDC0" authority="" box="[157,441,513,537]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="387" phylum="Arthropoda" rank="subGenus" subGenus="Coptophlebia">
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<emphasis id="B940153CFFC6FFD13FB5DD3FFF2DFDCE" bold="true" box="[157,233,513,537]" italics="true" pageId="30" pageNumber="369">EMPIS</emphasis>
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(
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<emphasis id="B940153CFFC6FFD13FD2DD3FFE75FDCE" bold="true" box="[250,433,513,537]" italics="true" pageId="30" pageNumber="369">COPTOPHLEBIA</emphasis>
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)
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</taxonomicName>
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<taxonomicName id="4C34B2ADFFC6FFD13EEADD3AFDBEFDCE" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[450,634,516,535]" pageId="30" pageNumber="369" rank="species" species="cameronensis" status="sp. nov.">CAMERONENSIS</taxonomicName>
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<emphasis id="B940153CFFC6FFD13DAADD3AFD22FDC0" bold="true" box="[642,742,513,537]" pageId="30" pageNumber="369">
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<taxonomicNameLabel id="A273A847FFC6FFD13DAADD3AFD22FDC0" box="[642,742,513,537]" pageId="30" pageNumber="369" rank="species">SP. NOV.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="8B8BC92EFFC6FFD13EB8DD1FFE30FDE0" blockId="30.[157,742,513,569]" box="[400,500,545,569]" pageId="30" pageNumber="369">
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(
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<figureCitation id="130FD5ABFFC6FFD13EB0DD1FFE28FDE0" box="[408,492,545,569]" captionStart="Figure 24" captionStartId="32.[142,220,883,902]" captionTargetBox="[306,1257,192,845]" captionTargetId="figure-497@32.[300,1260,183,851]" captionTargetPageId="32" captionText="Figure 24. The Empis (Coptophlebia) jacobsoni clade, E. (C.) cameronensis sp. nov. male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435314" httpUri="https://zenodo.org/record/5435314/files/figure.png" pageId="30" pageNumber="369">FIG. 24</figureCitation>
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)
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</paragraph>
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<paragraph id="8B8BC92EFFC6FFD13FA6DD6CFDAEFD7C" blockId="30.[142,758,593,677]" pageId="30" pageNumber="369">
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<emphasis id="B940153CFFC6FFD13FA6DD6CFEF7FDBF" box="[142,307,593,615]" italics="true" pageId="30" pageNumber="369">Type material:</emphasis>
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<materialsCitation id="3B5CC373FFC6FFD13E14DD6FFDB8FD5F" collectingDate="1985-04-07" collectionCode="AMS" collectorName="D. K. McAlpine" country="Malaysia" location="Cameron Highlands" pageId="30" pageNumber="369" specimenCount="1" typeStatus="holotype">
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<typeStatus id="548F778CFFC6FFD13E14DD6FFE61FDBE" box="[316,421,593,615]" pageId="30" pageNumber="369" type="holotype">Holotype</typeStatus>
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: male,
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<collectingCountry id="F32389BEFFC6FFD13D24DD6FFDB3FDBE" box="[524,631,593,615]" name="Malaysia" pageId="30" pageNumber="369">Malaysia</collectingCountry>
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,
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<location id="8EEB9FF5FFC6FFD13DA5DD6FFEC2FD5F" LSID="urn:lsid:plazi:treatment:039D7838FFC5FFD03C53D835FC71FB49:8EEB9FF5FFC6FFD13DA5DD6FFEC2FD5F" country="Malaysia" name="Cameron Highlands" pageId="30" pageNumber="369">Cameron Highlands</location>
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,
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<date id="FF8AEFEEFFC6FFD13E3BDD4EFEBDFD5F" box="[275,377,624,646]" pageId="30" pageNumber="369" value="1985-04-07">
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<collectingDate id="EFCE1606FFC6FFD13E3BDD4EFEBDFD5F" box="[275,377,624,646]" pageId="30" pageNumber="369" value="1985-04-07">7.iv.1985</collectingDate>
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</date>
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,
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<collectorName id="26C1ACF8FFC6FFD13EAEDD4EFDE3FD5F" box="[390,551,624,646]" pageId="30" pageNumber="369">D.K.McAlpine</collectorName>
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(
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<collectionCode id="ED2551EBFFC6FFD13D1FDD4EFDB0FD5F" box="[567,628,624,646]" pageId="30" pageNumber="369">AMS</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B5CC373FFC6FFD13DAFDD4EFDA2FD7C" collectingDate="1985-04-05" collectionCode="AMS" collectorName="D. K. McAlpine" country="Malaysia" location="Cameron Highlands" pageId="30" pageNumber="369" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<typeStatus id="548F778CFFC6FFD13DAFDD4EFD36FD5C" box="[647,754,624,645]" pageId="30" pageNumber="369" type="paratype">Paratype</typeStatus>
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||
:
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<specimenCount id="9D3202A7FFC6FFD13FA6DDB1FF36FD7C" box="[142,242,655,677]" pageId="30" pageNumber="369" type="female">1 female</specimenCount>
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, same data but
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<date id="FF8AEFEEFFC6FFD13E84DDB1FDD5FD7C" box="[428,529,655,677]" pageId="30" pageNumber="369" value="1985-04-05">
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<collectingDate id="EFCE1606FFC6FFD13E84DDB1FDD5FD7C" box="[428,529,655,677]" pageId="30" pageNumber="369" value="1985-04-05">5.iv.1985</collectingDate>
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</date>
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||
(
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<collectionCode id="ED2551EBFFC6FFD13D09DDB1FD9AFD7C" box="[545,606,655,677]" pageId="30" pageNumber="369">AMS</collectionCode>
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||
)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8B8BC92EFFC6FFD13FA6DD83FE89FD28" blockId="30.[142,756,701,753]" pageId="30" pageNumber="369">
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<emphasis id="B940153CFFC6FFD13FA6DD83FEC8FD0B" box="[142,268,701,722]" italics="true" pageId="30" pageNumber="369">Etymology:</emphasis>
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The name of the species is derived from the type-locality.
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</paragraph>
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<paragraph id="8B8BC92EFFC6FFD03FA6DC34FB15FDF2" blockId="30.[142,758,777,922]" lastBlockId="31.[826,1443,195,1168]" lastPageId="31" lastPageNumber="370" pageId="30" pageNumber="369">
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||
<emphasis id="B940153CFFC6FFD13FA6DC34FED2FCC6" box="[142,278,778,799]" italics="true" pageId="30" pageNumber="369">Description:</emphasis>
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Blackish-grey species of middle size. Fore tibia deformed with an anterodorsal row of
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<emphasis id="B940153CFFC6FFD13DABDC17FD50FCE7" box="[643,660,809,830]" italics="true" pageId="30" pageNumber="369">c.</emphasis>
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8 spinelike bristles on basal half. First mid tarsomere whitish. Sternite 7 with strong, long posterior bristles. MALE: HEAD. Occiput blackish to dusty grey. Bristles of ocellar triangle missing. Face black. Palpus black with 1 distinct bristle. Antenna black, first flagellomere conical, second and third flagellomeres missing. Labrum dark brown, length
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<emphasis id="B940153CFFC6FFD13BBBDE1EFB60FEEC" box="[1171,1188,288,309]" italics="true" pageId="30" pageNumber="369">c.</emphasis>
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1.5× head height. Labium black with numerous short bristles. Labella broken. THORAX blackish, scutum dusty grey to subshiny. All bristles black. Antepronotum with 2 short lateral bristles. Proepisternum and prosternum with 1 and 2 short bristles, respectively. Acrostichals missing. Dorsocentrals uniserial, ending with 2 strong, long bristles in prescutellar depression. Anterior and posterior spiracles blackish. LEGS dark brown-blackish. Fore tibia deformed at base, somewhat S-shaped in frontal view, with anterodorsal row of
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<emphasis id="B940153CFFC6FFD13BC4DD6DFB39FDB1" box="[1260,1277,595,616]" italics="true" pageId="30" pageNumber="369">c.</emphasis>
|
||
8 spine-like bristles on basal half; first fore tarsomere with 1 strong, long dorsal basally, 2 strong, long anterolaterals basally, several others strong, shorter anterolaterals apically; other tarsomeres with apical circlet of strong bristles. Mid femur with strong, long ventral bristles basally; mid tibia with dorsal row of 3 strong, long bristles (at basal third, at apical quarter, apically), several strong, long posteroventrals and ventrals except basally; first mid tarsomere whitish with 1 strong, long ventral at base and apical circlet of strong bristles especially 1 long posterodorsal; other tarsomeres with apical circlet of strong bristles. Hind femur with dorsal and ventral rows of short bristles; hind tibia with dorsal row of 5 strong, long bristles, 1 strong, long anterodorsal apically, ventrally covered with numerous bristles longer apically; hind tarsus somewhat swollen; first hind tarsomere with 6 strong, long dorsal bristles and antero- and posteroventral rows of strong spine-like bristles; other tarsomeres with dorsal pair of long bristles apically. WING (length = 3.25) clear with veins brownish except M1 almost not visible. A1 abbreviated. Halter black. ABDOMEN brown-blackish with first two tergites somewhat dusty, covered with distinct bristles, tergite 8 more strongly sclerotized anterolaterally. Strong, long bristles at base, at the posterior margin of sternites and tergites, sternite 7 with strong, long posteriors, sternite 8 with numerous strong, long posterior bristles. HYPOPYGIUM (
|
||
<figureCitation id="130FD5ABFFC7FFD03B3ADE45FBA5FE48" box="[1042,1121,379,401]" captionStart="Figure 24" captionStartId="32.[142,220,883,902]" captionTargetBox="[306,1257,192,845]" captionTargetId="figure-497@32.[300,1260,183,851]" captionTargetPageId="32" captionText="Figure 24. The Empis (Coptophlebia) jacobsoni clade, E. (C.) cameronensis sp. nov. male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435314" httpUri="https://zenodo.org/record/5435314/files/figure.png" pageId="31" pageNumber="370">Fig. 24</figureCitation>
|
||
).
|
||
<taxonomicName id="4C34B2ADFFC7FFD03B5BDE45FB06FE49" box="[1139,1218,379,400]" class="Insecta" family="Kateretidae" genus="Cercus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="345" phylum="Arthropoda" rank="genus">Cercus</taxonomicName>
|
||
with minute spines at tip. Epandrium with pair of small anterodorsal protuberances, dorsal row of strong, long bristles, several other strong, long bristles at tip and ventrally. Hypandrium well sclerotized, sickle-shaped in lateral view. Phallus long, slender, not recurved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<table id="F9343B8EFFC600303FA5DCE4FA4AF8B5" box="[141,1422,986,1900]" pageId="30" pageNumber="369">
|
||
<paragraph id="8B8BC92EFFC6FFD13E11DCD2FABBFB9F" blockId="30.[313,1252,1004,1028]" lastBlockId="30.[159,1407,1045,1886]" pageId="30" pageNumber="369">
|
||
KEY TO SPECIES OF THE
|
||
<taxonomicName id="4C34B2ADFFC6FFD13D7EDCD3FC2FFBDB" authorityName="De Meijere" authorityYear="1907" box="[598,1003,1004,1028]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="380" phylum="Arthropoda" rank="species" species="jacobsoni" subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13D7EDCD3FD5BFBDB" box="[598,671,1005,1028]" italics="true" pageId="30" pageNumber="369">EMPIS</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13D98DCD2FC98FBDB" box="[688,860,1004,1028]" italics="true" pageId="30" pageNumber="369">COPTOPHLEBIA</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13C45DCD1FC2FFBDB" box="[877,1003,1007,1026]" italics="true" pageId="30" pageNumber="369">JACOBSONI</emphasis>
|
||
</taxonomicName>
|
||
CLADE (MALES ONLY) 1. Coxae and femora yellowish ............................................................................................................................................... 2 Legs brownish except for the yellowish to whitish mid tarsus .......................................................................................3
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFC6FFD13FB7DB6EFABAFB01" blockId="30.[159,1407,1045,1886]" pageId="30" pageNumber="369">
|
||
2. Phallus with a small ventral notch at middle, epandrium with numerous ventral bristles (
|
||
<figureCitation id="130FD5ABFFC6FFD13B47DB6EFB7DFBBA" box="[1135,1209,1104,1123]" captionStart="Figure 32" captionStartId="38.[142,220,1854,1873]" captionTargetBox="[226,1337,1284,1821]" captionTargetId="figure-554@38.[221,1341,1278,1824]" captionTargetPageId="38" captionText="Figure 32. The Empis (Coptophlebia) jacobsoni clade, E. (C.) mengyangensis sp. nov. A, male hypopygium in lateral view. Scale bar = 0.1 mm. B, male hind leg (tibia and tarsus) in lateral view. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5435330" httpUri="https://zenodo.org/record/5435330/files/figure.png" pageId="30" pageNumber="369">Fig. 32</figureCitation>
|
||
), fore tibia slender (in lateral view) with anterodorsal spine-like bristles basally, not as stout as in
|
||
<figureCitation id="130FD5ABFFC6FFD13B24DB53FBB6FB59" box="[1036,1138,1133,1152]" captionStart="Figure 30" captionStartId="37.[826,904,1026,1045]" captionTargetBox="[829,1437,537,993]" captionTargetId="figure-318@37.[829,1439,537,994]" captionTargetPageId="37" captionText="Figure 30. The Empis (Coptophlebia) jacobsoni clade, male of E. (C.) menglunensis sp. nov. scanning electron micrograph of fore tibia. Scale bar = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.5435326" httpUri="https://zenodo.org/record/5435326/files/figure.png" pageId="30" pageNumber="369">Figure 30</figureCitation>
|
||
.......................................... ...........................................................................................................................
|
||
<emphasis id="B940153CFFC6FFD13CE2DBB5FAECFB47" bold="true" box="[970,1320,1162,1182]" pageId="30" pageNumber="369">
|
||
<taxonomicName id="4C34B2ADFFC6FFD13CE2DBB5FB0DFB44" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[970,1225,1162,1182]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="369" phylum="Arthropoda" rank="species" species="mengyangensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13CE2DBB5FC26FB47" bold="true" box="[970,994,1163,1182]" italics="true" pageId="30" pageNumber="369">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13CD9DBB4FBCCFB44" bold="true" box="[1009,1032,1162,1181]" italics="true" pageId="30" pageNumber="369">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13B30DBB4FB0DFB44" bold="true" box="[1048,1225,1162,1181]" italics="true" pageId="30" pageNumber="369">mengyangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273A847FFC6FFD13BF9DBB5FAECFB47" box="[1233,1320,1163,1182]" pageId="30" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="F32389BEFFC6FFD13A1FDBB4FABCFB44" box="[1335,1400,1162,1181]" name="China" pageId="30" pageNumber="369">China</collectingCountry>
|
||
) Phallus lacking a small ventral notch at middle, epandrium with widely spaced ventral bristles (
|
||
<figureCitation id="130FD5ABFFC6FFD13BE9DB96FACFFB62" box="[1217,1291,1192,1211]" captionStart="Figure 31" captionStartId="37.[162,240,1854,1873]" captionTargetBox="[235,1364,1152,1793]" captionTargetId="figure-351@37.[219,1371,1135,1822]" captionTargetPageId="37" captionText="Figure 31. The Empis (Coptophlebia) jacobsoni clade, male of E. (C.) menglunensis sp. nov. male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435328" httpUri="https://zenodo.org/record/5435328/files/figure.png" pageId="30" pageNumber="369">Fig. 31</figureCitation>
|
||
), fore tibia thick with stout anterodorsal spine-like bristles basally (
|
||
<figureCitation id="130FD5ABFFC6FFD13C3FDBFBFCA5FB01" box="[791,865,1221,1240]" captionStart="Figure 30" captionStartId="37.[826,904,1026,1045]" captionTargetBox="[829,1437,537,993]" captionTargetId="figure-318@37.[829,1439,537,994]" captionTargetPageId="37" captionText="Figure 30. The Empis (Coptophlebia) jacobsoni clade, male of E. (C.) menglunensis sp. nov. scanning electron micrograph of fore tibia. Scale bar = 200 µm." figureDoi="http://doi.org/10.5281/zenodo.5435326" httpUri="https://zenodo.org/record/5435326/files/figure.png" pageId="30" pageNumber="369">Fig. 30</figureCitation>
|
||
) .................
|
||
<emphasis id="B940153CFFC6FFD13CF3DBFBFAECFB01" bold="true" box="[987,1320,1221,1240]" pageId="30" pageNumber="369">
|
||
<taxonomicName id="4C34B2ADFFC6FFD13CF3DBFBFB0CFB01" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[987,1224,1221,1240]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="369" phylum="Arthropoda" rank="species" species="menglunensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13CF3DBFBFC36FB01" bold="true" box="[987,1010,1221,1240]" italics="true" pageId="30" pageNumber="369">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13B29DBFBFBDDFB01" bold="true" box="[1025,1049,1221,1240]" italics="true" pageId="30" pageNumber="369">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13B00DBFBFB0CFB01" bold="true" box="[1064,1224,1221,1240]" italics="true" pageId="30" pageNumber="369">menglunensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273A847FFC6FFD13BF9DBFBFAECFB01" box="[1233,1320,1221,1240]" pageId="30" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="F32389BEFFC6FFD13A1FDBFBFABCFB01" box="[1335,1400,1221,1240]" name="China" pageId="30" pageNumber="369">China</collectingCountry>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFC6FFD13FB7DBDDFABBFACA" blockId="30.[159,1407,1045,1886]" pageId="30" pageNumber="369">3. Dorsal protuberances of epandrium very small, indistinct ..............................................................................................4 Dorsal protuberances of epandrium distinct ..................................................................................................................... 7</paragraph>
|
||
<paragraph id="8B8BC92EFFC6FFD13FB7DA23FABBFA94" blockId="30.[159,1407,1045,1886]" pageId="30" pageNumber="369">
|
||
4. First fore tarsomere about as long as fore tibia (
|
||
<figureCitation id="130FD5ABFFC6FFD13D95DA23FCD2FAE9" box="[701,790,1309,1328]" captionStart="Figure 33" captionStartId="39.[162,240,1854,1873]" captionTargetBox="[336,1269,1308,1813]" captionTargetId="figure-571@39.[318,1278,1294,1825]" captionTargetPageId="39" captionText="Figure 33. The Empis (Coptophlebia) jacobsoni clade, E. (C.) nimbaensis sp. nov. A, male fore leg (femur, tibia and first tarsomere) in lateral view. Scale bar = 0.5 mm. B, male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435332" httpUri="https://zenodo.org/record/5435332/files/figure.png" pageId="30" pageNumber="369">Fig. 33A</figureCitation>
|
||
)................................
|
||
<emphasis id="B940153CFFC6FFD13CC2DA23FAD8FAE9" bold="true" box="[1002,1308,1309,1328]" pageId="30" pageNumber="369">
|
||
<taxonomicName id="4C34B2ADFFC6FFD13CC2DA23FB78FAE9" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[1002,1212,1309,1328]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="369" phylum="Arthropoda" rank="species" species="nimbaensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13CC2DA23FBC5FAE9" bold="true" box="[1002,1025,1309,1328]" italics="true" pageId="30" pageNumber="369">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13B38DA23FBECFAE9" bold="true" box="[1040,1064,1309,1328]" italics="true" pageId="30" pageNumber="369">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13B1FDA23FB78FAE9" bold="true" box="[1079,1212,1309,1328]" italics="true" pageId="30" pageNumber="369">nimbaensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273A847FFC6FFD13BECDA23FAD8FAE9" box="[1220,1308,1309,1328]" pageId="30" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="F32389BEFFC6FFD13A02DA23FAB3FAE9" box="[1322,1399,1309,1328]" name="Guinea-Bissau" pageId="30" pageNumber="369">Guinea</collectingCountry>
|
||
) First fore tarsomere shorter than fore tibia...................................................................................................................... 5
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFC6FFD13FB7DA66FABBFA7C" blockId="30.[159,1407,1045,1886]" pageId="30" pageNumber="369">
|
||
5.
|
||
<taxonomicName id="4C34B2ADFFC6FFD13F95DA66FEC1FAB2" box="[189,261,1368,1387]" class="Insecta" family="Kateretidae" genus="Cercus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="345" phylum="Arthropoda" rank="genus">Cercus</taxonomicName>
|
||
with a rounded posteroventral projection subapically (Daugeron, 2002: fig. 1)................................................... ...............................................................................................................
|
||
<taxonomicName id="4C34B2ADFFC6FFD13C57DA4AFB03FA50" authority="Daugeron" authorityName="Daugeron" authorityYear="2002" box="[895,1223,1395,1417]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="369" phylum="Arthropoda" rank="species" species="nondouensis" subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13C57DA4AFC5CFA50" box="[895,920,1396,1417]" italics="true" pageId="30" pageNumber="369">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13C8EDA4AFC7AFA50" box="[934,958,1396,1417]" italics="true" pageId="30" pageNumber="369">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13CE3DA4DFB9CFA51" box="[971,1112,1395,1416]" italics="true" pageId="30" pageNumber="369">nondouensis</emphasis>
|
||
Daugeron
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F32389BEFFC6FFD13BFDDA4BFAB3FA51" box="[1237,1399,1397,1416]" name="New Caledonia" pageId="30" pageNumber="369">New Caledonia</collectingCountry>
|
||
)
|
||
<taxonomicName id="4C34B2ADFFC6FFD13F95DAADFEC1FA7F" box="[189,261,1427,1446]" class="Insecta" family="Kateretidae" genus="Cercus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="345" phylum="Arthropoda" rank="genus">Cercus</taxonomicName>
|
||
without a rounded posteroventral projection subapically ...................................................................................6
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFC6FFD13FB7DA8EFABAF9C2" blockId="30.[159,1407,1045,1886]" pageId="30" pageNumber="369">
|
||
6. Epandrium rounded at tip with ventral margin bearing
|
||
<emphasis id="B940153CFFC6FFD13C3ADA8EFCE6FA1B" box="[786,802,1456,1474]" italics="true" pageId="30" pageNumber="369">c.</emphasis>
|
||
10 bristles (
|
||
<figureCitation id="130FD5ABFFC6FFD13C8FDA8EFC36FA1A" box="[935,1010,1456,1475]" captionStart="Figure 29" captionStartId="36.[142,220,1854,1873]" captionTargetBox="[369,1191,1287,1820]" captionTargetId="figure-550@36.[365,1197,1250,1826]" captionTargetPageId="36" captionText="Figure 29. The Empis (C.) jacobsoni clade, E. (C.) loici sp. nov. male hypopygium in lateral view (phallus broken, hypandrium missing). Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435324" httpUri="https://zenodo.org/record/5435324/files/figure.png" pageId="30" pageNumber="369">Fig. 29</figureCitation>
|
||
). ............................................................ ...............................................................................................................
|
||
<emphasis id="B940153CFFC6FFD13CA8DAF3FBA4FA39" bold="true" box="[896,1120,1485,1504]" pageId="30" pageNumber="369">
|
||
<taxonomicName id="4C34B2ADFFC6FFD13CA8DAF3FBC5FA39" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[896,1025,1485,1504]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="369" phylum="Arthropoda" rank="species" species="loici" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13CA8DAF3FC53FA39" bold="true" box="[896,919,1485,1504]" italics="true" pageId="30" pageNumber="369">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13C8EDAF3FC79FA39" bold="true" box="[934,957,1485,1504]" italics="true" pageId="30" pageNumber="369">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13CE5DAF3FBC5FA39" bold="true" box="[973,1025,1485,1504]" italics="true" pageId="30" pageNumber="369">loici</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273A847FFC6FFD13B20DAF3FBA4FA39" box="[1032,1120,1485,1504]" pageId="30" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="F32389BEFFC6FFD13B47DAF3FAB3FA39" box="[1135,1399,1485,1504]" name="Central African Republic" pageId="30" pageNumber="369">Central African Republic</collectingCountry>
|
||
) Epandrium somewhat lengthened, tip partially truncated with
|
||
<emphasis id="B940153CFFC6FFD13C75DAD5FCA9FA24" box="[861,877,1515,1533]" italics="true" pageId="30" pageNumber="369">c.</emphasis>
|
||
15 bristles on the ventral margin (
|
||
<figureCitation id="130FD5ABFFC6FFD13BCBDAD4FAE9FA27" box="[1251,1325,1514,1534]" captionStart="Figure 28" captionStartId="35.[162,240,1564,1583]" captionTargetBox="[335,1271,1049,1526]" captionTargetId="figure-199@35.[315,1275,1032,1543]" captionTargetPageId="35" captionText="Figure 28. The Empis (Coptophlebia) jacobsoni clade, E. (C.) lachaisei sp. nov. male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435322" httpUri="https://zenodo.org/record/5435322/files/figure.png" pageId="30" pageNumber="369">Fig. 28</figureCitation>
|
||
) ........... ............................................................................................................................
|
||
<emphasis id="B940153CFFC6FFD13CFED936FB2AF9C2" bold="true" box="[982,1262,1543,1563]" pageId="30" pageNumber="369">
|
||
<taxonomicName id="4C34B2ADFFC6FFD13CFED936FB4AF9C3" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[982,1166,1543,1563]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="369" phylum="Arthropoda" rank="species" species="lachaisei" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13CFED936FC29F9C2" bold="true" box="[982,1005,1544,1563]" italics="true" pageId="30" pageNumber="369">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13CD4D936FBD0F9C2" bold="true" box="[1020,1044,1544,1563]" italics="true" pageId="30" pageNumber="369">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13B0BD939FB4AF9C3" bold="true" box="[1059,1166,1543,1562]" italics="true" pageId="30" pageNumber="369">lachaisei</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273A847FFC6FFD13BBED936FB2AF9C2" box="[1174,1262,1544,1563]" pageId="30" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="F32389BEFFC6FFD13BD4D936FAB3F9C2" box="[1276,1399,1544,1563]" name="Ivory Coast" pageId="30" pageNumber="369">Ivory Coast</collectingCountry>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFC6FFD13FB7D918FABBF949" blockId="30.[159,1407,1045,1886]" pageId="30" pageNumber="369">
|
||
7. Fore tarsus swollen with last four tarsomeres more or less rounded (
|
||
<figureCitation id="130FD5ABFFC6FFD13CBAD91BFC19F9E1" box="[914,989,1573,1592]" captionStart="Figure 25" captionStartId="33.[162,240,1841,1860]" captionTargetBox="[270,1329,198,1797]" captionTargetId="figure-50@33.[249,1343,183,1815]" captionTargetPageId="33" captionText="Figure 25. The Empis (Coptophlebia) jacobsoni clade, E. (C.) jacobsoni De Meijere: habitus, male." figureDoi="http://doi.org/10.5281/zenodo.5435316" httpUri="https://zenodo.org/record/5435316/files/figure.png" pageId="30" pageNumber="369">Fig. 25</figureCitation>
|
||
), epandrium with 1 apical bristle much stronger than other apicals (
|
||
<figureCitation id="130FD5ABFFC6FFD13EC4D97CFDF9F98F" box="[492,573,1602,1622]" captionStart="Figure 27" captionStartId="35.[162,240,949,968]" captionTargetBox="[333,1268,198,913]" captionTargetId="figure-157@35.[318,1278,182,930]" captionTargetPageId="35" captionText="Figure 27. The Empis (Coptophlebia) jacobsoni clade, E. (C.) jacobsoni De Meijere: male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435320" httpUri="https://zenodo.org/record/5435320/files/figure.png" pageId="30" pageNumber="369">Figs 27</figureCitation>
|
||
,
|
||
<figureCitation id="130FD5ABFFC6FFD13D60D97DFDB6F98F" box="[584,626,1603,1622]" captionStart="Figure 34" captionStartId="40.[806,884,1031,1050]" captionTargetBox="[822,1404,198,996]" captionTargetId="figure-656@40.[803,1411,178,1008]" captionTargetPageId="40" captionText="Figure 34. The Empis (Coptophlebia) jacobsoni clade, male of E. (C.) padangensis sp. nov. A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435334" httpUri="https://zenodo.org/record/5435334/files/figure.png" pageId="30" pageNumber="369">34A</figureCitation>
|
||
)........................................................................................................................ 8 Fore tarsus a little swollen, not rounded, epandrium without an apical bristle much stronger than other apicals (
|
||
<figureCitation id="130FD5ABFFC6FFD13FEDD943FECAF949" box="[197,270,1661,1680]" captionStart="Figure 24" captionStartId="32.[142,220,883,902]" captionTargetBox="[306,1257,192,845]" captionTargetId="figure-497@32.[300,1260,183,851]" captionTargetPageId="32" captionText="Figure 24. The Empis (Coptophlebia) jacobsoni clade, E. (C.) cameronensis sp. nov. male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435314" httpUri="https://zenodo.org/record/5435314/files/figure.png" pageId="30" pageNumber="369">Fig. 24</figureCitation>
|
||
) ..........................................................................................................
|
||
<emphasis id="B940153CFFC6FFD13C94D943FACCF949" bold="true" box="[956,1288,1661,1680]" pageId="30" pageNumber="369">
|
||
<taxonomicName id="4C34B2ADFFC6FFD13C94D943FB6DF949" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[956,1193,1661,1680]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="369" phylum="Arthropoda" rank="species" species="cameronensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13C94D943FC17F949" bold="true" box="[956,979,1661,1680]" italics="true" pageId="30" pageNumber="369">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13CCBD943FC3EF949" bold="true" box="[995,1018,1661,1680]" italics="true" pageId="30" pageNumber="369">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13B22D943FB6DF949" bold="true" box="[1034,1193,1661,1680]" italics="true" pageId="30" pageNumber="369">cameronensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273A847FFC6FFD13B98D943FACCF949" box="[1200,1288,1661,1680]" pageId="30" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="F32389BEFFC6FFD13A3FD943FABCF949" box="[1303,1400,1661,1680]" name="Malaysia" pageId="30" pageNumber="369">Malaysia</collectingCountry>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8B8BC92EFFC6FFD13FB7D9A5FABAF884" blockId="30.[159,1407,1045,1886]" pageId="30" pageNumber="369">
|
||
8. Apical four fore tarsomeres with numerous long dorsal bristles (
|
||
<figureCitation id="130FD5ABFFC6FFD13C4BD9A4FC6AF974" box="[867,942,1690,1709]" captionStart="Figure 25" captionStartId="33.[162,240,1841,1860]" captionTargetBox="[270,1329,198,1797]" captionTargetId="figure-50@33.[249,1343,183,1815]" captionTargetPageId="33" captionText="Figure 25. The Empis (Coptophlebia) jacobsoni clade, E. (C.) jacobsoni De Meijere: habitus, male." figureDoi="http://doi.org/10.5281/zenodo.5435316" httpUri="https://zenodo.org/record/5435316/files/figure.png" pageId="30" pageNumber="369">Fig. 25</figureCitation>
|
||
), hind femur somewhat S-shaped (visible in dorsal view), epandrium posteriorly strongly modified in a long very massive projection bearing 1 very strong bristle apically, hypandrium not apically developed in a long claw-like projection (
|
||
<figureCitation id="130FD5ABFFC6FFD13CFED9EBFBE4F931" box="[982,1056,1749,1768]" captionStart="Figure 27" captionStartId="35.[162,240,949,968]" captionTargetBox="[333,1268,198,913]" captionTargetId="figure-157@35.[318,1278,182,930]" captionTargetPageId="35" captionText="Figure 27. The Empis (Coptophlebia) jacobsoni clade, E. (C.) jacobsoni De Meijere: male hypopygium in lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435320" httpUri="https://zenodo.org/record/5435320/files/figure.png" pageId="30" pageNumber="369">Fig. 27</figureCitation>
|
||
)...................................................... .....................................................................................................................
|
||
<emphasis id="B940153CFFC6FFD13C8ED9CDFB06F8DF" bold="true" box="[934,1218,1778,1798]" pageId="30" pageNumber="369">
|
||
<taxonomicName id="4C34B2ADFFC6FFD13C8ED9CDFBA7F8DC" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[934,1123,1778,1798]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="369" phylum="Arthropoda" rank="species" species="jacobsoni" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13C8ED9CDFC79F8DF" bold="true" box="[934,957,1779,1798]" italics="true" pageId="30" pageNumber="369">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13CE5D9CCFC20F8DC" bold="true" box="[973,996,1778,1797]" italics="true" pageId="30" pageNumber="369">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13CDCD9CCFBA7F8DC" bold="true" box="[1012,1123,1778,1797]" italics="true" pageId="30" pageNumber="369">jacobsoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273A847FFC6FFD13B42D9CDFB06F8DF" box="[1130,1218,1779,1798]" pageId="30" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="F32389BEFFC6FFD13BF9D9CCFAFFF8DC" box="[1233,1339,1778,1797]" name="Indonesia" pageId="30" pageNumber="369">Indonesia</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F007CCFFC6FFD13A6ED9CCFAB3F8DC" box="[1350,1399,1778,1797]" country="Indonesia" name="Jawa Timur" pageId="30" pageNumber="369">Java</collectingRegion>
|
||
) Apical four fore tarsomeres with long bristles only apically, hind femur not S-shaped, epandrium not posteriorly modified in a massive projection (
|
||
<figureCitation id="130FD5ABFFC6FFD13D09D813FDB8F899" box="[545,636,1837,1856]" captionStart="Figure 34" captionStartId="40.[806,884,1031,1050]" captionTargetBox="[822,1404,198,996]" captionTargetId="figure-656@40.[803,1411,178,1008]" captionTargetPageId="40" captionText="Figure 34. The Empis (Coptophlebia) jacobsoni clade, male of E. (C.) padangensis sp. nov. A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435334" httpUri="https://zenodo.org/record/5435334/files/figure.png" pageId="30" pageNumber="369">Fig. 34A</figureCitation>
|
||
), hypandrium apically developed in a long claw-like projection (
|
||
<figureCitation id="130FD5ABFFC6FFD13A37D813FABCF899" box="[1311,1400,1837,1856]" captionStart="Figure 34" captionStartId="40.[806,884,1031,1050]" captionTargetBox="[822,1404,198,996]" captionTargetId="figure-656@40.[803,1411,178,1008]" captionTargetPageId="40" captionText="Figure 34. The Empis (Coptophlebia) jacobsoni clade, male of E. (C.) padangensis sp. nov. A, epandrium and cercus in lateral view; B, hypandrium and phallus in lateral view. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5435334" httpUri="https://zenodo.org/record/5435334/files/figure.png" pageId="30" pageNumber="369">Fig. 34B</figureCitation>
|
||
) ........................................................................................................
|
||
<emphasis id="B940153CFFC6FFD13C7ED875FB53F887" bold="true" box="[854,1175,1866,1886]" pageId="30" pageNumber="369">
|
||
<taxonomicName id="4C34B2ADFFC6FFD13C7ED875FBF3F884" authority="Daugeron & Grootaert, 2005" authorityName="Daugeron & Grootaert" authorityYear="2005" box="[854,1079,1866,1886]" class="Insecta" family="Empididae" genus="Empis" kingdom="Animalia" order="Diptera" pageId="30" pageNumber="369" phylum="Arthropoda" rank="species" species="padangensis" status="sp. nov." subGenus="Coptophlebia">
|
||
<emphasis id="B940153CFFC6FFD13C7ED875FCAAF887" bold="true" box="[854,878,1867,1886]" italics="true" pageId="30" pageNumber="369">E.</emphasis>
|
||
(
|
||
<emphasis id="B940153CFFC6FFD13C55D874FC50F884" bold="true" box="[893,916,1866,1885]" italics="true" pageId="30" pageNumber="369">C.</emphasis>
|
||
)
|
||
<emphasis id="B940153CFFC6FFD13C8CD874FBF3F884" bold="true" box="[932,1079,1866,1885]" italics="true" pageId="30" pageNumber="369">padangensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A273A847FFC6FFD13B17D875FB53F887" box="[1087,1175,1867,1886]" pageId="30" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<collectingCountry id="F32389BEFFC6FFD13B8ED874FAD4F884" box="[1190,1296,1866,1885]" name="Indonesia" pageId="30" pageNumber="369">Indonesia</collectingCountry>
|
||
:
|
||
<collectingRegion id="49F007CCFFC6FFD13A33D874FABCF884" box="[1307,1400,1866,1885]" country="Indonesia" name="Sumatera Utara" pageId="30" pageNumber="369">Sumatra</collectingRegion>
|
||
)
|
||
</paragraph>
|
||
</table>
|
||
<subSubSection id="C32E9AA5FFC7FFD03C7BDD0AFC71FB49" pageId="31" pageNumber="370" type="description">
|
||
<paragraph id="8B8BC92EFFC7FFD03C7BDD0AFC71FB49" blockId="31.[826,1443,195,1168]" pageId="31" pageNumber="370">FEMALE: Similar to male except for the following characters: ocellar triangle with pair of distinct bristles. Lower ommatidia enlarged, frons black, shiny, as wide as face. Second and third flagellomeres black, aristiform. Two fine, short visible acrostichals. Fore tibia with dorsal pennation on apical half, short ventral pennation on apical two-thirds; first fore tarsomere with dorsal pennation except apically. Mid femur with long and short antero- and posterodorsal pennation, respectively, ventral pennation; mid tibia with flattened ventral bristles, short and long antero- and posterodorsal pennation, respectively; first mid tarsomere with dorsal pennation, densely covered with spine-like bristles ventrally. Hind femur and tibia with long and very short antero- and posterodorsal pennation, respectively, ventral pennation; hind tibia with very short posterodorsal and anteroventral pennation. First hind tarsomere with long anterodorsal pennation except apically, short posterodorsal pennation.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |