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<document id="8BD8B7B3CBDD3D9DB6523D0541EB361A" ID-DOI="10.5852/ejt.2024.957.2651" ID-ISSN="2118-9773" ID-Zenodo-Dep="13773385" ID-ZooBank="9957F2DF-3F31-4B58-84BC-2F8E0F45C6E3" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726582870537" checkinUser="plazi" docAuthor="Selnekovič, Dávid, Jäch, Manfred A. &amp; Kodada, Ján" docDate="2024" docId="038B5001DA3EE775FDF7FB71FD6F27B9" docLanguage="en" docName="EJT.2024.957.1-229.pdf" docOrigin="European Journal of Taxonomy 957" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2651/12247" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Elmomorphus calvus Selnekovič, Jäch &amp; Kodada, 2024, sp. nov." docType="treatment" docUuid="384803E1-F693-4257-94BB-F785C824F6B4" docUuidSource="ZooBank" docVersion="2" lastPageNumber="59" masterDocId="FFB22879DA08E74FFF9EFFBAFFE32440" masterDocTitle="Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)" masterLastPageNumber="229" masterPageNumber="1" pageNumber="55" updateTime="1726584008522" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="483236718E861AECAFCFD9BA53C55F2A">Taxonomic revision of the genus Elmomorphus Sharp, 1888. II. Redescription of the genus and review of the species from India, Nepal, Bhutan, Myanmar, China, Thailand, Laos, Cambodia, and Vietnam (Coleoptera: Dryopidae)</mods:title>
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<mods:namePart id="DB220F29AD448C9DED46B5C1EAFF9438">Selnekovič, Dávid</mods:namePart>
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<mods:affiliation id="BC7D6038E5154049C400FEFC44975887">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
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<mods:namePart id="C323F22742663574E1E342E8FB2D19BC">Jäch, Manfred A.</mods:namePart>
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<mods:affiliation id="762605AAD548A55690269AB0CDE470F5">Naturhistorisches Museum Wien, Burgring 7, A- 1010 Wien, Austria.</mods:affiliation>
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<mods:namePart id="E08EC71CA0AF717AC90A3DABD9F4DD3A">Kodada, Ján</mods:namePart>
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<mods:affiliation id="56E8CD67D12357C62529B5C3D8E212E3">Department of Zoology, Faculty of Natural Sciences, Comenius University in Bratislava, Ilkovičova 6, SK- 842 15 Bratislava, Slovakia.</mods:affiliation>
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<paragraph id="8B9DE117DA3EE779FDF7FB71FC2820A7" blockId="54.[379,1209,1227,1323]" box="[617,971,1227,1255]" pageId="54" pageNumber="55">
<heading id="D0D5567BDA3EE779FDF7FB71FC2820A7" box="[617,971,1227,1255]" centered="true" fontSize="11" level="2" pageId="54" pageNumber="55" reason="2">
<taxonomicName id="4C229A94DA3EE779FDF7FB71FC8920A6" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[617,874,1227,1254]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calvus" status="sp. nov.">
<emphasis id="B9563D05DA3EE779FDF7FB71FC8920A6" bold="true" box="[617,874,1227,1254]" italics="true" pageId="54" pageNumber="55">Elmomorphus calvus</emphasis>
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<taxonomicNameLabel id="A265807EDA3EE779FCEFFB77FC2820A7" box="[881,971,1229,1255]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C338B29CDA3EE779FD0FFAABFC40216B" box="[657,931,1297,1323]" pageId="54" pageNumber="55" type="description">
<paragraph id="8B9DE117DA3EE779FD0FFAABFC40216B" blockId="54.[379,1209,1227,1323]" box="[657,931,1297,1323]" pageId="54" pageNumber="55">
<figureCitation id="1319FD92DA3EE779FD0FFAABFCFB216B" box="[657,792,1297,1323]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="55.[189,232,1969,1995]" captionStartId-1="56.[189,232,1888,1914]" captionTargetBox-0="[189,1399,265,1928]" captionTargetBox-1="[296,1282,278,1832]" captionTargetId-0="figure-12@55.[189,1399,265,1928]" captionTargetId-1="figure-21@56.[277,1295,265,1839]" captionTargetPageId-0="55" captionTargetPageId-1="56" captionText-0="Fig. 23. Elmomorphus calvus sp. nov., holotype, male (NMW), TL: 3.25 mm." captionText-1="Fig. 24. Elmomorphus calvus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13773441" figureDoi-1="http://doi.org/10.5281/zenodo.13773445" httpUri-0="https://zenodo.org/record/13773441/files/figure.png" httpUri-1="https://zenodo.org/record/13773445/files/figure.png" pageId="54" pageNumber="55">Figs 23–24</figureCitation>
, 
<figureCitation id="1319FD92DA3EE779FCBDFAABFCB4216B" box="[803,855,1297,1323]" captionStart="Fig" captionStartId="64.[189,232,1829,1855]" captionTargetBox="[242,1354,287,1780]" captionTargetId="figure-40@64.[223,1369,265,1788]" captionTargetPageId="64" captionText="Fig. 28. Bursa copulatrix and vagina. A. Elmomorphus elmoides sp. nov., paratype from type locality (NMW). B. E. calvus sp. nov., paratype from type locality (CKB). C. E. depressus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773457" httpUri="https://zenodo.org/record/13773457/files/figure.png" pageId="54" pageNumber="55">28B</figureCitation>
, 
<figureCitation id="1319FD92DA3EE779FCFDFAABFC40216B" box="[867,931,1297,1323]" captionStart="Fig" captionStartId="217.[189,232,1872,1898]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-44@217.[189,1399,265,1830]" captionTargetPageId="217" captionText="Fig. 109. Distribution maps. A. Elmomorphus fusiformis sp. nov. B. E. elmoides sp. nov. C. E. calvus sp. nov. D. E. hongkong sp. nov. E. E. depressus sp. nov. F. E. sulcatus Kodada, Selnekovič &amp; Jäch sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773712" httpUri="https://zenodo.org/record/13773712/files/figure.png" pageId="54" pageNumber="55">109C</figureCitation>
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<subSubSection id="C338B29CDA3EE779FF23FAE8FC6523A8" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="8B9DE117DA3EE779FF23FAE8FE2A212C" blockId="54.[189,457,1362,1388]" box="[189,457,1362,1388]" pageId="54" pageNumber="55">
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<emphasis id="B9563D05DA3EE779FF23FAE8FE2A212C" bold="true" box="[189,457,1362,1388]" pageId="54" pageNumber="55">Differential diagnosis</emphasis>
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<taxonomicName id="4C229A94DA3EE779FF23FA3AFE5021DA" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[189,435,1408,1434]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calvus" status="sp. nov.">
<emphasis id="B9563D05DA3EE779FF23FA3AFE5021DA" box="[189,435,1408,1434]" italics="true" pageId="54" pageNumber="55">Elmomorphus calvus</emphasis>
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(
<figureCitation id="1319FD92DA3EE779FDBCFA3AFD9521DA" box="[546,630,1408,1434]" captionStart="Fig" captionStartId="55.[189,232,1969,1995]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@55.[189,1399,265,1928]" captionTargetPageId="55" captionText="Fig. 23. Elmomorphus calvus sp. nov., holotype, male (NMW), TL: 3.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13773441" httpUri="https://zenodo.org/record/13773441/files/figure.png" pageId="54" pageNumber="55">Fig. 23</figureCitation>
) is characterised by the presence of plastron on the frontoclypeus, the anterolateral portions of the vertex and pronotum, and the lateral sides of the elytra. The elytral punctation consists of small scattered punctures and large punctures arranged in nine well-impressed striae. The pronotum is strongly convex with distinctly rounded lateral sides. Males possess conspicuous long erect setae on the labrum and small clusters of setae on the prosternal process, metaventrite and ventrite 5. The aedeagus is large, the phallobase is robust and strongly curved (
<figureCitation id="1319FD92DA3EE779FBD3F997FB422208" box="[1101,1185,1581,1608]" captionStart="Fig" captionStartId="56.[189,232,1888,1914]" captionTargetBox="[296,1282,278,1832]" captionTargetId="figure-21@56.[277,1295,265,1839]" captionTargetPageId="56" captionText="Fig. 24. Elmomorphus calvus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773445" httpUri="https://zenodo.org/record/13773445/files/figure.png" pageId="54" pageNumber="55">Fig. 24</figureCitation>
). Bursa copulatrix without sclerotised spines (
<figureCitation id="1319FD92DA3EE779FE62F9EAFD84222A" box="[508,615,1616,1642]" captionStart="Fig" captionStartId="64.[189,232,1829,1855]" captionTargetBox="[242,1354,287,1780]" captionTargetId="figure-40@64.[223,1369,265,1788]" captionTargetPageId="64" captionText="Fig. 28. Bursa copulatrix and vagina. A. Elmomorphus elmoides sp. nov., paratype from type locality (NMW). B. E. calvus sp. nov., paratype from type locality (CKB). C. E. depressus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773457" httpUri="https://zenodo.org/record/13773457/files/figure.png" pageId="54" pageNumber="55">Fig. 28B</figureCitation>
). A similar combination of characters is also shared by 
<taxonomicName id="4C229A94DA3EE779FB70F9EBFA95222A" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1262,1398,1616,1642]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="elmoides" status="sp. nov.">
<emphasis id="B9563D05DA3EE779FB70F9EBFA95222A" box="[1262,1398,1616,1642]" italics="true" pageId="54" pageNumber="55">E. elmoides</emphasis>
</taxonomicName>
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, which differs in the smaller, TL: 
<specimenCount id="9D242A9EDA3EE779FD2BF9C9FD0222CD" box="[693,737,1651,1677]" count="1" pageId="54" pageNumber="55" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DA3EE779FD74F9C9FC7422CD" box="[746,919,1651,1677]" metricMagnitude="-3" metricUnit="m" metricValue="2.575" metricValueMax="2.68" metricValueMin="2.47" pageId="54" pageNumber="55" unit="mm" value="2.575" valueMax="2.68" valueMin="2.47">2.47–2.68 mm</quantity>
(2.55 ± 0.06, n=10), 
<specimenCount id="9D242A9EDA3EE779FB0BF9C9FB2222CD" box="[1173,1217,1651,1677]" count="1" pageId="54" pageNumber="55" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA3EE779FB54F9C8FA9422CD" box="[1226,1399,1650,1677]" metricMagnitude="-3" metricUnit="m" metricValue="2.795" metricValueMax="2.94" metricValueMin="2.65" pageId="54" pageNumber="55" unit="mm" value="2.795" valueMax="2.94" valueMin="2.65">2.65–2.94 mm</quantity>
(2.81 ± 0.09, n=10) in 
<taxonomicName id="4C229A94DA3EE779FE5AF92CFDAF22EF" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[452,588,1685,1711]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="elmoides">
<emphasis id="B9563D05DA3EE779FE5AF92CFDAF22EF" box="[452,588,1685,1711]" italics="true" pageId="54" pageNumber="55">E. elmoides</emphasis>
</taxonomicName>
versus TL: 
<specimenCount id="9D242A9EDA3EE779FD48F92CFCE122F0" box="[726,770,1686,1712]" count="1" pageId="54" pageNumber="55" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DA3EE779FC96F92FFC5122F0" box="[776,946,1685,1712]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" metricValueMax="3.35" metricValueMin="3.17" pageId="54" pageNumber="55" unit="mm" value="3.26" valueMax="3.35" valueMin="3.17">3.17–3.35 mm</quantity>
(3.25 ± 0.06, n=7), 
<specimenCount id="9D242A9EDA3EE779FB05F92CFB2422F0" box="[1179,1223,1686,1712]" count="1" pageId="54" pageNumber="55" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA3EE779FB53F92FFA9422F0" box="[1229,1399,1685,1712]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.45" metricValueMin="3.15" pageId="54" pageNumber="55" unit="mm" value="3.3" valueMax="3.45" valueMin="3.15">3.15–3.45 mm</quantity>
(3.30 ± 0.11, n =10) in 
<taxonomicName id="4C229A94DA3EE779FE56F903FDD62292" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[456,565,1720,1746]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calvus">
<emphasis id="B9563D05DA3EE779FE56F903FDD62292" box="[456,565,1720,1746]" italics="true" pageId="54" pageNumber="55">E. calvus</emphasis>
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, and less convex body (
<figureCitation id="1319FD92DA3EE779FCCDF902FC482292" box="[851,939,1720,1746]" captionStart="Fig" captionStartId="51.[189,232,1970,1996]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-16@51.[189,1399,265,1928]" captionTargetPageId="51" captionText="Fig. 21. Elmomorphus elmoides sp. nov., paratype, male from Buon Luoi, Vietnam (CKB), TL: 2.55 mm." figureDoi="http://doi.org/10.5281/zenodo.13773435" httpUri="https://zenodo.org/record/13773435/files/figure.png" pageId="54" pageNumber="55">Fig. 21</figureCitation>
), the pronotum being less convex with straight lateral sides in 
<taxonomicName id="4C229A94DA3EE779FE4DF961FDB822B5" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[467,603,1755,1781]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="elmoides">
<emphasis id="B9563D05DA3EE779FE4DF961FDB822B5" box="[467,603,1755,1781]" italics="true" pageId="54" pageNumber="55">E. elmoides</emphasis>
</taxonomicName>
versus strongly convex with rounded lateral sides in 
<taxonomicName id="4C229A94DA3EE779FB48F961FAA022B5" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1238,1347,1755,1781]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calvus">
<emphasis id="B9563D05DA3EE779FB48F961FAA022B5" box="[1238,1347,1755,1781]" italics="true" pageId="54" pageNumber="55">E. calvus</emphasis>
</taxonomicName>
, the pronotal plastron being present along the entire pronotal sides in 
<taxonomicName id="4C229A94DA3EE779FC4FF944FBBF2357" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[977,1116,1789,1815]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="elmoides">
<emphasis id="B9563D05DA3EE779FC4FF944FBBF2357" box="[977,1116,1789,1815]" italics="true" pageId="54" pageNumber="55">E. elmoides</emphasis>
</taxonomicName>
versus restricted to the anterior half in 
<taxonomicName id="4C229A94DA3EE779FEE5F89BFE09237A" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[379,490,1824,1850]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calvus">
<emphasis id="B9563D05DA3EE779FEE5F89BFE09237A" box="[379,490,1824,1850]" italics="true" pageId="54" pageNumber="55">E. calvus</emphasis>
</taxonomicName>
, the aedeagus with a slender and almost straight phallobase in 
<taxonomicName id="4C229A94DA3EE779FB75F89BFA95237A" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1259,1398,1824,1850]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="elmoides">
<emphasis id="B9563D05DA3EE779FB75F89BFA95237A" box="[1259,1398,1824,1850]" italics="true" pageId="54" pageNumber="55">E. elmoides</emphasis>
</taxonomicName>
(
<figureCitation id="1319FD92DA3EE779FF58F8F8FEC2231D" box="[198,289,1858,1885]" captionStart="Fig" captionStartId="52.[189,232,1896,1922]" captionTargetBox="[415,1181,285,1839]" captionTargetId="figure-25@52.[395,1210,264,1849]" captionTargetPageId="52" captionText="Fig. 22. Elmomorphus elmoides sp. nov., paratype from type locality (CKB), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773439" httpUri="https://zenodo.org/record/13773439/files/figure.png" pageId="54" pageNumber="55">Fig. 22</figureCitation>
) versus an expanded and strongly bent phallobase in 
<taxonomicName id="4C229A94DA3EE779FC58F8F9FBD4231C" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[966,1079,1858,1884]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calvus">
<emphasis id="B9563D05DA3EE779FC58F8F9FBD4231C" box="[966,1079,1858,1884]" italics="true" pageId="54" pageNumber="55">E. calvus</emphasis>
</taxonomicName>
, and the bursa copulatrix possessing large sclerotised spines in 
<taxonomicName id="4C229A94DA3EE779FDE5F8DCFCE7233F" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[635,772,1893,1919]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="elmoides">
<emphasis id="B9563D05DA3EE779FDE5F8DCFCE7233F" box="[635,772,1893,1919]" italics="true" pageId="54" pageNumber="55">E. elmoides</emphasis>
</taxonomicName>
(
<figureCitation id="1319FD92DA3EE779FC8AF8DFFC61233F" box="[788,898,1893,1919]" captionStart="Fig" captionStartId="64.[189,232,1829,1855]" captionTargetBox="[242,1354,287,1780]" captionTargetId="figure-40@64.[223,1369,265,1788]" captionTargetPageId="64" captionText="Fig. 28. Bursa copulatrix and vagina. A. Elmomorphus elmoides sp. nov., paratype from type locality (NMW). B. E. calvus sp. nov., paratype from type locality (CKB). C. E. depressus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773457" httpUri="https://zenodo.org/record/13773457/files/figure.png" pageId="54" pageNumber="55">Fig. 28A</figureCitation>
), while in 
<taxonomicName id="4C229A94DA3EE779FB9EF8DCFB8F233F" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[1024,1132,1893,1919]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calvus">
<emphasis id="B9563D05DA3EE779FB9EF8DCFB8F233F" box="[1024,1132,1893,1919]" italics="true" pageId="54" pageNumber="55">E. calvus</emphasis>
</taxonomicName>
, the spines are absent. 
<taxonomicName id="4C229A94DA3EE779FF20F832FE5623E2" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[190,437,1928,1954]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="calvus">
<emphasis id="B9563D05DA3EE779FF20F832FE5623E2" box="[190,437,1928,1954]" italics="true" pageId="54" pageNumber="55">Elmomorphus calvus</emphasis>
</taxonomicName>
may also resemble 
<taxonomicName id="4C229A94DA3EE779FD3FF833FCD523E2" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[673,822,1928,1954]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="depressus" status="sp. nov.">
<emphasis id="B9563D05DA3EE779FD3FF833FCD523E2" box="[673,822,1928,1954]" italics="true" pageId="54" pageNumber="55">E. depressus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA3EE779FCA3F832FC7A23E2" box="[829,921,1928,1954]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C229A94DA3EE779FC4DF833FBB023E2" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[979,1107,1928,1954]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="sulcatus" status="sp. nov.">
<emphasis id="B9563D05DA3EE779FC4DF833FC0723E2" box="[979,996,1929,1954]" italics="true" pageId="54" pageNumber="55">E</emphasis>
. 
<emphasis id="B9563D05DA3EE779FC6DF832FBB023E2" box="[1011,1107,1928,1954]" italics="true" pageId="54" pageNumber="55">sulcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA3EE779FBC2F832FB5923E2" box="[1116,1210,1928,1954]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
; however, these species differ in the broader and more convex body shape (
<figureCitation id="1319FD92DA3EE779FC14F810FC0B2385" box="[906,1000,1962,1989]" captionStart="Fig" captionStartId="63.[189,232,1970,1996]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@63.[189,1399,265,1928]" captionTargetPageId="63" captionText="Fig. 27. Elmomorphus depressus sp. nov., holotype, female (IAECAS), TL: 3.58 mm." figureDoi="http://doi.org/10.5281/zenodo.13773455" httpUri="https://zenodo.org/record/13773455/files/figure.png" pageId="54" pageNumber="55">Figs 27</figureCitation>
, 
<figureCitation id="1319FD92DA3EE779FC69F811FBF42385" box="[1015,1047,1963,1989]" captionStart="Fig" captionStartId="84.[189,232,1811,1837]" captionTargetBox="[196,1393,285,1746]" captionTargetId="figure-62@84.[189,1399,265,1759]" captionTargetPageId="84" captionText="Fig. 39. A. Elmomorphus prosternalis Hinton, 1935, holotype (BMNH), prosternal process and median part of metaventrite. B. E. montanus (Grouvelle, 1913), syntype, male (MNHN), prosternal process and median part of metaventrite. C. E. montanus, syntype (MNHN), aedeagus, lateral aspect. D. Same, ventral aspect, detail of parameres and penis. E. Same, ventral aspect. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773490" httpUri="https://zenodo.org/record/13773490/files/figure.png" pageId="54" pageNumber="55">39</figureCitation>
), the absence of conspicuous clusters of setae in males, and a differently shaped aedeagus.
</paragraph>
</subSubSection>
<caption id="DF5DB19FDA3FE778FF23F80BFBAE238C" ID-DOI="http://doi.org/10.5281/zenodo.13773441" ID-Zenodo-Dep="13773441" box="[189,1101,1969,1997]" httpUri="https://zenodo.org/record/13773441/files/figure.png" pageId="55" pageNumber="56" startId="55.[189,232,1969,1995]" targetBox="[189,1399,265,1928]" targetPageId="55">
<paragraph id="8B9DE117DA3FE778FF23F80BFBAE238C" blockId="55.[189,1101,1969,1997]" box="[189,1101,1969,1997]" pageId="55" pageNumber="56">
<emphasis id="B9563D05DA3FE778FF23F80BFEF9238C" bold="true" box="[189,282,1969,1996]" pageId="55" pageNumber="56">Fig. 23.</emphasis>
<taxonomicName id="4C229A94DA3FE778FEBEF808FDF4238C" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[288,535,1970,1996]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="55" pageNumber="56" phylum="Arthropoda" rank="species" species="calvus" status="sp. nov.">
<emphasis id="B9563D05DA3FE778FEBEF808FDF4238C" box="[288,535,1970,1996]" italics="true" pageId="55" pageNumber="56">Elmomorphus calvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA3FE778FD81F808FD9F238C" box="[543,636,1970,1996]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, male (NMW), TL: 3.25 mm.
</paragraph>
</caption>
<caption id="DF5DB19FDA30E777FF23F8DAFE7A23DE" ID-DOI="http://doi.org/10.5281/zenodo.13773445" ID-Zenodo-Dep="13773445" httpUri="https://zenodo.org/record/13773445/files/figure.png" pageId="56" pageNumber="57" startId="56.[189,232,1888,1914]" targetBox="[296,1282,278,1832]" targetPageId="56">
<paragraph id="8B9DE117DA30E777FF23F8DAFE7A23DE" blockId="56.[188,1399,1888,1950]" pageId="56" pageNumber="57">
<emphasis id="B9563D05DA30E777FF23F8DAFEFA233A" bold="true" box="[189,281,1888,1914]" pageId="56" pageNumber="57">Fig. 24.</emphasis>
<taxonomicName id="4C229A94DA30E777FE81F8DAFDF6233A" authorityName="Selnekovič &amp; Jäch &amp; Kodada" authorityYear="2024" box="[287,533,1888,1914]" class="Insecta" family="Dryopidae" genus="Elmomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="56" pageNumber="57" phylum="Arthropoda" rank="species" species="calvus" status="sp. nov.">
<emphasis id="B9563D05DA30E777FE81F8DAFDF6233A" box="[287,533,1888,1914]" italics="true" pageId="56" pageNumber="57">Elmomorphus calvus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265807EDA30E777FD85F8DAFD9B233A" box="[539,632,1888,1914]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
, holotype (NMW), aedeagus. 
<emphasis id="B9563D05DA30E777FC4BF8DAFC09233A" bold="true" box="[981,1002,1888,1914]" pageId="56" pageNumber="57">A</emphasis>
. Lateral aspect. 
<emphasis id="B9563D05DA30E777FB37F8DAFB5E233A" bold="true" box="[1193,1213,1888,1914]" pageId="56" pageNumber="57">B</emphasis>
. Ventral aspect. Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C338B29CDA31E776FF23FEB3FE8A2534" pageId="57" pageNumber="58" type="etymology">
<paragraph id="8B9DE117DA31E776FF23FEB3FEA72563" blockId="57.[189,324,265,291]" box="[189,324,265,291]" pageId="57" pageNumber="58">
<heading id="D0D5567BDA31E776FF23FEB3FEA72563" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B9563D05DA31E776FF23FEB3FEA72563" bold="true" box="[189,324,265,291]" pageId="57" pageNumber="58">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA31E776FF23FE8DFE8A2534" blockId="57.[189,1398,311,372]" pageId="57" pageNumber="58">The epithet is a Latin adjective in the nominative singular meaning ‘hairless’, referring to the glabrous dorsal surface.</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA31E776FF23FE26FD7827C9" pageId="57" pageNumber="58" type="materials_examined">
<paragraph id="8B9DE117DA31E776FF23FE26FE8F25F6" blockId="57.[189,364,412,438]" box="[189,364,412,438]" pageId="57" pageNumber="58">
<heading id="D0D5567BDA31E776FF23FE26FE8F25F6" bold="true" box="[189,364,412,438]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B9563D05DA31E776FF23FE26FE8F25F6" bold="true" box="[189,364,412,438]" pageId="57" pageNumber="58">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA31E776FF74FE71FEBF25A5" blockId="57.[189,1399,459,556]" box="[234,348,459,485]" pageId="57" pageNumber="58">
<emphasis id="B9563D05DA31E776FF74FE71FEBF25A5" bold="true" box="[234,348,459,485]" pageId="57" pageNumber="58">
<typeStatus id="54995FB5DA31E776FF74FE71FEBF25A5" box="[234,348,459,485]" pageId="57" pageNumber="58" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DA31E776FF23FE54FE4F266C" blockId="57.[189,1399,459,556]" pageId="57" pageNumber="58">
<materialsCitation id="3B4AEB4ADA31E776FF23FE54FE4F266C" collectingDate="1996-06-08" collectionCode="NMW" collectorName="Dembicky" country="Vietnam" location="Vinh Phuc Province" pageId="57" pageNumber="58" specimenCount="1" specimenCount-male="1" stateProvince="Vinh Phuc" typeStatus="holotype">
<collectingCountry id="F335A187DA31E776FF23FE54FEA72648" box="[189,324,494,520]" name="Vietnam" pageId="57" pageNumber="58">VIETNAM</collectingCountry>
– 
<emphasis id="B9563D05DA31E776FEF9FE54FD862648" bold="true" box="[359,613,493,520]" pageId="57" pageNumber="58">
<collectingRegion id="49E62FF5DA31E776FEF9FE54FD862648" box="[359,613,493,520]" country="Vietnam" name="Vinh Phuc" pageId="57" pageNumber="58">Vinh Phúc Province</collectingRegion>
</emphasis>
• 
<specimenCount id="9D242A9EDA31E776FD1AFE54FD7E2648" box="[644,669,494,520]" count="1" pageId="57" pageNumber="58" type="male">♂</specimenCount>
; “N-VIETNAM: Tam Dao (2) 1.- 
<date id="FF9CC7D7DA31E776FBB5FE54FB7F2648" box="[1067,1180,494,520]" pageId="57" pageNumber="58" value="1996-06-08">
<collectingDate id="EFD83E3FDA31E776FBB5FE54FB7F2648" box="[1067,1180,494,520]" pageId="57" pageNumber="58" value="1996-06-08">8.6.1996</collectingDate>
</date>
leg. 
<collectorName id="26D784C1DA31E776FB45FE54FAB62648" box="[1243,1365,494,520]" pageId="57" pageNumber="58">Dembicky</collectorName>
&amp; Pacholatko”; 
<collectionCode id="ED3379D2DA31E776FEC2FDA8FE4B266C" box="[348,424,530,556]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="57" pageNumber="58">NMW</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9DE117DA31E776FF74FDE9FE84262D" blockId="57.[189,1399,595,727]" box="[234,359,595,621]" pageId="57" pageNumber="58">
<emphasis id="B9563D05DA31E776FF74FDE9FE84262D" bold="true" box="[234,359,595,621]" pageId="57" pageNumber="58">
<typeStatus id="54995FB5DA31E776FF74FDE9FE84262D" box="[234,359,595,621]" pageId="57" pageNumber="58" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B9DE117DA31E776FF23FDCCFDB42697" blockId="57.[189,1399,595,727]" pageId="57" pageNumber="58">
<materialsCitation id="3B4AEB4ADA31E776FF23FDCCFDC126D0" box="[189,546,630,656]" country="China" location="Guangxi Province" pageId="57" pageNumber="58" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
<collectingCountry id="F335A187DA31E776FF23FDCCFEF926D0" box="[189,282,630,656]" name="China" pageId="57" pageNumber="58">CHINA</collectingCountry>
<emphasis id="B9563D05DA31E776FEBBFDCCFDC126D0" bold="true" box="[293,546,630,656]" pageId="57" pageNumber="58">
– 
<collectingRegion id="49E62FF5DA31E776FEA0FDCCFDC126D0" box="[318,546,630,656]" country="China" name="Guangxi" pageId="57" pageNumber="58">Guangxi Province</collectingRegion>
</emphasis>
</materialsCitation>
<emphasis id="B9563D05DA31E776FDB3FDCCFDD426D0" bold="true" box="[557,567,630,656]" pageId="57" pageNumber="58">•</emphasis>
<materialsCitation id="3B4AEB4ADA31E776FDDDFDCCFDB02697" collectingDate="2013-04-05" collectingDateMax="2013-04-09" collectingDateMin="2013-04-05" collectionCode="NMPC" collectorName="M. Fikacek &amp; J. Hajek &amp; J. Ruzicka" country="China" elevation="325" latitude="21.906666" location="National Forest" longLatPrecision="125" longitude="107.903336" municipality="National Forest Park" pageId="57" pageNumber="58" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="9D242A9EDA31E776FDDDFDCCFD6126D0" box="[579,642,630,656]" count="1" pageId="57" pageNumber="58" type="generic">1 ex.</specimenCount>
; “ 
<collectingCountry id="F335A187DA31E776FD02FDCCFD1D26D0" box="[668,766,630,656]" name="China" pageId="57" pageNumber="58">CHINA</collectingCountry>
, 
<collectingRegion id="49E62FF5DA31E776FC90FDCCFC7426D0" box="[782,919,630,656]" country="China" name="Guangxi" pageId="57" pageNumber="58">GUANGXI</collectingRegion>
A.R., 
<date id="FF9CC7D7DA31E776FC74FDCCFB9526D1" box="[1002,1142,630,657]" pageId="57" pageNumber="58" value="2013-04-05" valueMax="2013-04-09" valueMin="2013-04-05">
<collectingDate id="EFD83E3FDA31E776FC74FDCCFB9526D1" box="[1002,1142,630,657]" pageId="57" pageNumber="58" value="2013-04-05" valueMax="2013-04-09" valueMin="2013-04-05">5.-9.iv.2013</collectingDate>
</date>
SHIWANDA SHAN 
<collectingMunicipality id="6BF97B6DDA31E776FF23FD23FE5B26F3" box="[189,440,665,692]" pageId="57" pageNumber="58">
<location id="8EFDB7CCDA31E776FF23FD23FE9526F4" LSID="urn:lsid:plazi:treatment:038B5001DA3EE775FDF7FB71FD6F27B9:8EFDB7CCDA31E776FF23FD23FE9526F4" box="[189,374,665,692]" country="China" latitude="21.906666" longLatPrecision="125" longitude="107.903336" municipality="National Forest Park" name="National Forest" pageId="57" pageNumber="58" stateProvince="Guangxi">National Forest</location>
Park
</collectingMunicipality>
(forested river valley; in river) 
<geoCoordinate id="EE1687D0DA31E776FCDCFD22FC5E26F4" box="[834,957,664,692]" degrees="21" direction="north" minutes="54.4" orientation="latitude" pageId="57" pageNumber="58" precision="92" value="21.906666">21°54.4′N</geoCoordinate>
, 
<geoCoordinate id="EE1687D0DA31E776FC52FD22FBB126F4" box="[972,1106,664,692]" degrees="107" direction="east" minutes="54.2" orientation="longitude" pageId="57" pageNumber="58" precision="92" value="107.903336">107°54.2′E</geoCoordinate>
, 
<quantity id="4CDA4CF2DA31E776FBFCFD23FB0526F4" box="[1122,1254,665,692]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.6" metricValueMin="2.9" pageId="57" pageNumber="58" unit="m" value="325.0" valueMax="360.0" valueMin="290.0">
<elevation id="000F0624DA31E776FBFCFD23FB0526F4" box="[1122,1254,665,692]" metricMagnitude="2" metricUnit="m" metricValue="3.25" metricValueMax="3.6" metricValueMin="2.9" pageId="57" pageNumber="58" unit="m" value="325.0" valueMax="360.0" valueMin="290.0">290-360 m</elevation>
</quantity>
<collectorName id="26D784C1DA31E776FB6FFD23FA9126F3" box="[1265,1394,665,691]" pageId="57" pageNumber="58">M.Fikáček</collectorName>
, 
<collectorName id="26D784C1DA31E776FF23FD06FEF62696" box="[189,277,700,726]" pageId="57" pageNumber="58">J.Hájek</collectorName>
&amp; 
<collectorName id="26D784C1DA31E776FEA5FD06FE4D2697" box="[315,430,700,727]" pageId="57" pageNumber="58">J.Růžička</collectorName>
leg.”, 
<collectionCode id="ED3379D2DA31E776FE62FD07FDB02697" box="[508,595,701,727]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="57" pageNumber="58">NMPC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9DE117DA31E776FF23FD44FA952759" blockId="57.[189,1398,765,793]" box="[189,1398,765,793]" pageId="57" pageNumber="58">
<materialsCitation id="3B4AEB4ADA31E776FF23FD44FA922759" box="[189,1393,765,793]" collectingDate="1996-06-08" collectionCode="CKB, NMW" collectorName="Dembicky" country="Vietnam" location="Vinh Phuc Province" pageId="57" pageNumber="58" specimenCount="26" specimenCount-female="18" specimenCount-male="8" stateProvince="Vinh Phuc">
<collectingCountry id="F335A187DA31E776FF23FD44FEA72758" box="[189,324,766,792]" name="Vietnam" pageId="57" pageNumber="58">VIETNAM</collectingCountry>
<emphasis id="B9563D05DA31E776FED2FD44FD8F2758" bold="true" box="[332,620,765,792]" pageId="57" pageNumber="58">
– 
<collectingRegion id="49E62FF5DA31E776FEFFFD44FDB92758" box="[353,602,765,792]" country="Vietnam" name="Vinh Phuc" pageId="57" pageNumber="58">Vinh Phúc Province</collectingRegion>
•
</emphasis>
<specimenCount id="9D242A9EDA31E776FDEDFD44FD542759" box="[627,695,766,793]" count="8" pageId="57" pageNumber="58" type="male">8 ♂♂</specimenCount>
, 
<specimenCount id="9D242A9EDA31E776FD5AFD44FCF52759" box="[708,790,766,793]" count="18" pageId="57" pageNumber="58" type="female">18 ♀♀</specimenCount>
; same collection data as for holotype; 
<collectionCode id="ED3379D2DA31E776FB44FD44FAFA2758" box="[1242,1305,766,792]" pageId="57" pageNumber="58">CKB</collectionCode>
, 
<collectionCode id="ED3379D2DA31E776FABBFD45FA922759" box="[1317,1393,767,793]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="57" pageNumber="58">NMW</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9DE117DA31E776FF23FCFAFEBE2719" blockId="57.[189,349,831,858]" box="[189,349,831,858]" pageId="57" pageNumber="58">
<heading id="D0D5567BDA31E776FF23FCFAFEBE2719" bold="true" box="[189,349,831,858]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B9563D05DA31E776FF23FCFAFEBE2719" bold="true" box="[189,349,831,858]" pageId="57" pageNumber="58">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA31E776FF23FCD4FD7827C9" blockId="57.[189,667,878,905]" box="[189,667,878,905]" pageId="57" pageNumber="58">
<collectingCountry id="F335A187DA31E776FF23FCD4FEC027C8" box="[189,291,878,904]" name="Vietnam" pageId="57" pageNumber="58">Vietnam</collectingCountry>
, 
<collectingRegion id="49E62FF5DA31E776FEB0FCD4FDFF27C8" box="[302,540,878,904]" country="Vietnam" name="Vinh Phuc" pageId="57" pageNumber="58">Vinh Phúc Province</collectingRegion>
, Tam Dao.
</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA31E775FF23FC15FDE227C8" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58" type="description">
<paragraph id="8B9DE117DA31E776FF23FC15FEAD2789" blockId="57.[189,334,943,969]" box="[189,334,943,969]" pageId="57" pageNumber="58">
<heading id="D0D5567BDA31E776FF23FC15FEAD2789" bold="true" box="[189,334,943,969]" fontSize="11" level="3" pageId="57" pageNumber="58" reason="3">
<emphasis id="B9563D05DA31E776FF23FC15FEAD2789" bold="true" box="[189,334,943,969]" pageId="57" pageNumber="58">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA31E776FF23FC65FC5020C6" blockId="57.[189,1399,990,1158]" pageId="57" pageNumber="58">
Measurements (mm): TL: 
<specimenCount id="9D242A9EDA31E776FE71FC65FDF827B9" box="[495,539,991,1017]" count="1" pageId="57" pageNumber="58" type="male">♂♂</specimenCount>
3.17–3.35 (3.25 ±0.06, n=7), 
<specimenCount id="9D242A9EDA31E776FCE3FC65FC4A27B9" box="[893,937,991,1017]" count="1" pageId="57" pageNumber="58" type="female">♀♀</specimenCount>
3.15–3.45 (3.30 ± 0.11, n =10); PL: 
<specimenCount id="9D242A9EDA31E776FAD4FC65FA9527B9" box="[1354,1398,991,1017]" count="1" pageId="57" pageNumber="58" type="male">♂♂</specimenCount>
0.83–0.91 (0.86± 0.03, n= 7), 
<specimenCount id="9D242A9EDA31E776FDAEFBB8FDBF205C" box="[560,604,1026,1052]" count="1" pageId="57" pageNumber="58" type="female">♀♀</specimenCount>
0.83–0.91 (0.87± 0.03, n=10); PW: 
<specimenCount id="9D242A9EDA31E776FBB5FBB8FBB4205C" box="[1067,1111,1026,1052]" count="1" pageId="57" pageNumber="58" type="male">♂♂</specimenCount>
1.27–1.38 (1.33± 0.04, n= 7), 
<specimenCount id="9D242A9EDA31E776FE8DFB9FFEDC207F" box="[275,319,1061,1087]" count="1" pageId="57" pageNumber="58" type="female">♀♀</specimenCount>
1.29–1.45 (1.38± 0.06, n= 10); EL: 
<specimenCount id="9D242A9EDA31E776FC8CFB9FFCDD207F" box="[786,830,1061,1087]" count="1" pageId="57" pageNumber="58" type="male">♂♂</specimenCount>
2.34–2.47 (2.39± 0.04, n= 7), 
<specimenCount id="9D242A9EDA31E776FB58FB9FFB11207F" box="[1222,1266,1061,1087]" count="1" pageId="57" pageNumber="58" type="female">♀♀</specimenCount>
2.31–2.54 (2.43 ± 0.08, n=10); EW: 
<specimenCount id="9D242A9EDA31E776FE6DFBF3FDFC2023" box="[499,543,1097,1123]" count="1" pageId="57" pageNumber="58" type="male">♂♂</specimenCount>
1.46–1.59 (1.53±0.04, n=7), 
<specimenCount id="9D242A9EDA31E776FC0FFBF3FC5E2023" box="[913,957,1097,1123]" count="1" pageId="57" pageNumber="58" type="female">♀♀</specimenCount>
1.50–1.66 (1.59± 0.06, n= 10); PhL: 1.12–1.21 (1.17± 0.03, n =7); PrL: 0.40–0.49 (0.45 ±0.03, n =7).
</paragraph>
<paragraph id="8B9DE117DA31E776FF23FB17FC9D2137" blockId="57.[189,1399,1197,1400]" pageId="57" pageNumber="58">
Body ovate (
<figureCitation id="1319FD92DA31E776FECAFB17FE482087" box="[340,427,1197,1223]" captionStart="Fig" captionStartId="55.[189,232,1969,1995]" captionTargetBox="[189,1399,265,1928]" captionTargetId="figure-12@55.[189,1399,265,1928]" captionTargetPageId="55" captionText="Fig. 23. Elmomorphus calvus sp. nov., holotype, male (NMW), TL: 3.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13773441" httpUri="https://zenodo.org/record/13773441/files/figure.png" pageId="57" pageNumber="58">Fig. 23</figureCitation>
), widest behind midlength of elytra, strongly convex dorsally, highest point at first quarter of elytral length. Colouration black, except for reddish brown mouthparts, antennae, trochanters, and tarsi, and brown proximal portions of tibiae. Vestiture consisting of short decumbent yellowish setae. Plastron present on cranium (except for semicircular area on vertex), on anterolateral portions of pronotum, and on lateral elytral bands. Ventral surface with thin plastron setae, except on prosternal process, and median parts of metaventrite and ventrites 1–2.
</paragraph>
<paragraph id="8B9DE117DA31E776FF23FA25FE7C22CD" blockId="57.[189,1399,1439,1677]" pageId="57" pageNumber="58">
Head surface covered with plastron, except for semicircular area on vertex; with round setiferous punctures slightly smaller than eye facet, separated by 0.5–1.0 × facet diameter. Labrum transverse, anterior margin concave; surface microreticulate, with small punctures; males with conspicuous long setae, slightly shorter than terminal maxillary palpomere. Anterior margin of clypeus straight, with row of short dense setae. Eyes large, oval, protruding, interfacetal setae short; ID: 
<specimenCount id="9D242A9EDA31E776FB08F997FB212207" box="[1174,1218,1581,1607]" count="1" pageId="57" pageNumber="58" type="male">♂♂</specimenCount>
<quantity id="4CDA4CF2DA31E776FB55F996FA942207" box="[1227,1399,1580,1607]" metricMagnitude="-4" metricUnit="m" metricValue="4.95" metricValueMax="5.1" metricValueMin="4.8" pageId="57" pageNumber="58" unit="mm" value="0.495" valueMax="0.51" valueMin="0.48">0.48–0.51 mm</quantity>
(0.49 ± 0.01, n=7), 
<specimenCount id="9D242A9EDA31E776FE30F9EAFE39222A" box="[430,474,1616,1642]" count="1" pageId="57" pageNumber="58" type="female">♀♀</specimenCount>
<quantity id="4CDA4CF2DA31E776FE76F9F5FD7A222A" box="[488,665,1615,1642]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.4" metricValueMin="5.2" pageId="57" pageNumber="58" unit="mm" value="0.53" valueMax="0.54" valueMin="0.52">0.52–0.54 mm</quantity>
(0.52 ± 0.01, n= 4). Antennae short, 10-segmented, densely covered with setae.
</paragraph>
<paragraph id="8B9DE117DA31E775FF23F90FFDF325F3" blockId="57.[189,1399,1716,2025]" lastBlockId="58.[189,1399,265,435]" lastPageId="58" lastPageNumber="59" pageId="57" pageNumber="58">
Pronotum strongly convex, wider than long, lateral sides distinctly rounded, PW/PL: 
<specimenCount id="9D242A9EDA31E776FB54F90FFB15228F" box="[1226,1270,1717,1743]" count="1" pageId="57" pageNumber="58" type="male">♂♂</specimenCount>
1.46–1.63 (1.55± 0.05, n =7), 
<specimenCount id="9D242A9EDA31E776FE39F962FE3022B2" box="[423,467,1752,1778]" count="1" pageId="57" pageNumber="58" type="female">♀♀</specimenCount>
1.54–1.65 (1.58± 0.04, n=10); surface smooth with round punctures slightly wider than those on head, separated by about half a puncture diameter; plastron present anterolaterally. Lateral margins of prosternal process straight and slightly raised, with conspicuous cluster of long erect setae in males; median keel flat; posterior edge rounded. Lateral margins of metaventral process raised; metaventral disc depressed in males, with two clusters of long and erect setae posteriorly, in females flat and without such clusters; glabrous area usually widening posteriorly. Scutellum wider than long, smooth, with several setiferous punctures. Elytra strongly convex, widest behind middle, EL/EW: 
<specimenCount id="9D242A9EDA31E776FB51F816FB182386" box="[1231,1275,1964,1990]" count="1" pageId="57" pageNumber="58" type="male">♂♂</specimenCount>
1.52–1.61 (1.57 ± 0.03, n =7), 
<specimenCount id="9D242A9EDA31E776FE39F875FE3023A9" box="[423,467,1999,2025]" count="1" pageId="57" pageNumber="58" type="female">♀♀</specimenCount>
1.47–1.57 (1.52 ±0.03, n =10); small punctures scattered over entire surface, large punctures arranged in nine well impressed striae, strial punctures confluent; plastron confined to two lateral bands, each extending from elytral side to fourth elytral stria. Tibiae straight, protibia ca 1.3× as long as protarsus, PrTL/PL: 
<specimenCount id="9D242A9EDA32E775FDA8FEE8FD81252C" box="[566,610,338,364]" count="1" pageId="58" pageNumber="59" type="male">♂♂</specimenCount>
0.95–1.08 (1.04 ±0.05, n=7), 
<specimenCount id="9D242A9EDA32E775FC4AFEE8FBE3252C" box="[980,1024,338,364]" count="1" pageId="58" pageNumber="59" type="female">♀♀</specimenCount>
0.95–1.05 (0.99± 0.03, n =10). Protarsus with terminal tarsomere nearly as long as all preceding segments combined; foreclaws long and slender, strongly curved.
</paragraph>
<paragraph id="8B9DE117DA32E775FF23FE61FEDC268E" blockId="58.[189,1399,475,718]" pageId="58" pageNumber="59">
Ventrites with plastron, except for median portions of ventrites 1–2. Intercoxal process wide, lateral margins raised, admedian keels distinct. Ventrite 5 evenly convex in both sexes, in males with two clusters of long, erect setae before apex, in females with short longitudinal subapical keel. Aedeagus (
<figureCitation id="1319FD92DA32E775FF58FDF2FEC12622" box="[198,290,584,610]" captionStart="Fig" captionStartId="56.[189,232,1888,1914]" captionTargetBox="[296,1282,278,1832]" captionTargetId="figure-21@56.[277,1295,265,1839]" captionTargetPageId="56" captionText="Fig. 24. Elmomorphus calvus sp. nov., holotype (NMW), aedeagus. A. Lateral aspect. B. Ventral aspect. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773445" httpUri="https://zenodo.org/record/13773445/files/figure.png" pageId="58" pageNumber="59">Fig. 24</figureCitation>
): phallobase remarkably long and robust, strongly curved, PhL/PrL: 2.34–2.84 (2.62 ± 0.18, n= 7); parameres rather short, slightly curved ventrad, apices narrowly rounded (lateral aspect); penis curved ventrad apically; sclerotised fibula long and wide. Bursa copulatrix without sclerotised spines (
<figureCitation id="1319FD92DA32E775FF58FD0EFED2268E" box="[198,305,692,718]" captionStart="Fig" captionStartId="64.[189,232,1829,1855]" captionTargetBox="[242,1354,287,1780]" captionTargetId="figure-40@64.[223,1369,265,1788]" captionTargetPageId="64" captionText="Fig. 28. Bursa copulatrix and vagina. A. Elmomorphus elmoides sp. nov., paratype from type locality (NMW). B. E. calvus sp. nov., paratype from type locality (CKB). C. E. depressus sp. nov., holotype (IAECAS). Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13773457" httpUri="https://zenodo.org/record/13773457/files/figure.png" pageId="58" pageNumber="59">Fig. 28B</figureCitation>
).
</paragraph>
<paragraph id="8B9DE117DA32E775FF23FD4CFDD42750" blockId="58.[189,567,758,784]" box="[189,567,758,784]" pageId="58" pageNumber="59">
<heading id="D0D5567BDA32E775FF23FD4CFDD42750" bold="true" box="[189,567,758,784]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="B9563D05DA32E775FF23FD4CFDD42750" bold="true" box="[189,567,758,784]" pageId="58" pageNumber="59">Secondary sexual dimorphism</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA32E775FF23FC9FFDE227C8" blockId="58.[189,1399,805,904]" pageId="58" pageNumber="59">Males with conspicuous long erect setae on labrum, prosternal process, median part of metaventrite and on ventrite 5; metaventral disc depressed. In females, metaventral disc flat; ventrite 5 with short longitudinal subapical keel.</paragraph>
</subSubSection>
<subSubSection id="C338B29CDA32E775FF23FC0AFD6F27B9" box="[189,652,944,1017]" pageId="58" pageNumber="59" type="distribution">
<paragraph id="8B9DE117DA32E775FF23FC0AFEB5278A" blockId="58.[189,342,944,970]" box="[189,342,944,970]" pageId="58" pageNumber="59">
<heading id="D0D5567BDA32E775FF23FC0AFEB5278A" bold="true" box="[189,342,944,970]" fontSize="11" level="3" pageId="58" pageNumber="59" reason="3">
<emphasis id="B9563D05DA32E775FF23FC0AFEB5278A" bold="true" box="[189,342,944,970]" pageId="58" pageNumber="59">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8B9DE117DA32E775FF23FC65FD6F27B9" blockId="58.[189,652,991,1017]" box="[189,652,991,1017]" pageId="58" pageNumber="59">
<collectingCountry id="F335A187DA32E775FF23FC65FEE027B9" box="[189,259,991,1017]" name="China" pageId="58" pageNumber="59">China</collectingCountry>
(
<collectingRegion id="49E62FF5DA32E775FE8DFC65FE9D27B9" box="[275,382,991,1017]" country="China" name="Guangxi" pageId="58" pageNumber="59">Guangxi</collectingRegion>
), 
<collectingCountry id="F335A187DA32E775FE0FFC65FE1727B9" box="[401,500,991,1017]" name="Vietnam" pageId="58" pageNumber="59">Vietnam</collectingCountry>
(
<figureCitation id="1319FD92DA32E775FD9AFC65FD9C27B9" box="[516,639,991,1017]" captionStart="Fig" captionStartId="217.[189,232,1872,1898]" captionTargetBox="[254,1334,266,1829]" captionTargetId="figure-44@217.[189,1399,265,1830]" captionTargetPageId="217" captionText="Fig. 109. Distribution maps. A. Elmomorphus fusiformis sp. nov. B. E. elmoides sp. nov. C. E. calvus sp. nov. D. E. hongkong sp. nov. E. E. depressus sp. nov. F. E. sulcatus Kodada, Selnekovič &amp; Jäch sp. nov." figureDoi="http://doi.org/10.5281/zenodo.13773712" httpUri="https://zenodo.org/record/13773712/files/figure.png" pageId="58" pageNumber="59">Fig. 109C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>