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<document id="CC4688E8CD6E9BB51B719A2CF0A8FB8A" ID-CLB-Dataset="20340" ID-DOI="10.11646/zootaxa.5105.3.2" ID-GBIF-Dataset="4f25a761-8f7b-47d3-84c9-ad2917b90f74" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332731" ID-ZooBank="83013C70-69E4-42BD-8CDD-0FA320927290" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646597716981" checkinUser="plazi" docAuthor="Bhansali, Sneha &amp; Wesener, Thomas" docDate="2022" docId="03FBA627FFBCAD0FE389F8C5FD28F821" docLanguage="en" docName="zootaxa.5105.3.2.pdf" docOrigin="Zootaxa 5105 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zephronia erawani Bhansali &amp; Wesener 2022, new species" docType="treatment" docVersion="8" lastPageNumber="374" masterDocId="FFC2DE5FFFB1AD1EE31EFF84FF89FFAF" masterDocTitle="New Thai giant pill-millipede species, with new genetic barcoding data (Diplopoda Sphaerotheriida, Zephroniidae)" masterLastPageNumber="380" masterPageNumber="357" pageNumber="370" updateTime="1699317184639" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="C196293F5DE515C6A370C0C3F1C6303E">
<mods:title id="DEF5BA2DFB9CAAEADF81935BD85DA1BF">New Thai giant pill-millipede species, with new genetic barcoding data (Diplopoda Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name id="FF1974F950A50CFE8E87CD4E3D13C447" type="personal">
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<mods:roleTerm id="270025DAEADFD531E6C403708782FD4F">Author</mods:roleTerm>
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<mods:namePart id="A05744228939A99160C22D5C4D288DAA">Bhansali, Sneha</mods:namePart>
<mods:nameIdentifier id="99C13FAB8651A04CC919D62D267B614B" type="ORCID">0000-0002-5440-9958</mods:nameIdentifier>
<mods:affiliation id="895E51C63874E09E130000C5761B0549">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
<mods:nameIdentifier id="666843CE2DEDC9BF12844685483D85B0" type="email">sneharbhansali@gmail.com</mods:nameIdentifier>
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<mods:namePart id="12BF3D31E81FD776C795651B85FEDAB9">Wesener, Thomas</mods:namePart>
<mods:nameIdentifier id="2EDA8B84BC35238AC3D0945885A53D90" type="ORCID">0000-0002-2028-3541</mods:nameIdentifier>
<mods:affiliation id="6D18336B9E14F8995BE0E9C86F217989">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 160, D- 53113 Bonn, Germany. &amp; T. Wesener @ leibniz-zfmk. de; https: // orcid. org / 0000 - 0002 - 2028 - 3541</mods:affiliation>
<mods:nameIdentifier id="3F86D7645411E4509C47B851967F6B95" type="email">esener@leibniz-zfmk.de</mods:nameIdentifier>
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<mods:title id="351154B63707F194C5FEA51B769125E0">Zootaxa</mods:title>
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<mods:part id="398E6633197D4381AF9E620BA362108D">
<mods:date id="CD65ADF95320955D402A6293626CF6CE">2022</mods:date>
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<mods:number id="1D5DC0136A1EF9CBEEA4EE8D2C353EAD">2022-03-04</mods:number>
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<mods:number id="1A428D291B86F3B1DC0463973E9C76E5">5105</mods:number>
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<treatment id="03FBA627FFBCAD0FE389F8C5FD28F821" ID-DOI="http://doi.org/10.5281/zenodo.6333447" ID-GBIF-Taxon="193890959" ID-Zenodo-Dep="6333447" LSID="urn:lsid:plazi:treatment:03FBA627FFBCAD0FE389F8C5FD28F821" httpUri="http://treatment.plazi.org/id/03FBA627FFBCAD0FE389F8C5FD28F821" lastPageId="17" lastPageNumber="374" pageId="13" pageNumber="370">
<subSubSection id="C34844BAFFBCAD13E389F8C5FC92F8F3" box="[151,795,1857,1884]" pageId="13" pageNumber="370" type="nomenclature">
<paragraph id="8BED1731FFBCAD13E389F8C5FC92F8F3" blockId="13.[151,795,1857,1920]" box="[151,795,1857,1884]" pageId="13" pageNumber="370">
<heading id="D0A5A05DFFBCAD13E389F8C5FC92F8F3" bold="true" box="[151,795,1857,1884]" fontSize="11" level="1" pageId="13" pageNumber="370" reason="1">
<emphasis id="B926CB23FFBCAD13E389F8C5FC92F8F3" bold="true" box="[151,795,1857,1884]" pageId="13" pageNumber="370">
<taxonomicName id="4C526CB2FFBCAD13E389F8C5FDF6F8F3" ID-CoL="9BDJT" authority="Bhansali &amp;Wesener" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[151,639,1857,1884]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="370" phylum="Arthropoda" rank="species" species="erawani" status="sp. nov.">
<emphasis id="B926CB23FFBCAD13E389F8C5FE09F8F3" bold="true" box="[151,384,1857,1884]" italics="true" pageId="13" pageNumber="370">Zephronia erawani</emphasis>
Bhansali &amp;Wesener
</taxonomicName>
,
<taxonomicNameLabel id="A2157658FFBCAD13E195F8C6FC92F8F3" box="[651,795,1858,1884]" pageId="13" pageNumber="370" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBCAD13E389F8E2FE10F82F" box="[151,409,1894,1920]" pageId="13" pageNumber="370" type="description">
<paragraph id="8BED1731FFBCAD13E389F8E2FE10F82F" blockId="13.[151,795,1857,1920]" box="[151,409,1894,1920]" pageId="13" pageNumber="370">
<figureCitation id="13690BB4FFBCAD13E389F8E2FE9EF82F" box="[151,279,1894,1920]" captionStart="FIGURE 2" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[337,1247,181,1766]" captionTargetId="figure-22@6.[337,1249,181,1766]" captionTargetPageId="6" captionText="FIGURE 2. Map of Thailand with known distribution of Sphaerobelum spp. and Zephronia spp. Star represents new species. Diamonds represent new localities. Fluorescent Green = S. meridionalis Bhansali &amp; Wesener sp. nov.; Pink = S. aesculus; Brown = S. truncatum; Chrome yellow = Z. chrysomallos Bhansali &amp; Wesener sp. nov.; White = Z. erawani Bhansali &amp; Wesener sp. nov.; Black = Z. viridisoma; Red = Z. panhai; Dark green = Z. golovatchi; Purple = Z. siamensis; Orange = Z. enghoffi; Yellow = Z. lannaensis; Blue = Z. phrain. B. Habitus photograph of Z. chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♀ (ZFMK MYR8826)." figureDoi="http://doi.org/10.5281/zenodo.6332735" httpUri="https://zenodo.org/record/6332735/files/figure.png" pageId="13" pageNumber="370">Figures 2A</figureCitation>
,
<figureCitation id="13690BB4FFBCAD13E23CF8E2FEC9F82F" box="[290,320,1894,1920]" captionStart="FIGURE 12" captionStartId="16.[151,250,1442,1466]" captionTargetBox="[164,1423,489,1412]" captionTargetId="figure-162@16.[164,1423,489,1418]" captionTargetPageId="16" captionText="FIGURE 12. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), volume rendering based on microcomputed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cp = central pad; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; o = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Po = postfemur; Pre = prefemur; Pt = paratergite; Stp = stigmatic plate; St = stipites; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332755" httpUri="https://zenodo.org/record/6332755/files/figure.png" pageId="13" pageNumber="370">12</figureCitation>
,
<figureCitation id="13690BB4FFBCAD13E252F8E2FEE3F82F" box="[332,362,1894,1920]" captionStart="FIGURE 13" captionStartId="17.[151,250,1297,1321]" captionTargetBox="[151,1436,188,1273]" captionTargetId="figure-17@17.[151,1436,181,1273]" captionTargetPageId="17" captionText="FIGURE 13. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (ZMUC K56-9), drawings, A. Left leg 9, posterior view B. Second left coxa with gonopore, posterior view. C. Right anterior telopod, anterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332757" httpUri="https://zenodo.org/record/6332757/files/figure.png" pageId="13" pageNumber="370">13</figureCitation>
,
<figureCitation id="13690BB4FFBCAD13E268F8E2FE1CF82F" box="[374,405,1894,1920]" captionStart="FIGURE 14" captionStartId="18.[151,250,1492,1516]" captionTargetBox="[151,1436,185,1468]" captionTargetId="figure-22@18.[151,1436,181,1468]" captionTargetPageId="18" captionText="FIGURE 14. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), drawings, A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view. Abbreviations: ct = crenulated teeth; imf = immovable finger; ss = sclerotized spot. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332759" httpUri="https://zenodo.org/record/6332759/files/figure.png" pageId="13" pageNumber="370">14</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBCAD10E389F82AFADDFE04" lastPageId="14" lastPageNumber="371" pageId="13" pageNumber="370" type="diagnosis">
<paragraph id="8BED1731FFBCAD10E389F82AFC66FECC" blockId="13.[151,1436,1966,2028]" lastBlockId="14.[151,1437,149,680]" lastPageId="14" lastPageNumber="371" pageId="13" pageNumber="370">
<emphasis id="B926CB23FFBCAD13E389F82AFE83F867" bold="true" box="[151,266,1966,1992]" pageId="13" pageNumber="370">Diagnosis</emphasis>
. The position of the organ of Tömösváry at the brim and not in the antennal groove identifies this species as a member of the
<taxonomicName id="4C526CB2FFBCAD13E269F856FE62F843" authorityName="Gray" authorityYear="1832" box="[375,491,2002,2028]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B926CB23FFBCAD13E269F856FE62F843" box="[375,491,2002,2028]" italics="true" pageId="13" pageNumber="370">Zephronia</emphasis>
</taxonomicName>
senso-stricto group with which it also aligns weakly genetically. The sensory cones on the palpi of the gnathochilarium are arranged in a single field, not in separate clusters and the body is covered in a dense coat of golden setae like in several other Thai
<taxonomicName id="4C526CB2FFBFAD10E03AFF3DFC11FF7C" authorityName="Gray" authorityYear="1832" box="[804,920,185,211]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B926CB23FFBFAD10E03AFF3DFC11FF7C" box="[804,920,185,211]" italics="true" pageId="14" pageNumber="371">Zephronia</emphasis>
</taxonomicName>
. The greyish-brown color resembles those of
<taxonomicName id="4C526CB2FFBFAD10E389FF5AFEC2FF58" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[151,331,221,247]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBFAD10E389FF5AFEC2FF58" box="[151,331,221,247]" italics="true" pageId="14" pageNumber="371">Z. chrysomallos</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBFAD10E24AFF59FE27FF58" bold="true" box="[340,430,221,247]" pageId="14" pageNumber="371">
<taxonomicNameLabel id="A2157658FFBFAD10E24AFF59FE27FF58" box="[340,430,221,247]" pageId="14" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C526CB2FFBFAD10E2F9FF5AFDDDFF58" box="[487,596,221,247]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="B926CB23FFBFAD10E2F9FF5AFDDDFF58" box="[487,596,221,247]" italics="true" pageId="14" pageNumber="371">Z. panhai</emphasis>
</taxonomicName>
. The male second leg mesal coxal margin is extended into a membranous process (
<figureCitation id="13690BB4FFBFAD10E3E2FE85FEE8FEB4" box="[252,353,257,283]" captionStart="FIGURE 13" captionStartId="17.[151,250,1297,1321]" captionTargetBox="[151,1436,188,1273]" captionTargetId="figure-17@17.[151,1436,181,1273]" captionTargetPageId="17" captionText="FIGURE 13. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (ZMUC K56-9), drawings, A. Left leg 9, posterior view B. Second left coxa with gonopore, posterior view. C. Right anterior telopod, anterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332757" httpUri="https://zenodo.org/record/6332757/files/figure.png" pageId="14" pageNumber="371">Fig. 13B</figureCitation>
arrow), a character shared only with
<taxonomicName id="4C526CB2FFBFAD10E00FFE86FC2FFEB4" box="[785,934,257,283]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="B926CB23FFBFAD10E00FFE86FC2FFEB4" box="[785,934,257,283]" italics="true" pageId="14" pageNumber="371">Z. golovatchi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C526CB2FFBFAD10E0FEFE86FBD5FEB4" box="[992,1116,257,283]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="enghoffi">
<emphasis id="B926CB23FFBFAD10E0FEFE86FBD5FEB4" box="[992,1116,257,283]" italics="true" pageId="14" pageNumber="371">Z. enghoffi</emphasis>
</taxonomicName>
.
<taxonomicName id="4C526CB2FFBFAD10E775FE86FB6EFEB4" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[1131,1255,258,283]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="erawani" status="sp. nov.">
<emphasis id="B926CB23FFBFAD10E775FE86FB6EFEB4" box="[1131,1255,258,283]" italics="true" pageId="14" pageNumber="371">Z. erawani</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBFAD10E7F1FE85FAC2FEB4" bold="true" box="[1263,1355,257,283]" pageId="14" pageNumber="371">
<taxonomicNameLabel id="A2157658FFBFAD10E7F1FE85FAC2FEB4" box="[1263,1355,257,283]" pageId="14" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from both
<taxonomicName id="4C526CB2FFBFAD10E212FEA2FE16FE90" box="[268,415,293,319]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="golovatchi">
<emphasis id="B926CB23FFBFAD10E212FEA2FE16FE90" box="[268,415,293,319]" italics="true" pageId="14" pageNumber="371">Z. golovatchi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C526CB2FFBFAD10E2CAFEA2FDC5FE90" box="[468,588,293,319]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="enghoffi">
<emphasis id="B926CB23FFBFAD10E2CAFEA2FDC5FE90" box="[468,588,293,319]" italics="true" pageId="14" pageNumber="371">Z. enghoffi</emphasis>
</taxonomicName>
in the shorter marginal bristles of the endotergum which end well before the margin, while the bristles are projecting above the two in both other species.
</paragraph>
<paragraph id="8BED1731FFBFAD10E3D9FEEAFADDFE04" blockId="14.[151,1437,149,680]" pageId="14" pageNumber="371">
<taxonomicName id="4C526CB2FFBFAD10E3D9FEEAFEC9FE28" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[199,320,366,391]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="erawani" status="sp. nov.">
<emphasis id="B926CB23FFBFAD10E3D9FEEAFEC9FE28" box="[199,320,366,391]" italics="true" pageId="14" pageNumber="371">Z. erawani</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBFAD10E258FEE9FE16FE28" bold="true" box="[326,415,365,391]" pageId="14" pageNumber="371">
<taxonomicNameLabel id="A2157658FFBFAD10E258FEE9FE16FE28" box="[326,415,365,391]" pageId="14" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs genetically in the COI barcoding fragment from
<taxonomicName id="4C526CB2FFBFAD10E70EFEEAFBF2FE28" box="[1040,1147,365,391]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="B926CB23FFBFAD10E70EFEEAFBF2FE28" box="[1040,1147,365,391]" italics="true" pageId="14" pageNumber="371">Z. panhai</emphasis>
</taxonomicName>
by a p-distance of 11.6%, from
<taxonomicName id="4C526CB2FFBFAD10E3CBFE16FE0EFE04" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[213,391,401,427]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBFAD10E3CBFE16FE0EFE04" box="[213,391,401,427]" italics="true" pageId="14" pageNumber="371">Z. chrysomallos</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBFAD10E291FE15FE61FE04" bold="true" box="[399,488,401,427]" pageId="14" pageNumber="371">
<taxonomicNameLabel id="A2157658FFBFAD10E291FE15FE61FE04" box="[399,488,401,427]" pageId="14" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by a p-distance of 11.9% and from
<taxonomicName id="4C526CB2FFBFAD10E067FE16FB86FE04" box="[889,1039,401,427]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="lannaensis">
<emphasis id="B926CB23FFBFAD10E067FE16FB86FE04" box="[889,1039,401,427]" italics="true" pageId="14" pageNumber="371">Z. lannaensis</emphasis>
</taxonomicName>
by 12.6–13.3% (Sup. file 1).
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBFAD10E3D9FE30FD4DFE5C" pageId="14" pageNumber="371" type="etymology">
<paragraph id="8BED1731FFBFAD10E3D9FE30FD4DFE5C" blockId="14.[151,1437,149,680]" pageId="14" pageNumber="371">
<emphasis id="B926CB23FFBFAD10E3D9FE30FE17FE61" bold="true" box="[199,414,436,462]" pageId="14" pageNumber="371">Derivatio nominis</emphasis>
. Named after the
<typeStatus id="54E9A993FFBFAD10E173FE31FD14FE60" box="[621,669,437,463]" pageId="14" pageNumber="371">type</typeStatus>
locality, the Erawan waterfalls. Erawan is a mythological elephant, similar to the Hindi Airavata. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBFAD10E3D9FE78FAEBFD2C" pageId="14" pageNumber="371" type="materials_examined">
<paragraph id="8BED1731FFBFAD10E3D9FE78FDA7FDB8" blockId="14.[151,1437,149,680]" box="[199,558,508,535]" pageId="14" pageNumber="371">
<emphasis id="B926CB23FFBFAD10E3D9FE78FE20FDB9" bold="true" box="[199,425,508,534]" pageId="14" pageNumber="371">Material examined</emphasis>
(total:
<specimenCount id="9D54DCB8FFBFAD10E2E4FE79FDADFDB8" box="[506,548,509,535]" pageId="14" pageNumber="371" type="male">1 ♂</specimenCount>
)
</paragraph>
<paragraph id="8BED1731FFBFAD10E3D9FDA5FEB1FD94" blockId="14.[151,1437,149,680]" box="[199,312,545,571]" pageId="14" pageNumber="371">
<typeStatus id="54E9A993FFBFAD10E3D9FDA5FEBDFD94" box="[199,308,545,571]" pageId="14" pageNumber="371" type="holotype">Holotype</typeStatus>
:
</paragraph>
<paragraph id="8BED1731FFBFAD10E3D9FDC1FAEBFD2C" blockId="14.[151,1437,149,680]" pageId="14" pageNumber="371">
<materialsCitation id="3B3A1D6CFFBFAD10E3D9FDC1FAEBFD2C" ID-GBIF-Occurrence="3698753330" collectingDate="1990-06-06" collectorName="A. R. Rasmussen &amp; B. Helwigh." country="Thailand" elevation="377" latitude="14.358701" location="Erawan Waterfall" longLatPrecision="1" longitude="99.14062" municipality="Si Sawat District" pageId="14" pageNumber="371" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi" typeStatus="holotype">
<collectingCountry id="F34557A1FFBFAD10E3D9FDC1FED5FDF0" box="[199,348,581,607]" name="Thailand" pageId="14" pageNumber="371">THAILAND</collectingCountry>
:
<specimenCount id="9D54DCB8FFBFAD10E27AFDC1FE04FDF0" box="[356,397,581,607]" pageId="14" pageNumber="371" type="male">1 ♂</specimenCount>
,
<emphasis id="B926CB23FFBFAD10E289FDC1FDB4FDF0" bold="true" box="[407,573,581,607]" pageId="14" pageNumber="371">NHMD K56-9</emphasis>
,
<collectingRegion id="4996D9D3FFBFAD10E156FDC1FCD9FDF0" box="[584,848,581,607]" country="Thailand" name="Kanchanaburi" pageId="14" pageNumber="371">Kanchanaburi Province</collectingRegion>
,
<collectingMunicipality id="6B898D4BFFBFAD10E046FDC1FB9DFDF0" box="[856,1044,581,607]" pageId="14" pageNumber="371">Si Sawat District</collectingMunicipality>
,
<quantity id="4CAABAD4FFBFAD10E702FDC1FBE8FDF0" box="[1052,1121,581,608]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="14" pageNumber="371" unit="km" value="50.0">50 km</quantity>
W of
<collectingRegion id="4996D9D3FFBFAD10E780FDC1FAB7FDF0" box="[1182,1342,581,607]" country="Thailand" name="Kanchanaburi" pageId="14" pageNumber="371">Kanchanaburi</collectingRegion>
,
<location id="8E8D41EAFFBFAD10E659FDC1FE8BFD2C" LSID="urn:lsid:plazi:treatment:03FBA627FFBCAD0FE389F8C5FD28F821:8E8D41EAFFBFAD10E659FDC1FE8BFD2C" country="Thailand" latitude="14.358701" longLatPrecision="1" longitude="99.14062" municipality="Si Sawat District" name="Erawan Waterfall" pageId="14" pageNumber="371" stateProvince="Kanchanaburi">Erawan Waterfall</location>
, [probably:
<geoCoordinate id="EE6671F6FFBFAD10E296FDEDFDB8FD2C" box="[392,561,617,643]" degrees="14" direction="north" minutes="21" orientation="latitude" pageId="14" pageNumber="371" precision="1" seconds="31.32" value="14.358701">14°21'31.32&quot;N</geoCoordinate>
,
<geoCoordinate id="EE6671F6FFBFAD10E123FDEDFD55FD2C" box="[573,732,617,643]" degrees="99" direction="east" minutes="8" orientation="longitude" pageId="14" pageNumber="371" precision="1" seconds="26.23" value="99.14062">99° 8'26.23&quot;E</geoCoordinate>
,
<quantity id="4CAABAD4FFBFAD10E1F6FDEDFCBAFD2C" box="[744,819,616,643]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="14" pageNumber="371" unit="m" value="377.0">
<elevation id="007FF002FFBFAD10E1F6FDEDFCBAFD2C" box="[744,819,616,643]" metricMagnitude="2" metricUnit="m" metricValue="3.77" pageId="14" pageNumber="371" unit="m" value="377.0">377 m</elevation>
</quantity>
],
<date id="FFEC31F1FFBFAD10E058FDEDFC33FD2C" box="[838,954,617,643]" pageId="14" pageNumber="371" value="1990-06-06">
<collectingDate id="EFA8C819FFBFAD10E058FDEDFC33FD2C" box="[838,954,617,643]" pageId="14" pageNumber="371" value="1990-06-06">6.VI.1990</collectingDate>
</date>
, leg.
<collectorName id="26A772E7FFBFAD10E0EAFDEDFB3CFD2C" box="[1012,1205,617,643]" pageId="14" pageNumber="371">A. R. Rasmussen</collectorName>
&amp;
<collectorName id="26A772E7FFBFAD10E7C7FDEDFAEBFD2C" box="[1241,1378,617,643]" pageId="14" pageNumber="371">B. Helwigh.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBFAD0FE3D9FD08FBD8F88D" lastPageId="17" lastPageNumber="374" pageId="14" pageNumber="371" type="description">
<paragraph id="8BED1731FFBFAD10E3D9FD08FDB1FD07" blockId="14.[151,1437,149,680]" box="[199,568,652,680]" pageId="14" pageNumber="371">
<emphasis id="B926CB23FFBFAD10E3D9FD08FED8FD09" bold="true" box="[199,337,652,678]" pageId="14" pageNumber="371">Description</emphasis>
(based on
<typeStatus id="54E9A993FFBFAD10E2D2FD09FDB9FD07" box="[460,560,653,680]" pageId="14" pageNumber="371" type="holotype">holotype</typeStatus>
)
</paragraph>
<caption id="DF2D47B9FFBFAD10E389F8A7FF4FF840" ID-DOI="http://doi.org/10.5281/zenodo.6332751" ID-Zenodo-Dep="6332751" httpUri="https://zenodo.org/record/6332751/files/figure.png" pageId="14" pageNumber="371" startId="14.[151,250,1827,1851]" targetBox="[152,1436,707,1803]" targetPageId="14">
<paragraph id="8BED1731FFBFAD10E389F8A7FF4FF840" blockId="14.[151,1437,1827,2031]" pageId="14" pageNumber="371">
<emphasis id="B926CB23FFBFAD10E389F8A7FEA8F894" bold="true" box="[151,289,1827,1851]" pageId="14" pageNumber="371">FIGURE 10.</emphasis>
<taxonomicName id="4C526CB2FFBFAD10E236F8A7FD70F894" authority="Bhansali &amp; Wesener" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[296,761,1827,1851]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="14" pageNumber="371" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBFAD10E236F8A7FD96F894" box="[296,543,1827,1851]" italics="true" pageId="14" pageNumber="371">Zephronia chrysomallos</emphasis>
Bhansali &amp; Wesener
</taxonomicName>
<emphasis id="B926CB23FFBFAD10E01EF8A7FCD8F894" bold="true" box="[768,849,1827,1851]" pageId="14" pageNumber="371">
<taxonomicNameLabel id="A2157658FFBFAD10E01EF8A7FCD8F894" box="[768,849,1827,1851]" pageId="14" pageNumber="371" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂ (
<emphasis id="B926CB23FFBFAD10E0FCF8A7FB2CF894" bold="true" box="[994,1189,1827,1851]" pageId="14" pageNumber="371">ZFMK MYR8826</emphasis>
), drawings,
<emphasis id="B926CB23FFBFAD10E63AF8A7FAB4F894" bold="true" box="[1316,1341,1827,1851]" pageId="14" pageNumber="371">A.</emphasis>
First left coxa with stigmatic plate, posterior view.
<emphasis id="B926CB23FFBFAD10E150F8C3FDECF8F0" bold="true" box="[590,613,1863,1887]" pageId="14" pageNumber="371">B.</emphasis>
Left leg 9, posterior view.
<emphasis id="B926CB23FFBFAD10E098F8C3FC16F8F0" bold="true" box="[902,927,1863,1887]" pageId="14" pageNumber="371">C.</emphasis>
Second left coxa with gonopore, posterior view.
<emphasis id="B926CB23FFBFAD10E389F8EFFF39F82C" bold="true" box="[151,176,1899,1923]" pageId="14" pageNumber="371">D.</emphasis>
Right anterior telopod, lateral view.
<emphasis id="B926CB23FFBFAD10E125F8EFFDDBF82C" bold="true" box="[571,594,1899,1923]" pageId="14" pageNumber="371">E.</emphasis>
Right anterior telopod, posterior view.
<emphasis id="B926CB23FFBFAD10E0E6F8EFFB82F82C" bold="true" box="[1016,1035,1899,1923]" pageId="14" pageNumber="371">F.</emphasis>
Right anterior telopod, anterior view.
<emphasis id="B926CB23FFBFAD10E389F80BFEB9F808" bold="true" box="[151,304,1935,1959]" pageId="14" pageNumber="371">Abbreviations</emphasis>
: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines.
<emphasis id="B926CB23FFBFAD10E61DF837FAFAF864" bold="true" box="[1283,1395,1971,1995]" pageId="14" pageNumber="371">Scale bars</emphasis>
= 1 mm.
</paragraph>
</caption>
<caption id="DF2D47B9FFBEAD11E389FAB7FDB5FA3C" ID-DOI="http://doi.org/10.5281/zenodo.6332753" ID-Zenodo-Dep="6332753" httpUri="https://zenodo.org/record/6332753/files/figure.png" pageId="15" pageNumber="372" startId="15.[151,250,1331,1355]" targetBox="[153,1436,194,1307]" targetPageId="15">
<paragraph id="8BED1731FFBEAD11E389FAB7FDB5FA3C" blockId="15.[151,1437,1331,1427]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E389FAB7FE95FAE4" bold="true" box="[151,284,1331,1355]" pageId="15" pageNumber="372">FIGURE 11.</emphasis>
<taxonomicName id="4C526CB2FFBEAD11E23EFAB7FD6BFAE4" authority="Bhansali &amp; Wesener" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[288,738,1331,1355]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="372" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBEAD11E23EFAB7FD9AFAE4" box="[288,531,1331,1355]" italics="true" pageId="15" pageNumber="372">Zephronia chrysomallos</emphasis>
Bhansali &amp; Wesener
</taxonomicName>
<emphasis id="B926CB23FFBEAD11E1F8FAB7FCBDFAE4" bold="true" box="[742,820,1331,1355]" pageId="15" pageNumber="372">
<taxonomicNameLabel id="A2157658FFBEAD11E1F8FAB7FCBDFAE4" box="[742,820,1331,1355]" pageId="15" pageNumber="372" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂ (
<emphasis id="B926CB23FFBEAD11E0A4FAB7FBF4FAE4" bold="true" box="[954,1149,1331,1355]" pageId="15" pageNumber="372">NHMD MYR8826</emphasis>
), drawings,
<emphasis id="B926CB23FFBEAD11E7E6FAB7FA99FAE4" bold="true" box="[1272,1296,1331,1355]" pageId="15" pageNumber="372">A.</emphasis>
Left posterior telopod, anterior view.
<emphasis id="B926CB23FFBEAD11E29EFAD3FE11FAC0" bold="true" box="[384,408,1367,1391]" pageId="15" pageNumber="372">B.</emphasis>
Left posterior telopod, posterior view.
<emphasis id="B926CB23FFBEAD11E03CFAD3FC32FAC0" bold="true" box="[802,955,1367,1391]" pageId="15" pageNumber="372">Abbreviations</emphasis>
: ct = crenulated teeth; imf = immovable finger; ss = sclerotized spot.
<emphasis id="B926CB23FFBEAD11E26FFAFFFE56FA3C" bold="true" box="[369,479,1403,1427]" pageId="15" pageNumber="372">Scale bars</emphasis>
= 1 mm.
</paragraph>
</caption>
<paragraph id="8BED1731FFBEAD11E3D9FA45FD2DF9AF" blockId="15.[151,1437,1473,2040]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9FA45FF7EFA74" bold="true" box="[199,247,1473,1499]" pageId="15" pageNumber="372">Size</emphasis>
. Length:
<quantity id="4CAABAD4FFBEAD11E27BFA45FE45FA73" box="[357,460,1473,1500]" metricMagnitude="-2" metricUnit="m" metricValue="3.1399999999999997" pageId="15" pageNumber="372" unit="mm" value="31.4">31.4 mm</quantity>
. Width of thoracic shield
<quantity id="4CAABAD4FFBEAD11E1F0FA45FCDDFA73" box="[750,852,1473,1500]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" pageId="15" pageNumber="372" unit="mm" value="14.8">14.8 mm</quantity>
, of widest segment (6)
<quantity id="4CAABAD4FFBEAD11E743FA45FB4DFA73" box="[1117,1220,1473,1500]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="15" pageNumber="372" unit="mm" value="16.5">16.5 mm</quantity>
. Height of thoracic shield
<quantity id="4CAABAD4FFBEAD11E3FFFA61FEB0F9AF" box="[225,313,1509,1536]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="15" pageNumber="372" unit="mm" value="9.3">9.3 mm</quantity>
, of highest segment (6)
<quantity id="4CAABAD4FFBEAD11E159FA61FD29F9AF" box="[583,672,1509,1536]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="15" pageNumber="372" unit="mm" value="9.8">9.8 mm</quantity>
.
</paragraph>
<paragraph id="8BED1731FFBEAD11E3D9F98DFBA3F9C4" blockId="15.[151,1437,1473,2040]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9F98DFE92F98C" bold="true" box="[199,283,1545,1571]" pageId="15" pageNumber="372">Colour</emphasis>
. Faded after 30 years in 75% ethanol. Collum medium brown, head and legs with traces of green. Antenna green. Tergites and anal shield pale greyish-brown, darker brown towards posterior margin. Thoracic shield-like tergite with darker groove and margins. Anal shield colour like tergite.
</paragraph>
<paragraph id="8BED1731FFBEAD11E3D9F9F1FCA5F977" blockId="15.[151,1437,1473,2040]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9F9F1FE8EF920" bold="true" box="[199,263,1653,1679]" pageId="15" pageNumber="372">Head</emphasis>
. Number of ommatidia 95. Tömösváry organ placed on the rim of antennal groove. Antennae short, not reaching to center of head (
<figureCitation id="13690BB4FFBEAD11E2D5F91DFDDFF91C" box="[459,598,1689,1715]" captionStart="FIGURE 12" captionStartId="16.[151,250,1442,1466]" captionTargetBox="[164,1423,489,1412]" captionTargetId="figure-162@16.[164,1423,489,1418]" captionTargetPageId="16" captionText="FIGURE 12. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), volume rendering based on microcomputed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cp = central pad; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; o = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Po = postfemur; Pre = prefemur; Pt = paratergite; Stp = stigmatic plate; St = stipites; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332755" httpUri="https://zenodo.org/record/6332755/files/figure.png" pageId="15" pageNumber="372">Fig. 12A–D</figureCitation>
). Antennomere lengths: 1&gt;2=3=4=5&lt;&lt;6. Antennomer 6 laterally flattened, apically widened, axe-shaped, carrying 91/96 apical cones.
</paragraph>
<paragraph id="8BED1731FFBEAD11E3D9F965FD8DF954" blockId="15.[151,1437,1473,2040]" box="[199,516,1761,1788]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9F965FEDFF954" bold="true" box="[199,342,1761,1787]" pageId="15" pageNumber="372">Epipharynx</emphasis>
. Not dissected.
</paragraph>
<paragraph id="8BED1731FFBEAD11E3D9F881FABCF8B0" blockId="15.[151,1437,1473,2040]" box="[199,1333,1797,1824]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9F881FE1FF8B0" bold="true" box="[199,406,1797,1823]" pageId="15" pageNumber="372">Gnathochilarium</emphasis>
(
<figureCitation id="13690BB4FFBEAD11E2BBF881FD87F8B0" box="[421,526,1797,1823]" captionStart="FIGURE 12" captionStartId="16.[151,250,1442,1466]" captionTargetBox="[164,1423,489,1412]" captionTargetId="figure-162@16.[164,1423,489,1418]" captionTargetPageId="16" captionText="FIGURE 12. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), volume rendering based on microcomputed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cp = central pad; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; o = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Po = postfemur; Pre = prefemur; Pt = paratergite; Stp = stigmatic plate; St = stipites; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332755" httpUri="https://zenodo.org/record/6332755/files/figure.png" pageId="15" pageNumber="372">Fig. 12D</figureCitation>
). Gnathochilarium palpi with the sensory cones arranged in single field.
</paragraph>
<paragraph id="8BED1731FFBEAD11E3D9F8ADFE61F8EC" blockId="15.[151,1437,1473,2040]" box="[199,488,1833,1860]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9F8ADFEB3F8EC" bold="true" box="[199,314,1833,1859]" pageId="15" pageNumber="372">Mandible</emphasis>
. Not dissected.
</paragraph>
<paragraph id="8BED1731FFBEAD11E3D9F8C9FBECF8C8" blockId="15.[151,1437,1473,2040]" box="[199,1125,1869,1896]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9F8C9FEA9F8C8" bold="true" box="[199,288,1869,1895]" pageId="15" pageNumber="372">Collum</emphasis>
. Numerous setae, distributed towards margins, few setae in central region.
</paragraph>
<paragraph id="8BED1731FFBEAD11E3D9F8F5FA95F823" blockId="15.[151,1437,1473,2040]" box="[199,1308,1905,1932]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9F8F5FE09F824" bold="true" box="[199,384,1905,1931]" pageId="15" pageNumber="372">Thoracic shield</emphasis>
. Thoracic shield grooves shallow and wide with 2 or 3 weak ledges (
<figureCitation id="13690BB4FFBEAD11E798F8F5FA87F823" box="[1158,1294,1905,1932]" captionStart="FIGURE 12" captionStartId="16.[151,250,1442,1466]" captionTargetBox="[164,1423,489,1412]" captionTargetId="figure-162@16.[164,1423,489,1418]" captionTargetPageId="16" captionText="FIGURE 12. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), volume rendering based on microcomputed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cp = central pad; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; o = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Po = postfemur; Pre = prefemur; Pt = paratergite; Stp = stigmatic plate; St = stipites; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332755" httpUri="https://zenodo.org/record/6332755/files/figure.png" pageId="15" pageNumber="372">Fig. 12A, B</figureCitation>
).
</paragraph>
<paragraph id="8BED1731FFBEAD11E3D9F811FCB5F81F" blockId="15.[151,1437,1473,2040]" box="[199,828,1941,1968]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9F811FE3EF800" bold="true" box="[199,439,1941,1967]" pageId="15" pageNumber="372">First stigmatic plate</emphasis>
. Smaller than coxa, rounded apex.
</paragraph>
<paragraph id="8BED1731FFBEAD11E3D9F83DFC1BF857" blockId="15.[151,1437,1473,2040]" pageId="15" pageNumber="372">
<emphasis id="B926CB23FFBEAD11E3D9F83DFEAFF87C" bold="true" box="[199,294,1977,2003]" pageId="15" pageNumber="372">Tergites</emphasis>
. Tergites covered in short, dense setae visible at higher magnification, with dull orange skin-like surface. Paratergite tips weakly projecting backwards (
<figureCitation id="13690BB4FFBEAD11E1E3F859FC0DF857" box="[765,900,2013,2040]" captionStart="FIGURE 12" captionStartId="16.[151,250,1442,1466]" captionTargetBox="[164,1423,489,1412]" captionTargetId="figure-162@16.[164,1423,489,1418]" captionTargetPageId="16" captionText="FIGURE 12. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), volume rendering based on microcomputed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cp = central pad; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; o = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Po = postfemur; Pre = prefemur; Pt = paratergite; Stp = stigmatic plate; St = stipites; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332755" httpUri="https://zenodo.org/record/6332755/files/figure.png" pageId="15" pageNumber="372">Fig. 12A, B</figureCitation>
).
</paragraph>
<paragraph id="8BED1731FFA1AD0EE3D9FF12FAD4FF1E" blockId="16.[151,1437,150,465]" box="[199,1373,150,177]" pageId="16" pageNumber="373">
<emphasis id="B926CB23FFA1AD0EE3D9FF12FEB8FF1F" bold="true" box="[199,305,150,176]" pageId="16" pageNumber="373">Pleurites</emphasis>
(
<figureCitation id="13690BB4FFA1AD0EE25EFF13FE2EFF1E" box="[320,423,151,177]" captionStart="FIGURE 12" captionStartId="16.[151,250,1442,1466]" captionTargetBox="[164,1423,489,1412]" captionTargetId="figure-162@16.[164,1423,489,1418]" captionTargetPageId="16" captionText="FIGURE 12. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), volume rendering based on microcomputed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cp = central pad; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; o = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Po = postfemur; Pre = prefemur; Pt = paratergite; Stp = stigmatic plate; St = stipites; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332755" httpUri="https://zenodo.org/record/6332755/files/figure.png" pageId="16" pageNumber="373">Fig. 12C</figureCitation>
). Pleurite 1 projecting, tip well-rounded. Pleurite 2 well-rounded, weakly projecting.
</paragraph>
<paragraph id="8BED1731FFA1AD0EE3D9FF3FFE22FEB2" blockId="16.[151,1437,150,465]" pageId="16" pageNumber="373">
<emphasis id="B926CB23FFA1AD0EE3D9FF3FFED7FF7A" bold="true" box="[199,350,187,213]" pageId="16" pageNumber="373">Endotergum</emphasis>
. With a regular flat margin. Outer zone with three dense rows of irregular marginal setae, not extending beyond posterior margin, but reaching 4/5 of the outer area. Middle section with cuticular impressions. Inner area without setae.
</paragraph>
<paragraph id="8BED1731FFA1AD0EE3D9FEA3FC60FE26" blockId="16.[151,1437,150,465]" pageId="16" pageNumber="373">
<emphasis id="B926CB23FFA1AD0EE3D9FEA3FF76FEEE" bold="true" box="[199,255,295,321]" pageId="16" pageNumber="373">Legs</emphasis>
. Ventral spines on leg 1 2/2, at 2 4 /4, at 3 7/7. Leg 3 with single apical spine. Mid-body legs with 2 or 3 apical and 8–10 ventral spines (
<figureCitation id="13690BB4FFA1AD0EE10AFECFFDF4FECA" box="[532,637,331,357]" captionStart="FIGURE 13" captionStartId="17.[151,250,1297,1321]" captionTargetBox="[151,1436,188,1273]" captionTargetId="figure-17@17.[151,1436,181,1273]" captionTargetPageId="17" captionText="FIGURE 13. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (ZMUC K56-9), drawings, A. Left leg 9, posterior view B. Second left coxa with gonopore, posterior view. C. Right anterior telopod, anterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332757" httpUri="https://zenodo.org/record/6332757/files/figure.png" pageId="16" pageNumber="373">Fig. 13A</figureCitation>
). Inner margin of femur with 14 small triangular teeth, concentrated at apical half, but not excavated. Femur 1.7, tarsus 3.6 times longer than wide.
</paragraph>
<paragraph id="8BED1731FFA1AD0EE3D9FE16FB68FE7E" blockId="16.[151,1437,150,465]" pageId="16" pageNumber="373">
<emphasis id="B926CB23FFA1AD0EE3D9FE16FEDBFE03" bold="true" box="[199,338,402,428]" pageId="16" pageNumber="373">Anal shield</emphasis>
. Bell-shaped and like tergites covered with short dense setae visible at higher magnification. Underside with single black locking carina of medium length, located close to pleurite (
<figureCitation id="13690BB4FFA1AD0EE773FE33FB5DFE7E" box="[1133,1236,439,465]" captionStart="FIGURE 12" captionStartId="16.[151,250,1442,1466]" captionTargetBox="[164,1423,489,1412]" captionTargetId="figure-162@16.[164,1423,489,1418]" captionTargetPageId="16" captionText="FIGURE 12. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), volume rendering based on microcomputed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cp = central pad; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; o = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Po = postfemur; Pre = prefemur; Pt = paratergite; Stp = stigmatic plate; St = stipites; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332755" httpUri="https://zenodo.org/record/6332755/files/figure.png" pageId="16" pageNumber="373">Fig. 12C</figureCitation>
).
</paragraph>
<caption id="DF2D47B9FFA1AD0EE389FA26FF58F93D" ID-DOI="http://doi.org/10.5281/zenodo.6332755" ID-Zenodo-Dep="6332755" httpUri="https://zenodo.org/record/6332755/files/figure.png" pageId="16" pageNumber="373" startId="16.[151,250,1442,1466]" targetBox="[164,1423,489,1412]" targetPageId="16">
<paragraph id="8BED1731FFA1AD0EE389FA26FF58F93D" blockId="16.[151,1437,1442,1682]" pageId="16" pageNumber="373">
<emphasis id="B926CB23FFA1AD0EE389FA26FE96FA15" bold="true" box="[151,287,1442,1466]" pageId="16" pageNumber="373">FIGURE 12.</emphasis>
<taxonomicName id="4C526CB2FFA1AD0EE23AFA26FD31FA16" authority="Bhansali &amp; Wesener" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[292,696,1442,1466]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="373" phylum="Arthropoda" rank="species" species="erawani" status="sp. nov.">
<emphasis id="B926CB23FFA1AD0EE23AFA26FE6CFA15" box="[292,485,1442,1466]" italics="true" pageId="16" pageNumber="373">Zephronia erawani</emphasis>
Bhansali &amp; Wesener
</taxonomicName>
<emphasis id="B926CB23FFA1AD0EE1A3FA26FC85FA15" bold="true" box="[701,780,1442,1466]" pageId="16" pageNumber="373">
<taxonomicNameLabel id="A2157658FFA1AD0EE1A3FA26FC85FA15" box="[701,780,1442,1466]" pageId="16" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂ (
<emphasis id="B926CB23FFA1AD0EE089FA26FBA6FA15" bold="true" box="[919,1071,1442,1466]" pageId="16" pageNumber="373">NHMD K56-9</emphasis>
), volume rendering based on microcomputed tomography.
<emphasis id="B926CB23FFA1AD0EE296FA42FE28FA71" bold="true" box="[392,417,1478,1502]" pageId="16" pageNumber="373">A.</emphasis>
Habitus, lateral view.
<emphasis id="B926CB23FFA1AD0EE198FA42FD14FA71" bold="true" box="[646,669,1478,1502]" pageId="16" pageNumber="373">B.</emphasis>
Habitus, ventro-lateral view.
<emphasis id="B926CB23FFA1AD0EE0D2FA42FC6CFA71" bold="true" box="[972,997,1478,1502]" pageId="16" pageNumber="373">C.</emphasis>
Habitus, ventral view.
<emphasis id="B926CB23FFA1AD0EE7CCFA42FB62FA71" bold="true" box="[1234,1259,1478,1502]" pageId="16" pageNumber="373">D.</emphasis>
Habitus, anterior view.
<emphasis id="B926CB23FFA1AD0EE3CBFA6EFF64F9AD" bold="true" box="[213,237,1514,1538]" pageId="16" pageNumber="373">E.</emphasis>
Body-ring architecture, cross section trough midbody-ring.
<emphasis id="B926CB23FFA1AD0EE042FA6EFC7CF9AD" bold="true" box="[860,1013,1514,1538]" pageId="16" pageNumber="373">Abbreviations</emphasis>
: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cp = central pad; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; o = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Po = postfemur; Pre = prefemur; Pt = paratergite; Stp = stigmatic plate; St = stipites; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Ts = thoracic shield. Not to scale.
</paragraph>
</caption>
<paragraph id="8BED1731FFA1AD0EE3D9F944FC32F951" blockId="16.[151,1437,1728,2043]" pageId="16" pageNumber="373">
<emphasis id="B926CB23FFA1AD0EE3D9F944FEF4F975" bold="true" box="[199,381,1728,1754]" pageId="16" pageNumber="373">Male gonopore</emphasis>
. Located directly on mesal margin of coxa 2, covered posteriorly by a single semicircular sclerotized plate. Membrane rising mesally into a projection (
<figureCitation id="13690BB4FFA1AD0EE056F960FC26F951" box="[840,943,1764,1790]" captionStart="FIGURE 13" captionStartId="17.[151,250,1297,1321]" captionTargetBox="[151,1436,188,1273]" captionTargetId="figure-17@17.[151,1436,181,1273]" captionTargetPageId="17" captionText="FIGURE 13. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (ZMUC K56-9), drawings, A. Left leg 9, posterior view B. Second left coxa with gonopore, posterior view. C. Right anterior telopod, anterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332757" httpUri="https://zenodo.org/record/6332757/files/figure.png" pageId="16" pageNumber="373">Fig. 13B</figureCitation>
).
</paragraph>
<paragraph id="8BED1731FFA1AD0EE3D9F88CFBB5F855" blockId="16.[151,1437,1728,2043]" pageId="16" pageNumber="373">
<emphasis id="B926CB23FFA1AD0EE3D9F88CFE10F88D" bold="true" box="[199,409,1800,1826]" pageId="16" pageNumber="373">Anterior telopods</emphasis>
(
<figureCitation id="13690BB4FFA1AD0EE2B8F88CFDA2F88D" box="[422,555,1800,1826]" captionStart="FIGURE 13" captionStartId="17.[151,250,1297,1321]" captionTargetBox="[151,1436,188,1273]" captionTargetId="figure-17@17.[151,1436,181,1273]" captionTargetPageId="17" captionText="FIGURE 13. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (ZMUC K56-9), drawings, A. Left leg 9, posterior view B. Second left coxa with gonopore, posterior view. C. Right anterior telopod, anterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332757" httpUri="https://zenodo.org/record/6332757/files/figure.png" pageId="16" pageNumber="373">Fig. 13C–E</figureCitation>
). Syncoxite with few setae. Podomere 1 rectangular, as long as wide. Numerous setae, mainly at median margins. Podomere 2 slightly narrower than podomere 1. Process massive, well-rounded, apically slightly tapering, relatively short and wide (
<figureCitation id="13690BB4FFA1AD0EE015F8D4FC17F8C4" box="[779,926,1872,1899]" captionStart="FIGURE 13" captionStartId="17.[151,250,1297,1321]" captionTargetBox="[151,1436,188,1273]" captionTargetId="figure-17@17.[151,1436,181,1273]" captionTargetPageId="17" captionText="FIGURE 13. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (ZMUC K56-9), drawings, A. Left leg 9, posterior view B. Second left coxa with gonopore, posterior view. C. Right anterior telopod, anterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332757" httpUri="https://zenodo.org/record/6332757/files/figure.png" pageId="16" pageNumber="373">Fig. 13D, E</figureCitation>
). Process visible protruding and exceeding podomere
<quantity id="4CAABAD4FFA1AD0EE210F8F0FEB3F821" box="[270,314,1908,1934]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="16" pageNumber="373" unit="in" value="3.0">3 in</quantity>
anterior view (
<figureCitation id="13690BB4FFA1AD0EE2FBF8F0FDC4F821" box="[485,589,1908,1934]" captionStart="FIGURE 13" captionStartId="17.[151,250,1297,1321]" captionTargetBox="[151,1436,188,1273]" captionTargetId="figure-17@17.[151,1436,181,1273]" captionTargetPageId="17" captionText="FIGURE 13. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (ZMUC K56-9), drawings, A. Left leg 9, posterior view B. Second left coxa with gonopore, posterior view. C. Right anterior telopod, anterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332757" httpUri="https://zenodo.org/record/6332757/files/figure.png" pageId="16" pageNumber="373">Fig. 13D</figureCitation>
). Podomere 3 cylindrical, with a black spot at the anterior margin visible in posterior view. Podomere 4 partially fused to podomere 3, suture can be seen in anterior view (
<figureCitation id="13690BB4FFA1AD0EE7D5F81CFAB8F81D" box="[1227,1329,1944,1970]" captionStart="FIGURE 13" captionStartId="17.[151,250,1297,1321]" captionTargetBox="[151,1436,188,1273]" captionTargetId="figure-17@17.[151,1436,181,1273]" captionTargetPageId="17" captionText="FIGURE 13. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (ZMUC K56-9), drawings, A. Left leg 9, posterior view B. Second left coxa with gonopore, posterior view. C. Right anterior telopod, anterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332757" httpUri="https://zenodo.org/record/6332757/files/figure.png" pageId="16" pageNumber="373">Fig. 13C</figureCitation>
), while it is completely fused in posterior, lateral and mesal views. Podomere 4 small, narrowing at the apex. Meso-anterior margin with one spine and a black spot at apex visible in posterior and lateral view.
</paragraph>
<caption id="DF2D47B9FFA0AD0FE389FA95FD1EFA16" ID-DOI="http://doi.org/10.5281/zenodo.6332757" ID-Zenodo-Dep="6332757" httpUri="https://zenodo.org/record/6332757/files/figure.png" pageId="17" pageNumber="374" startId="17.[151,250,1297,1321]" targetBox="[151,1436,188,1273]" targetPageId="17">
<paragraph id="8BED1731FFA0AD0FE389FA95FD1EFA16" blockId="17.[151,1437,1297,1466]" pageId="17" pageNumber="374">
<emphasis id="B926CB23FFA0AD0FE389FA95FEA9FA86" bold="true" box="[151,288,1297,1321]" pageId="17" pageNumber="374">FIGURE 13.</emphasis>
<taxonomicName id="4C526CB2FFA0AD0FE238FA95FD34FA86" authority="Bhansali &amp; Wesener" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[294,701,1297,1322]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="374" phylum="Arthropoda" rank="species" species="erawani" status="sp. nov.">
<emphasis id="B926CB23FFA0AD0FE238FA95FE61FA86" box="[294,488,1297,1321]" italics="true" pageId="17" pageNumber="374">Zephronia erawani</emphasis>
Bhansali &amp; Wesener
</taxonomicName>
<emphasis id="B926CB23FFA0AD0FE1DDFA96FC9AFA85" bold="true" box="[707,787,1298,1322]" pageId="17" pageNumber="374">
<taxonomicNameLabel id="A2157658FFA0AD0FE1DDFA96FC9AFA85" box="[707,787,1298,1322]" pageId="17" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂ (
<emphasis id="B926CB23FFA0AD0FE0BFFA95FBBFFA86" bold="true" box="[929,1078,1297,1321]" pageId="17" pageNumber="374">ZMUC K56-9</emphasis>
), drawings,
<emphasis id="B926CB23FFA0AD0FE7ABFA95FB44FA86" bold="true" box="[1205,1229,1297,1321]" pageId="17" pageNumber="374">A.</emphasis>
Left leg 9, posterior view
<emphasis id="B926CB23FFA0AD0FE3CFFAB2FF61FAE1" bold="true" box="[209,232,1334,1358]" pageId="17" pageNumber="374">B.</emphasis>
Second left coxa with gonopore, posterior view.
<emphasis id="B926CB23FFA0AD0FE1F7FAB1FC8BFAE2" bold="true" box="[745,770,1333,1357]" pageId="17" pageNumber="374">C.</emphasis>
Right anterior telopod, anterior view.
<emphasis id="B926CB23FFA0AD0FE78FFAB2FB23FAE1" bold="true" box="[1169,1194,1334,1358]" pageId="17" pageNumber="374">D.</emphasis>
Right anterior telopod, lateral view.
<emphasis id="B926CB23FFA0AD0FE206FADEFEA6FADD" bold="true" box="[280,303,1370,1394]" pageId="17" pageNumber="374">E.</emphasis>
Right anterior telopod, posterior view.
<emphasis id="B926CB23FFA0AD0FE1A4FADDFCDAFADE" bold="true" box="[698,851,1369,1393]" pageId="17" pageNumber="374">Abbreviations</emphasis>
: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines.
<emphasis id="B926CB23FFA0AD0FE2D2FA25FDB2FA15" bold="true" box="[460,571,1441,1466]" pageId="17" pageNumber="374">Scale bars</emphasis>
= 1 mm.
</paragraph>
</caption>
<paragraph id="8BED1731FFA0AD0FE3D9FA63FBD8F88D" blockId="17.[151,1437,1511,1934]" pageId="17" pageNumber="374">
<emphasis id="B926CB23FFA0AD0FE3D9FA63FE2FF9AD" bold="true" box="[199,422,1511,1538]" pageId="17" pageNumber="374">Posterior telopods</emphasis>
(
<figureCitation id="13690BB4FFA0AD0FE2A4FA6CFD9AF9AD" box="[442,531,1512,1538]" captionStart="FIGURE 14" captionStartId="18.[151,250,1492,1516]" captionTargetBox="[151,1436,185,1468]" captionTargetId="figure-22@18.[151,1436,181,1468]" captionTargetPageId="18" captionText="FIGURE 14. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), drawings, A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view. Abbreviations: ct = crenulated teeth; imf = immovable finger; ss = sclerotized spot. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332759" httpUri="https://zenodo.org/record/6332759/files/figure.png" pageId="17" pageNumber="374">Fig. 14</figureCitation>
). Syncoxite inner horns (not drawn): apex well rounded, apically diverging. Podomere 1 with short setae distributed in median and lateral margin, median margin glabrous. Podomere 2 with long and wide immovable finger, 2.5 times longer than wide. Podomere 2 with numerous setae visible in anterior view, almost entirely glabrous in posterior view. Immovable finger slightly narrow anteriorly and straight. Apical process with sclerotized round spots in anterior view (
<figureCitation id="13690BB4FFA0AD0FE00EF9FCFCF2F93D" box="[784,891,1656,1682]" captionStart="FIGURE 14" captionStartId="18.[151,250,1492,1516]" captionTargetBox="[151,1436,185,1468]" captionTargetId="figure-22@18.[151,1436,181,1468]" captionTargetPageId="18" captionText="FIGURE 14. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), drawings, A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view. Abbreviations: ct = crenulated teeth; imf = immovable finger; ss = sclerotized spot. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332759" httpUri="https://zenodo.org/record/6332759/files/figure.png" pageId="17" pageNumber="374">Fig. 14A</figureCitation>
, arrow). Membrane toward podomere 3 with a two-tipped membranous lobe. Podomere 3 setose in anterior view, with more setae concentrated medially. With row of 15 large, crenulated teeth on posterior side (
<figureCitation id="13690BB4FFA0AD0FE1CFF944FCBEF975" box="[721,823,1728,1754]" captionStart="FIGURE 14" captionStartId="18.[151,250,1492,1516]" captionTargetBox="[151,1436,185,1468]" captionTargetId="figure-22@18.[151,1436,181,1468]" captionTargetPageId="18" captionText="FIGURE 14. Zephronia erawani Bhansali &amp; Wesener sp. nov., holotype ♂ (NHMD K56-9), drawings, A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view. Abbreviations: ct = crenulated teeth; imf = immovable finger; ss = sclerotized spot. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332759" httpUri="https://zenodo.org/record/6332759/files/figure.png" pageId="17" pageNumber="374">Fig. 14B</figureCitation>
, arrow). Large membranous ledge on podomere 3 with single spine present basal region. Podomere 4 glabrous on both sides, slightly curved towards immovable finger. Membranous lobe with 2 long spines. Immovable finger shorter than movable finger.
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFA0AD0FE3D9F8AFFD28F821" pageId="17" pageNumber="374" type="distribution">
<paragraph id="8BED1731FFA0AD0FE3D9F8AFFED0F8EA" blockId="17.[151,1437,1511,1934]" box="[199,345,1835,1861]" pageId="17" pageNumber="374">
<emphasis id="B926CB23FFA0AD0FE3D9F8AFFED0F8EA" bold="true" box="[199,345,1835,1861]" pageId="17" pageNumber="374">Distribution</emphasis>
</paragraph>
<paragraph id="8BED1731FFA0AD0FE3D9F8D4FD28F821" blockId="17.[151,1437,1511,1934]" pageId="17" pageNumber="374">
<taxonomicName id="4C526CB2FFA0AD0FE3D9F8D4FEC9F8C6" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[199,320,1872,1897]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="374" phylum="Arthropoda" rank="species" species="erawani" status="sp. nov.">
<emphasis id="B926CB23FFA0AD0FE3D9F8D4FEC9F8C6" box="[199,320,1872,1897]" italics="true" pageId="17" pageNumber="374">Z. erawani</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFA0AD0FE258F8D4FE16F8C5" bold="true" box="[326,415,1872,1898]" pageId="17" pageNumber="374">
<taxonomicNameLabel id="A2157658FFA0AD0FE258F8D4FE16F8C5" box="[326,415,1872,1898]" pageId="17" pageNumber="374" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently only known from the
<typeStatus id="54E9A993FFA0AD0FE007F8D4FCC0F8C5" box="[793,841,1872,1898]" pageId="17" pageNumber="374">type</typeStatus>
locality (
<figureCitation id="13690BB4FFA0AD0FE0ACF8D4FB83F8C5" box="[946,1034,1872,1898]" captionStart="FIGURE 2" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[337,1247,181,1766]" captionTargetId="figure-22@6.[337,1249,181,1766]" captionTargetPageId="6" captionText="FIGURE 2. Map of Thailand with known distribution of Sphaerobelum spp. and Zephronia spp. Star represents new species. Diamonds represent new localities. Fluorescent Green = S. meridionalis Bhansali &amp; Wesener sp. nov.; Pink = S. aesculus; Brown = S. truncatum; Chrome yellow = Z. chrysomallos Bhansali &amp; Wesener sp. nov.; White = Z. erawani Bhansali &amp; Wesener sp. nov.; Black = Z. viridisoma; Red = Z. panhai; Dark green = Z. golovatchi; Purple = Z. siamensis; Orange = Z. enghoffi; Yellow = Z. lannaensis; Blue = Z. phrain. B. Habitus photograph of Z. chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♀ (ZFMK MYR8826)." figureDoi="http://doi.org/10.5281/zenodo.6332735" httpUri="https://zenodo.org/record/6332735/files/figure.png" pageId="17" pageNumber="374">Fig. 2A</figureCitation>
) and might be microendemic, as are other, closely related species in nearby forests.
</paragraph>
</subSubSection>
</treatment>
</document>
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