141 lines
11 KiB
XML
141 lines
11 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.72.744" ID-GBIF-Dataset="9b029a3f-f31b-421f-a62e-fb4dd8ff6731" ID-PMC="PMC3083004" ID-Pensoft-Pub="1313-2970-72-23" ID-PubMed="21594103" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-72-23" ModsDocOrigin="ZooKeys 72" ModsDocTitle="Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451250861919" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak" docDate="2010" docId="671A1BF25BB2073DDE9AC8CAF93C9D30" docLanguage="en" docName="ZooKeys 72: 23-68" docOrigin="ZooKeys 72" docSource="http://dx.doi.org/10.3897/zookeys.72.744" docTitle="Tylopus parajeekeli Likhitrakarn, Golovatch, Prateepasen & Panha, 2010, sp. n." docType="treatment" docVersion="5" lastPageNumber="33" masterDocId="D7146C3215665021FFDFFFACFF8CFFD0" masterDocTitle="Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="68" masterPageNumber="23" pageNumber="32" updateTime="1668165214761" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prateepasen, Rujiporn</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="volume">
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<mods:number>72</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>23</mods:start>
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<mods:end>68</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.72.744</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.72.744</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-72-23</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159360929" LSID="urn:lsid:zoobank.org:act:703DB743-0898-4B3A-8D37-A4DD31FE7CD1" httpUri="http://treatment.plazi.org/id/671A1BF25BB2073DDE9AC8CAF93C9D30" lastPageId="10" lastPageNumber="33" pageId="9" pageNumber="32">
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<subSubSection pageId="9" pageNumber="32" type="nomenclature">
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<paragraph pageId="9" pageNumber="32">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:703DB743-0898-4B3A-8D37-A4DD31FE7CD1" class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus parajeekeli" order="Polydesmida" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="parajeekeli">Tylopus parajeekeli</taxonomicName>
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<taxonomicNameLabel pageId="9" pageNumber="32">sp. n.</taxonomicNameLabel>
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Figs 1315
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="32" type="holotype">
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<paragraph pageId="9" pageNumber="32">Holotype</paragraph>
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<paragraph pageId="9" pageNumber="32">♂ (CUMZ), Thailand, Chiang Mai Province, Chom Thong District, Doi Inthanon National Park, summit, 2520 m, 18°34'29N, 98°28'48E, 12.10.2009, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="32" type="paratype">
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<paragraph pageId="9" pageNumber="32">Paratype:</paragraph>
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<paragraph pageId="9" pageNumber="32">1 ♂ (CUMZ), same locality, together with holotype.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="32" type="name">
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<paragraph pageId="9" pageNumber="32">Name:</paragraph>
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<paragraph pageId="9" pageNumber="32">
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To emphasize the close resemblance to
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus jeekeli" order="Polydesmida" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="jeekeli">Tylopus jeekeli</taxonomicName>
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Golovatch & Enghoff, 1993.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="32" type="diagnosis">
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<paragraph pageId="9" pageNumber="32">Diagnosis:</paragraph>
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<paragraph pageId="9" pageNumber="32">
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Very similar to
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus jeekeli" order="Polydesmida" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="jeekeli">Tylopus jeekeli</taxonomicName>
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, especially as regards its gonopod conformation, but differs in the paraterga lying much lower (at ca 1/3 versus 1/4-1/5 midbody height), in the caudal corners of the paraterga protruding behind the rear tergal contour already from segment 16 (versus segment 2), and also in gonopod spine z being much smaller and placed closer to the base of spine h.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="33" pageId="9" pageNumber="32" type="description">
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<paragraph pageId="9" pageNumber="32">Description:</paragraph>
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<paragraph pageId="9" pageNumber="32">
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Length 31 mm (holotype) or 31.5 mm (♂), width of midbody pro- and metazona 2.4 and 3.2 mm (holotype) or 2.3 and 3.4 mm (♂), respectively. Coloration of live animals and alcohol material uniformly blackish-brown (Fig. 13
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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); calluses of paraterga a little lighter, brown; antennomeres 1-5 light brown to yellowish, legs and venter light brown to grey-yellowish (Fig. 13
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="9" pageNumber="32">
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All characters as in
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus bispinosus" order="Polydesmida" pageId="9" pageNumber="32" phylum="Arthropoda" rank="species" species="bispinosus">Tylopus bispinosus</taxonomicName>
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sp. n., except as follows.
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</paragraph>
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<paragraph pageId="10" pageNumber="33">
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<pageBreakToken pageId="10" pageNumber="33" start="start">Antennae</pageBreakToken>
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rather short and slender, reaching behind segment 3 dorsally. In width, head = segment 3 <collum <segments 2 and 4 <5-16; thereafter body gradually and gently tapering towards telson (Fig. 1B).
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</paragraph>
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<paragraph pageId="10" pageNumber="33">
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Paraterga on collum like large rounded flaps (Fig. 13B, C). Following paraterga lying at about 1/3 midbody height, evidently declined ventrolaterally, subhorizontal only on a few posteriormost segments, mostly pointed caudally, subrectangular until segment 15, thereafter increasingly well protruding behind rear tergal contour (Fig. 13
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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). Metaterga with 2(3)+2(3) and 3
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<normalizedToken originalValue="–5+3–">-5+3-</normalizedToken>
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5 long setae arranged in two transverse rows. Axial line present on both halves of metaterga. Transverse sulcus present on segments 5-18, very finely beaded at bottom (Fig. 13B, F, G). Stricture between pro- and metazona finely striolate (Fig. 13B, D). Epiproct tip broad and emarginate (Fig. 13G, H). Hypoproct semi-circular, both caudal setae widely separated (Fig. 13H). Pleurosternal carinae as complete ridges on segments 2-4, thereafter retained until segment 17 mostly as a small caudal tooth (Fig. 13
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<normalizedToken originalValue="C–F">C-F</normalizedToken>
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).
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</paragraph>
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<paragraph pageId="10" pageNumber="33">A low, only slightly divided, setose lobe between coxae 4 (Fig. 13I, J). Legs relatively short, ca 1.6-1.7 times as long as midbody height (Fig. 13K). Femora evidently bulged laterally (Fig. 13K); all postgonopodal legs except two last pairs with an evident adenostyle in parabasal 1/3 of each postfemur and tibia; tarsal brushes missing; all telopoditomeres except tarsi with dense adpressed pilosity (Fig. 13K).</paragraph>
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<paragraph pageId="10" pageNumber="33">Gonopods (Figs 14, 15) with lobe l well-demarcated; spine h small, but elongate, not bifid; spine z very small, dentiform, placed at base of spine h.</paragraph>
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<caption pageId="10" pageNumber="33">
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<paragraph pageId="10" pageNumber="33">
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Figure 13.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus parajeekeli" order="Polydesmida" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="parajeekeli">Tylopus parajeekeli</taxonomicName>
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sp. n., ♂ holotype (A) and ♂ paratype (
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<normalizedToken originalValue="B–K">B-K</normalizedToken>
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). A habitus, live coloration B, C anterior part of body, dorsal and lateral views, respectively D, E segments 10 and 11, dorsal and lateral views, respectively F, G, H posterior part of body, lateral, dorsal and ventral views, respectively I, J sternal cones between coxae 4, subcaudal and sublateral views, respectively K midbody leg.
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</paragraph>
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</caption>
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<caption pageId="10" pageNumber="33">
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<paragraph pageId="10" pageNumber="33">
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Figure 14.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus parajeekeli" order="Polydesmida" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="parajeekeli">Tylopus parajeekeli</taxonomicName>
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sp. n., ♂ paratype. A, B right gonopod, mesal and lateral views, respectively
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<normalizedToken originalValue="C–F">C-F</normalizedToken>
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distal part of right gonopod, submesal, sublateral, subcaudal and suboral views, respectively. Scale bar: 0.2 mm.
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</paragraph>
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</caption>
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<caption pageId="10" pageNumber="33">
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<paragraph pageId="10" pageNumber="33">
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Figure 15.
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<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Tylopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tylopus parajeekeli" order="Polydesmida" pageId="10" pageNumber="33" phylum="Arthropoda" rank="species" species="parajeekeli">Tylopus parajeekeli</taxonomicName>
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sp. n., ♂ paratype. A, B right gonopod, lateral and mesal views, respectively. Scale bar: 0.5 mm.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |