304 lines
36 KiB
XML
304 lines
36 KiB
XML
<document id="129CFCD903B44B5E3F781B68A7C72506" ID-CLB-Dataset="45293" ID-DOI="10.11646/zootaxa.3223.1.4" ID-GBIF-Dataset="8088247b-16b1-48e8-b65f-2690084e90f4" ID-ISSN="1175-5326" ID-Zenodo-Dep="210833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460449353620" checkinUser="plazi" docAuthor="Jia, Fenglong, Zhao, Shuang & Fery, Hans" docDate="2012" docId="952587BC1C75FF9AFF32FCD86022FE78" docLanguage="en" docName="zt03223p067.pdf" docOrigin="Zootaxa 3223 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.3223.1.4" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hydroporus polaris Fall 1923" docType="treatment" docVersion="12" lastPageNumber="64" masterDocId="691CFFC41C7DFF93FFA5FFC1647FFF8C" masterDocTitle="Hydroporus sejilashan sp. n., a new diving beetle of the acutangulus - complex from Xizang, China (Qinghai-Tibet Plateau), and notes on other taxa of the genus (Coleoptera, Dytiscidae, Hydroporinae)" masterLastPageNumber="67" masterPageNumber="55" pageNumber="63" updateTime="1698338074243" updateUser="ExternalLinkService">
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<mods:title id="76B686AD092982CCC16FC41A72390B2B">Hydroporus sejilashan sp. n., a new diving beetle of the acutangulus - complex from Xizang, China (Qinghai-Tibet Plateau), and notes on other taxa of the genus (Coleoptera, Dytiscidae, Hydroporinae)</mods:title>
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<mods:namePart id="A220964FCA3F19CC104C2FD7E840F997">Jia, Fenglong</mods:namePart>
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<mods:namePart id="0C3F1D4B29481DE50F355637C7B5BECB">Zhao, Shuang</mods:namePart>
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<mods:namePart id="7363C4C3C4A1F2082D5B5077866E5672">Fery, Hans</mods:namePart>
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<mods:title id="7DF9F3D820FC48130CE36A18868BB8EB">Zootaxa</mods:title>
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<mods:date id="643451AF48F362572D61CC644434203E">2012</mods:date>
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<mods:identifier id="9341DE4562C126CA91DB59ADFB9FAD8F" type="DOI">10.11646/zootaxa.3223.1.4</mods:identifier>
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<treatment id="952587BC1C75FF9AFF32FCD86022FE78" ID-DOI="http://doi.org/10.5281/zenodo.5914741" ID-GBIF-Taxon="119510187" ID-Zenodo-Dep="5914741" LSID="urn:lsid:plazi:treatment:952587BC1C75FF9AFF32FCD86022FE78" httpUri="http://treatment.plazi.org/id/952587BC1C75FF9AFF32FCD86022FE78" lastPageId="9" lastPageNumber="64" pageId="8" pageNumber="63">
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<subSubSection id="559665211C75FF9BFF32FCD8667CFCBE" box="[151,515,792,819]" pageId="8" pageNumber="63" type="nomenclature">
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<paragraph id="1D3336AA1C75FF9BFF32FCD8667CFCBE" blockId="8.[151,515,792,819]" box="[151,515,792,819]" pageId="8" pageNumber="63">
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<heading id="467B81C61C75FF9BFF32FCD8667CFCBE" bold="true" box="[151,515,792,819]" fontSize="11" level="1" pageId="8" pageNumber="63" reason="1">
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<taxonomicName id="DA8C4D291C75FF9BFF32FCD8667CFCBE" ID-CoL="6MJH4" authority="Fall, 1923" authorityName="Fall" authorityYear="1923" box="[151,515,792,819]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
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<emphasis id="2FF8EAB81C75FF9BFF32FCD8667CFCBE" bold="true" box="[151,515,792,819]" pageId="8" pageNumber="63">
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<emphasis id="2FF8EAB81C75FF9BFF32FCD865FDFCBF" bold="true" box="[151,386,793,819]" italics="true" pageId="8" pageNumber="63">Hydroporus polaris</emphasis>
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<bibRefCitation id="791D4B5B1C75FF9BFE2CFCD9667CFCBE" author="Fall" box="[393,515,792,818]" pageId="8" pageNumber="63" refString="Fall, H. C. (1923) A revision of the North American species of Hydroporus and Agaporus. Privately printed, 129 pp." type="book" year="1923">Fall, 1923</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="559665211C75FF9AFF32FC986022FE78" lastPageId="9" lastPageNumber="64" pageId="8" pageNumber="63" type="discussion">
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<paragraph id="1D3336AA1C75FF9BFF32FC98612CFA01" blockId="8.[151,1437,857,2033]" pageId="8" pageNumber="63">
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<bibRefCitation id="791D4B5B1C75FF9BFF32FC986541FCFD" author="Fall" box="[151,318,857,881]" pageId="8" pageNumber="63" refString="Fall, H. C. (1923) A revision of the North American species of Hydroporus and Agaporus. Privately printed, 129 pp." type="book" year="1923">Fall (1923: 92)</bibRefCitation>
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described this species from two males and one female from Bernard Harbour (Nunavut,
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<collectingCountry id="659B763A1C75FF9BFABEFC98610DFCFD" box="[1307,1394,857,881]" name="Canada" pageId="8" pageNumber="63">Canada</collectingCountry>
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, ca.
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<geoCoordinate id="78B8506D1C75FF9BFF32FCBC6487FC19" box="[151,248,893,917]" direction="north" orientation="latitude" pageId="8" pageNumber="63" precision="55" value="68.788">68.788N</geoCoordinate>
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<geoCoordinate id="78B8506D1C75FF9BFF5AFCBC6516FC19" box="[255,361,893,917]" direction="west" orientation="longitude" pageId="8" pageNumber="63" precision="555" value="-114.82">114.82W</geoCoordinate>
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; square
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<quantity id="DA749B4F1C75FF9BFE66FCBC6591FC19" box="[451,494,893,917]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="8" pageNumber="63" unit="in" value="9.0">9 in</quantity>
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<figureCitation id="85B72A2F1C75FF9BFE50FCBC6638FC19" box="[501,583,893,917]" captionStart="FIGURE 13" captionStartId="10.[151,250,932,955]" captionTargetBox="[264,1322,233,911]" captionTargetId="figure@10.[264,1322,233,911]" captionTargetPageId="10" captionText="FIGURE 13. Distribution of: Hydroporus acutangulus (circles), Hydroporus polaris (squares), Hydroporus tuvaensis (triangle), Hydroporus tibetanus (stars), and Hydroporus sejilashan sp. n. (encircled star); the circle with question mark refers to the type locality of Hydroporus sumakovi (see text). Red squares indicate capitals of countries / provinces or other important and well-known cities." httpUri="https://zenodo.org/record/210837/files/figure.png" pageId="8" pageNumber="63">Fig. 13</figureCitation>
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). In the same work
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<bibRefCitation id="791D4B5B1C75FF9BFC80FCBC67B3FC19" author="Fall" box="[805,972,893,917]" pageId="8" pageNumber="63" refString="Fall, H. C. (1923) A revision of the North American species of Hydroporus and Agaporus. Privately printed, 129 pp." type="book" year="1923">Fall (1923: 93)</bibRefCitation>
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described
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<taxonomicName id="DA8C4D291C75FF9BFBE3FCBC611DFC19" box="[1094,1378,893,917]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="subvirescens">
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<emphasis id="2FF8EAB81C75FF9BFBE3FCBC611DFC19" box="[1094,1378,893,917]" italics="true" pageId="8" pageNumber="63">Hydroporus subvirescens</emphasis>
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</taxonomicName>
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after specimens collected at Cape Collinson (Alaska,
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<collectingCountry id="659B763A1C75FF9BFD0AFC606695FC35" box="[687,746,929,953]" name="United States of America" pageId="8" pageNumber="63">USA</collectingCountry>
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, ca.
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<geoCoordinate id="78B8506D1C75FF9BFCBEFC606703FC35" box="[795,892,929,953]" direction="north" orientation="latitude" pageId="8" pageNumber="63" precision="55" value="69.986">69.986N</geoCoordinate>
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; square
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<quantity id="DA749B4F1C75FF9BFBF0FC606000FC35" box="[1109,1151,929,953]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="8" pageNumber="63" unit="in" value="8.0">8 in</quantity>
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<figureCitation id="85B72A2F1C75FF9BFB20FC6060A9FC35" box="[1157,1238,929,953]" captionStart="FIGURE 13" captionStartId="10.[151,250,932,955]" captionTargetBox="[264,1322,233,911]" captionTargetId="figure@10.[264,1322,233,911]" captionTargetPageId="10" captionText="FIGURE 13. Distribution of: Hydroporus acutangulus (circles), Hydroporus polaris (squares), Hydroporus tuvaensis (triangle), Hydroporus tibetanus (stars), and Hydroporus sejilashan sp. n. (encircled star); the circle with question mark refers to the type locality of Hydroporus sumakovi (see text). Red squares indicate capitals of countries / provinces or other important and well-known cities." httpUri="https://zenodo.org/record/210837/files/figure.png" pageId="8" pageNumber="63">Fig. 13</figureCitation>
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). He provided for this taxon the following features: "with evident viridi-aeneous surface lustre" on the elytra and a "distinctly alutaceous and dull" upper surface. Additionally, he wrote: "Front tarsi of male moderately dilated, its anterior claw not appreciably different from the posterior." We have studied the male
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<typeStatus id="C23788081C75FF9BFC72FBCC6046FBA9" box="[983,1081,1037,1061]" pageId="8" pageNumber="63" type="holotype">holotype</typeStatus>
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and the female
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<typeStatus id="C23788081C75FF9BFB53FBCF6129FBAA" box="[1270,1366,1038,1062]" pageId="8" pageNumber="63" type="paratype">paratype</typeStatus>
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of
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<taxonomicName id="DA8C4D291C75FF9BFA24FBCF6499FBC5" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
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<emphasis id="2FF8EAB81C75FF9BFA24FBCF6499FBC5" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
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</taxonomicName>
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as well as the male
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<typeStatus id="C23788081C75FF9BFE7CFBF06644FBC5" box="[473,571,1073,1097]" pageId="8" pageNumber="63" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="DA8C4D291C75FF9BFDC3FBF36766FBC5" box="[614,793,1073,1097]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="subvirescens">
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<emphasis id="2FF8EAB81C75FF9BFDC3FBF36766FBC5" box="[614,793,1073,1097]" italics="true" pageId="8" pageNumber="63">H. subvirescens</emphasis>
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</taxonomicName>
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(all in CNC). To our great surprise, the
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<typeStatus id="C23788081C75FF9BFB51FBF06129FBC5" box="[1268,1366,1073,1097]" pageId="8" pageNumber="63" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="DA8C4D291C75FF9BFA24FBF36499FBE1" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
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<emphasis id="2FF8EAB81C75FF9BFA24FBF36499FBE1" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
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</taxonomicName>
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has a "distinctly alutaceous and dull" upper surface with a well visible metallic sheen on the elytra (sic!), while the
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<typeStatus id="C23788081C75FF9BFEAEFBB86512FB1D" box="[267,365,1145,1169]" pageId="8" pageNumber="63" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="DA8C4D291C75FF9BFE3DFBBB6633FB1D" box="[408,588,1145,1169]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="subvirescens">
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<emphasis id="2FF8EAB81C75FF9BFE3DFBBB6633FB1D" box="[408,588,1145,1169]" italics="true" pageId="8" pageNumber="63">H. subvirescens</emphasis>
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</taxonomicName>
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has a shiny (although reticulate) surface and does not show any metallic sheen. In addition, both male
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<typeStatus id="C23788081C75FF9BFE40FB5C662DFB39" box="[485,594,1181,1205]" pageId="8" pageNumber="63" type="holotype">holotypes</typeStatus>
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have the anterior protarsal claws clearly strongly curved near the base and in the more distal part straight. While the appearance of the shape of the protarsal claws depends somewhat on their orientation and can be misinterpreted, this is not true of the reticulation and sheen of the dorsal surface. Thus, we must assume that either Fall himself or any person who dealt with these
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<typeStatus id="C23788081C75FF9BFBA4FACB6043FAAE" box="[1025,1084,1290,1314]" pageId="8" pageNumber="63">types</typeStatus>
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after Fall has mixed up both
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<typeStatus id="C23788081C75FF9BFF32FAEC657BFAC9" box="[151,260,1325,1349]" pageId="8" pageNumber="63" type="holotype">holotypes</typeStatus>
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by mistake. But we made yet another interesting observation: the dull appearance and the metallic sheen can be easily removed mechanically, for instance by means of a brush with hard hairs (tested in a very small area of the elytra). We agree with
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<bibRefCitation id="791D4B5B1C75FF9BFE1AFAB766E7FA01" author="Larson" box="[447,664,1397,1421]" pageId="8" pageNumber="63" refString="Larson, D. J., Alarie, Y. & Roughley, R. E. (2000) Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. Ottawa: NRC Research Press, 982 pp." type="book" year="2000">
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Larson
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<emphasis id="2FF8EAB81C75FF9BFDB1FAB7663AFA01" box="[532,581,1397,1421]" italics="true" pageId="8" pageNumber="63">et al</emphasis>
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. (2000
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</bibRefCitation>
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: 348) who have synonymised
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<taxonomicName id="DA8C4D291C75FF9BFC4EFAB760E4FA01" box="[1003,1179,1397,1421]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="subvirescens">
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<emphasis id="2FF8EAB81C75FF9BFC4EFAB760E4FA01" box="[1003,1179,1397,1421]" italics="true" pageId="8" pageNumber="63">H. subvirescens</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="DA8C4D291C75FF9BFB7EFAB76133FA01" box="[1243,1356,1397,1421]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
|
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<emphasis id="2FF8EAB81C75FF9BFB7EFAB76133FA01" box="[1243,1356,1397,1421]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="1D3336AA1C75FF9BFF62FA586142F905" blockId="8.[151,1437,857,2033]" pageId="8" pageNumber="63">
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One of the main differences between
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<taxonomicName id="DA8C4D291C75FF9BFDDAFA5B668BFA3D" box="[639,756,1433,1457]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
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<emphasis id="2FF8EAB81C75FF9BFDDAFA5B668BFA3D" box="[639,756,1433,1457]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="DA8C4D291C75FF9BFC97FA5B679FFA3D" box="[818,992,1433,1457]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="acutangulus">
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<emphasis id="2FF8EAB81C75FF9BFC97FA5B679FFA3D" box="[818,992,1433,1457]" italics="true" pageId="8" pageNumber="63">H. acutangulus</emphasis>
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</taxonomicName>
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is the reticulation which is present in males and females of the former on the entire upper and ventral surface, while only present in some females of the latter. The colouration of the upper surface is varying between lighter and darker brown in
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<taxonomicName id="DA8C4D291C75FF9BFB2FFA236084FA75" box="[1162,1275,1505,1529]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
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<emphasis id="2FF8EAB81C75FF9BFB2FFA2360E1FA75" box="[1162,1182,1506,1529]" italics="true" pageId="8" pageNumber="63">H</emphasis>
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.
|
||
<emphasis id="2FF8EAB81C75FF9BFB09FA206084FA75" box="[1196,1275,1505,1529]" italics="true" pageId="8" pageNumber="63">polaris</emphasis>
|
||
</taxonomicName>
|
||
, but generally more brownish than in the mostly blackish
|
||
<taxonomicName id="DA8C4D291C75FF9BFDDEF9C76759F991" box="[635,806,1541,1565]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="2FF8EAB81C75FF9BFDDEF9C76759F991" box="[635,806,1541,1565]" italics="true" pageId="8" pageNumber="63">H. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
(compare
|
||
<figureCitation id="85B72A2F1C75FF9BFC38F9C46077F991" box="[925,1032,1541,1565]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1824,1847]" captionTargetBox="[284,1307,543,1722]" captionTargetId="figure@2.[264,1323,523,1809]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: (1) Hydroporus acutangulus, (2) Hydroporus polaris, (3) Hydroporus tuvaensis, (4) Hydroporus tibetanus (lectotype), (5) Hydroporus sejilashan sp. n." httpUri="https://zenodo.org/record/210834/files/figure.png" pageId="8" pageNumber="63">Figs. 1–2</figureCitation>
|
||
). Additionally, we want to report an observation that we have not found in the literature: the shape of the metacoxal lines is rather variable in
|
||
<taxonomicName id="DA8C4D291C75FF9BFA86F9EB61EBF9CD" box="[1315,1428,1577,1601]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C75FF9BFA86F9EB61EBF9CD" box="[1315,1428,1577,1601]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
; sometimes, they are only slightly impressed far before reaching the posterior margin of the metaventrite and often vanish there. If they reach that margin, they are in some specimens converging anteriorly, and not parallel.
|
||
</paragraph>
|
||
<paragraph id="1D3336AA1C75FF9BFF62F95461E3F87D" blockId="8.[151,1437,857,2033]" pageId="8" pageNumber="63">
|
||
<bibRefCitation id="791D4B5B1C75FF9BFF62F9546668F921" author="Nilsson" box="[199,535,1685,1709]" pageId="8" pageNumber="63" refString="Nilsson, A. N. & Holmen M. (1995) The aquatic Adephaga (Coleoptera) of Fennoscandia and Denmark. II. Dytiscidae. Fauna Entomologica Scandinavica, 32, 1 - 192." type="journal article" year="1995">Nilsson & Holmen (1995: 48)</bibRefCitation>
|
||
suspected that
|
||
<taxonomicName id="DA8C4D291C75FF9BFD61F9576710F921" box="[708,879,1685,1709]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="2FF8EAB81C75FF9BFD61F9576710F921" box="[708,879,1685,1709]" italics="true" pageId="8" pageNumber="63">H. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DA8C4D291C75FF9BFC00F9576068F921" box="[933,1047,1685,1709]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C75FF9BFC00F9576068F921" box="[933,1047,1685,1709]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
might be conspecific and reported
|
||
<taxonomicName id="DA8C4D291C75FF9BFF32F97B6576F95D" box="[151,265,1721,1745]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C75FF9BFF32F97B6576F95D" box="[151,265,1721,1745]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
from the "extreme north-east of Siberia (Wrangel Island, coll. G. Lafer)" (square
|
||
<quantity id="DA749B4F1C75FF9BFB02F97B60ACF95D" box="[1191,1235,1721,1745]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="8" pageNumber="63" unit="in" value="7.0">7 in</quantity>
|
||
<figureCitation id="85B72A2F1C75FF9BFB7EF9786152F95D" box="[1243,1325,1721,1745]" captionStart="FIGURE 13" captionStartId="10.[151,250,932,955]" captionTargetBox="[264,1322,233,911]" captionTargetId="figure@10.[264,1322,233,911]" captionTargetPageId="10" captionText="FIGURE 13. Distribution of: Hydroporus acutangulus (circles), Hydroporus polaris (squares), Hydroporus tuvaensis (triangle), Hydroporus tibetanus (stars), and Hydroporus sejilashan sp. n. (encircled star); the circle with question mark refers to the type locality of Hydroporus sumakovi (see text). Red squares indicate capitals of countries / provinces or other important and well-known cities." httpUri="https://zenodo.org/record/210837/files/figure.png" pageId="8" pageNumber="63">Fig. 13</figureCitation>
|
||
) which is "identical with Nearctic material".
|
||
<bibRefCitation id="791D4B5B1C75FF9BFDE6F91F6746F979" author="Larson" box="[579,825,1757,1781]" pageId="8" pageNumber="63" refString="Larson, D. J., Alarie, Y. & Roughley, R. E. (2000) Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. Ottawa: NRC Research Press, 982 pp." type="book" year="2000">
|
||
Larson
|
||
<emphasis id="2FF8EAB81C75FF9BFD04F91F66A3F979" box="[673,732,1757,1781]" italics="true" pageId="8" pageNumber="63">et al</emphasis>
|
||
. (2000
|
||
</bibRefCitation>
|
||
: 350) listed several characters which separate
|
||
<taxonomicName id="DA8C4D291C75FF9BFA24F91F655FF895" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="2FF8EAB81C75FF9BFA24F91F655FF895" italics="true" pageId="8" pageNumber="63">H. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="DA8C4D291C75FF9BFEC3F8C365A6F895" box="[358,473,1793,1817]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C75FF9BFEC3F8C365A6F895" box="[358,473,1793,1817]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
and conclude: "Unless it is shown that these characters vary clinally across northern Asia, the differences are sufficient to justify the recognition of two species." We would refrain from confirming the thesis that
|
||
<taxonomicName id="DA8C4D291C75FF9BFEAAF88B65FFF8ED" box="[271,384,1865,1889]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C75FF9BFEAAF88B65FFF8ED" box="[271,384,1865,1889]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DA8C4D291C75FF9BFE12F88B661DF8ED" box="[439,610,1865,1889]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="2FF8EAB81C75FF9BFE12F88B661DF8ED" box="[439,610,1865,1889]" italics="true" pageId="8" pageNumber="63">H. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
are not conspecific, we can confirm, however, most statements of
|
||
<bibRefCitation id="791D4B5B1C75FF9BFAEBF88B6562F809" author="Larson" pageId="8" pageNumber="63" refString="Larson, D. J., Alarie, Y. & Roughley, R. E. (2000) Predaceous diving beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. Ottawa: NRC Research Press, 982 pp." type="book" year="2000">
|
||
Larson
|
||
<emphasis id="2FF8EAB81C75FF9BFF32F8AF64B6F809" box="[151,201,1901,1925]" italics="true" pageId="8" pageNumber="63">et al</emphasis>
|
||
. (2000
|
||
</bibRefCitation>
|
||
: 349, 350), except one:
|
||
<taxonomicName id="DA8C4D291C75FF9BFD88F8AF66A6F809" box="[557,729,1901,1925]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="2FF8EAB81C75FF9BFD88F8AF66A6F809" box="[557,729,1901,1925]" italics="true" pageId="8" pageNumber="63">H. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
appears not to be broader than
|
||
<taxonomicName id="DA8C4D291C75FF9BFBE5F8AF60CDF809" box="[1088,1202,1901,1925]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C75FF9BFBE5F8AF60CDF809" box="[1088,1202,1901,1925]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
, on the contrary, the latter species appears to be broader, and this is due chiefly to the more rounded body shape of
|
||
<taxonomicName id="DA8C4D291C75FF9BFB1DF8536156F825" box="[1208,1321,1937,1961]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C75FF9BFB1DF8536156F825" box="[1208,1321,1937,1961]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<figureCitation id="85B72A2F1C75FF9BFAC0F85064BCF841" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1824,1847]" captionTargetBox="[284,1307,543,1722]" captionTargetId="figure@2.[264,1323,523,1809]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: (1) Hydroporus acutangulus, (2) Hydroporus polaris, (3) Hydroporus tuvaensis, (4) Hydroporus tibetanus (lectotype), (5) Hydroporus sejilashan sp. n." httpUri="https://zenodo.org/record/210834/files/figure.png" pageId="8" pageNumber="63">Figs. 1–2</figureCitation>
|
||
). Our measurements yielded for
|
||
<taxonomicName id="DA8C4D291C75FF9BFD97F87766DCF841" box="[562,675,1973,1997]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="63" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C75FF9BFD97F87766DCF841" box="[562,675,1973,1997]" italics="true" pageId="8" pageNumber="63">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
the following values: TL:
|
||
<quantity id="DA749B4F1C75FF9BFC6BF8746027F841" box="[974,1112,1973,1997]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="8" pageNumber="63" unit="mm" value="3.25" valueMax="3.4" valueMin="3.1">3.1–3.4 mm</quantity>
|
||
, MW:
|
||
<quantity id="DA749B4F1C75FF9BFB01F8746151F841" box="[1188,1326,1973,1997]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="8" pageNumber="63" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 mm</quantity>
|
||
, TL/MW: 1.88–2.00 (
|
||
<bibRefCitation id="791D4B5B1C75FF9BFEB8F8186151F87D" author="Pederzani" box="[285,1326,2009,2033]" pageId="8" pageNumber="63" refString="Pederzani, F. (2001) Hydroporus tuvaensis n. sp. from Tuva Republic (Russia) and notes on the Hydroporus acutangulus - polaris species-complex (Coleoptera Dytiscidae). Atti dell'Accademia Roveretana degli Agiati, a. 251 (2001), s. VIII, v. 1 (B), 233 - 240." type="journal article" year="2001">
|
||
Pederzani 2001: 238 provided similar values: TL:
|
||
<quantity id="DA749B4F1C75FF9BFC23F818604BF87D" box="[902,1076,2009,2033]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.35" metricValueMin="3.15" pageId="8" pageNumber="63" unit="mm" value="3.25" valueMax="3.35" valueMin="3.15">3.15–3.35 mm</quantity>
|
||
, TL/MW: 1.88–2.00
|
||
</bibRefCitation>
|
||
). The
|
||
<emphasis id="2FF8EAB81C75FF9BFA24F81B61E3F87D" box="[1409,1436,2010,2033]" italics="true" pageId="8" pageNumber="63">H.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="1D3336AA1C74FF9AFF32FF596731FF58" blockId="9.[151,1436,151,500]" pageId="9" pageNumber="64">
|
||
<taxonomicName id="DA8C4D291C74FF9AFF32FF59655FFF3C" box="[151,288,152,176]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="2FF8EAB81C74FF9AFF32FF59655FFF3C" box="[151,288,152,176]" italics="true" pageId="9" pageNumber="64">acutangulus</emphasis>
|
||
</taxonomicName>
|
||
studied are distinctly smaller: TL: 2.8–3.0 mm, MW:
|
||
<quantity id="DA749B4F1C74FF9AFC30FF566050FF23" box="[917,1071,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.475" metricValueMax="1.5" metricValueMin="1.45" pageId="9" pageNumber="64" unit="mm" value="1.475" valueMax="1.5" valueMin="1.45">1.45–1.5 mm</quantity>
|
||
, TL/MW: 1.93–2.00 (
|
||
<bibRefCitation id="791D4B5B1C74FF9AFA89FF56673DFF58" author="Pederzani" pageId="9" pageNumber="64" refString="Pederzani, F. (2001) Hydroporus tuvaensis n. sp. from Tuva Republic (Russia) and notes on the Hydroporus acutangulus - polaris species-complex (Coleoptera Dytiscidae). Atti dell'Accademia Roveretana degli Agiati, a. 251 (2001), s. VIII, v. 1 (B), 233 - 240." type="journal article" year="2001">
|
||
Pederzani 2001: 238 provided: TL:
|
||
<quantity id="DA749B4F1C74FF9AFE17FF7D6627FF58" box="[434,600,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="3.065" metricValueMax="3.23" metricValueMin="2.9" pageId="9" pageNumber="64" unit="mm" value="3.065" valueMax="3.23" valueMin="2.9">2.90–3.23 mm</quantity>
|
||
, TL/MW: 1.83–2.07
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1D3336AA1C74FF9AFF62FF21600EFE20" blockId="9.[151,1436,151,500]" pageId="9" pageNumber="64">
|
||
We want to add that at least those
|
||
<taxonomicName id="DA8C4D291C74FF9AFDEDFF20668DFF74" box="[584,754,224,248]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="2FF8EAB81C74FF9AFDEDFF20668DFF74" box="[584,754,224,248]" italics="true" pageId="9" pageNumber="64">H. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
from northern Europe and the Nearctic
|
||
<taxonomicName id="DA8C4D291C74FF9AFB13FF206157FF74" box="[1206,1320,224,248]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C74FF9AFB13FF206157FF74" box="[1206,1320,224,248]" italics="true" pageId="9" pageNumber="64">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
which we have studied appear externally absolutely different, the
|
||
<taxonomicName id="DA8C4D291C74FF9AFCACFEC467CAFE90" box="[777,949,260,284]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="2FF8EAB81C74FF9AFCACFEC467CAFE90" box="[777,949,260,284]" italics="true" pageId="9" pageNumber="64">H. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
being smaller and distinctly more elongate because the sides of the body are rather parallel, while the
|
||
<taxonomicName id="DA8C4D291C74FF9AFC85FEE867EFFECC" box="[800,912,296,320]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="polaris">
|
||
<emphasis id="2FF8EAB81C74FF9AFC85FEE867EFFECC" box="[800,912,296,320]" italics="true" pageId="9" pageNumber="64">H. polaris</emphasis>
|
||
</taxonomicName>
|
||
have evenly rounded sides and at a first glance look like a
|
||
<taxonomicName id="DA8C4D291C74FF9AFEB1FE8D6754FEE8" authority="Gschwendtner, 1923" authorityName="Gschwendtner" authorityYear="1923" box="[276,811,332,356]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="transgrediens">
|
||
<emphasis id="2FF8EAB81C74FF9AFEB1FE8D6644FEE8" box="[276,571,332,356]" italics="true" pageId="9" pageNumber="64">Hydroporus transgrediens</emphasis>
|
||
Gschwendtner, 1923
|
||
</taxonomicName>
|
||
, or a small
|
||
<taxonomicName id="DA8C4D291C74FF9AFC09FE8D6130FEE8" authority="Fabricius, 1782" authorityName="Fabricius" authorityYear="1782" box="[940,1359,332,356]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="planus">
|
||
<emphasis id="2FF8EAB81C74FF9AFC09FE8D60FBFEE8" box="[940,1156,332,356]" italics="true" pageId="9" pageNumber="64">Hydroporus planus</emphasis>
|
||
(Fabricius, 1782)
|
||
</taxonomicName>
|
||
. If one compares the habitus photos of both species (
|
||
<figureCitation id="85B72A2F1C74FF9AFD2AFEAE6687FE0B" box="[655,760,367,391]" captionStart="FIGURES 1 – 5" captionStartId="2.[151,264,1824,1847]" captionTargetBox="[284,1307,543,1722]" captionTargetId="figure@2.[264,1323,523,1809]" captionTargetPageId="2" captionText="FIGURES 1 – 5. Habitus of: (1) Hydroporus acutangulus, (2) Hydroporus polaris, (3) Hydroporus tuvaensis, (4) Hydroporus tibetanus (lectotype), (5) Hydroporus sejilashan sp. n." httpUri="https://zenodo.org/record/210834/files/figure.png" pageId="9" pageNumber="64">Figs. 1–2</figureCitation>
|
||
) it is hardly believable that both taxa shall belong to one and the same species and that characters shall vary clinally so strongly across northern Asia.
|
||
</paragraph>
|
||
<paragraph id="1D3336AA1C74FF9AFF62FE766022FE78" blockId="9.[151,1436,151,500]" pageId="9" pageNumber="64">
|
||
Like
|
||
<taxonomicName id="DA8C4D291C74FF9AFEA1FE7865CDFE5C" box="[260,434,440,464]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="acutangulus">
|
||
<emphasis id="2FF8EAB81C74FF9AFEA1FE7865CDFE5C" box="[260,434,440,464]" italics="true" pageId="9" pageNumber="64">H. acutangulus</emphasis>
|
||
</taxonomicName>
|
||
, this species can be also easily distinguished from
|
||
<taxonomicName id="DA8C4D291C74FF9AFBA9FE7860DBFE5C" box="[1036,1188,440,464]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="64" phylum="Arthropoda" rank="species" species="sejilashan" status="sp. nov.">
|
||
<emphasis id="2FF8EAB81C74FF9AFBA9FE7860DBFE5C" box="[1036,1188,440,464]" italics="true" pageId="9" pageNumber="64">H. sejilashan</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="34CB57C31C74FF9AFB0BFE796091FE43" box="[1198,1262,440,463]" pageId="9" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
|
||
by the arcuate anterior protarsal claws (males) and, additionally, by the colouration of the upper side.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |