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<document id="CE53E2F28D64B14C5031F97DC3AF18C0" ID-CLB-Dataset="20271" ID-DOI="10.5852/ejt.2022.799.1681" ID-GBIF-Dataset="f53d5465-7162-45d6-892b-dfc0b8d99789" ID-ISSN="2118-9773" ID-Zenodo-Dep="6341270" ID-ZooBank="F52DBAB0-38C7-400F-9BA1-E2D8E6B19E7E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1646831675572" checkinUser="felipe" docAuthor="Aubrechtová, Martina &amp; Korn, Dieter" docDate="2022" docId="03FE5868FFAB3D05DD72FE8BFD2BFB3D" docLanguage="en" docName="ejt-799_1-108.pdf" docOrigin="European Journal of Taxonomy 799 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.799.1681" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Trilacinoceras hunanense Lai &amp; Qi 1977" docType="treatment" docVersion="10" lastPageNumber="95" masterDocId="FFC72010FFF53D5BDF7EFF83FFE3FFD1" masterDocTitle="Taxonomy and ontogeny of the Lituitida (Cephalopoda) from Orthoceratite Limestone erratics (Middle Ordovician)" masterLastPageNumber="108" masterPageNumber="1" pageNumber="95" updateTime="1699317703155" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="750B55A2B712E070DD2D55A75AB01199">Taxonomy and ontogeny of the Lituitida (Cephalopoda) from Orthoceratite Limestone erratics (Middle Ordovician)</mods:title>
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<mods:namePart id="16E576892968BD861A3245D8C1BE88BB">Aubrechtová, Martina</mods:namePart>
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<mods:affiliation id="7018572536445BB7FFCED79BA8D7BF45">Institute of Geology and Palaeontology, Faculty of Science, Charles University Prague, Albertov 6, Prague, 12843, Czech Republic. &amp; Institute of Geology, Czech Academy of Sciences, Rozvojová 269, Prague, 16500, Czech Republic.</mods:affiliation>
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<mods:namePart id="1B014CEE9E890572D025F1F5F5E56CB6">Korn, Dieter</mods:namePart>
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<mods:affiliation id="3E8337F9F3C81A06871BF9831F25B1CC">Museum für Naturkunde, Leibniz-Institut für Evolutions- und Biodiversitätsforschung, Invalidenstrasse 43, 10115 Berlin, Germany.</mods:affiliation>
<mods:nameIdentifier id="43AA7131044C8E66CE2092334C6562E8" type="email">dieter.korn@mfn.berlin</mods:nameIdentifier>
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<mods:title id="65ACF8F53084EFF729CCF60B6EFFD05A">European Journal of Taxonomy</mods:title>
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<mods:date id="D471A3EDC8A010CD960612BAA6A13B54">2022</mods:date>
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<treatment id="03FE5868FFAB3D05DD72FE8BFD2BFB3D" ID-DOI="http://doi.org/10.5281/zenodo.6343283" ID-GBIF-Taxon="193941456" ID-Zenodo-Dep="6343283" LSID="urn:lsid:plazi:treatment:03FE5868FFAB3D05DD72FE8BFD2BFB3D" httpUri="http://treatment.plazi.org/id/03FE5868FFAB3D05DD72FE8BFD2BFB3D" lastPageNumber="95" pageId="94" pageNumber="95">
<subSubSection id="C34DBAF5FFAB3D05DD72FE8BFBCBFEF2" box="[524,1064,264,291]" pageId="94" pageNumber="95" type="nomenclature">
<paragraph id="8BE8E97EFFAB3D05DD72FE8BFBCBFEF2" blockId="94.[524,1064,264,326]" box="[524,1064,264,291]" pageId="94" pageNumber="95">
<heading id="D0A05E12FFAB3D05DD72FE8BFBCBFEF2" box="[524,1064,264,291]" centered="true" fontSize="11" level="2" pageId="94" pageNumber="95" reason="2">
<taxonomicName id="4C5792FDFFAB3D05DD72FE8BFBCBFEF2" authority="Lai &amp; Qi, 1977" authorityName="Lai &amp; Qi" authorityYear="1977" box="[524,1064,264,291]" class="Cephalopoda" family="Lituitidae" genus="Trilacinoceras" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Orthocerida" pageId="94" pageNumber="95" phylum="Mollusca" rank="species" species="hunanense">
<emphasis id="B923356CFFAB3D05DD72FE8BFD58FEF2" bold="true" box="[524,699,264,291]" italics="true" pageId="94" pageNumber="95">Trilacinoceras</emphasis>
cf.
<emphasis id="B923356CFFAB3D05DD99FE8BFC8EFEF2" bold="true" box="[743,877,264,291]" italics="true" pageId="94" pageNumber="95">hunanense</emphasis>
Lai &amp; Qi, 1977
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34DBAF5FFAB3D05DD07FEAFFC58FE97" box="[633,955,300,326]" pageId="94" pageNumber="95" type="description">
<paragraph id="8BE8E97EFFAB3D05DD07FEAFFC58FE97" blockId="94.[524,1064,264,326]" box="[633,955,300,326]" pageId="94" pageNumber="95">
<figureCitation id="136CF5FBFFAB3D05DD07FEAFFD05FE97" box="[633,742,300,326]" captionStart="Fig" captionStartId="92.[189,232,1733,1759]" captionTargetBox="[225,1361,278,1639]" captionTargetId="figure-15@92.[206,1381,268,1653]" captionTargetPageId="92" captionText="Fig. 69. Species of Trilacinoceras Sweet, 1958. A. Trilacinoceras filix sp. nov., holotype MB.C.30557 (Neben &amp; Krueger 1961 Coll.) from the Upper Grey Orthoceratite Limestone of Rerik (Ostsee) (Mecklenburg-Vorpommern); previously figured by Neben &amp; Krueger (1971: pl. 31 fig. 4). B. Trilacinoceras cf. hunanense Lai &amp; Qi, 1977, specimen MB.C.29655 (Bottke Coll.) from the Datianba Formation of Songtao County (Guizhou, South China). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6341424" httpUri="https://zenodo.org/record/6341424/files/figure.png" pageId="94" pageNumber="95">Figs 69B</figureCitation>
,
<figureCitation id="136CF5FBFFAB3D05DD8CFEAFFCF1FE97" box="[754,786,300,326]" captionStart="Fig" captionStartId="94.[189,232,1941,1967]" captionTargetBox="[262,1325,1422,1879]" captionTargetId="figure-334@94.[261,1327,1420,1881]" captionTargetPageId="94" captionText="Fig. 71. Trilacinoceras cf. hunanense Lai &amp; Qi, 1977. A. Coiling outline of specimen MB.C.29655. B–E. Conch proportions and coiling rates of specimen MB.C.29655. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6341428" httpUri="https://zenodo.org/record/6341428/files/figure.png" pageId="94" pageNumber="95">71</figureCitation>
,
<tableCitation id="C6D5DCC5FFAB3D05DC60FEAFFC58FE97" box="[798,955,300,326]" captionStart-0="Table 12" captionStart-1="Table 13" captionStartId-0="90.[189,259,265,291]" captionStartId-1="93.[189,259,265,291]" captionTargetPageId-0="90" captionTargetPageId-1="93" captionText-0="Table 12. Conch measurements (in mm), ratios and rates of coiled conch parts of Trilacinoceras Sweet, 1958." captionText-1="Table 13. Conch measurements (in mm), and expansion rates of uncoiled conch parts of Trilacinoceras Sweet, 1958." pageId="94" pageNumber="95">Tables 12–13</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34DBAF5FFAB3D05DFC3FEEEFBAAFE66" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="8BE8E97EFFAB3D05DFC3FEEEFE49FE56" blockId="94.[189,426,365,391]" box="[189,426,365,391]" pageId="94" pageNumber="95">
<heading id="D0A05E12FFAB3D05DFC3FEEEFE49FE56" bold="true" box="[189,426,365,391]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="B923356CFFAB3D05DFC3FEEEFE49FE56" bold="true" box="[189,426,365,391]" pageId="94" pageNumber="95">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BE8E97EFFAB3D05DFC3FE1FFBAAFE66" blockId="94.[189,1097,412,439]" box="[189,1097,412,439]" pageId="94" pageNumber="95">
<materialsCitation id="3B3FE323FFAB3D05DFC3FE1FFBA7FE66" ID-GBIF-Occurrence="3700114366" box="[189,1092,412,439]" collectionCode="MB" collectorName="Bottke" country="China" location="Songtao" municipality="Songtao" pageId="94" pageNumber="95" specimenCode="MB.C.29655" specimenCount="1" stateProvince="Guizhou">
<collectingCountry id="F340A9EEFFAB3D05DFC3FE1FFEFBFE67" box="[189,280,412,438]" name="China" pageId="94" pageNumber="95">CHINA</collectingCountry>
•
<specimenCount id="9D5122F7FFAB3D05DE4FFE1FFE65FE66" box="[305,390,412,439]" pageId="94" pageNumber="95" type="generic">1 spec.</specimenCount>
;
<collectingRegion id="4993279CFFAB3D05DEECFE1FFE19FE67" box="[402,506,412,438]" country="China" name="Guizhou" pageId="94" pageNumber="95">Guizhou</collectingRegion>
,
<collectingMunicipality id="6B8C7304FFAB3D05DD78FE1FFD89FE67" box="[518,618,412,438]" pageId="94" pageNumber="95">Songtao</collectingMunicipality>
;
<geologicalTimeScale id="F97769D5FFAB3D05DD0BFE1FFD1EFE67" box="[629,765,412,438]" pageId="94" pageNumber="95">Ordovician</geologicalTimeScale>
;
<collectorName id="26A28CA8FFAB3D05DC77FE1FFCB4FE67" box="[777,855,412,438]" pageId="94" pageNumber="95">Bottke</collectorName>
Coll.;
<specimenCode id="DBF14105FFAB3D05DCD6FE1FFBA7FE66" box="[936,1092,412,439]" pageId="94" pageNumber="95">MB.C.29655</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34DBAF5FFAB3D05DFC3FE5EFC5FFC1D" pageId="94" pageNumber="95" type="description">
<paragraph id="8BE8E97EFFAB3D05DFC3FE5EFEADFE26" blockId="94.[189,334,477,503]" box="[189,334,477,503]" pageId="94" pageNumber="95">
<heading id="D0A05E12FFAB3D05DFC3FE5EFEADFE26" bold="true" box="[189,334,477,503]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="B923356CFFAB3D05DFC3FE5EFEADFE26" bold="true" box="[189,334,477,503]" pageId="94" pageNumber="95">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BE8E97EFFAB3D05DFC3FD8FFC5FFC1D" blockId="94.[188,1401,524,973]" pageId="94" pageNumber="95">
Specimen MB.C.29655 (
<figureCitation id="136CF5FBFFAB3D05DE90FD8FFDBEFDF6" box="[494,605,524,551]" captionStart="Fig" captionStartId="92.[189,232,1733,1759]" captionTargetBox="[225,1361,278,1639]" captionTargetId="figure-15@92.[206,1381,268,1653]" captionTargetPageId="92" captionText="Fig. 69. Species of Trilacinoceras Sweet, 1958. A. Trilacinoceras filix sp. nov., holotype MB.C.30557 (Neben &amp; Krueger 1961 Coll.) from the Upper Grey Orthoceratite Limestone of Rerik (Ostsee) (Mecklenburg-Vorpommern); previously figured by Neben &amp; Krueger (1971: pl. 31 fig. 4). B. Trilacinoceras cf. hunanense Lai &amp; Qi, 1977, specimen MB.C.29655 (Bottke Coll.) from the Datianba Formation of Songtao County (Guizhou, South China). Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6341424" httpUri="https://zenodo.org/record/6341424/files/figure.png" pageId="94" pageNumber="95">Fig. 69B</figureCitation>
) is a
<quantity id="4CAF449BFFAB3D05DDCEFD8FFCF4FDF6" box="[688,791,524,551]" metricMagnitude="-1" metricUnit="m" metricValue="3.26" pageId="94" pageNumber="95" unit="mm" value="326.0">326 mm</quantity>
long conch, which is partially embedded in the surrounding dark-brown limestone. The coiled part has a diameter of
<quantity id="4CAF449BFFAB3D05DC9FFDACFBD0FD9B" box="[993,1075,559,586]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="94" pageNumber="95" unit="mm" value="36.0">36 mm</quantity>
and consists of at least 2.25 tightly coiled volutions (initial chamber is not preserved); the whorl expansion rate is 2.10 (
<figureCitation id="136CF5FBFFAB3D05DA70FDD1FA8BFDBC" box="[1294,1384,594,621]" captionStart="Fig" captionStartId="94.[189,232,1941,1967]" captionTargetBox="[262,1325,1422,1879]" captionTargetId="figure-334@94.[261,1327,1420,1881]" captionTargetPageId="94" captionText="Fig. 71. Trilacinoceras cf. hunanense Lai &amp; Qi, 1977. A. Coiling outline of specimen MB.C.29655. B–E. Conch proportions and coiling rates of specimen MB.C.29655. Scale bar units = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6341428" httpUri="https://zenodo.org/record/6341428/files/figure.png" pageId="94" pageNumber="95">Fig. 71</figureCitation>
). The uncoiled part is relatively strongly curved and expands with an angle of 4°. Ornament is preserved along the whole length of the specimen and is slightly irregular in distance. In the coiled part, the ornament consists of growth lines and raised lirae; in inner whorls, the ornament elements are straight, later, a shallow sinus is developed ventrolaterally. In the rest of the conch, narrow annuli (up to
<quantity id="4CAF449BFFAB3D05DA60FD5CFA94FD2B" box="[1310,1399,735,762]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="94" pageNumber="95" unit="mm" value="2.7">2.7 mm</quantity>
in distance) and lirae (up to
<quantity id="4CAF449BFFAB3D05DD63FC80FD98FCCC" box="[541,635,771,797]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="94" pageNumber="95" unit="mm" value="0.3">0.3 mm</quantity>
apart) are present; the distance and thickness of the ornament elements is irregular; the conch is slightly constricted, where the ornament elements are the finest. The ornament elements in the backcoiled part have a rectiradiate direction but towards the straight part, they become rursiradiate; a marked dorsal projection and a sinus between the mid-flank and the ventrolateral projection is developed. The adapertural part of the specimen might represent the body chamber but terminal aperture is not preserved and suture lines are not visible.
</paragraph>
</subSubSection>
<subSubSection id="C34DBAF5FFAB3D05DFC3FC77FD2BFB3D" pageId="94" pageNumber="95" type="discussion">
<paragraph id="8BE8E97EFFAB3D05DFC3FC77FECCFBDF" blockId="94.[189,303,1012,1038]" box="[189,303,1012,1038]" pageId="94" pageNumber="95">
<heading id="D0A05E12FFAB3D05DFC3FC77FECCFBDF" bold="true" box="[189,303,1012,1038]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="B923356CFFAB3D05DFC3FC77FECCFBDF" bold="true" box="[189,303,1012,1038]" pageId="94" pageNumber="95">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE8E97EFFAB3D05DFC3FBA1FD2BFB3D" blockId="94.[188,1399,1058,1261]" pageId="94" pageNumber="95">
<taxonomicName id="4C5792FDFFAB3D05DFC3FBA1FDFFFBED" authorityName="Lai &amp; Qi" authorityYear="1977" box="[189,540,1058,1084]" class="Cephalopoda" family="Lituitidae" genus="Trilacinoceras" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Orthocerida" pageId="94" pageNumber="95" phylum="Mollusca" rank="species" species="hunanense">
<emphasis id="B923356CFFAB3D05DFC3FBA1FE8BFBED" box="[189,360,1058,1084]" italics="true" pageId="94" pageNumber="95">Trilacinoceras</emphasis>
cf.
<emphasis id="B923356CFFAB3D05DEE1FBA1FDFFFBED" box="[415,540,1058,1084]" italics="true" pageId="94" pageNumber="95">hunanense</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5792FDFFAB3D05DDB6FBA0FC84FBED" authorityName="Lai &amp; Qi" authorityYear="1977" box="[712,871,1058,1084]" class="Cephalopoda" family="Lituitidae" genus="Trilacinoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="94" pageNumber="95" phylum="Mollusca" rank="species" species="hunanense">
<emphasis id="B923356CFFAB3D05DDB6FBA0FC84FBED" box="[712,871,1058,1084]" italics="true" pageId="94" pageNumber="95">T. hunanense</emphasis>
</taxonomicName>
in a smaller diameter of the coiled conch (
<quantity id="4CAF449BFFAB3D05DFBBFBC5FEFFFBB1" box="[197,284,1094,1120]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="94" pageNumber="95" unit="mm" value="36.0">36 mm</quantity>
vs
<quantity id="4CAF449BFFAB3D05DE37FBC5FE43FBB1" box="[329,416,1094,1120]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="94" pageNumber="95" unit="mm" value="40.0">40 mm</quantity>
) and a more strongly curved (vs straight) uncoiled part (Xiang Fang, pers. com.).
<taxonomicName id="4C5792FDFFAB3D05DFC1FBEAFE3BFB52" authority="Aubrechtová &amp; Korn, 2022" authorityName="Aubrechtová &amp; Korn" authorityYear="2022" box="[191,472,1129,1155]" class="Cephalopoda" family="Lituitidae" genus="Trilacinoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthocerida" pageId="94" pageNumber="95" phylum="Mollusca" rank="species" species="knoefleri" status="sp. nov.">
<emphasis id="B923356CFFAB3D05DFC1FBEAFE3BFB52" box="[191,472,1129,1155]" italics="true" pageId="94" pageNumber="95">Trilacinoceras knoefleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2108817FFAB3D05DEA2FBEAFDD5FB52" box="[476,566,1129,1155]" pageId="94" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished by its smaller expansion angle and coarser, rectiradiate ornament of the backcoiled part. The rursiradiate direction of ornament elements in the straight conch part of
<taxonomicName id="4C5792FDFFAB3D05DE64FB2CFD9BFB18" authorityName="Lai &amp; Qi" authorityYear="1977" box="[282,632,1199,1225]" class="Cephalopoda" family="Lituitidae" genus="Trilacinoceras" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Animalia" order="Orthocerida" pageId="94" pageNumber="95" phylum="Mollusca" rank="species" species="hunanense">
<emphasis id="B923356CFFAB3D05DE64FB2CFE26FB18" box="[282,453,1199,1225]" italics="true" pageId="94" pageNumber="95">Trilacinoceras</emphasis>
cf.
<emphasis id="B923356CFFAB3D05DE85FB2CFD9BFB18" box="[507,632,1199,1225]" italics="true" pageId="94" pageNumber="95">hunanense</emphasis>
</taxonomicName>
distinguishes the Chinese specimen from all specimens in the collection that come from European regions.
</paragraph>
</subSubSection>
</treatment>
</document>
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