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<document id="030EC8778DAE4B0B5B10EEA99AA57E51" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFF8E73D032FAFECDE1F4FA6A" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium fisheri Wesener 2009, new species" docType="treatment" docVersion="4" lastPageNumber="129" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="126" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="A1FF977A3F42CAFA8602ECE3171644A6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DB8CCF56BF2EAF6152662318DB957632">
<mods:title id="A2687B49808E26F0E859DCD15CEA8AF1">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
</mods:titleInfo>
<mods:name id="A29D2BCD040434AF869126AE3645C9A1" type="personal">
<mods:role id="B240792027E715DAAD87F0567CA60056">
<mods:roleTerm id="592E98EF04B733FE9D07161B99A6F61F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F52C0C9B97767E1023EA62870F2D14D">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="F840C56FA4C1920E8FC68F6200E2B262">text</mods:typeOfResource>
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<mods:titleInfo id="45C6722A00E695C10FC1F6D455C8F31C">
<mods:title id="82AC5E238393E20FBA315B42851113A4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DDB682CFDED37753AAC92C6F05CEF0C7">
<mods:date id="929F5BE6C6020305009507AFF0BD77CA">2009</mods:date>
<mods:detail id="E450D1B48B14614B79ED929D16A17776" type="pubDate">
<mods:number id="84EF1F7D919B38E37AC7467A75171EEC">2009-05-11</mods:number>
</mods:detail>
<mods:detail id="169CD7C86260DC3B5D428759C75109A8" type="volume">
<mods:number id="84E0C6D1879DDED68322495A8287E6F3">2097</mods:number>
</mods:detail>
<mods:detail id="2E1179AA8338DE3A0A2AB0B90B079F1D" type="issue">
<mods:number id="B343BEC44194D60D986B4B79D41CF54B">1</mods:number>
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<mods:extent id="CC7E8AE764DBF728710CBDDB6A5B08B5" unit="page">
<mods:start id="AE554BDFA5CF61B8E521FCF91CFBC49C">1</mods:start>
<mods:end id="651DD858508952B63F807D351836424E">134</mods:end>
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<mods:location id="094DDAAA94B34D557DC50C5047F75D0E">
<mods:url id="D7DDCAA7FA21892E83E9694D47473BA8">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
</mods:location>
<mods:classification id="089E4899CBE0763DFB110E91F62E600E">journal article</mods:classification>
<mods:identifier id="B3E2F1FAB5107806B78E0CCC0D11E14C" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="37F696AE1C2C11705527479A29BCD099" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="56390BB902939B2CB421DF025F8C6E77" type="Zenodo-Dep">5316536</mods:identifier>
<mods:identifier id="AEE3B6B34F6C46AF24B01322498BC105" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
</mods:mods>
<treatment id="03FFBE1DFF8E73D032FAFECDE1F4FA6A" ID-DOI="http://doi.org/10.5281/zenodo.5325539" ID-Zenodo-Dep="5325539" LSID="urn:lsid:plazi:treatment:03FFBE1DFF8E73D032FAFECDE1F4FA6A" httpUri="http://treatment.plazi.org/id/03FFBE1DFF8E73D032FAFECDE1F4FA6A" lastPageId="128" lastPageNumber="129" pageId="125" pageNumber="126">
<subSubSection id="C34C5C80FF8E732D32FAFECDE13DFE48" box="[151,570,375,401]" pageId="125" pageNumber="126" type="nomenclature">
<paragraph id="8BE90F0BFF8E732D32FAFECDE13DFE48" blockId="125.[151,570,375,401]" box="[151,570,375,401]" pageId="125" pageNumber="126">
<heading id="D0A1B867FF8E732D32FAFECDE13DFE48" bold="true" box="[151,570,375,401]" fontSize="11" level="1" pageId="125" pageNumber="126" reason="1">
<emphasis id="B922D319FF8E732D32FAFECDE13DFE48" bold="true" box="[151,570,375,401]" pageId="125" pageNumber="126">
<taxonomicName id="4C567488FF8E732D32FAFECDE29BFE48" ID-CoL="5D7GZ" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,412,375,401]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="B922D319FF8E732D32FAFECDE29BFE48" bold="true" box="[151,412,375,401]" italics="true" pageId="125" pageNumber="126">Zoosphaerium fisheri</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2116E62FF8E732D33C6FECDE13DFE48" box="[427,570,375,401]" pageId="125" pageNumber="126" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF8E732D32FAFE7AE116FDDB" pageId="125" pageNumber="126" type="etymology">
<paragraph id="8BE90F0BFF8E732D32FAFE7AE695FE02" blockId="125.[151,1436,448,2022]" box="[151,1426,448,475]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32FAFE7AE287FE03" bold="true" box="[151,384,448,474]" pageId="125" pageNumber="126">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FF8E732D33E5FE7BE2D4FE03" authorityName="Wesener" authorityYear="2009" box="[392,467,449,474]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="B922D319FF8E732D33E5FE7BE2D4FE03" box="[392,467,449,474]" italics="true" pageId="125" pageNumber="126">fisheri</emphasis>
</taxonomicName>
, after Brian L. Fisher (CAS), who collected numerous millipedes on
<collectingCountry id="F3414F9BFF8E732D3692FE7BE688FE02" box="[1279,1423,449,475]" name="Madagascar" pageId="125" pageNumber="126">Madagascar</collectingCountry>
.
</paragraph>
<paragraph id="8BE90F0BFF8E732D32ABFE5DE116FDDB" blockId="125.[151,1436,448,2022]" box="[198,529,487,514]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32ABFE5DE2A7FDD8" bold="true" box="[198,416,487,513]" pageId="125" pageNumber="126">Studied material:</emphasis>
4 M,
<specimenCount id="9D50C482FF8E732D338BFE52E116FDDB" box="[486,529,488,514]" pageId="125" pageNumber="126" type="generic">4 F.</specimenCount>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF8E732D32ABFDB4E0AEFDAF" pageId="125" pageNumber="126" type="materials_examined">
<paragraph id="8BE90F0BFF8E732D32ABFDB4E02CFD96" blockId="125.[151,1436,448,2022]" pageId="125" pageNumber="126">
<materialsCitation id="3B3E0556FF8E732D32ABFDB4E02CFD96" collectionCode="SMF" collectorName="G. Osche" country="Madagascar" latitude="-18.9" location="Madagascar" longLatPrecision="1273" longitude="47.916668" pageId="125" pageNumber="126" specimenCount="1" typeStatus="holotype">
<emphasis id="B922D319FF8E732D32ABFDB4E23FFDF1" bold="true" box="[198,312,526,552]" pageId="125" pageNumber="126">
<typeStatus id="54EDB1A9FF8E732D32ABFDB4E23FFDF1" box="[198,312,526,552]" pageId="125" pageNumber="126" type="holotype">Holotype</typeStatus>
</emphasis>
: 1 M (broken, probably
<quantity id="4CAEA2EEFF8E732D3004FDB4E1B9FDF1" box="[617,702,526,552]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="125" pageNumber="126" unit="mm" value="25.0">25 mm</quantity>
long),
<emphasis id="B922D319FF8E732D3177FDB4E052FDF1" bold="true" box="[794,853,526,552]" pageId="125" pageNumber="126">
<collectionCode id="ED4797CEFF8E732D3177FDB4E052FDF1" box="[794,853,526,552]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="125" pageNumber="126">SMF</collectionCode>
</emphasis>
.
<emphasis id="B922D319FF8E732D310AFDB5E70CFDF1" box="[871,1035,527,552]" italics="true" pageId="125" pageNumber="126">Locus typicus</emphasis>
:
<collectingCountry id="F3414F9BFF8E732D3673FDB4E7AAFDF1" box="[1054,1197,526,552]" name="Madagascar" pageId="125" pageNumber="126">Madagascar</collectingCountry>
, Mandraka, coll.
<collectorName id="26A36ADDFF8E732D37EEFDB4E3E4FD96" pageId="125" pageNumber="126">G. Osche</collectorName>
, possibly Mandraka, rainforest,
<geoCoordinate id="EE6269CCFF8E732D3031FD8FE1BDFD96" box="[604,698,565,591]" degrees="18" direction="south" minutes="54" orientation="latitude" pageId="125" pageNumber="126" precision="925" value="-18.9">18°54'S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFF8E732D30ABFD8FE020FD96" box="[710,807,565,591]" degrees="47" direction="east" minutes="55" orientation="longitude" pageId="125" pageNumber="126" precision="925" value="47.916668">47°55'E</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFF8E732D32ABFDE1E0AEFDAF" blockId="125.[151,1436,448,2022]" box="[198,937,603,630]" pageId="125" pageNumber="126">
<materialsCitation id="3B3E0556FF8E732D32ABFDE1E0A2FDAF" box="[198,933,603,630]" collectionCode="SMF" collectorName="G. Osche" country="Madagascar" latitude="-18.9" location="Madagascar" longLatPrecision="1273" longitude="47.916668" pageId="125" pageNumber="126" specimenCount="1" typeStatus="paratype">
<emphasis id="B922D319FF8E732D32ABFDE1E244FDAC" bold="true" box="[198,323,603,629]" pageId="125" pageNumber="126">
<typeStatus id="54EDB1A9FF8E732D32ABFDE1E244FDAC" box="[198,323,603,629]" pageId="125" pageNumber="126" type="paratype">Paratypes</typeStatus>
</emphasis>
: 3 M, 4 F, (all broken)
<emphasis id="B922D319FF8E732D3038FDE1E196FDAC" bold="true" box="[597,657,603,629]" pageId="125" pageNumber="126">
<collectionCode id="ED4797CEFF8E732D3038FDE1E196FDAC" box="[597,657,603,629]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="125" pageNumber="126">SMF</collectionCode>
</emphasis>
, same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF8E732D32ABFD38E0FFFD45" box="[198,1016,642,668]" pageId="125" pageNumber="126" type="distribution">
<paragraph id="8BE90F0BFF8E732D32ABFD38E0FFFD45" blockId="125.[151,1436,448,2022]" box="[198,1016,642,668]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32ABFD38E26EFD45" bold="true" box="[198,361,642,668]" pageId="125" pageNumber="126">Distribution:</emphasis>
Only known from the rainforest of Mandraka (Fig. 77).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF8E732D32ABFD12E669FB39" pageId="125" pageNumber="126" type="diagnosis">
<paragraph id="8BE90F0BFF8E732D32ABFD12E152FB9F" blockId="125.[151,1436,448,2022]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32ABFD12E24FFD1B" bold="true" box="[198,328,680,706]" pageId="125" pageNumber="126">Diagnosis:</emphasis>
Relatively small, up to
<quantity id="4CAEA2EEFF8E732D300AFD13E1BCFD1A" box="[615,699,681,707]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="125" pageNumber="126" unit="mm" value="30.0">30 mm</quantity>
long. Color (changed by alcohol) black, anterior parts of anal shield and tergites 3–12 brown, antennae and legs green. Anterior part of tergites and anal shield in both sexes conspicuously covered with long hair. First antennomere with groove, antennomeres 1–5 with sclerotized teeth (Fig. 87B), disc with 16–22 apical cones (Fig. 87A). First stigmatic plate with prominent, wide lobe (Figs 86A, B). Movable finger of posterior telopods weakly curved, apically only slightly tapering, with three thin spines (Fig. 86F); posterior side with
<emphasis id="B922D319FF8E732D30FBFCD0E1D6FC5A" box="[662,721,874,899]" italics="true" pageId="125" pageNumber="126">circa</emphasis>
11 sclerotized, crenulated teeth (Fig. 86G). Fixed finger of unusual shape, tip curved and more slender than rest of finger. Third podomere of anterior telopods laterally of cavity with row of 8–10 large, sclerotized, crenulated teeth (Fig. 86E). One stridulation rib on male harp (Fig. 86D) and 1 or 2 on each side of female washboard. Two same-sized locking carinae on anal shield. Operculum of vulva slightly extending beyond coxa, on apical margin medially notched, elongated into two rounded, same-sized lobes (Fig. 86C).
</paragraph>
<paragraph id="8BE90F0BFF8E732D32ABFBE8E669FB39" blockId="125.[151,1436,448,2022]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32ABFBE8E28BFBB5" bold="true" box="[198,396,1106,1132]" pageId="125" pageNumber="126">Similar species:</emphasis>
The anterior and posterior telopods of
<emphasis id="B922D319FF8E732D310AFBE9E76BFBB5" box="[871,1132,1107,1132]" italics="true" pageId="125" pageNumber="126">
<taxonomicName id="4C567488FF8E732D310AFBE9E76FFBB5" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[871,1128,1107,1132]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">Zoosphaerium fisheri</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FF8E732D361BFBE8E7BDFBB5" bold="true" box="[1142,1210,1106,1132]" pageId="125" pageNumber="126">
<taxonomicNameLabel id="A2116E62FF8E732D361BFBE8E7BDFBB5" box="[1142,1210,1106,1132]" pageId="125" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
are of a shape not displayed among other known giant-pill millipede species. The posterior telopods are quite similar to those of
<taxonomicName id="4C567488FF8E732D32FAFB1AE240FB60" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,327,1184,1209]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="analavelona" status="sp. nov.">
<emphasis id="B922D319FF8E732D32FAFB1AE240FB60" box="[151,327,1184,1209]" italics="true" pageId="125" pageNumber="126">Z. analavelona</emphasis>
</taxonomicName>
<emphasis id="B922D319FF8E732D3320FB25E288FB60" bold="true" box="[333,399,1183,1209]" pageId="125" pageNumber="126">
<taxonomicNameLabel id="A2116E62FF8E732D3320FB25E288FB60" box="[333,399,1183,1209]" pageId="125" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
but
<emphasis id="B922D319FF8E732D33AEFB1AE134FB60" box="[451,563,1184,1209]" italics="true" pageId="125" pageNumber="126">
<taxonomicName id="4C567488FF8E732D33AEFB1AE128FB60" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[451,559,1184,1209]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="125" pageNumber="126" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">Z. fisheri</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FF8E732D3057FB25E17BFB60" bold="true" box="[570,636,1183,1209]" pageId="125" pageNumber="126">
<taxonomicNameLabel id="A2116E62FF8E732D3057FB25E17BFB60" box="[570,636,1183,1209]" pageId="125" pageNumber="126" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
differs strongly from the latter in the shape of the female operculum (Fig. 86C), number of apical cones on the antenna and the third podomere of anterior telopods (Fig. 86E).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF8E73D032ABFB57E1F4FA6A" lastPageId="128" lastPageNumber="129" pageId="125" pageNumber="126" type="description">
<paragraph id="8BE90F0BFF8E732D32ABFB57E3FFFA8C" blockId="125.[151,1436,448,2022]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32ABFB57E267FADE" bold="true" box="[198,352,1261,1287]" pageId="125" pageNumber="126">Description:</emphasis>
Body length: Males (
<specimenCount id="9D50C482FF8E732D301BFB57E015FADE" box="[630,786,1261,1287]" pageId="125" pageNumber="126" type="generic">4 specimens</specimenCount>
): width of thoracic shield up to
<quantity id="4CAEA2EEFF8E732D36C6FB57E61AFADE" box="[1195,1309,1261,1287]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="125" pageNumber="126" unit="mm" value="10.8">10.8 mm</quantity>
, height of thoracic shield up to
<quantity id="4CAEA2EEFF8E732D33E3FAAEE2ECFAF7" box="[398,491,1300,1326]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="125" pageNumber="126" unit="mm" value="6.2">6.2 mm</quantity>
. Females (
<specimenCount id="9D50C482FF8E732D3004FAAEE007FAF7" box="[617,768,1300,1326]" pageId="125" pageNumber="126" type="generic">4 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFF8E732D31C5FAAEE0F9FAF7" box="[936,1022,1300,1326]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="125" pageNumber="126" unit="mm" value="30.0">30 mm</quantity>
, width up to
<quantity id="4CAEA2EEFF8E732D36F5FAAEE605FAF7" box="[1176,1282,1300,1326]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="125" pageNumber="126" unit="mm" value="14.7">14.7 mm</quantity>
, height up to
<quantity id="4CAEA2EEFF8E732D32FAFA81E3F3FA8C" box="[151,244,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="125" pageNumber="126" unit="mm" value="7.7">7.7 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFF8E732D32AAFADBE06AFAA2" blockId="125.[151,1436,448,2022]" box="[199,877,1377,1403]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAFADBE229FAA3" box="[199,302,1377,1402]" italics="true" pageId="125" pageNumber="126">Habitus:</emphasis>
Tergites covered anteriorly with numerous hairs.
</paragraph>
<paragraph id="8BE90F0BFF8E732D32AAFA32E3E7FA36" blockId="125.[151,1436,448,2022]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAFA32E256FA78" box="[199,337,1416,1441]" italics="true" pageId="125" pageNumber="126">Coloration:</emphasis>
In most specimens faded by light and alcohol. In best preserved specimens, head, collum, thoracic shield and tergites black, anterior parts of anal shield and tergites 3–12 brown, antenna and legs green.
</paragraph>
<paragraph id="8BE90F0BFF8E732D32AAFA46E12BF9CF" blockId="125.[151,1436,448,2022]" box="[199,556,1532,1558]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAFA46E217F9CC" box="[199,272,1532,1557]" italics="true" pageId="125" pageNumber="126">Head:</emphasis>
Eyes with 50–55 ocelli.
</paragraph>
<paragraph id="8BE90F0BFF8E732D32AAF999E3DEF969" blockId="125.[151,1436,448,2022]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAF999E238F9E5" box="[199,319,1571,1596]" italics="true" pageId="125" pageNumber="126">Antennae:</emphasis>
Length of antennomeres: 1&lt;2=3=4=5&lt;6 (Fig. 87B), sixth antennomere cylindrical, bearing disc with 16–22 apical cones (Fig. 87A). First antennomere of greatest width, short and cylindrical, with groove and 1 or 2 rows of
<emphasis id="B922D319FF8E732D33A2F9CAE1B7F950" box="[463,688,1648,1673]" italics="true" pageId="125" pageNumber="126">sensilla basiconica</emphasis>
(Fig. 87B). Sclerotized teeth at base of antennomeres 1–5 (Fig. 87B).
</paragraph>
<paragraph id="8BE90F0BFF8E732D32AAF907E026F927" blockId="125.[151,1436,448,2022]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAF907E230F90F" box="[199,311,1725,1750]" italics="true" pageId="125" pageNumber="126">Mandible</emphasis>
: Six rows of pectinate lamellae, number of teeth declining apically to proximally (Fig. 87C). Condylus with two steps near apical margin (Fig. 87C).
</paragraph>
<paragraph id="8BE90F0BFF8E732D32AAF8B1E70FF8FC" blockId="125.[151,1436,448,2022]" box="[199,1032,1803,1829]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAF8B1E296F8FD" box="[199,401,1803,1828]" italics="true" pageId="125" pageNumber="126">Gnathochilarium</emphasis>
: Lateral of palpi four sensory cones, located together.
</paragraph>
<paragraph id="8BE90F0BFF8E732D32AAF88BE74CF892" blockId="125.[151,1436,448,2022]" box="[199,1099,1841,1867]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAF88BE220F893" box="[199,295,1841,1866]" italics="true" pageId="125" pageNumber="126">Collum:</emphasis>
Only on anterior and posterior margin with some long, isolated hair.
</paragraph>
<paragraph id="8BE90F0BFF8E732D32AAF8E2E1C5F8AB" blockId="125.[151,1436,448,2022]" box="[199,706,1880,1906]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAF8E2E279F8A8" box="[199,382,1880,1905]" italics="true" pageId="125" pageNumber="126">Thoracic shield</emphasis>
: Surface similar to tergites.
</paragraph>
<paragraph id="8BE90F0BFF8E732D32AAF8C5E00AF866" blockId="125.[151,1436,448,2022]" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAF8C5E224F841" box="[199,291,1919,1944]" italics="true" pageId="125" pageNumber="126">Tergites</emphasis>
: Smooth, covered on anterior half with numerous long hairs, posterior margin with 1 or 2 rows of long hairs. Paratergites projecting slightly posteriorly.
</paragraph>
<paragraph id="8BE90F0BFF8E732E32AAF876E1D6FF01" blockId="125.[151,1436,448,2022]" lastBlockId="126.[151,1436,152,487]" lastPageId="126" lastPageNumber="127" pageId="125" pageNumber="126">
<emphasis id="B922D319FF8E732D32AAF876E267F83C" box="[199,352,1996,2021]" italics="true" pageId="125" pageNumber="126">Endotergum:</emphasis>
Inner section with broad spines and few setae (Fig. 88D). Between marginal ridge and inner area single row of large, circular cuticular impressions (Fig. 88D). Externally two rows of marginal bristles. Bristles not extending beyond tergite (Fig. 88D).
</paragraph>
<paragraph id="8BE90F0BFF8D732E32AAFF5FE716FEFF" blockId="126.[151,1436,152,487]" pageId="126" pageNumber="127">
<emphasis id="B922D319FF8D732E32AAFF5FE25FFF27" box="[199,344,229,254]" italics="true" pageId="126" pageNumber="127">Anal shield:</emphasis>
Well-rounded, neither bell-shaped nor tapered. In both sexes covered with numerous long hairs. Ventral side of anal shield with two same-sized black locking carinae.
</paragraph>
<paragraph id="8BE90F0BFF8D732E32AAFE88E60EFE43" blockId="126.[151,1436,152,487]" pageId="126" pageNumber="127">
<emphasis id="B922D319FF8D732E32AAFE88E20FFE92" box="[199,264,306,331]" italics="true" pageId="126" pageNumber="127">Legs:</emphasis>
Tarsi of first pair of legs with two to four, second with
<emphasis id="B922D319FF8D732E31F5FE88E0D4FE92" box="[920,979,306,331]" italics="true" pageId="126" pageNumber="127">circa</emphasis>
five and third with
<emphasis id="B922D319FF8D732E36D0FE88E7FFFE92" box="[1213,1272,306,331]" italics="true" pageId="126" pageNumber="127">circa</emphasis>
seven ventral spines. First two leg pairs with weakly curved claws and without apical spine. Tarsi of legs 4–21 with curved claws, 10–12 ventral spines and one apical spine. Femur 2.1 and tarsi 4.3 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFF8D732E32AAFE1DE3FBFE3E" blockId="126.[151,1436,152,487]" pageId="126" pageNumber="127">
<emphasis id="B922D319FF8D732E32AAFE1DE28BFE19" box="[199,396,423,448]" italics="true" pageId="126" pageNumber="127">Stigmatic plates:</emphasis>
First lobe well-rounded, very wide and long, reaching up to apical end of prefemur (Figs 87A, B).
</paragraph>
<paragraph id="8BE90F0BFF8D732E32FAF999E7EFF931" blockId="126.[151,1436,1571,1769]" pageId="126" pageNumber="127">
<emphasis id="B922D319FF8D732E32FAF999E22AF9E2" bold="true" box="[151,301,1571,1595]" pageId="126" pageNumber="127">FIGURE 086</emphasis>
.
<taxonomicName id="4C567488FF8D732E3357F99EE118F9E2" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[314,543,1572,1595]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="126" pageNumber="127" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="B922D319FF8D732E3357F99EE118F9E2" box="[314,543,1572,1595]" italics="true" pageId="126" pageNumber="127">Zoosphaerium fisheri</emphasis>
</taxonomicName>
<emphasis id="B922D319FF8D732E3048F999E166F9E2" bold="true" box="[549,609,1571,1595]" pageId="126" pageNumber="127">
<taxonomicNameLabel id="A2116E62FF8D732E3048F999E166F9E2" box="[549,609,1571,1595]" pageId="126" pageNumber="127" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, A, B, D–G
<typeStatus id="54EDB1A9FF8D732E3085F99EE04FF9E2" box="[744,840,1572,1595]" pageId="126" pageNumber="127" type="holotype">holotype</typeStatus>
, C female
<typeStatus id="54EDB1A9FF8D732E31D4F99EE710F9E2" box="[953,1047,1572,1595]" pageId="126" pageNumber="127" type="paratype">paratype</typeStatus>
,
<emphasis id="B922D319FF8D732E364FF999E732F9E2" bold="true" box="[1058,1077,1571,1595]" pageId="126" pageNumber="127">A</emphasis>
: first left coxa and prefemur with first stigmatic plate, posterior view;
<emphasis id="B922D319FF8D732E307AF9FCE12EF987" bold="true" box="[535,553,1606,1630]" pageId="126" pageNumber="127">B</emphasis>
: first stigmatic plate, anterior view;
<emphasis id="B922D319FF8D732E31C6F9FCE0B9F987" bold="true" box="[939,958,1606,1630]" pageId="126" pageNumber="127">C</emphasis>
: second left coxa with vulva, posterior view;
<emphasis id="B922D319FF8D732E32FAF9D3E3ADF958" bold="true" box="[151,170,1641,1665]" pageId="126" pageNumber="127">D</emphasis>
: right anterior telopod, anterior view;
<emphasis id="B922D319FF8D732E303FF9D3E163F958" bold="true" box="[594,612,1641,1665]" pageId="126" pageNumber="127">E</emphasis>
: last two podomeres of anterior telopod, lateral view;
<emphasis id="B922D319FF8D732E36D1F9D3E7CBF958" bold="true" box="[1212,1228,1641,1665]" pageId="126" pageNumber="127">F</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FF8D732E335CF931E241F97A" bold="true" box="[305,326,1675,1699]" pageId="126" pageNumber="127">G</emphasis>
: posterior telopod, posterior view. Abbreviations:
<emphasis id="B922D319FF8D732E3133F931E082F97A" bold="true" box="[862,901,1675,1699]" pageId="126" pageNumber="127">cav</emphasis>
= cavity;
<emphasis id="B922D319FF8D732E3183F931E711F97A" bold="true" box="[1006,1046,1675,1699]" pageId="126" pageNumber="127">cr-t</emphasis>
= crenulated teeth;
<emphasis id="B922D319FF8D732E3684F931E60EF97A" bold="true" box="[1257,1289,1675,1699]" pageId="126" pageNumber="127">Cx</emphasis>
= coxa;
<emphasis id="B922D319FF8D732E3709F931E681F97A" bold="true" box="[1380,1414,1675,1699]" pageId="126" pageNumber="127">EP</emphasis>
= external plate of vulva;
<emphasis id="B922D319FF8D732E33CFF914E2BBF91F" bold="true" box="[418,444,1710,1734]" pageId="126" pageNumber="127">IP</emphasis>
= mesal plate of vulva;
<emphasis id="B922D319FF8D732E30BEF914E1F6F91F" bold="true" box="[723,753,1710,1734]" pageId="126" pageNumber="127">ml</emphasis>
= membranous lobe;
<emphasis id="B922D319FF8D732E318AF914E0FBF91F" bold="true" box="[999,1020,1710,1734]" pageId="126" pageNumber="127">O</emphasis>
= operculum;
<emphasis id="B922D319FF8D732E36CEF914E7CCF91F" bold="true" box="[1187,1227,1710,1734]" pageId="126" pageNumber="127">Pre</emphasis>
= prefemur;
<emphasis id="B922D319FF8D732E370CF914E684F91F" bold="true" box="[1377,1411,1710,1734]" pageId="126" pageNumber="127">s-p</emphasis>
= sclerotized spot;
<emphasis id="B922D319FF8D732E3321F96BE27CF930" bold="true" box="[332,379,1745,1769]" pageId="126" pageNumber="127">scl-t</emphasis>
= sclerotized teeth;
<emphasis id="B922D319FF8D732E303EF96BE172F930" bold="true" box="[595,629,1745,1769]" pageId="126" pageNumber="127">SR</emphasis>
= stridulation rib;
<emphasis id="B922D319FF8D732E3150F96BE054F930" bold="true" box="[829,851,1745,1769]" pageId="126" pageNumber="127">St</emphasis>
= stigmatic plate. Scale bars =
<quantity id="4CAEA2EEFF8D732E36CEF96BE7E3F931" box="[1187,1252,1745,1768]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="126" pageNumber="127" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFF8D732E32AAF8A0E661F871" blockId="126.[151,1436,1818,2038]" pageId="126" pageNumber="127">
<emphasis id="B922D319FF8D732E32AAF8A0E2F2F8EA" box="[199,501,1818,1843]" italics="true" pageId="126" pageNumber="127">Female sexual characters</emphasis>
: Washboard consisting of short, but broad and well-developed stridulation ribs, with 1 or 2 ribs on each half. Vulva large, covering more than 3/4 of coxal surface. Operculum large, protruding above apical coxa margin. Apical margin of operculum medially notched, forming two same-sized, well-rounded processes (Fig. 87C). Mesal plate long, reaching up to half of operculum height (Fig. 87C).
</paragraph>
<paragraph id="8BE90F0BFF8D732E32AAF80FE127F82F" blockId="126.[151,1436,1818,2038]" pageId="126" pageNumber="127">
<emphasis id="B922D319FF8D732E32AAF80FE2E6F817" box="[199,481,1973,1998]" italics="true" pageId="126" pageNumber="127">Male sexual characters</emphasis>
: Male gonopore covered with large, sclerotized, undivided and rounded plate. Apical part of plate membranous.
</paragraph>
<paragraph id="8BE90F0BFF8C732F32AAFF22E252FEAA" blockId="127.[151,1436,152,371]" pageId="127" pageNumber="128">
<emphasis id="B922D319FF8C732F32AAFF22E290FF68" box="[199,407,152,177]" italics="true" pageId="127" pageNumber="128">Anterior telopods</emphasis>
(Figs 86D, E): First podomere of rectangular shape, width slightly larger than height. Harp with single stridulation rib, rib long and well-developed. Second podomere process lobe-like and little curved with rounded edges, protruding to one third of third podomeres height, apically with sclerotized spots. Third podomere cavity mesally with up to three thin, sclerotized spines, in basal half with numerous small sclerotized spots, laterally with 8–10 crenulated, sclerotized teeth. Third podomere apically with one sclerotized spot.
</paragraph>
<paragraph id="8BE90F0BFF8C732F32FAF839E297F839" blockId="127.[151,1436,1923,2017]" pageId="127" pageNumber="128">
<emphasis id="B922D319FF8C732F32FAF839E22AF842" bold="true" box="[151,301,1923,1947]" pageId="127" pageNumber="128">FIGURE 087</emphasis>
.
<taxonomicName id="4C567488FF8C732F3357F83EE118F842" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[314,543,1924,1947]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="127" pageNumber="128" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="B922D319FF8C732F3357F83EE118F842" box="[314,543,1924,1947]" italics="true" pageId="127" pageNumber="128">Zoosphaerium fisheri</emphasis>
</taxonomicName>
<emphasis id="B922D319FF8C732F3048F839E166F842" bold="true" box="[549,609,1923,1947]" pageId="127" pageNumber="128">
<taxonomicNameLabel id="A2116E62FF8C732F3048F839E166F842" box="[549,609,1923,1947]" pageId="127" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FF8C732F3003F83EE1CAF842" box="[622,717,1924,1947]" pageId="127" pageNumber="128" type="paratype">paratype</typeStatus>
, SEM,
<emphasis id="B922D319FF8C732F3176F839E029F842" bold="true" box="[795,814,1923,1947]" pageId="127" pageNumber="128">A</emphasis>
: antenna, apical view on disc;
<emphasis id="B922D319FF8C732F361AF839E78EF842" bold="true" box="[1143,1161,1923,1947]" pageId="127" pageNumber="128">B</emphasis>
: antenna, lateral view;
<emphasis id="B922D319FF8C732F37ECF839E693F842" bold="true" box="[1409,1428,1923,1947]" pageId="127" pageNumber="128">C</emphasis>
: right mandible, mesal view. Abbreviations:
<emphasis id="B922D319FF8C732F3002F81CE191F867" bold="true" box="[623,662,1958,1982]" pageId="127" pageNumber="128">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FF8C732F3127F81CE06DF867" bold="true" box="[842,874,1958,1982]" pageId="127" pageNumber="128">Co</emphasis>
= condylus;
<emphasis id="B922D319FF8C732F3195F81CE711F867" bold="true" box="[1016,1046,1958,1982]" pageId="127" pageNumber="128">eT</emphasis>
= external tooth;
<emphasis id="B922D319FF8C732F36BAF81CE7FBF867" bold="true" box="[1239,1276,1958,1982]" pageId="127" pageNumber="128">mp</emphasis>
= molar plate;
<emphasis id="B922D319FF8C732F32FAF873E3BFF838" bold="true" box="[151,184,1993,2017]" pageId="127" pageNumber="128">pL</emphasis>
= pectinate lamella.
</paragraph>
<paragraph id="8BE90F0BFF7373D032AAFF22E04AFE43" blockId="128.[151,1436,152,410]" pageId="128" pageNumber="129">
<emphasis id="B922D319FF7373D032AAFF22E2A1FF68" box="[199,422,152,177]" italics="true" pageId="128" pageNumber="129">Posterior telopods</emphasis>
(Figs 86F, G): Movable finger elongated with well-rounded tip, length 3.0 times greater than width, apically only slightly tapering. Hollowed-out inner margin with three sclerotized spines, on posterior side with 10–11 small, crenulated, sclerotized teeth. Movable finger slightly longer than fixed finger. Fixed finger basally broad, with 1 or 2 short, sclerotized spines, apically strongly curved, from curvature towards apical tip much more slender than basally, margin apically covered with sclerotized spots. First podomere glabrous, second and third podomere covered on both sides with hair, especially at lateral margins. Inner horns of syncoxite with pointed, slender tip.
</paragraph>
<paragraph id="8BE90F0BFF7373D032FAFAECE1F4FA6A" blockId="128.[151,1436,1366,1459]" pageId="128" pageNumber="129">
<emphasis id="B922D319FF7373D032FAFAECE23DFAB7" bold="true" box="[151,314,1366,1390]" pageId="128" pageNumber="129">FIGURE 088</emphasis>
. Endoterga of midbody tergites,
<typeStatus id="54EDB1A9FF7373D030B1FAECE041FAB4" box="[732,838,1366,1389]" pageId="128" pageNumber="129" type="holotype">holotypes</typeStatus>
,
<emphasis id="B922D319FF7373D03133FAECE076FAB7" bold="true" box="[862,881,1366,1390]" pageId="128" pageNumber="129">A</emphasis>
:
<taxonomicName id="4C567488FF7373D031E0FAEDE7E6FAB7" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[909,1249,1367,1390]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="micropiligerum" status="sp. nov.">
<emphasis id="B922D319FF7373D031E0FAEDE7E6FAB7" box="[909,1249,1367,1390]" italics="true" pageId="128" pageNumber="129">Zoosphaerium micropiligerum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF7373D03698FAECE639FAB7" bold="true" box="[1269,1342,1366,1390]" pageId="128" pageNumber="129">
<taxonomicNameLabel id="A2116E62FF7373D03698FAECE639FAB7" box="[1269,1342,1366,1390]" pageId="128" pageNumber="129" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FF7373D03734FAECE66CFAB7" bold="true" box="[1369,1387,1366,1390]" pageId="128" pageNumber="129">B</emphasis>
:
<taxonomicName id="4C567488FF7373D037EBFAEDE24DFA49" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="pseudopiligerum" status="sp. nov.">
<emphasis id="B922D319FF7373D037EBFAEDE24DFA49" italics="true" pageId="128" pageNumber="129">Z. pseudopiligerum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF7373D0333EFAC3E296FA48" bold="true" box="[339,401,1401,1425]" pageId="128" pageNumber="129">
<taxonomicNameLabel id="A2116E62FF7373D0333EFAC3E296FA48" box="[339,401,1401,1425]" pageId="128" pageNumber="129" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FF7373D033CCFAC3E2B3FA48" bold="true" box="[417,436,1401,1425]" pageId="128" pageNumber="129">C</emphasis>
:
<taxonomicName id="4C567488FF7373D033A8FAC3E16FFA49" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[453,616,1401,1424]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="analavelona" status="sp. nov.">
<emphasis id="B922D319FF7373D033A8FAC3E16FFA49" box="[453,616,1401,1424]" italics="true" pageId="128" pageNumber="129">Z. analavelona</emphasis>
</taxonomicName>
<emphasis id="B922D319FF7373D0301DFAC3E1A9FA48" bold="true" box="[624,686,1401,1425]" pageId="128" pageNumber="129">
<taxonomicNameLabel id="A2116E62FF7373D0301DFAC3E1A9FA48" box="[624,686,1401,1425]" pageId="128" pageNumber="129" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FF7373D030D2FAC3E1D5FA48" bold="true" box="[703,722,1401,1425]" pageId="128" pageNumber="129">D</emphasis>
:
<taxonomicName id="4C567488FF7373D0308FFAC3E042FA49" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[738,837,1401,1424]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="128" pageNumber="129" phylum="Arthropoda" rank="species" species="fisheri" status="sp. nov.">
<emphasis id="B922D319FF7373D0308FFAC3E042FA49" box="[738,837,1401,1424]" italics="true" pageId="128" pageNumber="129">Z. fisheri</emphasis>
</taxonomicName>
<emphasis id="B922D319FF7373D03123FAC3E08BFA48" bold="true" box="[846,908,1401,1425]" pageId="128" pageNumber="129">
<taxonomicNameLabel id="A2116E62FF7373D03123FAC3E08BFA48" box="[846,908,1401,1425]" pageId="128" pageNumber="129" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Abbreviations:
<emphasis id="B922D319FF7373D03651FAC3E748FA48" bold="true" box="[1084,1103,1401,1425]" pageId="128" pageNumber="129">ci</emphasis>
= cuticular impressions;
<emphasis id="B922D319FF7373D0370BFAC3E684FA48" bold="true" box="[1382,1411,1401,1425]" pageId="128" pageNumber="129">IA</emphasis>
= inner area;
<emphasis id="B922D319FF7373D03362FA21E233FA6A" bold="true" box="[271,308,1435,1459]" pageId="128" pageNumber="129">mb</emphasis>
= marginal bristles;
<emphasis id="B922D319FF7373D0307CFA21E134FA6A" bold="true" box="[529,563,1435,1459]" pageId="128" pageNumber="129">mr</emphasis>
= marginal ridge.
</paragraph>
</subSubSection>
</treatment>
</document>
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