treatments-xml/data/03/FB/A6/03FBA627FFBBAD13E389FA11FBB9F95F.xml

<document id="00E866973272C10E4CD800662D8D5E13" ID-CLB-Dataset="20340" ID-DOI="10.11646/zootaxa.5105.3.2" ID-GBIF-Dataset="4f25a761-8f7b-47d3-84c9-ad2917b90f74" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332731" ID-ZooBank="83013C70-69E4-42BD-8CDD-0FA320927290" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646597716981" checkinUser="plazi" docAuthor="Bhansali, Sneha &amp; Wesener, Thomas" docDate="2022" docId="03FBA627FFBBAD13E389FA11FBB9F95F" docLanguage="en" docName="zootaxa.5105.3.2.pdf" docOrigin="Zootaxa 5105 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zephronia chrysomallos Bhansali &amp; Wesener 2022, new species" docType="treatment" docVersion="8" lastPageNumber="370" masterDocId="FFC2DE5FFFB1AD1EE31EFF84FF89FFAF" masterDocTitle="New Thai giant pill-millipede species, with new genetic barcoding data (Diplopoda Sphaerotheriida, Zephroniidae)" masterLastPageNumber="380" masterPageNumber="357" pageNumber="367" updateTime="1699317184639" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A705592E4CC802C8336DC51DDCE3A102">New Thai giant pill-millipede species, with new genetic barcoding data (Diplopoda Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:namePart id="2215B8238308E9125E4180B28B2B8083">Bhansali, Sneha</mods:namePart>
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<mods:affiliation id="6AAB3D156F74CF2D9924975F9B87916E">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:namePart id="40A0D0919BDB4CFEECCB73D73ED9EA4E">Wesener, Thomas</mods:namePart>
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<mods:affiliation id="2FFE69ABDF22D2DE6F3A5329B7C28297">Zoological Research Museum Alexander Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 160, D- 53113 Bonn, Germany. &amp; T. Wesener @ leibniz-zfmk. de; https: // orcid. org / 0000 - 0002 - 2028 - 3541</mods:affiliation>
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<treatment id="03FBA627FFBBAD13E389FA11FBB9F95F" ID-DOI="http://doi.org/10.5281/zenodo.6333445" ID-GBIF-Taxon="193890960" ID-Zenodo-Dep="6333445" LSID="urn:lsid:plazi:treatment:03FBA627FFBBAD13E389FA11FBB9F95F" httpUri="http://treatment.plazi.org/id/03FBA627FFBBAD13E389FA11FBB9F95F" lastPageId="13" lastPageNumber="370" pageId="10" pageNumber="367">
<subSubSection id="C34844BAFFBBAD14E389FA11FCE9FA1F" box="[151,864,1429,1456]" pageId="10" pageNumber="367" type="nomenclature">
<paragraph id="8BED1731FFBBAD14E389FA11FCE9FA1F" blockId="10.[151,864,1429,1493]" box="[151,864,1429,1456]" pageId="10" pageNumber="367">
<heading id="D0A5A05DFFBBAD14E389FA11FCE9FA1F" bold="true" box="[151,864,1429,1456]" fontSize="11" level="1" pageId="10" pageNumber="367" reason="1">
<emphasis id="B926CB23FFBBAD14E389FA11FCE9FA1F" bold="true" box="[151,864,1429,1456]" pageId="10" pageNumber="367">
<taxonomicName id="4C526CB2FFBBAD14E389FA11FD4DFA1F" ID-CoL="9BDJV" authority="Bhansali &amp; Wesener" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[151,708,1429,1456]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBBAD14E389FA11FE37FA1F" bold="true" box="[151,446,1429,1456]" italics="true" pageId="10" pageNumber="367">Zephronia chrysomallos</emphasis>
Bhansali &amp; Wesener
</taxonomicName>
, 
<taxonomicNameLabel id="A2157658FFBBAD14E1CEFA12FCE9FA1F" box="[720,864,1430,1456]" pageId="10" pageNumber="367" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="C34844BAFFBBAD14E389FA3EFD98FA7B" box="[151,529,1466,1493]" pageId="10" pageNumber="367" type="description">
<paragraph id="8BED1731FFBBAD14E389FA3EFD98FA7B" blockId="10.[151,864,1429,1493]" box="[151,529,1466,1493]" pageId="10" pageNumber="367">
<figureCitation id="13690BB4FFBBAD14E389FA3EFEB1FA7B" box="[151,312,1466,1492]" captionStart="FIGURE 2" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[337,1247,181,1766]" captionTargetId="figure-22@6.[337,1249,181,1766]" captionTargetPageId="6" captionText="FIGURE 2. Map of Thailand with known distribution of Sphaerobelum spp. and Zephronia spp. Star represents new species. Diamonds represent new localities. Fluorescent Green = S. meridionalis Bhansali &amp; Wesener sp. nov.; Pink = S. aesculus; Brown = S. truncatum; Chrome yellow = Z. chrysomallos Bhansali &amp; Wesener sp. nov.; White = Z. erawani Bhansali &amp; Wesener sp. nov.; Black = Z. viridisoma; Red = Z. panhai; Dark green = Z. golovatchi; Purple = Z. siamensis; Orange = Z. enghoffi; Yellow = Z. lannaensis; Blue = Z. phrain. B. Habitus photograph of Z. chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♀ (ZFMK MYR8826)." figureDoi="http://doi.org/10.5281/zenodo.6332735" httpUri="https://zenodo.org/record/6332735/files/figure.png" pageId="10" pageNumber="367">Figures 2A–B</figureCitation>
, 
<figureCitation id="13690BB4FFBBAD14E25DFA3EFEEFFA7B" box="[323,358,1466,1492]" captionStart="FIGURE 6" captionStartId="10.[151,250,974,998]" captionTargetBox="[151,1436,181,950]" captionTargetId="figure-22@10.[151,1436,181,950]" captionTargetPageId="10" captionText="FIGURE 6. Endoterga of midbody tergites, scanning electron micrographs. A. Sphaerobelum meridionalis Bhansali &amp; Wesener sp. nov., holotype ♂ (MHNG 4B-2). B. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826). Abbreviations: Ci = cuticular impression; Ia = inner area; Ma = middle area; Oa = outer area." figureDoi="http://doi.org/10.5281/zenodo.6332743" httpUri="https://zenodo.org/record/6332743/files/figure.png" pageId="10" pageNumber="367">6B</figureCitation>
, 
<figureCitation id="13690BB4FFBBAD14E26CFA3FFE0BFA7A" box="[370,386,1467,1493]" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="10" pageNumber="367">7</figureCitation>
, 
<figureCitation id="13690BB4FFBBAD14E290FA3EFE14FA7B" box="[398,413,1466,1492]" captionStart="FIGURE 8" captionStartId="12.[151,250,1241,1265]" captionTargetBox="[151,1437,181,1216]" captionTargetId="figure-22@12.[151,1437,181,1217]" captionTargetPageId="12" captionText="FIGURE 8. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Left antennae, lateral view. B. Left sixth antennomere with antennal disc. C. Left antenna, disc. D. Gnathochilarium, apical view. E. Left lateral palpi. F. Left central pad, two types of sensory cones on central pad, detail. Abbreviations: ac = apical cone; ad = antennal disc; Cp = central pad; Hyp = hypopharynx; Ip = inner palpi; LL = lamellae lingulales; Sc = sensory cone; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332747" httpUri="https://zenodo.org/record/6332747/files/figure.png" pageId="10" pageNumber="367">8</figureCitation>
, 
<figureCitation id="13690BB4FFBBAD14E2B4FA3EFE33FA7B" box="[426,442,1466,1492]" captionStart="FIGURE 9" captionStartId="13.[151,250,707,731]" captionTargetBox="[152,1436,181,683]" captionTargetId="figure-17@13.[151,1436,181,684]" captionTargetPageId="13" captionText="FIGURE 9. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Gnathochilarium, ventral view. B. Right mandible, mesal view. Abbreviations: C = condyles; Cp = cental pad; eT = external tooth; Ip = inner palpi; iT = internal combined tooth; LL = lamellae linguales; Mn = mentum; pL = pectinate lamellae; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332749" httpUri="https://zenodo.org/record/6332749/files/figure.png" pageId="10" pageNumber="367">9</figureCitation>
, 
<figureCitation id="13690BB4FFBBAD14E2D8FA3EFE6DFA7B" box="[454,484,1466,1492]" captionStart="FIGURE 10" captionStartId="14.[151,250,1827,1851]" captionTargetBox="[152,1436,707,1803]" captionTargetId="figure-240@14.[151,1436,703,1803]" captionTargetPageId="14" captionText="FIGURE 10. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), drawings, A. First left coxa with stigmatic plate, posterior view. B. Left leg 9, posterior view. C. Second left coxa with gonopore, posterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. F. Right anterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332751" httpUri="https://zenodo.org/record/6332751/files/figure.png" pageId="10" pageNumber="367">10</figureCitation>
, 
<figureCitation id="13690BB4FFBBAD14E2EEFA3EFD87FA7B" box="[496,526,1466,1492]" captionStart="FIGURE 11" captionStartId="15.[151,250,1331,1355]" captionTargetBox="[153,1436,194,1307]" captionTargetId="figure-17@15.[151,1436,181,1307]" captionTargetPageId="15" captionText="FIGURE 11. Zephroniachrysomallos Bhansali &amp;Wesener sp. nov., holotype ♂ (NHMD MYR8826), drawings, A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view. Abbreviations: ct = crenulated teeth; imf = immovable finger; ss = sclerotized spot. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332753" httpUri="https://zenodo.org/record/6332753/files/figure.png" pageId="10" pageNumber="367">11</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBBAD14E389F986FE5FF85F" pageId="10" pageNumber="367" type="diagnosis">
<paragraph id="8BED1731FFBBAD14E389F986FE5FF85F" blockId="10.[151,1437,1538,2033]" pageId="10" pageNumber="367">
<emphasis id="B926CB23FFBBAD14E389F986FE83F9B3" bold="true" box="[151,266,1538,1564]" pageId="10" pageNumber="367">Diagnosis</emphasis>
. The position of the organ of Tömösváry at the brim and not in the antennal groove identifies this species as a member of the 
<taxonomicName id="4C526CB2FFBBAD14E271F9A2FE6AF9EF" authorityName="Gray" authorityYear="1832" box="[367,483,1574,1600]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B926CB23FFBBAD14E271F9A2FE6AF9EF" box="[367,483,1574,1600]" italics="true" pageId="10" pageNumber="367">Zephronia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2157658FFBBAD14E2F7F9A3FDFAF9EF" box="[489,627,1574,1601]" pageId="10" pageNumber="367" sensu="stricto">sensu stricto</taxonomicNameLabel>
group, with which it also aligns weakly genetically (
<figureCitation id="13690BB4FFBBAD14E7A3F9A2FB77F9EF" box="[1213,1278,1574,1600]" captionStart="FIGURE 1" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[212,1373,179,1877]" captionTargetId="figure-17@5.[212,1374,179,1883]" captionTargetPageId="5" captionText="FIGURE 1. Maximum likelihood tree based on the COI sequence after 1000 bootstrap replicates analyzed with the General Time Reversible Model. Numbers on branches indicate bootstrap support. Codon positions included were 1st+2nd+3rd. All positions with less than 5% site coverage were eliminated. The tree is drawn to scale, with branch length indicating genetic distance. Orange and green box represent new species of Zephronia and Sphaerobelum, respectively." figureDoi="http://doi.org/10.5281/zenodo.6332733" httpUri="https://zenodo.org/record/6332733/files/figure.png" pageId="10" pageNumber="367">Fig. 1</figureCitation>
). The sensory cones on the palpi of the gnathochilarium are arranged in a single field, not in separate clusters (
<figureCitation id="13690BB4FFBBAD14E7FEF9CEFAB1F9CB" box="[1248,1336,1610,1636]" captionStart="FIGURE 8" captionStartId="12.[151,250,1241,1265]" captionTargetBox="[151,1437,181,1216]" captionTargetId="figure-22@12.[151,1437,181,1217]" captionTargetPageId="12" captionText="FIGURE 8. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Left antennae, lateral view. B. Left sixth antennomere with antennal disc. C. Left antenna, disc. D. Gnathochilarium, apical view. E. Left lateral palpi. F. Left central pad, two types of sensory cones on central pad, detail. Abbreviations: ac = apical cone; ad = antennal disc; Cp = central pad; Hyp = hypopharynx; Ip = inner palpi; LL = lamellae lingulales; Sc = sensory cone; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332747" httpUri="https://zenodo.org/record/6332747/files/figure.png" pageId="10" pageNumber="367">Fig. 8D</figureCitation>
) and the body is covered in a dense coat of golden setae like in several other Thai 
<taxonomicName id="4C526CB2FFBBAD14E0F9F9EAFBD2F927" authorityName="Gray" authorityYear="1832" box="[999,1115,1646,1672]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="genus">
<emphasis id="B926CB23FFBBAD14E0F9F9EAFBD2F927" box="[999,1115,1646,1672]" italics="true" pageId="10" pageNumber="367">Zephronia</emphasis>
</taxonomicName>
. The greyish brown tergites color resembles those of 
<taxonomicName id="4C526CB2FFBBAD14E2A7F917FDBCF903" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[441,565,1683,1708]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="erawani" status="sp. nov.">
<emphasis id="B926CB23FFBBAD14E2A7F917FDBCF903" box="[441,565,1683,1708]" italics="true" pageId="10" pageNumber="367">Z. erawani</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBBAD14E120F916FD13F903" bold="true" box="[574,666,1682,1708]" pageId="10" pageNumber="367">
<taxonomicNameLabel id="A2157658FFBBAD14E120F916FD13F903" box="[574,666,1682,1708]" pageId="10" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C526CB2FFBBAD14E1CBF917FCCAF903" box="[725,835,1682,1708]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="B926CB23FFBBAD14E1CBF917FCCAF903" box="[725,835,1682,1708]" italics="true" pageId="10" pageNumber="367">Z. panhai</emphasis>
</taxonomicName>
(
<figureCitation id="13690BB4FFBBAD14E04AF916FC26F903" box="[852,943,1682,1708]" captionStart="FIGURE 2" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[337,1247,181,1766]" captionTargetId="figure-22@6.[337,1249,181,1766]" captionTargetPageId="6" captionText="FIGURE 2. Map of Thailand with known distribution of Sphaerobelum spp. and Zephronia spp. Star represents new species. Diamonds represent new localities. Fluorescent Green = S. meridionalis Bhansali &amp; Wesener sp. nov.; Pink = S. aesculus; Brown = S. truncatum; Chrome yellow = Z. chrysomallos Bhansali &amp; Wesener sp. nov.; White = Z. erawani Bhansali &amp; Wesener sp. nov.; Black = Z. viridisoma; Red = Z. panhai; Dark green = Z. golovatchi; Purple = Z. siamensis; Orange = Z. enghoffi; Yellow = Z. lannaensis; Blue = Z. phrain. B. Habitus photograph of Z. chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♀ (ZFMK MYR8826)." figureDoi="http://doi.org/10.5281/zenodo.6332735" httpUri="https://zenodo.org/record/6332735/files/figure.png" pageId="10" pageNumber="367">Fig. 2B</figureCitation>
). The male second leg mesal coxal margin is not extended into a membranous process, a character shared only with 
<taxonomicName id="4C526CB2FFBBAD14E0C2F933FBFAF97F" box="[988,1139,1718,1744]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="lannaensis">
<emphasis id="B926CB23FFBBAD14E0C2F933FBFAF97F" box="[988,1139,1718,1744]" italics="true" pageId="10" pageNumber="367">Z. lannaensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C526CB2FFBBAD14E7B2F933FA90F97F" box="[1196,1305,1718,1744]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="B926CB23FFBBAD14E7B2F933FA90F97F" box="[1196,1305,1718,1744]" italics="true" pageId="10" pageNumber="367">Z. panhai</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C526CB2FFBBAD14E637F932FEA5F95B" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBBAD14E637F932FEA5F95B" italics="true" pageId="10" pageNumber="367">Zephronia chrysomallos</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBBAD14E22DF95EFE04F95B" bold="true" box="[307,397,1754,1780]" pageId="10" pageNumber="367">
<taxonomicNameLabel id="A2157658FFBBAD14E22DF95EFE04F95B" box="[307,397,1754,1780]" pageId="10" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C526CB2FFBBAD14E13FF95FFD3FF95B" box="[545,694,1754,1780]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="lannaensis">
<emphasis id="B926CB23FFBBAD14E13FF95FFD3FF95B" box="[545,694,1754,1780]" italics="true" pageId="10" pageNumber="367">Z. lannaensis</emphasis>
</taxonomicName>
in the color (grey versus brown), telopod characters as well as the endotergum. 
<taxonomicName id="4C526CB2FFBBAD14E235F97BFE57F8B7" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[299,478,1790,1816]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBBAD14E235F97BFE57F8B7" box="[299,478,1790,1816]" italics="true" pageId="10" pageNumber="367">Z. chrysomallos</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBBAD14E2F8F97AFDC8F8B7" bold="true" box="[486,577,1790,1816]" pageId="10" pageNumber="367">
<taxonomicNameLabel id="A2157658FFBBAD14E2F8F97AFDC8F8B7" box="[486,577,1790,1816]" pageId="10" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from 
<taxonomicName id="4C526CB2FFBBAD14E1C9F97BFCCAF8B7" box="[727,835,1790,1816]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="B926CB23FFBBAD14E1C9F97BFCCAF8B7" box="[727,835,1790,1816]" italics="true" pageId="10" pageNumber="367">Z. panhai</emphasis>
</taxonomicName>
in being much larger in size (
<quantity id="4CAABAD4FFBBAD14E78BF97AFB6FF8B6" box="[1173,1254,1790,1817]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="367" unit="mm" value="40.0">40 mm</quantity>
versus 
<quantity id="4CAABAD4FFBBAD14E623F97AFA19F8B6" box="[1341,1424,1790,1817]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="367" unit="mm" value="20.0">20 mm</quantity>
), the higher number of ventral spines on the walking legs (10–12 versus 7–9), the higher number of apical cones on the antenna (78–88 versus around 50), the higher number of rows of marginal bristles on the endotergum (3 versus 2) and the shape of the first pleurite, which is weakly extended and well-rounded in 
<taxonomicName id="4C526CB2FFBBAD14E747F8EFFA84F82B" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[1113,1293,1898,1924]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBBAD14E747F8EFFA84F82B" box="[1113,1293,1898,1924]" italics="true" pageId="10" pageNumber="367">Z. chrysomallos</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBBAD14E60BF8EEFAF9F82B" bold="true" box="[1301,1392,1898,1924]" pageId="10" pageNumber="367">
<taxonomicNameLabel id="A2157658FFBBAD14E60BF8EEFAF9F82B" box="[1301,1392,1898,1924]" pageId="10" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
but with a long sharp process in 
<taxonomicName id="4C526CB2FFBBAD14E2C4F80BFDCFF807" box="[474,582,1934,1960]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="B926CB23FFBBAD14E2C4F80BFDCFF807" box="[474,582,1934,1960]" italics="true" pageId="10" pageNumber="367">Z. panhai</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C526CB2FFBBAD14E14AF80BFC8EF807" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[596,775,1934,1960]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBBAD14E14AF80BFC8EF807" box="[596,775,1934,1960]" italics="true" pageId="10" pageNumber="367">Z. chrysomallos</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBBAD14E010F80AFCE1F807" bold="true" box="[782,872,1934,1960]" pageId="10" pageNumber="367">
<taxonomicNameLabel id="A2157658FFBBAD14E010F80AFCE1F807" box="[782,872,1934,1960]" pageId="10" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs genetically in the COI barcoding fragment from 
<taxonomicName id="4C526CB2FFBBAD14E3CAF837FEB6F863" box="[212,319,1970,1996]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="panhai">
<emphasis id="B926CB23FFBBAD14E3CAF837FEB6F863" box="[212,319,1970,1996]" italics="true" pageId="10" pageNumber="367">Z. panhai</emphasis>
</taxonomicName>
by a p-distance of 10.8%, from 
<taxonomicName id="4C526CB2FFBBAD14E1BAF837FC94F863" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[676,797,1971,1996]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="erawani" status="sp. nov.">
<emphasis id="B926CB23FFBBAD14E1BAF837FC94F863" box="[676,797,1971,1996]" italics="true" pageId="10" pageNumber="367">Z. erawani</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBBAD14E03DF836FCF5F863" bold="true" box="[803,892,1970,1996]" pageId="10" pageNumber="367">
<taxonomicNameLabel id="A2157658FFBBAD14E03DF836FCF5F863" box="[803,892,1970,1996]" pageId="10" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by a p-distance of 11.9% and from 
<taxonomicName id="4C526CB2FFBBAD14E616F837FA15F863" box="[1288,1436,1970,1996]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="367" phylum="Arthropoda" rank="species" species="lannaensis">
<emphasis id="B926CB23FFBBAD14E616F837FA15F863" box="[1288,1436,1970,1996]" italics="true" pageId="10" pageNumber="367">Z. lannaensis</emphasis>
</taxonomicName>
by 14.8–15.2% (Sup. file 1).
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBAAD15E3D9FF12FE9DFF57" pageId="11" pageNumber="368" type="etymology">
<paragraph id="8BED1731FFBAAD15E3D9FF12FE9DFF57" blockId="11.[151,1437,150,969]" pageId="11" pageNumber="368">
<emphasis id="B926CB23FFBAAD15E3D9FF12FE17FF1F" bold="true" box="[199,414,150,176]" pageId="11" pageNumber="368">Derivatio nominis</emphasis>
. 
<taxonomicName id="4C526CB2FFBAAD15E2B2FF12FD1EFF1F" authority=", Greek" authorityName="Greek" box="[428,663,150,176]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="368" phylum="Arthropoda" rank="species" species="chrysomallos">chrysomallos, Greek</taxonomicName>
, meaning Golden Wool, and also the flying sheep ram Chrysomallos from Greek mythology, which was sacrificed to Zeus in order to obtain the famous Golden Fleece. Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBAAD15E3D9FE86FBC9FDCF" pageId="11" pageNumber="368" type="materials_examined">
<paragraph id="8BED1731FFBAAD15E3D9FE86FDB2FEB2" blockId="11.[151,1437,150,969]" box="[199,571,258,285]" pageId="11" pageNumber="368">
<emphasis id="B926CB23FFBAAD15E3D9FE86FE20FEB3" bold="true" box="[199,425,258,284]" pageId="11" pageNumber="368">Material examined</emphasis>
(total: 
<specimenCount id="9D54DCB8FFBAAD15E2EAFE86FDBAFEB2" box="[500,563,258,285]" pageId="11" pageNumber="368" type="male">7 ♂♂</specimenCount>
)
</paragraph>
<paragraph id="8BED1731FFBAAD15E3D9FEA2FEA6FEEF" blockId="11.[151,1437,150,969]" box="[199,303,294,320]" pageId="11" pageNumber="368">
<typeStatus id="54E9A993FFBAAD15E3D9FEA2FEA6FEEF" box="[199,303,294,320]" pageId="11" pageNumber="368" type="holotype">Holotype</typeStatus>
</paragraph>
<paragraph id="8BED1731FFBAAD15E3D9FECEFD98FE26" blockId="11.[151,1437,150,969]" pageId="11" pageNumber="368">
<materialsCitation id="3B3A1D6CFFBAAD15E3D9FECEFD98FE26" ID-GBIF-Occurrence="3698753313" collectionCode="ZFMK" country="Thailand" elevation="150" latitude="14.236667" location="Sai Yok Noi Waterfall" longLatPrecision="21" longitude="99.06278" municipality="Sai Yok District" pageId="11" pageNumber="368" specimenCode="MYR8826" specimenCount="151" specimenCount-male="1" stateProvince="Kanchanaburi" typeStatus="holotype">
<collectingCountry id="F34557A1FFBAAD15E3D9FECEFED5FECB" box="[199,348,330,356]" name="Thailand" pageId="11" pageNumber="368">THAILAND</collectingCountry>
: 
<specimenCount id="9D54DCB8FFBAAD15E272FECEFE17FECA" box="[364,414,330,357]" pageId="11" pageNumber="368" type="male">1 ♂</specimenCount>
, 
<emphasis id="B926CB23FFBAAD15E2B1FECEFD03FECB" bold="true" box="[431,650,330,356]" pageId="11" pageNumber="368">
<collectionCode id="ED438FF4FFBAAD15E2B1FECEFD8AFECB" box="[431,515,330,356]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="368" type="Museum">ZFMK</collectionCode>
<specimenCode id="DBF4BF4AFFBAAD15E111FECEFD03FECB" box="[527,650,330,356]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="368" type="Museum">MYR8826</specimenCode>
</emphasis>
, 
<collectingRegion id="4996D9D3FFBAAD15E180FECEFC24FECB" box="[670,941,330,356]" country="Thailand" name="Kanchanaburi" pageId="11" pageNumber="368">Kanchanaburi Province</collectingRegion>
, 
<collectingMunicipality id="6B898D4BFFBAAD15E0A0FECEFBF7FECB" box="[958,1150,330,356]" pageId="11" pageNumber="368">Sai Yok District</collectingMunicipality>
, 
<location id="8E8D41EAFFBAAD15E791FECEFA10FECB" LSID="urn:lsid:plazi:treatment:03FBA627FFBBAD13E389FA11FBB9F95F:8E8D41EAFFBAAD15E791FECEFA10FECB" box="[1167,1433,330,356]" country="Thailand" latitude="14.236667" longLatPrecision="21" longitude="99.06278" municipality="Sai Yok District" name="Sai Yok Noi Waterfall" pageId="11" pageNumber="368" stateProvince="Kanchanaburi">Sai Yok Noi Waterfall</location>
, 
<geoCoordinate id="EE6671F6FFBAAD15E389FEEAFEADFE26" box="[151,292,366,393]" degrees="14" direction="north" minutes="14" orientation="latitude" pageId="11" pageNumber="368" precision="15" seconds="12" value="14.236667">14°14'12'' N</geoCoordinate>
, 
<geoCoordinate id="EE6671F6FFBAAD15E231FEEAFE3EFE26" box="[303,439,366,393]" degrees="99" direction="east" minutes="03" orientation="longitude" pageId="11" pageNumber="368" precision="15" seconds="46" value="99.06278">99°03'46'' E</geoCoordinate>
, 
<specimenCount id="9D54DCB8FFBAAD15E2DDFEEAFD98FE26" box="[451,529,366,393]" pageId="11" pageNumber="368" type="generic">
<quantity id="4CAABAD4FFBAAD15E2DDFEEAFD84FE26" box="[451,525,366,393]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="368" unit="m" value="150.0">
<elevation id="007FF002FFBAAD15E2DDFEEAFD84FE26" box="[451,525,366,393]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="368" unit="m" value="150.0">150 m</elevation>
</quantity>
.
</specimenCount>
</materialsCitation>
</paragraph>
<paragraph id="8BED1731FFBAAD15E3D9FE16FEB6FE03" blockId="11.[151,1437,150,969]" box="[199,319,402,428]" pageId="11" pageNumber="368">
<emphasis id="B926CB23FFBAAD15E3D9FE16FEB6FE03" bold="true" box="[199,319,402,428]" pageId="11" pageNumber="368">
<typeStatus id="54E9A993FFBAAD15E3D9FE16FEB6FE03" box="[199,319,402,428]" pageId="11" pageNumber="368" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BED1731FFBAAD15E3D9FE32FE34FE5B" blockId="11.[151,1437,150,969]" pageId="11" pageNumber="368">
<materialsCitation id="3B3A1D6CFFBAAD15E3D9FE32FBFAFE7F" ID-GBIF-Occurrence="3698753325" accessionNumber="MYR11366" box="[199,1139,438,465]" collectionCode="ZFMK, CT" country="Thailand" elevation="150" latitude="14.236667" location="Sai Yok Noi Waterfall" longLatPrecision="21" longitude="99.06278" municipality="Sai Yok District" pageId="11" pageNumber="368" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi" typeStatus="paratype">
<collectingCountry id="F34557A1FFBAAD15E3D9FE32FED5FE7F" box="[199,348,438,464]" name="Thailand" pageId="11" pageNumber="368">THAILAND</collectingCountry>
: 
<specimenCount id="9D54DCB8FFBAAD15E278FE32FE18FE7E" box="[358,401,438,465]" pageId="11" pageNumber="368" type="male">1 ♂</specimenCount>
, 
<emphasis id="B926CB23FFBAAD15E283FE32FDF6FE7F" bold="true" box="[413,639,438,464]" pageId="11" pageNumber="368">
<collectionCode id="ED438FF4FFBAAD15E283FE32FE78FE7F" box="[413,497,438,464]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="368" type="Museum">ZFMK</collectionCode>
MYR11366
</emphasis>
, 
<collectionCode id="ED438FF4FFBAAD15E195FE32FD27FE7F" box="[651,686,438,464]" pageId="11" pageNumber="368">CT</collectionCode>
scan voucher, same data as for holotype
</materialsCitation>
; 
<materialsCitation id="3B3A1D6CFFBAAD15E763FE32FE30FE5B" ID-GBIF-Occurrence="3698753308" accessionNumber="MYR11365" collectionCode="ZFMK" country="Thailand" elevation="150" latitude="14.236667" location="Sai Yok Noi Waterfall" longLatPrecision="21" longitude="99.06278" municipality="Sai Yok District" pageId="11" pageNumber="368" specimenCount="1" specimenCount-male="1" stateProvince="Kanchanaburi" typeStatus="paratype">
<specimenCount id="9D54DCB8FFBAAD15E763FE32FB21FE7E" box="[1149,1192,438,465]" pageId="11" pageNumber="368" type="male">1 ♂</specimenCount>
, 
<emphasis id="B926CB23FFBAAD15E7AAFE32FA1FFE7F" bold="true" box="[1204,1430,438,464]" pageId="11" pageNumber="368">
<collectionCode id="ED438FF4FFBAAD15E7AAFE32FA81FE7F" box="[1204,1288,438,464]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="11" pageNumber="368" type="Museum">ZFMK</collectionCode>
MYR11365
</emphasis>
, same data as for holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BED1731FFBAAD15E3D9FE7AFEF3FDB7" blockId="11.[151,1437,150,969]" box="[199,378,510,536]" pageId="11" pageNumber="368">
<emphasis id="B926CB23FFBAAD15E3D9FE7AFEF3FDB7" bold="true" box="[199,378,510,536]" pageId="11" pageNumber="368">Other material</emphasis>
</paragraph>
<paragraph id="8BED1731FFBAAD15E3D9FDA6FBC9FDCF" blockId="11.[151,1437,150,969]" pageId="11" pageNumber="368">
<materialsCitation id="3B3A1D6CFFBAAD15E3D9FDA6FBB2FDCF" ID-GBIF-Occurrence="3698753326" collectingDate="2009-07-11" collectorName="Sai Yok Noi Waterfall &amp; Khao Phang WF" country="Thailand" elevation="160" latitude="14.239167" location="Sai Yok District" longLatPrecision="21" longitude="99.05778" pageId="11" pageNumber="368" specimenCount="4" specimenCount-male="4" stateProvince="Kanchanaburi">
<collectingCountry id="F34557A1FFBAAD15E3D9FDA6FED5FD93" box="[199,348,546,572]" name="Thailand" pageId="11" pageNumber="368">THAILAND</collectingCountry>
: 
<specimenCount id="9D54DCB8FFBAAD15E277FDA6FE25FD92" box="[361,428,546,573]" pageId="11" pageNumber="368" type="male">4 ♂♂</specimenCount>
, 
<emphasis id="B926CB23FFBAAD15E2A4FDA6FD99FD93" bold="true" box="[442,528,546,572]" pageId="11" pageNumber="368">CUMZ</emphasis>
, 
<collectingRegion id="4996D9D3FFBAAD15E13EFDA6FCA5FD93" box="[544,812,546,572]" country="Thailand" name="Kanchanaburi" pageId="11" pageNumber="368">Kanchanaburi Province</collectingRegion>
, 
<location id="8E8D41EAFFBAAD15E024FDA6FC7DFD93" LSID="urn:lsid:plazi:treatment:03FBA627FFBBAD13E389FA11FBB9F95F:8E8D41EAFFBAAD15E024FDA6FC7DFD93" box="[826,1012,546,572]" country="Thailand" latitude="14.239167" longLatPrecision="21" longitude="99.05778" name="Sai Yok District" pageId="11" pageNumber="368" stateProvince="Kanchanaburi">Sai Yok District</location>
, 
<collectorName id="26A772E7FFBAAD15E71FFDA6FB74FD93" box="[1025,1277,546,572]" pageId="11" pageNumber="368">Sai Yok Noi Waterfall</collectorName>
(
<collectorName id="26A772E7FFBAAD15E611FDA6FF4CFDCE" pageId="11" pageNumber="368">Khao Phang WF</collectorName>
), 
<geoCoordinate id="EE6671F6FFBAAD15E3C7FDC2FEECFDCE" box="[217,357,582,609]" degrees="14" direction="north" minutes="14" orientation="latitude" pageId="11" pageNumber="368" precision="15" seconds="21" value="14.239167">14°14'21'' N</geoCoordinate>
, 
<geoCoordinate id="EE6671F6FFBAAD15E26EFDC2FE71FDCE" box="[368,504,582,609]" degrees="99" direction="east" minutes="03" orientation="longitude" pageId="11" pageNumber="368" precision="15" seconds="28" value="99.05778">99°03'28'' E</geoCoordinate>
, ca. 
<elevation id="007FF002FFBAAD15E135FDC2FD26FDCF" box="[555,687,582,609]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="368" unit="m" value="160.0">
<quantity id="4CAABAD4FFBAAD15E135FDC2FDFBFDCE" box="[555,626,582,609]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="368" unit="m" value="160.0">160 m</quantity>
a.s.l.
</elevation>
, 
<date id="FFEC31F1FFBAAD15E1A5FDC2FCCFFDCF" box="[699,838,582,608]" pageId="11" pageNumber="368" value="2009-07-11">
<collectingDate id="EFA8C819FFBAAD15E1A5FDC2FCCFFDCF" box="[699,838,582,608]" pageId="11" pageNumber="368" value="2009-07-11">11.VII.2009</collectingDate>
</date>
, leg. ASRU members
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBAAD13E3D9FDEEFC84F92B" lastPageId="13" lastPageNumber="370" pageId="11" pageNumber="368" type="description">
<paragraph id="8BED1731FFBAAD15E3D9FDEEFDB1FD2A" blockId="11.[151,1437,150,969]" box="[199,568,618,645]" pageId="11" pageNumber="368">
<emphasis id="B926CB23FFBAAD15E3D9FDEEFED8FD2B" bold="true" box="[199,337,618,644]" pageId="11" pageNumber="368">Description</emphasis>
(based on 
<typeStatus id="54E9A993FFBAAD15E2D2FDEEFDB9FD2A" box="[460,560,618,645]" pageId="11" pageNumber="368" type="holotype">holotype</typeStatus>
)
</paragraph>
<paragraph id="8BED1731FFBAAD15E3D9FD0AFCA1FD62" blockId="11.[151,1437,150,969]" pageId="11" pageNumber="368">
<emphasis id="B926CB23FFBAAD15E3D9FD0AFF7EFD07" bold="true" box="[199,247,654,680]" pageId="11" pageNumber="368">Size</emphasis>
. Length: ca. 
<quantity id="4CAABAD4FFBAAD15E28BFD0AFE62FD06" box="[405,491,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="11" pageNumber="368" unit="mm" value="40.0">40 mm</quantity>
. Width of thoracic shield 
<quantity id="4CAABAD4FFBAAD15E003FD0AFC0EFD06" box="[797,903,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="11" pageNumber="368" unit="mm" value="16.3">16.3 mm</quantity>
, of widest segment (8th) 
<quantity id="4CAABAD4FFBAAD15E7ABFD0AFAA9FD06" box="[1205,1312,654,681]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="11" pageNumber="368" unit="mm" value="16.9">16.9 mm</quantity>
. Height of thoracic shield 
<quantity id="4CAABAD4FFBAAD15E25CFD36FE13FD62" box="[322,410,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="11" pageNumber="368" unit="mm" value="9.4">9.4 mm</quantity>
, of highest segment (8th) 
<quantity id="4CAABAD4FFBAAD15E1A0FD36FCADFD62" box="[702,804,690,717]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="368" unit="mm" value="11.5">11.5 mm</quantity>
.
</paragraph>
<paragraph id="8BED1731FFBAAD15E3D9FD52FA1EFC97" blockId="11.[151,1437,150,969]" pageId="11" pageNumber="368">
<emphasis id="B926CB23FFBAAD15E3D9FD52FE82FD5F" bold="true" box="[199,267,726,752]" pageId="11" pageNumber="368">Color</emphasis>
(from living specimens). Head, collum and anterior margin of thoracic shield dark brown to black. Antenna dark brown, legs pale brown. Tergites and anal shield greyish-brown (fading to light brown in ethanol), darker towards posterior margin. Thoracic shield-like tergite with darker anterior and posterior margins. (
<figureCitation id="13690BB4FFBAAD15E62FFC9AFA00FC97" box="[1329,1417,798,824]" captionStart="FIGURE 2" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[337,1247,181,1766]" captionTargetId="figure-22@6.[337,1249,181,1766]" captionTargetPageId="6" captionText="FIGURE 2. Map of Thailand with known distribution of Sphaerobelum spp. and Zephronia spp. Star represents new species. Diamonds represent new localities. Fluorescent Green = S. meridionalis Bhansali &amp; Wesener sp. nov.; Pink = S. aesculus; Brown = S. truncatum; Chrome yellow = Z. chrysomallos Bhansali &amp; Wesener sp. nov.; White = Z. erawani Bhansali &amp; Wesener sp. nov.; Black = Z. viridisoma; Red = Z. panhai; Dark green = Z. golovatchi; Purple = Z. siamensis; Orange = Z. enghoffi; Yellow = Z. lannaensis; Blue = Z. phrain. B. Habitus photograph of Z. chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♀ (ZFMK MYR8826)." figureDoi="http://doi.org/10.5281/zenodo.6332735" httpUri="https://zenodo.org/record/6332735/files/figure.png" pageId="11" pageNumber="368">Fig. 2B</figureCitation>
).
</paragraph>
<paragraph id="8BED1731FFBAAD15E3D9FCC6FD28FC67" blockId="11.[151,1437,150,969]" pageId="11" pageNumber="368">
<emphasis id="B926CB23FFBAAD15E3D9FCC6FE8EFCF3" bold="true" box="[199,263,834,860]" pageId="11" pageNumber="368">Head</emphasis>
(
<figureCitation id="13690BB4FFBAAD15E207FCC6FEFFFCF2" box="[281,374,834,861]" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="11" pageNumber="368">Fig. 7D</figureCitation>
). Number of ommatidia more than 90. Tömösváry organ placed on rim of antennal groove. Antennae (
<figureCitation id="13690BB4FFBAAD15E208FCE2FE1CFC2F" box="[278,405,870,896]" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="11" pageNumber="368">Fig. 7A–D</figureCitation>
) short, axe-shaped (
<figureCitation id="13690BB4FFBAAD15E19BFCE2FC8CFC2F" box="[645,773,870,896]" captionStart="FIGURE 8" captionStartId="12.[151,250,1241,1265]" captionTargetBox="[151,1437,181,1216]" captionTargetId="figure-22@12.[151,1437,181,1217]" captionTargetPageId="12" captionText="FIGURE 8. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Left antennae, lateral view. B. Left sixth antennomere with antennal disc. C. Left antenna, disc. D. Gnathochilarium, apical view. E. Left lateral palpi. F. Left central pad, two types of sensory cones on central pad, detail. Abbreviations: ac = apical cone; ad = antennal disc; Cp = central pad; Hyp = hypopharynx; Ip = inner palpi; LL = lamellae lingulales; Sc = sensory cone; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332747" httpUri="https://zenodo.org/record/6332747/files/figure.png" pageId="11" pageNumber="368">Fig. 8A–C</figureCitation>
), not reaching to center of head (
<figureCitation id="13690BB4FFBAAD15E788FCE2FB7AFC2F" box="[1174,1267,870,896]" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="11" pageNumber="368">Fig. 7C</figureCitation>
), antennomere lengths: 6&gt;1&gt;5=4=3=2. Antennomere 1 basally with scales (
<figureCitation id="13690BB4FFBAAD15E023FC0EFC11FC0B" box="[829,920,906,932]" captionStart="FIGURE 8" captionStartId="12.[151,250,1241,1265]" captionTargetBox="[151,1437,181,1216]" captionTargetId="figure-22@12.[151,1437,181,1217]" captionTargetPageId="12" captionText="FIGURE 8. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Left antennae, lateral view. B. Left sixth antennomere with antennal disc. C. Left antenna, disc. D. Gnathochilarium, apical view. E. Left lateral palpi. F. Left central pad, two types of sensory cones on central pad, detail. Abbreviations: ac = apical cone; ad = antennal disc; Cp = central pad; Hyp = hypopharynx; Ip = inner palpi; LL = lamellae lingulales; Sc = sensory cone; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332747" httpUri="https://zenodo.org/record/6332747/files/figure.png" pageId="11" pageNumber="368">Fig. 8A</figureCitation>
). Antennomere 6 apically wide, and flattened, axe-shaped, with 78/88 apical cones (
<figureCitation id="13690BB4FFBAAD15E122FC2AFD1DFC67" box="[572,660,942,968]" captionStart="FIGURE 8" captionStartId="12.[151,250,1241,1265]" captionTargetBox="[151,1437,181,1216]" captionTargetId="figure-22@12.[151,1437,181,1217]" captionTargetPageId="12" captionText="FIGURE 8. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Left antennae, lateral view. B. Left sixth antennomere with antennal disc. C. Left antenna, disc. D. Gnathochilarium, apical view. E. Left lateral palpi. F. Left central pad, two types of sensory cones on central pad, detail. Abbreviations: ac = apical cone; ad = antennal disc; Cp = central pad; Hyp = hypopharynx; Ip = inner palpi; LL = lamellae lingulales; Sc = sensory cone; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332747" httpUri="https://zenodo.org/record/6332747/files/figure.png" pageId="11" pageNumber="368">Fig. 8C</figureCitation>
).
</paragraph>
<caption id="DF2D47B9FFBAAD15E389F8A1FB64F85E" ID-DOI="http://doi.org/10.5281/zenodo.6332745" ID-Zenodo-Dep="6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="11" pageNumber="368" startId="11.[151,250,1829,1853]" targetBox="[151,1431,992,1806]" targetPageId="11">
<paragraph id="8BED1731FFBAAD15E389F8A1FB64F85E" blockId="11.[151,1437,1829,2034]" pageId="11" pageNumber="368">
<emphasis id="B926CB23FFBAAD15E389F8A1FE9AF891" bold="true" box="[151,275,1829,1854]" pageId="11" pageNumber="368">FIGURE 7.</emphasis>
<taxonomicName id="4C526CB2FFBAAD15E204F8A1FD6EF892" authority="Bhansali &amp; Wesener" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[282,743,1829,1854]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="368" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBAAD15E204F8A1FD99F892" box="[282,528,1829,1853]" italics="true" pageId="11" pageNumber="368">Zephronia chrysomallos</emphasis>
Bhansali &amp; Wesener
</taxonomicName>
<emphasis id="B926CB23FFBAAD15E1F3F8A2FCB7F891" bold="true" box="[749,830,1830,1854]" pageId="11" pageNumber="368">
<taxonomicNameLabel id="A2157658FFBAAD15E1F3F8A2FCB7F891" box="[749,830,1830,1854]" pageId="11" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype ♂ (ZFMK 
<emphasis id="B926CB23FFBAAD15E706F8A1FB1AF892" bold="true" box="[1048,1171,1829,1853]" pageId="11" pageNumber="368">MYR11366</emphasis>
), volume rendering based on micro-computed tomography. 
<emphasis id="B926CB23FFBAAD15E2F7F8CDFD8BF8CE" bold="true" box="[489,514,1865,1889]" pageId="11" pageNumber="368">A.</emphasis>
Habitus, lateral view. 
<emphasis id="B926CB23FFBAAD15E1FAF8CEFD72F8CD" bold="true" box="[740,763,1866,1890]" pageId="11" pageNumber="368">B.</emphasis>
Habitus, ventro-lateral view. 
<emphasis id="B926CB23FFBAAD15E739F8CDFBB6F8CE" bold="true" box="[1063,1087,1865,1889]" pageId="11" pageNumber="368">C.</emphasis>
Habitus, ventral view. 
<emphasis id="B926CB23FFBAAD15E636F8CEFAC8F8CD" bold="true" box="[1320,1345,1866,1890]" pageId="11" pageNumber="368">D.</emphasis>
Habitus, anterior view. 
<emphasis id="B926CB23FFBAAD15E236F8EAFEC9F829" bold="true" box="[296,320,1902,1926]" pageId="11" pageNumber="368">E.</emphasis>
Body-ring architecture, cross section trough midbody-ring. 
<emphasis id="B926CB23FFBAAD15E0BEF8E9FBB0F82A" bold="true" box="[928,1081,1901,1925]" pageId="11" pageNumber="368">Abbreviations</emphasis>
: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale.
</paragraph>
</caption>
<caption id="DF2D47B9FFBDAD12E389FB5DFDC2FA2E" ID-DOI="http://doi.org/10.5281/zenodo.6332747" ID-Zenodo-Dep="6332747" httpUri="https://zenodo.org/record/6332747/files/figure.png" pageId="12" pageNumber="369" startId="12.[151,250,1241,1265]" targetBox="[151,1437,181,1216]" targetPageId="12">
<paragraph id="8BED1731FFBDAD12E389FB5DFDC2FA2E" blockId="12.[151,1437,1241,1409]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E389FB5DFE91FB5E" bold="true" box="[151,280,1241,1265]" pageId="12" pageNumber="369">FIGURE 8.</emphasis>
<taxonomicName id="4C526CB2FFBDAD12E23AFB5DFC8CFB5F" authority="Bhansali &amp; Wesener" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[292,773,1241,1265]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="12" pageNumber="369" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBDAD12E23AFB5DFD96FB5E" box="[292,543,1241,1265]" italics="true" pageId="12" pageNumber="369">Zephronia chrysomallos</emphasis>
Bhansali &amp; Wesener
</taxonomicName>
<emphasis id="B926CB23FFBDAD12E00EFB5DFCEFFB5E" bold="true" box="[784,870,1241,1265]" pageId="12" pageNumber="369">
<taxonomicNameLabel id="A2157658FFBDAD12E00EFB5DFCEFFB5E" box="[784,870,1241,1265]" pageId="12" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂ (
<emphasis id="B926CB23FFBDAD12E71AFB5DFB43FB5E" bold="true" box="[1028,1226,1241,1265]" pageId="12" pageNumber="369">ZFMK MYR8826</emphasis>
), scanning electron micrographs. 
<emphasis id="B926CB23FFBDAD12E237FB79FECBFABA" bold="true" box="[297,322,1277,1301]" pageId="12" pageNumber="369">A.</emphasis>
Left antennae, lateral view. 
<emphasis id="B926CB23FFBDAD12E19AFB79FD12FABA" bold="true" box="[644,667,1277,1301]" pageId="12" pageNumber="369">B.</emphasis>
Left sixth antennomere with antennal disc. 
<emphasis id="B926CB23FFBDAD12E796FB79FB29FABA" bold="true" box="[1160,1184,1277,1301]" pageId="12" pageNumber="369">C.</emphasis>
Left antenna, disc. 
<emphasis id="B926CB23FFBDAD12E69AFB79FA15FABA" bold="true" box="[1412,1436,1277,1301]" pageId="12" pageNumber="369">D.</emphasis>
Gnathochilarium, apical view. 
<emphasis id="B926CB23FFBDAD12E2C7FAA5FE79FA96" bold="true" box="[473,496,1313,1337]" pageId="12" pageNumber="369">E.</emphasis>
Left lateral palpi. 
<emphasis id="B926CB23FFBDAD12E1A4FAA5FD44FA96" bold="true" box="[698,717,1313,1337]" pageId="12" pageNumber="369">F.</emphasis>
Left central pad, two types of sensory cones on central pad, detail. 
<emphasis id="B926CB23FFBDAD12E389FAC1FEB9FAF2" bold="true" box="[151,304,1349,1373]" pageId="12" pageNumber="369">Abbreviations</emphasis>
: ac = apical cone; ad = antennal disc; Cp = central pad; Hyp = hypopharynx; Ip = inner palpi; LL = lamellae lingulales; Sc = sensory cone; St = stipites.
</paragraph>
</caption>
<paragraph id="8BED1731FFBDAD12E3D9FA2BFDFCFA66" blockId="12.[151,1437,1455,2022]" box="[199,629,1455,1482]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E3D9FA2BFEDFFA66" bold="true" box="[199,342,1455,1481]" pageId="12" pageNumber="369">Epipharynx</emphasis>
. With a large inner tooth.
</paragraph>
<paragraph id="8BED1731FFBDAD12E3D9FA57FF4CF9F6" blockId="12.[151,1437,1455,2022]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E3D9FA57FE1FFA42" bold="true" box="[199,406,1491,1517]" pageId="12" pageNumber="369">Gnathochilarium</emphasis>
(
<figureCitation id="13690BB4FFBDAD12E2B4FA57FD85FA41" box="[426,524,1491,1518]" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="12" pageNumber="369">Figs 7D</figureCitation>
, 
<figureCitation id="13690BB4FFBDAD12E102FA57FDD5FA42" box="[540,604,1491,1517]" captionStart="FIGURE 8" captionStartId="12.[151,250,1241,1265]" captionTargetBox="[151,1437,181,1216]" captionTargetId="figure-22@12.[151,1437,181,1217]" captionTargetPageId="12" captionText="FIGURE 8. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Left antennae, lateral view. B. Left sixth antennomere with antennal disc. C. Left antenna, disc. D. Gnathochilarium, apical view. E. Left lateral palpi. F. Left central pad, two types of sensory cones on central pad, detail. Abbreviations: ac = apical cone; ad = antennal disc; Cp = central pad; Hyp = hypopharynx; Ip = inner palpi; LL = lamellae lingulales; Sc = sensory cone; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332747" httpUri="https://zenodo.org/record/6332747/files/figure.png" pageId="12" pageNumber="369">8D–F</figureCitation>
, 
<figureCitation id="13690BB4FFBDAD12E173FA57FD1BFA42" box="[621,658,1491,1517]" captionStart="FIGURE 9" captionStartId="13.[151,250,707,731]" captionTargetBox="[152,1436,181,683]" captionTargetId="figure-17@13.[151,1436,181,684]" captionTargetPageId="13" captionText="FIGURE 9. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Gnathochilarium, ventral view. B. Right mandible, mesal view. Abbreviations: C = condyles; Cp = cental pad; eT = external tooth; Ip = inner palpi; iT = internal combined tooth; LL = lamellae linguales; Mn = mentum; pL = pectinate lamellae; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332749" httpUri="https://zenodo.org/record/6332749/files/figure.png" pageId="12" pageNumber="369">9A</figureCitation>
). Lamellae linguales with numerous long setae, more towards the margins. Stipes and mentum with numerous long setae in a regular pattern (
<figureCitation id="13690BB4FFBDAD12E0EAFA73FBC6F9BE" box="[1012,1103,1527,1553]" captionStart="FIGURE 9" captionStartId="13.[151,250,707,731]" captionTargetBox="[152,1436,181,683]" captionTargetId="figure-17@13.[151,1436,181,684]" captionTargetPageId="13" captionText="FIGURE 9. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Gnathochilarium, ventral view. B. Right mandible, mesal view. Abbreviations: C = condyles; Cp = cental pad; eT = external tooth; Ip = inner palpi; iT = internal combined tooth; LL = lamellae linguales; Mn = mentum; pL = pectinate lamellae; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332749" httpUri="https://zenodo.org/record/6332749/files/figure.png" pageId="12" pageNumber="369">Fig. 9A</figureCitation>
). Gnathochilarium palpi with sensory cones arranged in single field (
<figureCitation id="13690BB4FFBDAD12E143F99FFD3EF99A" box="[605,695,1563,1589]" captionStart="FIGURE 8" captionStartId="12.[151,250,1241,1265]" captionTargetBox="[151,1437,181,1216]" captionTargetId="figure-22@12.[151,1437,181,1217]" captionTargetPageId="12" captionText="FIGURE 8. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Left antennae, lateral view. B. Left sixth antennomere with antennal disc. C. Left antenna, disc. D. Gnathochilarium, apical view. E. Left lateral palpi. F. Left central pad, two types of sensory cones on central pad, detail. Abbreviations: ac = apical cone; ad = antennal disc; Cp = central pad; Hyp = hypopharynx; Ip = inner palpi; LL = lamellae lingulales; Sc = sensory cone; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332747" httpUri="https://zenodo.org/record/6332747/files/figure.png" pageId="12" pageNumber="369">Fig. 8E</figureCitation>
). 'Pillows' of central pads with 
<specimenCount id="9D54DCB8FFBDAD12E73AF998FB1AF999" box="[1060,1171,1564,1590]" pageId="12" pageNumber="369" type="generic" typeStatus="types">two types</specimenCount>
of sensory cones (
<figureCitation id="13690BB4FFBDAD12E671F99FFF31F9F6" captionStart="FIGURE 8" captionStartId="12.[151,250,1241,1265]" captionTargetBox="[151,1437,181,1216]" captionTargetId="figure-22@12.[151,1437,181,1217]" captionTargetPageId="12" captionText="FIGURE 8. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Left antennae, lateral view. B. Left sixth antennomere with antennal disc. C. Left antenna, disc. D. Gnathochilarium, apical view. E. Left lateral palpi. F. Left central pad, two types of sensory cones on central pad, detail. Abbreviations: ac = apical cone; ad = antennal disc; Cp = central pad; Hyp = hypopharynx; Ip = inner palpi; LL = lamellae lingulales; Sc = sensory cone; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332747" httpUri="https://zenodo.org/record/6332747/files/figure.png" pageId="12" pageNumber="369">Fig. 8F</figureCitation>
).
</paragraph>
<paragraph id="8BED1731FFBDAD12E3D9F9E7FD61F90E" blockId="12.[151,1437,1455,2022]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E3D9F9E7FEB3F9D2" bold="true" box="[199,314,1635,1661]" pageId="12" pageNumber="369">Mandible</emphasis>
(
<figureCitation id="13690BB4FFBDAD12E255F9E7FE2CF9D2" box="[331,421,1635,1661]" captionStart="FIGURE 9" captionStartId="13.[151,250,707,731]" captionTargetBox="[152,1436,181,683]" captionTargetId="figure-17@13.[151,1436,181,684]" captionTargetPageId="13" captionText="FIGURE 9. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), scanning electron micrographs. A. Gnathochilarium, ventral view. B. Right mandible, mesal view. Abbreviations: C = condyles; Cp = cental pad; eT = external tooth; Ip = inner palpi; iT = internal combined tooth; LL = lamellae linguales; Mn = mentum; pL = pectinate lamellae; St = stipites." figureDoi="http://doi.org/10.5281/zenodo.6332749" httpUri="https://zenodo.org/record/6332749/files/figure.png" pageId="12" pageNumber="369">Fig. 9B</figureCitation>
). External tooth undivided, inner tooth with 3 lobes. Pectinate lamellae with 6 or 7 rows. Condylus at anterior margin with two distinct ridges.
</paragraph>
<paragraph id="8BED1731FFBDAD12E3D9F92FFA15F96A" blockId="12.[151,1437,1455,2022]" box="[199,1436,1707,1734]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E3D9F92FFEA9F96A" bold="true" box="[199,288,1707,1733]" pageId="12" pageNumber="369">Collum</emphasis>
(
<figureCitation id="13690BB4FFBDAD12E231F92FFE27F96A" box="[303,430,1707,1733]" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="12" pageNumber="369">Figs 7A, C</figureCitation>
). With few setae distributed towards the margins, more concentrated towards lateral ends.
</paragraph>
<paragraph id="8BED1731FFBDAD12E3D9F94BFABAF946" blockId="12.[151,1437,1455,2022]" box="[199,1331,1743,1770]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E3D9F94BFE09F946" bold="true" box="[199,384,1743,1769]" pageId="12" pageNumber="369">Thoracic shield</emphasis>
. Thoracic shield grooves wide and shallow (
<figureCitation id="13690BB4FFBDAD12E06AF94BFC7AF946" box="[884,1011,1743,1769]" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="12" pageNumber="369">Figs 7A–C</figureCitation>
), without sclerotized ledges.
</paragraph>
<paragraph id="8BED1731FFBDAD12E3D9F977FB1FF8A1" blockId="12.[151,1437,1455,2022]" box="[199,1174,1779,1806]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E3D9F977FE3EF8A2" bold="true" box="[199,439,1779,1805]" pageId="12" pageNumber="369">First stigmatic plate</emphasis>
(
<figureCitation id="13690BB4FFBDAD12E2D8F977FDA7F8A2" box="[454,558,1779,1805]" captionStart="FIGURE 10" captionStartId="14.[151,250,1827,1851]" captionTargetBox="[152,1436,707,1803]" captionTargetId="figure-240@14.[151,1436,703,1803]" captionTargetPageId="14" captionText="FIGURE 10. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), drawings, A. First left coxa with stigmatic plate, posterior view. B. Left leg 9, posterior view. C. Second left coxa with gonopore, posterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. F. Right anterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332751" httpUri="https://zenodo.org/record/6332751/files/figure.png" pageId="12" pageNumber="369">Fig. 10A</figureCitation>
). Smaller than coxa, with rounded sub-triangular apex.
</paragraph>
<paragraph id="8BED1731FFBDAD12E3D9F893FBB3F8F9" blockId="12.[151,1437,1455,2022]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E3D9F893FEAFF89E" bold="true" box="[199,294,1815,1841]" pageId="12" pageNumber="369">Tergites</emphasis>
. Tergites densely covered in short, dense setae visible at higher magnification, with dull orange skinlike surface (
<figureCitation id="13690BB4FFBDAD12E236F8BFFE09F8FA" box="[296,384,1851,1877]" captionStart="FIGURE 2" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[337,1247,181,1766]" captionTargetId="figure-22@6.[337,1249,181,1766]" captionTargetPageId="6" captionText="FIGURE 2. Map of Thailand with known distribution of Sphaerobelum spp. and Zephronia spp. Star represents new species. Diamonds represent new localities. Fluorescent Green = S. meridionalis Bhansali &amp; Wesener sp. nov.; Pink = S. aesculus; Brown = S. truncatum; Chrome yellow = Z. chrysomallos Bhansali &amp; Wesener sp. nov.; White = Z. erawani Bhansali &amp; Wesener sp. nov.; Black = Z. viridisoma; Red = Z. panhai; Dark green = Z. golovatchi; Purple = Z. siamensis; Orange = Z. enghoffi; Yellow = Z. lannaensis; Blue = Z. phrain. B. Habitus photograph of Z. chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♀ (ZFMK MYR8826)." figureDoi="http://doi.org/10.5281/zenodo.6332735" httpUri="https://zenodo.org/record/6332735/files/figure.png" pageId="12" pageNumber="369">Fig. 2B</figureCitation>
). Paratergite tips only weakly projecting backwards (
<figureCitation id="13690BB4FFBDAD12E0CAF8BFFBA4F8F9" box="[980,1069,1851,1878]" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="12" pageNumber="369">Fig. 7B</figureCitation>
).
</paragraph>
<paragraph id="8BED1731FFBDAD12E3D9F8DBFADDF8D6" blockId="12.[151,1437,1455,2022]" box="[199,1364,1887,1914]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E3D9F8DBFEB8F8D6" bold="true" box="[199,305,1887,1913]" pageId="12" pageNumber="369">Pleurites</emphasis>
. Pleurite 1 well-rounded, weakly projecting. Pleurite 2 well-rounded, not projecting (
<figureCitation id="13690BB4FFBDAD12E7F1F8DBFACEF8D6" box="[1263,1351,1887,1913]" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="12" pageNumber="369">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="8BED1731FFBDAD12E3D9F807FCBDF84A" blockId="12.[151,1437,1455,2022]" pageId="12" pageNumber="369">
<emphasis id="B926CB23FFBDAD12E3D9F807FED7F832" bold="true" box="[199,350,1923,1949]" pageId="12" pageNumber="369">Endotergum</emphasis>
(
<figureCitation id="13690BB4FFBDAD12E273F807FE4CF831" box="[365,453,1923,1950]" captionStart="FIGURE 6" captionStartId="10.[151,250,974,998]" captionTargetBox="[151,1436,181,950]" captionTargetId="figure-22@10.[151,1436,181,950]" captionTargetPageId="10" captionText="FIGURE 6. Endoterga of midbody tergites, scanning electron micrographs. A. Sphaerobelum meridionalis Bhansali &amp; Wesener sp. nov., holotype ♂ (MHNG 4B-2). B. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826). Abbreviations: Ci = cuticular impression; Ia = inner area; Ma = middle area; Oa = outer area." figureDoi="http://doi.org/10.5281/zenodo.6332743" httpUri="https://zenodo.org/record/6332743/files/figure.png" pageId="12" pageNumber="369">Fig. 6B</figureCitation>
). With a regular flat margin. Outer area with two dense rows of marginal setae, reaching 4/5 of outer area. Middle section with a single row of rounded cuticular impression, distance between impression 2–3 times their diameter. Inner area without setae or spines.
</paragraph>
<caption id="DF2D47B9FFBCAD13E389FD47FEF3FCE7" ID-DOI="http://doi.org/10.5281/zenodo.6332749" ID-Zenodo-Dep="6332749" httpUri="https://zenodo.org/record/6332749/files/figure.png" pageId="13" pageNumber="370" startId="13.[151,250,707,731]" targetBox="[152,1436,181,683]" targetPageId="13">
<paragraph id="8BED1731FFBCAD13E389FD47FEF3FCE7" blockId="13.[151,1437,707,840]" pageId="13" pageNumber="370">
<emphasis id="B926CB23FFBCAD13E389FD47FE91FD74" bold="true" box="[151,280,707,731]" pageId="13" pageNumber="370">FIGURE 9.</emphasis>
<taxonomicName id="4C526CB2FFBCAD13E23AFD40FC8CFD74" authority="Bhansali &amp; Wesener" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[292,773,708,732]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="370" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBCAD13E23AFD40FD96FD73" box="[292,543,708,732]" italics="true" pageId="13" pageNumber="370">Zephronia chrysomallos</emphasis>
Bhansali &amp; Wesener
</taxonomicName>
<emphasis id="B926CB23FFBCAD13E00EFD40FCEFFD73" bold="true" box="[784,870,708,732]" pageId="13" pageNumber="370">
<taxonomicNameLabel id="A2157658FFBCAD13E00EFD40FCEFFD73" box="[784,870,708,732]" pageId="13" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂ (
<emphasis id="B926CB23FFBCAD13E71AFD40FB43FD74" bold="true" box="[1028,1226,707,732]" pageId="13" pageNumber="370">ZFMK MYR8826</emphasis>
), scanning electron micrographs. 
<emphasis id="B926CB23FFBCAD13E23BFD63FEB4FD50" bold="true" box="[293,317,743,767]" pageId="13" pageNumber="370">A.</emphasis>
Gnathochilarium, ventral view. 
<emphasis id="B926CB23FFBCAD13E193FD6CFD63FD50" bold="true" box="[653,746,743,768]" pageId="13" pageNumber="370">B. Right</emphasis>
mandible, mesal view. 
<emphasis id="B926CB23FFBCAD13E0FFFD63FBF3FD50" bold="true" box="[993,1146,743,767]" pageId="13" pageNumber="370">Abbreviations</emphasis>
: C = condyles; Cp = cental pad; eT = external tooth; Ip = inner palpi; iT = internal combined tooth; LL = lamellae linguales; Mn = mentum; pL = pectinate lamellae; St = stipites.
</paragraph>
</caption>
<paragraph id="8BED1731FFBCAD13E3D9FCF2FB8DFC77" blockId="13.[151,1437,886,1776]" pageId="13" pageNumber="370">
<emphasis id="B926CB23FFBCAD13E3D9FCF2FF76FC3F" bold="true" box="[199,255,886,912]" pageId="13" pageNumber="370">Legs</emphasis>
. Ventral spines on leg 1 2/2, at 2 4 /4, at 3 5/5. Leg 3 with single apical spine. Mid-body legs with 3 or 4 apical and 10–12 ventral spines (
<figureCitation id="13690BB4FFBCAD13E114FC1EFDFBFC1B" box="[522,626,922,948]" captionStart="FIGURE 10" captionStartId="14.[151,250,1827,1851]" captionTargetBox="[152,1436,707,1803]" captionTargetId="figure-240@14.[151,1436,703,1803]" captionTargetPageId="14" captionText="FIGURE 10. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), drawings, A. First left coxa with stigmatic plate, posterior view. B. Left leg 9, posterior view. C. Second left coxa with gonopore, posterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. F. Right anterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332751" httpUri="https://zenodo.org/record/6332751/files/figure.png" pageId="13" pageNumber="370">Fig. 10B</figureCitation>
). Inner margin of femur with 12–15 small triangular teeth, concentrated at apical half, but not excavated. Femur 1.5, tarsus 3.3 times longer than wide.
</paragraph>
<paragraph id="8BED1731FFBCAD13E3D9FC66FF41FBEB" blockId="13.[151,1437,886,1776]" pageId="13" pageNumber="370">
<emphasis id="B926CB23FFBCAD13E3D9FC66FEC4FC53" bold="true" box="[199,333,994,1020]" pageId="13" pageNumber="370">Anal shield</emphasis>
. Well-rounded and like tergites covered with short dense setae visible at higher magnification (
<figureCitation id="13690BB4FFBCAD13E671FC66FF32FB8F" captionStart="FIGURE 2" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[337,1247,181,1766]" captionTargetId="figure-22@6.[337,1249,181,1766]" captionTargetPageId="6" captionText="FIGURE 2. Map of Thailand with known distribution of Sphaerobelum spp. and Zephronia spp. Star represents new species. Diamonds represent new localities. Fluorescent Green = S. meridionalis Bhansali &amp; Wesener sp. nov.; Pink = S. aesculus; Brown = S. truncatum; Chrome yellow = Z. chrysomallos Bhansali &amp; Wesener sp. nov.; White = Z. erawani Bhansali &amp; Wesener sp. nov.; Black = Z. viridisoma; Red = Z. panhai; Dark green = Z. golovatchi; Purple = Z. siamensis; Orange = Z. enghoffi; Yellow = Z. lannaensis; Blue = Z. phrain. B. Habitus photograph of Z. chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♀ (ZFMK MYR8826)." figureDoi="http://doi.org/10.5281/zenodo.6332735" httpUri="https://zenodo.org/record/6332735/files/figure.png" pageId="13" pageNumber="370">Fig. 2B</figureCitation>
). Ventral side with dark-colored medium-sized locking carina placed slightly closer to pleurite than margin (
<figureCitation id="13690BB4FFBCAD13E671FB82FF32FBEB" captionStart="FIGURE 7" captionStartId="11.[151,250,1829,1853]" captionTargetBox="[151,1431,992,1806]" captionTargetId="figure-270@11.[151,1436,992,1806]" captionTargetPageId="11" captionText="FIGURE 7. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., paratype ♂ (ZFMK MYR11366), volume rendering based on micro-computed tomography. A. Habitus, lateral view. B. Habitus, ventro-lateral view. C. Habitus, ventral view. D. Habitus, anterior view. E. Body-ring architecture, cross section trough midbody-ring. Abbreviations: Ant = Antennae; As = anal shield; Cl = clypeus; Col = collum; Cx = coxa; Fe = femur; Gc = gnathochilarium; Go = gonopore; Ip = inner palpi; Lc = locking carina; O = ommatidia; Pl = pleurite; Pl1= pleurite 1; Pl2= pleurite 2; Pre = prefemur; Pt = paratergite; St = stigmatic plate; Ta = tarsus; Te = tergite; Tg = thoracic shield groove; Ti = tibia; Tr = tracheal apodeme; Ts = thoracic shield. Not to scale." figureDoi="http://doi.org/10.5281/zenodo.6332745" httpUri="https://zenodo.org/record/6332745/files/figure.png" pageId="13" pageNumber="370">Fig. 7C</figureCitation>
).
</paragraph>
<paragraph id="8BED1731FFBCAD13E3D9FBCAFC23FB23" blockId="13.[151,1437,886,1776]" pageId="13" pageNumber="370">
<emphasis id="B926CB23FFBCAD13E3D9FBCAFE69FBC7" bold="true" box="[199,480,1102,1128]" pageId="13" pageNumber="370">Male sexual characters</emphasis>
. Male gonopore consisting of large membranous opening located directly at mesal margin of coxa 2 (
<figureCitation id="13690BB4FFBCAD13E27BFBF6FE45FB23" box="[357,460,1138,1164]" captionStart="FIGURE 10" captionStartId="14.[151,250,1827,1851]" captionTargetBox="[152,1436,707,1803]" captionTargetId="figure-240@14.[151,1436,703,1803]" captionTargetPageId="14" captionText="FIGURE 10. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), drawings, A. First left coxa with stigmatic plate, posterior view. B. Left leg 9, posterior view. C. Second left coxa with gonopore, posterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. F. Right anterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332751" httpUri="https://zenodo.org/record/6332751/files/figure.png" pageId="13" pageNumber="370">Fig. 10C</figureCitation>
). Coxae 2 without membranous processes.
</paragraph>
<paragraph id="8BED1731FFBCAD13E3D9FB12FC7DFACB" blockId="13.[151,1437,886,1776]" pageId="13" pageNumber="370">
<emphasis id="B926CB23FFBCAD13E3D9FB12FE15FB1F" bold="true" box="[199,412,1174,1200]" pageId="13" pageNumber="370">Anterior telopods</emphasis>
(
<figureCitation id="13690BB4FFBCAD13E2B2FB12FDB3FB1F" box="[428,570,1174,1200]" captionStart="FIGURE 10" captionStartId="14.[151,250,1827,1851]" captionTargetBox="[152,1436,707,1803]" captionTargetId="figure-240@14.[151,1436,703,1803]" captionTargetPageId="14" captionText="FIGURE 10. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), drawings, A. First left coxa with stigmatic plate, posterior view. B. Left leg 9, posterior view. C. Second left coxa with gonopore, posterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. F. Right anterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332751" httpUri="https://zenodo.org/record/6332751/files/figure.png" pageId="13" pageNumber="370">Figs 10D–F</figureCitation>
). Syncoxite setaceous. Podomere 1 rectangular, as long as wide. Few setae, mainly at margins and in apical half. Podomere 
<quantity id="4CAABAD4FFBCAD13E1ADFB3EFD56FB7B" box="[691,735,1210,1236]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="13" pageNumber="370" unit="in" value="2.0">2 in</quantity>
anterior view as wide as but slightly narrower as podomere 1. Process massive, visible in anterior view (
<figureCitation id="13690BB4FFBCAD13E171FB5AFD5AFB57" box="[623,723,1246,1272]" captionStart="FIGURE 10" captionStartId="14.[151,250,1827,1851]" captionTargetBox="[152,1436,707,1803]" captionTargetId="figure-240@14.[151,1436,703,1803]" captionTargetPageId="14" captionText="FIGURE 10. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), drawings, A. First left coxa with stigmatic plate, posterior view. B. Left leg 9, posterior view. C. Second left coxa with gonopore, posterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. F. Right anterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332751" httpUri="https://zenodo.org/record/6332751/files/figure.png" pageId="13" pageNumber="370">Fig. 10E</figureCitation>
), relatively short and wide. Process protruding above basal half of podomere 3, triangular (
<figureCitation id="13690BB4FFBCAD13E2DBFA86FDC5FAB3" box="[453,588,1282,1308]" captionStart="FIGURE 10" captionStartId="14.[151,250,1827,1851]" captionTargetBox="[152,1436,707,1803]" captionTargetId="figure-240@14.[151,1436,703,1803]" captionTargetPageId="14" captionText="FIGURE 10. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), drawings, A. First left coxa with stigmatic plate, posterior view. B. Left leg 9, posterior view. C. Second left coxa with gonopore, posterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. F. Right anterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332751" httpUri="https://zenodo.org/record/6332751/files/figure.png" pageId="13" pageNumber="370">Figs 10E, F</figureCitation>
). Podomere 3 cylindrical, 1.4 longer than wide, with one spine in the mesal membranous region, apically with 4 crenulated teeth (
<figureCitation id="13690BB4FFBCAD13E1E2FAA2FCEFFAEF" box="[764,870,1318,1344]" captionStart="FIGURE 10" captionStartId="14.[151,250,1827,1851]" captionTargetBox="[152,1436,707,1803]" captionTargetId="figure-240@14.[151,1436,703,1803]" captionTargetPageId="14" captionText="FIGURE 10. Zephronia chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♂ (ZFMK MYR8826), drawings, A. First left coxa with stigmatic plate, posterior view. B. Left leg 9, posterior view. C. Second left coxa with gonopore, posterior view. D. Right anterior telopod, lateral view. E. Right anterior telopod, posterior view. F. Right anterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; Cx = coxa; Fe = femur; Go = gonopore; Po = postfemur; Pre = prefemur; Sp-p = second podomere process; St = stigmatic plate; Syn = syncoxite; Ta = tarsus; Ti = tibia; vs = ventral spines. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332751" httpUri="https://zenodo.org/record/6332751/files/figure.png" pageId="13" pageNumber="370">Fig. 10D</figureCitation>
). Podomere 4 clearly separated from podomere 3, conical bearing a black spot at apex. Meso-posterior margin with two spines.
</paragraph>
<paragraph id="8BED1731FFBCAD13E3D9FAEAFC84F92B" blockId="13.[151,1437,886,1776]" pageId="13" pageNumber="370">
<emphasis id="B926CB23FFBCAD13E3D9FAEAFE2FFA27" bold="true" box="[199,422,1390,1416]" pageId="13" pageNumber="370">Posterior telopods</emphasis>
(
<figureCitation id="13690BB4FFBCAD13E2A4FAEAFD98FA27" box="[442,529,1390,1416]" captionStart="FIGURE 11" captionStartId="15.[151,250,1331,1355]" captionTargetBox="[153,1436,194,1307]" captionTargetId="figure-17@15.[151,1436,181,1307]" captionTargetPageId="15" captionText="FIGURE 11. Zephroniachrysomallos Bhansali &amp;Wesener sp. nov., holotype ♂ (NHMD MYR8826), drawings, A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view. Abbreviations: ct = crenulated teeth; imf = immovable finger; ss = sclerotized spot. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332753" httpUri="https://zenodo.org/record/6332753/files/figure.png" pageId="13" pageNumber="370">Fig. 11</figureCitation>
). Syncoxite inner horns (not drawn): apex well rounded, apically diverging. Podomere 1 rectangular, as wide as long, with short setae. Podomere 2 with long and narrow immovable finger, 5 times longer than wide. Immovable finger slightly wider posteriorly and apically very slightly curved towards movable finger. Immovable finger glabrous, remaining part of podomere 2 setose. Membranous lobe bearing 2 elongated processes, fused at base. Podomere 
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posterior view glabrous, in anterior view basal half setose. With a row of 17 or 18 large crenulated square teeth on posterior side (
<figureCitation id="13690BB4FFBCAD13E077F9A6FC44F993" box="[873,973,1570,1596]" captionStart="FIGURE 11" captionStartId="15.[151,250,1331,1355]" captionTargetBox="[153,1436,194,1307]" captionTargetId="figure-17@15.[151,1436,181,1307]" captionTargetPageId="15" captionText="FIGURE 11. Zephroniachrysomallos Bhansali &amp;Wesener sp. nov., holotype ♂ (NHMD MYR8826), drawings, A. Left posterior telopod, anterior view. B. Left posterior telopod, posterior view. Abbreviations: ct = crenulated teeth; imf = immovable finger; ss = sclerotized spot. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6332753" httpUri="https://zenodo.org/record/6332753/files/figure.png" pageId="13" pageNumber="370">Fig. 11B</figureCitation>
). Membranous ledge on podomere 3 with single spine. Podomere 4 on both sides glabrous, slightly curved towards immovable finger, membranous lobe with 2 spines. Immovable finger shorter than movable finger.
</paragraph>
</subSubSection>
<subSubSection id="C34844BAFFBCAD13E3D9F90AFBB9F95F" pageId="13" pageNumber="370" type="distribution">
<paragraph id="8BED1731FFBCAD13E3D9F90AFED0F907" blockId="13.[151,1437,886,1776]" box="[199,345,1678,1704]" pageId="13" pageNumber="370">
<emphasis id="B926CB23FFBCAD13E3D9F90AFED0F907" bold="true" box="[199,345,1678,1704]" pageId="13" pageNumber="370">Distribution</emphasis>
</paragraph>
<paragraph id="8BED1731FFBCAD13E3D9F937FBB9F95F" blockId="13.[151,1437,886,1776]" pageId="13" pageNumber="370">
<taxonomicName id="4C526CB2FFBCAD13E3D9F937FEF0F963" authority="Bhansali &amp; Wesener, 2022" authorityName="Bhansali &amp; Wesener" authorityYear="2022" box="[199,377,1714,1740]" class="Diplopoda" family="Zephroniidae" genus="Zephronia" kingdom="Animalia" order="Sphaerotheriida" pageId="13" pageNumber="370" phylum="Arthropoda" rank="species" species="chrysomallos" status="sp. nov.">
<emphasis id="B926CB23FFBCAD13E3D9F937FEF0F963" box="[199,377,1714,1740]" italics="true" pageId="13" pageNumber="370">Z. chrysomallos</emphasis>
</taxonomicName>
<emphasis id="B926CB23FFBCAD13E261F936FE5EF963" bold="true" box="[383,471,1714,1740]" pageId="13" pageNumber="370">
<taxonomicNameLabel id="A2157658FFBCAD13E261F936FE5EF963" box="[383,471,1714,1740]" pageId="13" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is currently only known from the 
<typeStatus id="54E9A993FFBCAD13E051F936FCF6F963" box="[847,895,1714,1740]" pageId="13" pageNumber="370">type</typeStatus>
locality near the Sai Yok Noi Waterfall (
<figureCitation id="13690BB4FFBCAD13E623F936FA1CF963" box="[1341,1429,1714,1740]" captionStart="FIGURE 2" captionStartId="6.[151,250,1791,1815]" captionTargetBox="[337,1247,181,1766]" captionTargetId="figure-22@6.[337,1249,181,1766]" captionTargetPageId="6" captionText="FIGURE 2. Map of Thailand with known distribution of Sphaerobelum spp. and Zephronia spp. Star represents new species. Diamonds represent new localities. Fluorescent Green = S. meridionalis Bhansali &amp; Wesener sp. nov.; Pink = S. aesculus; Brown = S. truncatum; Chrome yellow = Z. chrysomallos Bhansali &amp; Wesener sp. nov.; White = Z. erawani Bhansali &amp; Wesener sp. nov.; Black = Z. viridisoma; Red = Z. panhai; Dark green = Z. golovatchi; Purple = Z. siamensis; Orange = Z. enghoffi; Yellow = Z. lannaensis; Blue = Z. phrain. B. Habitus photograph of Z. chrysomallos Bhansali &amp; Wesener sp. nov., holotype ♀ (ZFMK MYR8826)." figureDoi="http://doi.org/10.5281/zenodo.6332735" httpUri="https://zenodo.org/record/6332735/files/figure.png" pageId="13" pageNumber="370">Fig. 2A</figureCitation>
) and might be microendemic, as are other, closely related species in nearby forests.
</paragraph>
</subSubSection>
</treatment>
</document>