280 lines
50 KiB
XML
280 lines
50 KiB
XML
<document id="A3DDF1A23D059581194A52B749FA57CC" ID-DOI="10.11646/zootaxa.5027.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="5449733" ID-ZooBank="0B8E6B3B-3F14-4959-AB39-ADF60E98EE43" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630481132125" checkinUser="plazi" docAuthor="Arce-Pérez, Roberto, Arriaga-Varela, Emmanuel, Novelo-Gutiérrez, Rodolfo & Navarrete-Heredia, José L." docDate="2021" docId="03FB87D9FFC986325DFDE2B4FA1CF92D" docLanguage="en" docName="zootaxa.5027.3.5.pdf" docOrigin="Zootaxa 5027 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hydrophilus (Dibolocelus) Bedel 1891" docType="key" docVersion="3" lastPageNumber="405" masterDocId="FFC2FFA1FFDB86205D6AE36FFF80FFC8" masterDocTitle="Giant water scavenger beetles Hydrophilus subgenus Dibolocelus (Coleoptera Hydrophilidae) from Mexico with description of two new species" masterLastPageNumber="407" masterPageNumber="387" pageNumber="405" updateTime="1699280670662" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D1D1B9BF93056EC199AE6363964C7CFD">Giant water scavenger beetles Hydrophilus subgenus Dibolocelus (Coleoptera Hydrophilidae) from Mexico with description of two new species</mods:title>
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<mods:name id="44189C88E4587EFEA6C852FC49BECADE" type="personal">
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<mods:namePart id="9EDDDC596D112E36DB7205F7B2CE1277">Arce-Pérez, Roberto</mods:namePart>
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<mods:affiliation id="595A0E2F41DD65BD743B9F8BF141FA87">Instituto de Ecología, A. C., Red de Biodiversidad y Sistemática, Carretera Antigua a Coatepec, 351, El Haya, 91070, Xalapa, Veracruz, Mexico.</mods:affiliation>
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<mods:namePart id="961700E243675A0DB265063785116DC3">Arriaga-Varela, Emmanuel</mods:namePart>
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<mods:nameIdentifier id="E09B983C4B71D5C9563AAEB0B9497258" type="ORCID">0000-0002-3336-8528</mods:nameIdentifier>
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<mods:affiliation id="BC5163B1097C62BFAEDAAF5DF557EE72">Centro de Estudios en Zoología, CUCBA, Universidad de Guadalajara, Apdo. postal 134, Zapopan, Jalisco, Mexico glenusmx @ gmail. com; https: // orcid. org / 0000 - 0002 - 3336 - 8528</mods:affiliation>
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<mods:nameIdentifier id="F36D150FA74832868506B0F5108D08E2" type="email">glenusmx@gmail.com</mods:nameIdentifier>
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<mods:namePart id="C702A0B5C96906DD66760542A8863689">Novelo-Gutiérrez, Rodolfo</mods:namePart>
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<mods:affiliation id="8F9807939ACFCD5774C6514D1B0330CD">Instituto de Ecología, A. C., Red de Biodiversidad y Sistemática, Carretera Antigua a Coatepec, 351, El Haya, 91070, Xalapa, Veracruz, Mexico.</mods:affiliation>
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<mods:namePart id="71FFFC775F5E2DA8656D56CCF7F66FA6">Navarrete-Heredia, José L.</mods:namePart>
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<mods:nameIdentifier id="86F77D6D34F65D032FFED896B381580D" type="ORCID">0000-0002-3336-8528</mods:nameIdentifier>
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<mods:affiliation id="FAC2EA2D68F5FF80F91BAD69955B2A0A">Centro de Estudios en Zoología, CUCBA, Universidad de Guadalajara, Apdo. postal 134, Zapopan, Jalisco, Mexico glenusmx @ gmail. com; https: // orcid. org / 0000 - 0002 - 3336 - 8528</mods:affiliation>
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<mods:nameIdentifier id="2E03248D4DDDAF91B0C53153582C183A" type="email">glenusmx@gmail.com</mods:nameIdentifier>
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<mods:title id="EC7E27A4D7E8E89FE4D0D194AF5FC239">Zootaxa</mods:title>
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<mods:date id="DFEB49EF56B1AABA375EA4D59F74C4BF">2021</mods:date>
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<mods:number id="4363D4ABC6C0E830C5A512E5D32E59A2">2021-09-01</mods:number>
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<mods:number id="1CA05B9E9C67B1826E5E3BB87D913A50">5027</mods:number>
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<mods:classification id="C65AC6B76215C767F21B41930CD65E38">journal article</mods:classification>
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<mods:identifier id="A43149D6B9ADB88E0241F7E77BE18860" type="DOI">10.11646/zootaxa.5027.3.5</mods:identifier>
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<mods:identifier id="F3DA1BFF1ACA2831BFEBE9C23F47314C" type="ISSN">1175-5326</mods:identifier>
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<subSubSection id="C3486544FFC986325DFDE2B4FB8AFD4D" pageId="18" pageNumber="405" type="nomenclature">
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<paragraph id="8BED36CFFFC986325DFDE2B4FB22FE3C" blockId="18.[151,1186,474,501]" box="[151,1186,474,501]" pageId="18" pageNumber="405">
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<heading id="D0A581A3FFC986325DFDE2B4FB22FE3C" bold="true" box="[151,1186,474,501]" fontSize="11" level="1" pageId="18" pageNumber="405" reason="1">
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<emphasis id="B926EADDFFC986325DFDE2B4FB22FE3C" bold="true" box="[151,1186,474,501]" pageId="18" pageNumber="405">
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Key for the species of
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<taxonomicName id="4C524D4CFFC986325CC3E2B5FD6BFE3C" authority="" authorityName="Bedel" authorityYear="1891" box="[425,747,474,501]" class="Liliopsida" family="Restionaceae" genus="Hydrophilus" kingdom="Plantae" order="Poales" pageId="18" pageNumber="405" phylum="Tracheophyta" rank="subGenus" subGenus="Dibolocelus">
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<emphasis id="B926EADDFFC986325CC3E2B5FDC1FE3D" bold="true" box="[425,577,474,501]" italics="true" pageId="18" pageNumber="405">Hydrophilus</emphasis>
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(
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<emphasis id="B926EADDFFC986325F38E2B5FD61FE3D" bold="true" box="[594,737,474,501]" italics="true" pageId="18" pageNumber="405">Dibolocelus</emphasis>
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)
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</taxonomicName>
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from
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<collectingCountry id="F345765FFFC986325E5DE2B5FC12FE3C" box="[823,914,474,500]" name="Mexico" pageId="18" pageNumber="405">Mexico</collectingCountry>
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and Central America
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BED36CFFFC986325DFDE14CFB8AFD4D" blockId="18.[151,1436,547,645]" pageId="18" pageNumber="405">
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<emphasis id="B926EADDFFC986325DFDE14CFF56FDF5" bold="true" box="[151,214,547,573]" pageId="18" pageNumber="405">Note.</emphasis>
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The first characters to appear in the couplets are those shared by both sexes when available. However, the best characters for specific identification are those found in male specimens. We recommend following this key with caution if only female specimens that are not associated to males are available.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3486544FFC986325DFDE1DFFA1CF92D" pageId="18" pageNumber="405" type="key">
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<keyStep id="30A62E6AFFC986325DFDE1DFFA1CFC67" pageId="18" pageNumber="405">
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<paragraph id="8BED36CFFFC986325DFDE1DFFA1CFCD5" blockId="18.[151,1437,688,798]" pageId="18" pageNumber="405">
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<keyLead id="30A395FAFFC986325DFDE1DFFA1CFCD5" pageId="18" pageNumber="405">
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1. Sixth antennomere long and oval, yellowish brown, almost completely covering the first article of antennal club (
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<figureCitation id="13692A4AFFC986325876E1DFFAECFD0D" box="[1308,1388,688,710]" captionStart="FIGURES 19–24" captionStartId="9.[151,264,1841,1865]" captionTargetBox="[162,1428,464,1806]" captionTargetId="figure-134@9.[151,1436,453,1817]" captionTargetPageId="9" captionText="FIGURES 19–24. Dorsal and ventral view of Hydrophilus (Dibolocelus) spp. 19–20 H. (D.) violaceonitens status restored; 21–22 H. (D.) pollens Sharp; 23–24 H. (D.) cf. purpurascens (Régimbart)." figureDoi="http://doi.org/10.5281/zenodo.5449743" httpUri="https://zenodo.org/record/5449743/files/figure.png" pageId="18" pageNumber="405">Figs. 23</figureCitation>
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,
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<figureCitation id="13692A4AFFC98632581CE1DFFA11FD0E" box="[1398,1425,688,710]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">33</figureCitation>
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); abdominal ventrites with vestiture leaving a slender oval glabrous area at mid-line in ventrite 4 and broader sub-triangular one in ventrite 5 (
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<figureCitation id="13692A4AFFC986325C0AE185FE52FCC8" box="[352,466,746,768]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">Figs. 31, 35</figureCitation>
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) in both sexes. Males with robust (inflated) maxillary palpi, third palpomere with a deep broad ventral excavation (
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<figureCitation id="13692A4AFFC986325C3DE067FE4BFCD6" box="[343,459,776,798]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">Figs. 29, 33</figureCitation>
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), females with narrow palpi, without ventral excavation................................... 2
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</keyLead>
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</paragraph>
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<paragraph id="8BED36CFFFC986325DFDE02DFA1CFC67" blockId="18.[151,1437,834,944]" pageId="18" pageNumber="405">
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<keyLead id="30A395FAFFC986325DFDE02DFA1CFC67" pageId="18" pageNumber="405">
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1’ Sixth antennomere short and oval, dark brown, not covering the first article of antennal club (
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<figureCitation id="13692A4AFFC98632591CE02DFB60FC9F" box="[1142,1248,834,856]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Figs. 7, 11</figureCitation>
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, |5, 25); abdominal ventrites with vestiture leaving a wide oval glabrous area at mid-line in ventrites 3–4; and a broader sub-rectangular one in ventrite
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both sexes (
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<figureCitation id="13692A4AFFC986325CA5E012FDC4FC5B" box="[463,580,893,915]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1577,1601]" captionTargetBox="[179,1416,189,1540]" captionTargetId="figure-18@3.[164,1423,179,1552]" captionTargetPageId="3" captionText="FIGURES 1–6. Dorsal and ventral view of Hydrophilus (Dibolocelus) spp. 1–2 H. (D.) ovatus Gemminger & Harold; 3–4 H. (D.) pseudovatus sp. nov.; 5–6 H. (D.) nucleoensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5449737" httpUri="https://zenodo.org/record/5449737/files/figure.png" pageId="18" pageNumber="405">Figs. 2, 4, 6</figureCitation>
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); males with maxillary palpi narrow (
|
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<figureCitation id="13692A4AFFC986325EDAE012FC76FC5B" box="[944,1014,893,915]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">Fig. 25</figureCitation>
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) or slightly flattened towards apex (
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<figureCitation id="13692A4AFFC98632583EE012FE9DFC78" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Figs. 7, 11, 15</figureCitation>
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) in both sexes.................................................................................. 3
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<keyLead id="30A395FAFFC986325DFDE0BAFA1DFBCF" pageId="18" pageNumber="405">
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2 (1). Anterior margin of mentum truncated and straight in both sexes (
|
||
<figureCitation id="13692A4AFFC986325E2DE0BAFC0EFC23" box="[839,910,981,1003]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">Fig. 33</figureCitation>
|
||
); male genitalia with broad parameres, slightly directed outward, with broad subtriangular apex (
|
||
<figureCitation id="13692A4AFFC986325F03E09DFD57FBC0" box="[617,727,1010,1032]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">Fig. 36 a, b</figureCitation>
|
||
)...........................
|
||
<emphasis id="B926EADDFFC986325952E09CFBD0FBC0" box="[1080,1104,1011,1032]" italics="true" pageId="18" pageNumber="405">H.</emphasis>
|
||
(
|
||
<emphasis id="B926EADDFFC986325937E09CFBEEFBC0" box="[1117,1134,1011,1032]" italics="true" pageId="18" pageNumber="405">D</emphasis>
|
||
.)
|
||
<taxonomicName id="4C524D4CFFC9863259E8E09DFA1DFBCF" ID-CoL="9P6TL" authority="Regimbart" authorityName="Regimbart" box="[1154,1437,1009,1032]" class="Insecta" family="Hydrophilidae" genus="Dibolocelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="405" phylum="Arthropoda" rank="species" species="purpurascens">
|
||
<emphasis id="B926EADDFFC9863259E8E09DFA1DFBCF" bold="true" box="[1154,1437,1009,1032]" pageId="18" pageNumber="405">
|
||
cf.
|
||
<emphasis id="B926EADDFFC9863259CBE09CFAA8FBC0" bold="true" box="[1185,1320,1011,1032]" italics="true" pageId="18" pageNumber="405">purpurascens</emphasis>
|
||
Régimbart
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BED36CFFFC986325DFDE742FA1DFBA8" blockId="18.[151,1437,1069,1120]" pageId="18" pageNumber="405">
|
||
<keyLead id="30A395FAFFC986325DFDE742FA1DFBA8" pageId="18" pageNumber="405">
|
||
2’ Anterior margin of mentum rounded, unknown female (
|
||
<figureCitation id="13692A4AFFC986325F95E742FCC8FB8B" box="[767,840,1069,1091]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">Fig. 29</figureCitation>
|
||
); male genitalia with parameres comparatively narrow, markedly directed inwards, with slender round apex. (
|
||
<figureCitation id="13692A4AFFC986325FDDE725FCA5FBA8" box="[695,805,1098,1120]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">Fig. 32 a, b</figureCitation>
|
||
).................................
|
||
<taxonomicName id="4C524D4CFFC9863259AFE724FA1DFBA8" authority="Sharp" authorityName="Sharp" box="[1221,1437,1098,1120]" class="Liliopsida" family="Restionaceae" genus="Hydrophilus" kingdom="Plantae" order="Poales" pageId="18" pageNumber="403" phylum="Tracheophyta" rank="species" species="pollens" subGenus="Dibolocelus">
|
||
<emphasis id="B926EADDFFC9863259AFE724FA1DFBA8" bold="true" box="[1221,1437,1098,1120]" pageId="18" pageNumber="405">
|
||
<emphasis id="B926EADDFFC9863259AFE724FB58FBA8" bold="true" box="[1221,1240,1099,1120]" italics="true" pageId="18" pageNumber="405">H</emphasis>
|
||
. (
|
||
<emphasis id="B926EADDFFC986325986E724FB7DFBA8" bold="true" box="[1260,1277,1099,1120]" italics="true" pageId="18" pageNumber="405">D</emphasis>
|
||
.)
|
||
<emphasis id="B926EADDFFC98632587BE725FAD8FBA8" bold="true" box="[1297,1368,1098,1120]" italics="true" pageId="18" pageNumber="405">pollens</emphasis>
|
||
Sharp
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A62E6AFFC986325DFDE7EAFA1CFAE5" pageId="18" pageNumber="405">
|
||
<paragraph id="8BED36CFFFC986325DFDE7EAFA1CFB1D" blockId="18.[151,1437,1157,1238]" pageId="18" pageNumber="405">
|
||
<keyLead id="30A395FAFFC986325DFDE7EAFA1CFB1D" pageId="18" pageNumber="405">
|
||
3 (1’). 3 (1’). Male with maxillary palpomeres 2 and 3 distinctly flattened and wide towards apex (
|
||
<figureCitation id="13692A4AFFC986325909E7EAFB4CFB53" box="[1123,1228,1157,1179]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Figs. 7, 11</figureCitation>
|
||
), slightly flattened in female; male with anterior tibiae internal spur sub-rectangular and with apex truncate (
|
||
<figureCitation id="13692A4AFFC986325948E7CDFB0AFB70" box="[1058,1162,1186,1208]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Figs. 8, 12</figureCitation>
|
||
), sub-triangular and acute in female.............................................................................................. 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BED36CFFFC986325DFDE795FA1CFAE5" blockId="18.[151,1437,1274,1326]" pageId="18" pageNumber="405">
|
||
<keyLead id="30A395FAFFC986325DFDE795FA1CFAE5" pageId="18" pageNumber="405">
|
||
3’ Male and female with maxillary palpomeres 2 and 3 distinctly slender, only slightly flattened (
|
||
<figureCitation id="13692A4AFFC98632590AE795FB30FAD8" box="[1120,1200,1274,1296]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Figs. 15</figureCitation>
|
||
,
|
||
<figureCitation id="13692A4AFFC9863259D3E795FB54FAD8" box="[1209,1236,1274,1296]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">25</figureCitation>
|
||
); males with anterior tibiae internal spur sub-rectangular with apex truncate straight (
|
||
<figureCitation id="13692A4AFFC986325E2EE677FC09FAE6" box="[836,905,1304,1326]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Fig. 16</figureCitation>
|
||
) or markedly bifid (
|
||
<figureCitation id="13692A4AFFC986325921E677FB11FAE6" box="[1099,1169,1304,1326]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">Fig. 26</figureCitation>
|
||
).................... 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A62E6AFFC986325DFDE63DFA1CF9FD" pageId="18" pageNumber="405">
|
||
<paragraph id="8BED36CFFFC986325DFDE63DFA1CFA6A" blockId="18.[151,1437,1362,1443]" pageId="18" pageNumber="405">
|
||
<keyLead id="30A395FAFFC986325DFDE63DFA1CFA6A" pageId="18" pageNumber="405">
|
||
4 (3). Metaventral spine short, not surpassing the posterior 3/4 of second abdominal ventrite (
|
||
<figureCitation id="13692A4AFFC98632594FE63DFBE9FAAF" box="[1061,1129,1362,1384]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1577,1601]" captionTargetBox="[179,1416,189,1540]" captionTargetId="figure-18@3.[164,1423,179,1552]" captionTargetPageId="3" captionText="FIGURES 1–6. Dorsal and ventral view of Hydrophilus (Dibolocelus) spp. 1–2 H. (D.) ovatus Gemminger & Harold; 3–4 H. (D.) pseudovatus sp. nov.; 5–6 H. (D.) nucleoensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5449737" httpUri="https://zenodo.org/record/5449737/files/figure.png" pageId="18" pageNumber="405">Figs. 4</figureCitation>
|
||
,
|
||
<figureCitation id="13692A4AFFC986325919E63DFB0DFAA0" box="[1139,1165,1362,1384]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">13</figureCitation>
|
||
); third palpomere very wide and flattened with its inner margin slightly curved (
|
||
<figureCitation id="13692A4AFFC986325FB0E61FFCA2FA4E" box="[730,802,1392,1414]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Fig. 11</figureCitation>
|
||
). Males with fifth protarsomere somewhat flattened with a small internal laminar expansion, shorter than previous tarsomeres together (
|
||
<figureCitation id="13692A4AFFC986325EE2E6E2FC50FA6B" box="[904,976,1421,1443]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Fig. 12</figureCitation>
|
||
);............
|
||
<emphasis id="B926EADDFFC9863259EEE6E2FA1CFA6A" bold="true" box="[1156,1436,1420,1443]" pageId="18" pageNumber="405">
|
||
<taxonomicName id="4C524D4CFFC9863259EEE6E2FAC9FA6A" authority="Arce-Pérez & Arriaga-Varela & Novelo-Gutiérrez & Navarrete-Heredia, 2021" authorityName="Arce-Pérez & Arriaga-Varela & Novelo-Gutiérrez & Navarrete-Heredia" authorityYear="2021" box="[1156,1353,1420,1443]" class="Liliopsida" family="Restionaceae" genus="Hydrophilus" kingdom="Plantae" order="Poales" pageId="18" pageNumber="405" phylum="Tracheophyta" rank="species" species="pseudovatus" status="sp. nov." subGenus="Dibolocelus">
|
||
<emphasis id="B926EADDFFC9863259EEE6E2FB1DFA6A" bold="true" box="[1156,1181,1421,1442]" italics="true" pageId="18" pageNumber="405">H.</emphasis>
|
||
(
|
||
<emphasis id="B926EADDFFC9863259C1E6E2FB42FA6A" bold="true" box="[1195,1218,1421,1442]" italics="true" pageId="18" pageNumber="405">D.</emphasis>
|
||
)
|
||
<emphasis id="B926EADDFFC9863259BAE6E3FAC9FA6A" bold="true" box="[1232,1353,1420,1442]" italics="true" pageId="18" pageNumber="405">pseudovatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21557A6FFC98632583AE6E2FA1CFA6A" box="[1360,1436,1421,1442]" pageId="18" pageNumber="405" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BED36CFFFC986325DFDE6A7FA1CF9FD" blockId="18.[151,1437,1480,1589]" pageId="18" pageNumber="405">
|
||
<keyLead id="30A395FAFFC986325DFDE6A7FA1CF9FD" pageId="18" pageNumber="405">
|
||
4’ Metaventral spine long, surpassing the posterior margin of second abdominal ventrite (
|
||
<figureCitation id="13692A4AFFC986325956E6A7FB02FA15" box="[1084,1154,1480,1502]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1577,1601]" captionTargetBox="[179,1416,189,1540]" captionTargetId="figure-18@3.[164,1423,179,1552]" captionTargetPageId="3" captionText="FIGURES 1–6. Dorsal and ventral view of Hydrophilus (Dibolocelus) spp. 1–2 H. (D.) ovatus Gemminger & Harold; 3–4 H. (D.) pseudovatus sp. nov.; 5–6 H. (D.) nucleoensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5449737" httpUri="https://zenodo.org/record/5449737/files/figure.png" pageId="18" pageNumber="405">Figs. 2</figureCitation>
|
||
,
|
||
<figureCitation id="13692A4AFFC9863259E4E6A7FB1CFA16" box="[1166,1180,1480,1502]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">9</figureCitation>
|
||
); third palpomere slightly wide and flat, compared to
|
||
<taxonomicName id="4C524D4CFFC986325C81E689FD2FFA32" authorityName="Arce-Pérez & Arriaga-Varela & Novelo-Gutiérrez & Navarrete-Heredia" authorityYear="2021" box="[491,687,1509,1531]" class="Liliopsida" family="Restionaceae" genus="Hydrophilus" kingdom="Plantae" order="Poales" pageId="18" pageNumber="405" phylum="Tracheophyta" rank="species" species="pseudovatus" subGenus="Dibolocelus">
|
||
<emphasis id="B926EADDFFC986325C81E689FD83FA33" box="[491,515,1510,1531]" italics="true" pageId="18" pageNumber="405">H.</emphasis>
|
||
(
|
||
<emphasis id="B926EADDFFC986325F7BE689FDA9FA33" box="[529,553,1510,1531]" italics="true" pageId="18" pageNumber="405">D.</emphasis>
|
||
)
|
||
<emphasis id="B926EADDFFC986325F5CE68AFD2FFA32" box="[566,687,1509,1530]" italics="true" pageId="18" pageNumber="405">pseudovatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B926EADDFFC986325FDAE68AFD34FA32" box="[688,692,1509,1530]" italics="true" pageId="18" pageNumber="405">,</emphasis>
|
||
with its inner margin slightly straight (
|
||
<figureCitation id="13692A4AFFC986325944E68AFBE9FA32" box="[1070,1129,1509,1531]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Fig. 7</figureCitation>
|
||
). Males with fifth protarsomere subcylindrical, without a laminar expansion, longer than previous tarsomeres together (
|
||
<figureCitation id="13692A4AFFC986325942E56DFBE0F9D0" box="[1064,1120,1538,1560]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Fig. 8</figureCitation>
|
||
)..............................................................................................
|
||
<taxonomicName id="4C524D4CFFC98632594DE54FFA1CF9FD" ID-CoL="9JD9F" authority="Gemminger & Harold" authorityName="Gemminger & Harold" box="[1063,1436,1567,1589]" class="Insecta" family="Hydrophilidae" genus="Dibolocelus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="405" phylum="Arthropoda" rank="species" species="ovatus" subGenus="Dibolocelus">
|
||
<emphasis id="B926EADDFFC98632594DE54FFA1CF9FD" bold="true" box="[1063,1436,1567,1589]" pageId="18" pageNumber="405">
|
||
<emphasis id="B926EADDFFC98632594DE54FFBC0F9FD" bold="true" box="[1063,1088,1568,1589]" italics="true" pageId="18" pageNumber="405">H.</emphasis>
|
||
(
|
||
<emphasis id="B926EADDFFC986325924E54FFBE5F9FD" bold="true" box="[1102,1125,1568,1589]" italics="true" pageId="18" pageNumber="405">D.</emphasis>
|
||
)
|
||
<emphasis id="B926EADDFFC986325919E54FFB33F9FD" bold="true" box="[1139,1203,1568,1589]" italics="true" pageId="18" pageNumber="405">ovatus</emphasis>
|
||
Gemminger & Harold
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A62E6AFFC986325DFDE535FA1CF92D" pageId="18" pageNumber="405">
|
||
<paragraph id="8BED36CFFFC986325DFDE535FA1CF945" blockId="18.[151,1436,1626,1678]" pageId="18" pageNumber="405">
|
||
<keyLead id="30A395FAFFC986325DFDE535FA1CF945" pageId="18" pageNumber="405">
|
||
5 (3’) Males with fifth protarsomere as long as the basal together, flat and with evident internal laminar expansion; internal spur of anterior tibiae of males sub-rectangular with apex truncate (
|
||
<figureCitation id="13692A4AFFC986325E75E517FCE5F946" box="[799,869,1656,1678]" captionStart="FIGURES 7–18" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[177,1413,184,1798]" captionTargetId="figure-25@4.[164,1423,181,1806]" captionTargetPageId="4" captionText="FIGURES 7–18. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 7-10 H. (D.) ovatus Gemminger & Harold; 7 ventral view of head (arrow: third labial palpomere); 8 protibia and tarsi, 9 ventral view of abdomen; 10a aedeagus ventral; 10b aedeagus dorsal; 11–14 H. (D.) pseudovatus sp. nov.; 11 ventral view of head(arrow: third labial palpomere); 12 protibia and tarsi, 13 ventral view of abdomen; 14a aedeagus ventral; 14b aedeagus dorsal; 15–18 H. (D.) nucleoensis sp. nov.; 15 ventral view of head (arrow: third labial palpomere); 16 protibia and tarsi, 17 ventral view of abdomen; 18a aedeagus ventral; 18b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449739" httpUri="https://zenodo.org/record/5449739/files/figure.png" pageId="18" pageNumber="405">Fig. 16</figureCitation>
|
||
).......................
|
||
<emphasis id="B926EADDFFC9863259E4E517FA1CF945" bold="true" box="[1166,1436,1655,1677]" pageId="18" pageNumber="405">
|
||
<taxonomicName id="4C524D4CFFC9863259E4E517FACEF945" authority="Arce-Pérez & Arriaga-Varela & Novelo-Gutiérrez & Navarrete-Heredia, 2021" authorityName="Arce-Pérez & Arriaga-Varela & Novelo-Gutiérrez & Navarrete-Heredia" authorityYear="2021" box="[1166,1358,1655,1677]" class="Liliopsida" family="Restionaceae" genus="Hydrophilus" kingdom="Plantae" order="Poales" pageId="18" pageNumber="405" phylum="Tracheophyta" rank="species" species="nucleoensis" status="sp. nov." subGenus="Dibolocelus">
|
||
<emphasis id="B926EADDFFC9863259E4E517FB27F945" bold="true" box="[1166,1191,1656,1677]" italics="true" pageId="18" pageNumber="405">H.</emphasis>
|
||
(
|
||
<emphasis id="B926EADDFFC9863259DFE517FB4CF945" bold="true" box="[1205,1228,1656,1677]" italics="true" pageId="18" pageNumber="405">D.</emphasis>
|
||
)
|
||
<emphasis id="B926EADDFFC9863259B0E518FACEF945" bold="true" box="[1242,1358,1655,1677]" italics="true" pageId="18" pageNumber="405">nucleoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21557A6FFC98632583EE517FA1CF945" box="[1364,1436,1656,1677]" pageId="18" pageNumber="405" rank="species">sp. nov</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BED36CFFFC986325DFDE5DDFA1CF92D" blockId="18.[151,1436,1714,1766]" pageId="18" pageNumber="405">
|
||
<keyLead id="30A395FAFFC986325DFDE5DDFA1CF92D" pageId="18" pageNumber="405">
|
||
5’ Males with fifth protarsomere shorter than previous tarsomeres together, wide and without internal expansion; internal spur of anterior tibiae of males with bifid apex (
|
||
<figureCitation id="13692A4AFFC986325F0EE5BFFD2AF92E" box="[612,682,1744,1766]" captionStart="FIGURES 25–36" captionStartId="10.[151,264,1782,1806]" captionTargetBox="[159,1436,191,1748]" captionTargetId="figure-25@10.[151,1436,181,1758]" captionTargetPageId="10" captionText="FIGURES 25–36. Morphological structures and male genitalia of Hydrophilus (Dibolocelus) spp. 25–28 H. (D.) violaceonitens sp. nov.; 25 ventral view of head (arrow: third labial palpomere); 26 protibia and tarsi, 27 ventral view of abdomen; 28a aedeagus ventral; 28b aedeagus dorsal; 29–32 H. (D.) pollens Sharp; 29 ventral view of head (arrow: third labial palpomere); 30 protibia and tarsi, 31 ventral view of abdomen; 32 aedeagus ventral; 33–36 H. (D.) cf. purpurascens (Régimbart); 33 ventral view of head (arrow: third labial palpomere); 34 protibia and tarsi (arrow: subrectangular row of hard setae); 35 ventral view of abdomen; 36a aedeagus ventral; 36b aedeagus dorsal." figureDoi="http://doi.org/10.5281/zenodo.5449747" httpUri="https://zenodo.org/record/5449747/files/figure.png" pageId="18" pageNumber="405">Fig. 26</figureCitation>
|
||
)............................
|
||
<taxonomicName id="4C524D4CFFC986325978E5BFFA1CF92D" authority="Jacquelin du Val" box="[1042,1436,1743,1765]" class="Liliopsida" family="Restionaceae" genus="Hydrophilus" kingdom="Plantae" order="Poales" pageId="18" pageNumber="405" phylum="Tracheophyta" rank="species" species="violaceonitens" subGenus="Dibolocelus">
|
||
<emphasis id="B926EADDFFC986325978E5BFFA1CF92D" bold="true" box="[1042,1436,1743,1765]" pageId="18" pageNumber="405">
|
||
<emphasis id="B926EADDFFC986325978E5BFFBABF92D" bold="true" box="[1042,1067,1744,1765]" italics="true" pageId="18" pageNumber="405">H.</emphasis>
|
||
(
|
||
<emphasis id="B926EADDFFC986325953E5BFFBD0F92D" bold="true" box="[1081,1104,1744,1765]" italics="true" pageId="18" pageNumber="405">D.</emphasis>
|
||
)
|
||
<emphasis id="B926EADDFFC986325934E5A0FB6BF92D" bold="true" box="[1118,1259,1743,1765]" italics="true" pageId="18" pageNumber="405">violaceonitens</emphasis>
|
||
Jacquelin du Val
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |