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<document id="C235F67C53AD5EFC0165CAF675DB34B8" ID-DOI="10.5281/zenodo.280458" ID-GBIF-Dataset="33e496c3-1dbf-48d5-be9b-88a64c478422" ID-ISSN="1175-5326" ID-Zenodo-Dep="280458" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460458190444" checkinUser="plazi" docAuthor="Komai, Tomoyuki, Tsuchida, Shinji &amp; Segonzac, Michel" docDate="2012" docId="03FB87A27D43DC26F483F8B3FD49FDFD" docLanguage="en" docName="zt03241p063.pdf" docOrigin="Zootaxa 3241" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lebbeus shinkaiae Komai, Tsuchida &amp; Segonzac, 2012, sp. nov." docType="treatment" docVersion="10" lastPageNumber="51" masterDocId="FFC2FFDA7D48DC36F414FFABFFC8FFEA" masterDocTitle="Records of species of the hippolytid genus Lebbeus White, 1847 (Crustacea: Decapoda: Caridea) from hydrothermal vents in the Pacific Ocean, with descriptions of three new species" masterLastPageNumber="63" masterPageNumber="35" pageNumber="46" updateTime="1698302915336" updateUser="plazi">
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<mods:title id="77CA77D9FF65FA190EA5621D573E65A7">Records of species of the hippolytid genus Lebbeus White, 1847 (Crustacea: Decapoda: Caridea) from hydrothermal vents in the Pacific Ocean, with descriptions of three new species</mods:title>
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<mods:namePart id="3FD2F74C95478FBA1D1FFD4900FBCC31">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="C403C1FFECF834BD1FB49479AB1B5637">Tsuchida, Shinji</mods:namePart>
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<mods:namePart id="AF3FF4514008BAF6DF3A53E8AEAF99C0">Segonzac, Michel</mods:namePart>
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<mods:date id="CD21331613B88C4A67FC27FAE0B7FDC6">2012</mods:date>
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<treatment id="03FB87A27D43DC26F483F8B3FD49FDFD" ID-DOI="http://doi.org/10.5281/zenodo.5490702" ID-GBIF-Taxon="119511589" ID-Zenodo-Dep="5490702" LSID="urn:lsid:plazi:treatment:03FB87A27D43DC26F483F8B3FD49FDFD" httpUri="http://treatment.plazi.org/id/03FB87A27D43DC26F483F8B3FD49FDFD" lastPageId="16" lastPageNumber="51" pageId="11" pageNumber="46">
<subSubSection id="C348653F7D43DC3DF483F8B3FE99F8B8" pageId="11" pageNumber="46" type="nomenclature">
<paragraph id="8BED36B47D43DC3DF483F8B3FE15F8D8" blockId="11.[151,477,1816,1874]" box="[151,477,1816,1842]" pageId="11" pageNumber="46">
<heading id="D0A581D87D43DC3DF483F8B3FE15F8D8" bold="true" box="[151,477,1816,1842]" fontSize="11" level="1" pageId="11" pageNumber="46" reason="1">
<emphasis id="B926EAA67D43DC3DF483F8B3FE15F8D8" bold="true" box="[151,477,1816,1842]" pageId="11" pageNumber="46">
<taxonomicName id="4C524D377D43DC3DF483F8B3FEB0F8D8" box="[151,376,1816,1842]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="shinkaiae" status="sp. nov.">
<emphasis id="B926EAA67D43DC3DF483F8B3FEB0F8D8" bold="true" box="[151,376,1816,1842]" italics="true" pageId="11" pageNumber="46">Lebbeus shinkaiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557DD7D43DC3DF594F8B3FE15F8D8" box="[384,477,1816,1842]" pageId="11" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36B47D43DC3DF483F891FE99F8B8" blockId="11.[151,477,1816,1874]" box="[151,337,1850,1874]" pageId="11" pageNumber="46">
(
<figureCitation id="13692A317D43DC3DF48BF891FEE8F8B8" box="[159,288,1850,1874]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStart-2="FIGURE 12" captionStart-3="FIGURE 13" captionStartId-0="12.[151,250,1366,1389]" captionStartId-1="13.[151,250,1896,1919]" captionStartId-2="14.[151,250,1852,1875]" captionStartId-3="15.[151,250,843,866]" captionTargetBox-0="[156,1429,558,1341]" captionTargetBox-1="[175,1407,554,1873]" captionTargetBox-2="[166,1404,219,1795]" captionTargetBox-3="[179,1410,214,802]" captionTargetId-0="figure@12.[151,1436,553,1345]" captionTargetId-1="figure@13.[151,1436,527,1887]" captionTargetId-2="figure@14.[151,1436,193,1831]" captionTargetId-3="figure@15.[151,1436,193,822]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionTargetPageId-2="14" captionTargetPageId-3="15" captionText-0="FIGURE 10. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Body (thoracic appendages and pleopods omitted) in lateral view. Scale bar: 5 mm." captionText-1="FIGURE 11. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for A C, E, F; 1 mm for D." captionText-2="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." captionText-3="FIGURE 13. Lebbeus shinkaiae sp. nov., paratype, male (cl 9.0 mm), JAMSTEC 072424 072432. A, carapace and cephalic appendages, lateral view; B, endopod of left first pleopod, dorsal (anterior) view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 2 mm for A; 1 mm for B, C." httpUri-0="https://zenodo.org/record/280467/files/figure.png" httpUri-1="https://zenodo.org/record/280468/files/figure.png" httpUri-2="https://zenodo.org/record/280469/files/figure.png" httpUri-3="https://zenodo.org/record/280470/files/figure.png" pageId="11" pageNumber="46">Figs 1013</figureCitation>
,
<figureCitation id="13692A317D43DC3DF53FF891FE80F8B8" box="[299,328,1850,1874]" captionStart="FIGURE 18" captionStartId="21.[151,250,1875,1898]" captionTargetBox="[252,1340,193,1861]" captionTargetId="figure@21.[252,1341,193,1862]" captionTargetPageId="21" captionText="FIGURE 18. Photographs taken in situ. A, Lebbeus laurentae Wicksten, 2010, one individual is seen on a tube of Riftia pachyptila Jones, 1981, EPR 13 ° N, taken during the HOPE 99 cruise (The blue color of the tubes of R. pachyptira is artificial, resulting from a growth experiment); B, Lebbeus shinkaiae sp. nov., photograph taken at Hatoma Knoll, Okinawa Trough, 1493 m deep, two individuals are seen together with individuals of Alvinocaris longirostris Kikuchi and Ohta, 1995 and Shinkaia crosnieri Baba &amp; Williams, 1998." httpUri="https://zenodo.org/record/280475/files/figure.png" pageId="11" pageNumber="46">18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C348653F7D43DC3DF483F828FE76F83C" pageId="11" pageNumber="46" type="reference_group">
<paragraph id="8BED36B47D43DC3DF483F828FE76F83C" blockId="11.[151,1436,1922,2006]" pageId="11" pageNumber="46">
<treatmentCitationGroup id="AB42119A7D43DC3DF483F828FE76F83C" pageId="11" pageNumber="46">
<taxonomicName id="4C524D377D43DC3DF483F828FD71F872" authority="Kikuchi &amp; Ohta 1995: 779" authorityName="Kikuchi &amp; Ohta" authorityPageNumber="779" authorityYear="1995" box="[151,697,1922,1945]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="washingtonianus">
<emphasis id="B926EAA67D43DC3DF483F828FE55F873" box="[151,413,1923,1945]" italics="true" pageId="11" pageNumber="46">Lebbeus washingtonianus</emphasis>
:
<treatmentCitation id="0AF310A57D43DC3DF5BDF829FD71F872" author="Kikuchi" box="[425,697,1922,1944]" page="779" pageId="11" pageNumber="46" year="1995">
<bibRefCitation id="EFC34B457D43DC3DF5BDF829FD4FF872" author="Kikuchi" box="[425,647,1922,1944]" pageId="11" pageNumber="46" refString="Kikuchi, T. &amp; Ohta, S. (1995) Two caridean shrimps of the families Bresiliidae and Hippolytidae from a hydrothermal field on the Iheya Ridge, off the Ryukyu Islands, Japan. Journal of Crustacean Biology, 15, 771 - 785." type="journal article" year="1995">Kikuchi &amp; Ohta 1995</bibRefCitation>
: 779
</treatmentCitation>
</taxonomicName>
, figs 812;
<treatmentCitation id="0AF310A57D43DC3DF73AF829FBE2F872" author="Fujikura" box="[814,1066,1922,1945]" page="234" pageId="11" pageNumber="46" year="1995">
<bibRefCitation id="EFC34B457D43DC3DF73AF829FC30F872" author="Fujikura" box="[814,1016,1922,1945]" pageId="11" pageNumber="46" refString="Fujikura, K., Hashimoto, J., Fujiwara, Y., Okutani, T. (1995) Community ecology of the chemosynthetic community at Off Hatsushima Site, Sagami Bay, Japan. JAMSTEC Journal of Deep Sea Research, 11, 227 - 241." type="journal article" year="1995">
Fujikura
<emphasis id="B926EAA67D43DC3DF79EF82FFC7EF873" box="[906,950,1923,1945]" italics="true" pageId="11" pageNumber="46">et al</emphasis>
. 1995
</bibRefCitation>
: 234
</treatmentCitation>
;
<treatmentCitation id="0AF310A57D43DC3DF020F829FA85F872" author="Watabe" box="[1076,1357,1922,1944]" page="32" pageId="11" pageNumber="46" year="2000">
<bibRefCitation id="EFC34B457D43DC3DF020F829FAEFF872" author="Watabe" box="[1076,1319,1922,1944]" pageId="11" pageNumber="46" refString="Watabe, H. &amp; Miyake, Y. (2000) Decapod fauna of the hydrothermally active and adjacent fields on the Hatoma Knoll, southern Japan. JAMSTEC Journal of Deep-sea Research, 17, 29 - 34." type="journal article" year="2000">Watabe &amp; Miyake 2000</bibRefCitation>
: 32
</treatmentCitation>
; Komai
<emphasis id="B926EAA67D43DC3DF4D3F809FF3EF85D" box="[199,246,1953,1975]" italics="true" pageId="11" pageNumber="46">et al</emphasis>
. 2004: 121;
<treatmentCitation id="0AF310A57D43DC3DF569F80AFD64F85D" ID-DOI="http://doi.org/10.5281/zenodo.5490540" ID-Zenodo-Dep="5490540" author="Martin" box="[381,684,1953,1975]" httpUri="http://treatment.plazi.org/id/03D96F29FFBFFF9AFF5EFCF9FBC30B7B" page="471" pageId="11" pageNumber="46" year="2005">
<bibRefCitation id="EFC34B457D43DC3DF569F80AFDB1F85D" author="Martin" box="[381,633,1953,1975]" pageId="11" pageNumber="46" refString="Martin, J. W. &amp; Haney, T. (2005) Decapod crustaceans from hydrothermal vents and cold seeps: a review through 2005. Zoological Journal of the Linnean Societ y, 145, 445 - 522." type="journal article" year="2005">Martin and Haney 2005</bibRefCitation>
: 471
</treatmentCitation>
(part); Hashimoto 2007: 198, unnumbered fig. Not
<taxonomicName id="4C524D377D43DC3DF0DAF80AFE71F83C" authority="Rathbun, 1902" authorityName="Rathbun" authorityYear="1902" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="washingtonianus">
<emphasis id="B926EAA67D43DC3DF0DAF80AFEC2F83C" italics="true" pageId="11" pageNumber="46">Lebbeus washingtonianus</emphasis>
(Rathbun, 1902)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C348653F7D44DC3AF483FF3CFB1BFE46" pageId="12" pageNumber="47" type="materials_examined">
<paragraph id="8BED36B47D44DC3AF483FF3CFB1BFE46" blockId="12.[151,1436,151,500]" pageId="12" pageNumber="47">
<emphasis id="B926EAA67D44DC3AF483FF3CFE4DFF5A" bold="true" box="[151,389,151,176]" pageId="12" pageNumber="47">Material examined.</emphasis>
<typeStatus id="54E988167D44DC3AF584FF3CFE35FF45" box="[400,509,151,175]" pageId="12" pageNumber="47" type="holotype">Holotype</typeStatus>
: DS
<taxonomicName id="4C524D377D44DC3AF62FFF33FD1CFF45" box="[571,724,151,176]" class="Polychaeta" family="Chrysopetalidae" genus="Shinkai" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="47" phylum="Annelida" rank="genus">
<emphasis id="B926EAA67D44DC3AF62FFF33FD47FF5A" box="[571,655,152,176]" italics="true" pageId="12" pageNumber="47">Shinkai</emphasis>
2000
</taxonomicName>
, dive #1182, Hatoma Knoll, Okinawa Trough,
<geoCoordinate id="EE6650737D44DC3AF117FF3CFA50FF45" box="[1283,1432,151,175]" direction="north" orientation="latitude" pageId="12" pageNumber="47" precision="1" value="24.853933">24°51.236N</geoCoordinate>
,
<geoCoordinate id="EE6650737D44DC3AF483FF17FEFEFF3E" box="[151,310,188,212]" direction="east" orientation="longitude" pageId="12" pageNumber="47" precision="1" value="123.84333">123°50.600E</geoCoordinate>
,
<quantity id="4CAA9B517D44DC3AF555FF17FE50FF3E" box="[321,408,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.474" pageId="12" pageNumber="47" unit="m" value="1474.0">1474 m</quantity>
,
<date id="FFEC10747D44DC3AF5B0FF17FDF5FF3E" box="[420,573,188,213]" pageId="12" pageNumber="47" value="2000-05-19">19 May 2000</date>
, female (cl
<quantity id="4CAA9B517D44DC3AF6ABFF17FCEFFF3E" box="[703,807,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="12" pageNumber="47" unit="mm" value="18.2">18.2 mm</quantity>
), NSMT-Cr 21982 (ex- JAMSTEC 028214).
<typeStatus id="54E988167D44DC3AF483FF4BFEC0FF12" box="[151,264,224,248]" pageId="12" pageNumber="47" type="paratype">Paratypes</typeStatus>
: DS
<taxonomicName id="4C524D377D44DC3AF529FF4BFE1BFF1D" box="[317,467,223,248]" class="Polychaeta" family="Chrysopetalidae" genus="Shinkai" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="47" phylum="Annelida" rank="genus">
<emphasis id="B926EAA67D44DC3AF529FF4BFE59FF12" box="[317,401,224,248]" italics="true" pageId="12" pageNumber="47">Shinkai</emphasis>
2000
</taxonomicName>
, dive #1187, Hatoma Knoll,
<geoCoordinate id="EE6650737D44DC3AF703FF74FC64FF1D" box="[791,940,223,247]" direction="north" orientation="latitude" pageId="12" pageNumber="47" precision="1" value="24.8536">24°51.216N</geoCoordinate>
,
<geoCoordinate id="EE6650737D44DC3AF7A3FF74FB9FFF1D" box="[951,1111,223,247]" direction="east" orientation="longitude" pageId="12" pageNumber="47" precision="1" value="123.84278">123°50.567E</geoCoordinate>
,
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,
<date id="FFEC10747D44DC3AF0D1FF74FA96FF1D" box="[1221,1374,223,248]" pageId="12" pageNumber="47" value="2000-05-26">26 May 2000</date>
, trap,
<specimenCount id="9D54FD3D7D44DC3AF483FEAFFF30FEF6" box="[151,248,260,284]" pageId="12" pageNumber="47" type="female">1 female</specimenCount>
(cl 13.0 mm), JAMSTEC 028210; dive #1273, Yonaguni Knoll No. 4,
<geoCoordinate id="EE6650737D44DC3AF00DFEAFFB66FEF6" box="[1049,1198,260,284]" direction="north" orientation="latitude" pageId="12" pageNumber="47" precision="1" value="24.848534">24°50.912N</geoCoordinate>
,
<geoCoordinate id="EE6650737D44DC3AF0ADFEAFFA90FEF6" box="[1209,1368,260,284]" direction="east" orientation="longitude" pageId="12" pageNumber="47" precision="1" value="122.700714">122°42.043E</geoCoordinate>
,
<quantity id="4CAA9B517D44DC3AF170FEAFFF67FED5" metricMagnitude="3" metricUnit="m" metricValue="1.339" pageId="12" pageNumber="47" unit="m" value="1339.0">1339 m</quantity>
,
<date id="FFEC10747D44DC3AF4AAFE8CFE95FED5" box="[190,349,295,320]" pageId="12" pageNumber="47" value="2001-05-24">24 May 2001</date>
, slurp gun,
<specimenCount id="9D54FD3D7D44DC3AF5FDFE8CFDFFFED5" box="[489,567,295,319]" pageId="12" pageNumber="47" type="male">1 male</specimenCount>
(cl 7.0 mm),
<specimenCount id="9D54FD3D7D44DC3AF6C9FE8CFC8AFED5" box="[733,834,295,319]" pageId="12" pageNumber="47" type="female">1 female</specimenCount>
(cl ca.
<quantity id="4CAA9B517D44DC3AF78BFE8CFC34FED5" box="[927,1020,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="12" pageNumber="47" unit="mm" value="9.5">9.5 mm</quantity>
), JAMSTEC 043589043594; ROV
<emphasis id="B926EAA67D44DC3AF483FEE7FE8AFE8E" box="[151,322,332,356]" italics="true" pageId="12" pageNumber="47">Hyper-Dolphin</emphasis>
, dive #697,
<geoCoordinate id="EE6650737D44DC3AF5D8FEE7FDA9FE8E" box="[460,609,332,356]" direction="north" orientation="latitude" pageId="12" pageNumber="47" precision="1" value="28.391634">28°23.498N</geoCoordinate>
, 127°38°370E,
<quantity id="4CAA9B517D44DC3AF70EFEE7FCABFE8E" box="[794,867,332,356]" metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" pageId="12" pageNumber="47" unit="m" value="691.0">691 m</quantity>
,
<date id="FFEC10747D44DC3AF77AFEE7FBCFFE8E" box="[878,1031,332,356]" pageId="12" pageNumber="47" value="2007-06-20">20 June 2007</date>
,
<specimenCount id="9D54FD3D7D44DC3AF005FEE7FBAFFE8E" box="[1041,1127,332,356]" pageId="12" pageNumber="47" type="male">2 males</specimenCount>
(cl 7.0, 9.0 mm),
<specimenCount id="9D54FD3D7D44DC3AF124FEE7FA54FE8E" box="[1328,1436,332,356]" pageId="12" pageNumber="47" type="female">3 females</specimenCount>
(cl
<quantity id="4CAA9B517D44DC3AF4A8FEC4FE9DFE6D" box="[188,341,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" metricValueMax="13.3" metricValueMin="6.3" pageId="12" pageNumber="47" unit="mm" value="9.8" valueMax="13.3" valueMin="6.3">6.313.3 mm</quantity>
), JAMSTEC
<date id="FFEC10747D44DC3AF5F9FEC4FD8BFE6D" box="[493,579,367,391]" pageId="12" pageNumber="47" value="24-07-24">072424</date>
<date id="FFEC10747D44DC3AF645FEC4FD6FFE6D" box="[593,679,367,391]" pageId="12" pageNumber="47" value="32-07-24">072432</date>
; dive #700, Minami-Ensei Knoll,
<geoCoordinate id="EE6650737D44DC3AF032FEC4FB73FE6D" box="[1062,1211,367,391]" direction="north" orientation="latitude" pageId="12" pageNumber="47" precision="1" value="28.389116">28°23.347N</geoCoordinate>
,
<geoCoordinate id="EE6650737D44DC3AF0D3FEC4FAAEFE6D" box="[1223,1382,367,391]" direction="east" orientation="longitude" pageId="12" pageNumber="47" precision="1" value="127.640114">127°38.407E</geoCoordinate>
,
<quantity id="4CAA9B517D44DC3AF166FEC4FF67FE46" metricMagnitude="2" metricUnit="m" metricValue="7.12" pageId="12" pageNumber="47" unit="m" value="712.0">712 m</quantity>
,
<date id="FFEC10747D44DC3AF4AFFE3FFE9AFE46" box="[187,338,404,428]" pageId="12" pageNumber="47" value="2007-06-22">22 June 2007</date>
,
<specimenCount id="9D54FD3D7D44DC3AF54AFE3FFE61FE46" box="[350,425,404,428]" pageId="12" pageNumber="47" type="male">1 male</specimenCount>
(cl 8.0 mm),
<specimenCount id="9D54FD3D7D44DC3AF656FE3FFD66FE46" box="[578,686,404,428]" pageId="12" pageNumber="47" type="female">2 females</specimenCount>
(cl 10.0,
<quantity id="4CAA9B517D44DC3AF70CFE3FFC48FE46" box="[792,896,404,428]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="12" pageNumber="47" unit="mm" value="11.6">11.6 mm</quantity>
), JAMSTEC
<date id="FFEC10747D44DC3AF002FE3FFBA3FE46" box="[1046,1131,404,428]" pageId="12" pageNumber="47" value="71-07-25">072571</date>
<date id="FFEC10747D44DC3AF06DFE3FFB07FE46" box="[1145,1231,404,428]" pageId="12" pageNumber="47" value="75-07-25">072575</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C348653F7D44DC26F4DDFE1CFB23FF45" lastPageId="16" lastPageNumber="51" pageId="12" pageNumber="47" type="description">
<paragraph id="8BED36B47D44DC3AF4DDFE1CFF15FE1E" blockId="12.[151,1436,151,500]" pageId="12" pageNumber="47">
<emphasis id="B926EAA67D44DC3AF4DDFE1CFE92FE3A" bold="true" box="[201,346,439,464]" pageId="12" pageNumber="47">Description.</emphasis>
<emphasis id="B926EAA67D44DC3AF576FE13FE09FE3A" box="[354,449,440,464]" italics="true" pageId="12" pageNumber="47">Females</emphasis>
. Body (
<figureCitation id="13692A317D44DC3AF609FE1CFDA7FE25" box="[541,623,439,463]" captionStart="FIGURE 10" captionStartId="12.[151,250,1366,1389]" captionTargetBox="[156,1429,558,1341]" captionTargetId="figure@12.[151,1436,553,1345]" captionTargetPageId="12" captionText="FIGURE 10. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Body (thoracic appendages and pleopods omitted) in lateral view. Scale bar: 5 mm." httpUri="https://zenodo.org/record/280467/files/figure.png" pageId="12" pageNumber="47">Fig. 10</figureCitation>
) moderately robust for genus; integument moderately firm, surface glabrous.
</paragraph>
<caption id="DF2D663C7D44DC3AF483FAFDFABAFA61" httpUri="https://zenodo.org/record/280467/files/figure.png" pageId="12" pageNumber="47" targetBox="[156,1429,558,1341]" targetPageId="12">
<paragraph id="8BED36B47D44DC3AF483FAFDFABAFA61" blockId="12.[151,1436,1366,1419]" pageId="12" pageNumber="47">
<emphasis id="B926EAA67D44DC3AF483FAFDFEE8FA81" bold="true" box="[151,288,1366,1389]" pageId="12" pageNumber="47">FIGURE 10.</emphasis>
<taxonomicName id="4C524D377D44DC3AF53CFAFDFE2DFA86" box="[296,485,1366,1388]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="shinkaiae" status="sp. nov.">
<emphasis id="B926EAA67D44DC3AF53CFAFDFE2DFA86" box="[296,485,1366,1388]" italics="true" pageId="12" pageNumber="47">Lebbeus shinkaiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557DD7D44DC3AF5F8FAFCFDF6FA86" box="[492,574,1367,1388]" pageId="12" pageNumber="47" rank="species">
<emphasis id="B926EAA67D44DC3AF5F8FAFCFDF6FA86" bold="true" box="[492,574,1367,1388]" pageId="12" pageNumber="47">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, female (cl 18.2 mm), DS
<taxonomicName id="4C524D377D44DC3AF7AFFAFDFB89FA86" box="[955,1089,1366,1388]" class="Polychaeta" family="Chrysopetalidae" genus="Shinkai" kingdom="Animalia" order="Phyllodocida" pageId="12" pageNumber="47" phylum="Annelida" rank="genus">
<emphasis id="B926EAA67D44DC3AF7AFFAFDFB89FA86" box="[955,1089,1366,1388]" italics="true" pageId="12" pageNumber="47">Shinkai 2000</emphasis>
</taxonomicName>
, dive #1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Body (thoracic appendages and pleopods omitted) in lateral view. Scale bar: 5 mm.
</paragraph>
</caption>
<paragraph id="8BED36B47D44DC3AF4DDFA18FA8EF8D8" blockId="12.[151,1437,1458,2022]" pageId="12" pageNumber="47">
Rostrum (
<figureCitation id="13692A317D44DC3AF52DFA19FE5EFA20" box="[313,406,1458,1482]" captionStart="FIGURE 10" captionStartId="12.[151,250,1366,1389]" captionTargetBox="[156,1429,558,1341]" captionTargetId="figure@12.[151,1436,553,1345]" captionTargetPageId="12" captionText="FIGURE 10. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Body (thoracic appendages and pleopods omitted) in lateral view. Scale bar: 5 mm." httpUri="https://zenodo.org/record/280467/files/figure.png" pageId="12" pageNumber="47">Figs. 10</figureCitation>
,
<figureCitation id="13692A317D44DC3AF5B6FA19FE74FA20" box="[418,444,1458,1482]" captionStart="FIGURE 11" captionStartId="13.[151,250,1896,1919]" captionTargetBox="[175,1407,554,1873]" captionTargetId="figure@13.[151,1436,527,1887]" captionTargetPageId="13" captionText="FIGURE 11. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for A C, E, F; 1 mm for D." httpUri="https://zenodo.org/record/280468/files/figure.png" pageId="12" pageNumber="47">11</figureCitation>
A, B) straight, directed forward or slightly descending, falling short of or reaching distal margin of first segment of antennular peduncle, 0.30.4 times carapace length; dorsal margin armed with 610 teeth, including 35 teeth on rostrum proper and 35 postrostral teeth; ventral margin armed with 13 small teeth clustered subterminally, ventral lamina slightly developed. Carapace (
<figureCitation id="13692A317D44DC3AF7DCF9B5FBE2F9DC" box="[968,1066,1566,1590]" captionStart="FIGURE 10" captionStartId="12.[151,250,1366,1389]" captionTargetBox="[156,1429,558,1341]" captionTargetId="figure@12.[151,1436,553,1345]" captionTargetPageId="12" captionText="FIGURE 10. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Body (thoracic appendages and pleopods omitted) in lateral view. Scale bar: 5 mm." httpUri="https://zenodo.org/record/280467/files/figure.png" pageId="12" pageNumber="47">Figs. 10</figureCitation>
,
<figureCitation id="13692A317D44DC3AF028F9B5FB9EF9DC" box="[1084,1110,1566,1590]" captionStart="FIGURE 11" captionStartId="13.[151,250,1896,1919]" captionTargetBox="[175,1407,554,1873]" captionTargetId="figure@13.[151,1436,527,1887]" captionTargetPageId="13" captionText="FIGURE 11. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for A C, E, F; 1 mm for D." httpUri="https://zenodo.org/record/280468/files/figure.png" pageId="12" pageNumber="47">11</figureCitation>
A, B) with low but distinct postrostral median carina extending to at least midlength of carapace, sloping toward rostral base; postrostral teeth relatively small, posteriormost one arising at 0.10.2 of carapace length; dorsal margin in lateral view gently convex; supraorbital tooth moderately strong, slightly upturned, arising anterior to level of rostral base, reaching or slightly overreaching tip of antennal tooth; orbital region depressed, orbital margin with convexity, base of eyestalk located between this convexity and suborbital lobe; deep U- or V-shaped notch present below base of supraorbital tooth; suborbital lobe well developed, triangular, reaching distal margin of antennal tooth; anterolateral margin between antennal and pterygostomial teeth strongly sinuous with shallow excavation below antennal tooth.
</paragraph>
<paragraph id="8BED36B47D44DC3BF4DDF895FB27FF3E" blockId="12.[151,1437,1458,2022]" lastBlockId="13.[151,1436,151,500]" lastPageId="13" lastPageNumber="48" pageId="12" pageNumber="47">
Abdomen (
<figureCitation id="13692A317D44DC3AF552F895FE50F8BC" box="[326,408,1854,1878]" captionStart="FIGURE 10" captionStartId="12.[151,250,1366,1389]" captionTargetBox="[156,1429,558,1341]" captionTargetId="figure@12.[151,1436,553,1345]" captionTargetPageId="12" captionText="FIGURE 10. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Body (thoracic appendages and pleopods omitted) in lateral view. Scale bar: 5 mm." httpUri="https://zenodo.org/record/280467/files/figure.png" pageId="12" pageNumber="47">Fig. 10</figureCitation>
) dorsally rounded. Second somite with deep transverse groove, posteriorly forming low but distinct ridge. Pleura of anterior three somites broadly rounded; fourth pleuron unarmed or armed with small posteroventral tooth; fifth pleuron with moderately strong posteroventral tooth. Sixth somite about 1.6 times longer than fifth somite and 1.6 times longer than deep, bearing small posteroventral tooth; posterolateral process terminating in small tooth. Telson (
<figureCitation id="13692A317D44DC3AF5FFF865FDF2F80C" box="[491,570,1998,2022]" captionStart="FIGURE 11" captionStartId="13.[151,250,1896,1919]" captionTargetBox="[175,1407,554,1873]" captionTargetId="figure@13.[151,1436,527,1887]" captionTargetPageId="13" captionText="FIGURE 11. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for A C, E, F; 1 mm for D." httpUri="https://zenodo.org/record/280468/files/figure.png" pageId="12" pageNumber="47">Fig. 11</figureCitation>
C) 1.61.7 times longer than sixth somite, tapering to slighlty convex posterior margin, bearing 57 (rarely 4) dorsolateral spines on each side; posterior margin with 2 pairs of lateral spines (mesial pair longer), 56 spiniform setulose setae and several longer thin plumose setae (
<figureCitation id="13692A317D45DC3BF06FFF17FB02FF3E" box="[1147,1226,188,212]" captionStart="FIGURE 11" captionStartId="13.[151,250,1896,1919]" captionTargetBox="[175,1407,554,1873]" captionTargetId="figure@13.[151,1436,527,1887]" captionTargetPageId="13" captionText="FIGURE 11. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for A C, E, F; 1 mm for D." httpUri="https://zenodo.org/record/280468/files/figure.png" pageId="13" pageNumber="48">Fig. 11</figureCitation>
D).
</paragraph>
<paragraph id="8BED36B47D45DC3BF4DDFF4BFD14FEF6" blockId="13.[151,1436,151,500]" pageId="13" pageNumber="48">
Eye (
<figureCitation id="13692A317D45DC3BF513FF74FE92FF1D" box="[263,346,223,247]" captionStart="FIGURE 11" captionStartId="13.[151,250,1896,1919]" captionTargetBox="[175,1407,554,1873]" captionTargetId="figure@13.[151,1436,527,1887]" captionTargetPageId="13" captionText="FIGURE 11. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for A C, E, F; 1 mm for D." httpUri="https://zenodo.org/record/280468/files/figure.png" pageId="13" pageNumber="48">Fig. 11</figureCitation>
A, B) subpyriform with stalk slightly narrowing proximally; cornea not dilated, its maximum width about 0.15 of carapace length; ocellus absent.
</paragraph>
<paragraph id="8BED36B47D45DC3BF4DDFE8CFBCBFE1E" blockId="13.[151,1436,151,500]" pageId="13" pageNumber="48">
Antennular peduncle (
<figureCitation id="13692A317D45DC3BF5D3FE8CFDDFFED5" box="[455,535,295,319]" captionStart="FIGURE 11" captionStartId="13.[151,250,1896,1919]" captionTargetBox="[175,1407,554,1873]" captionTargetId="figure@13.[151,1436,527,1887]" captionTargetPageId="13" captionText="FIGURE 11. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for A C, E, F; 1 mm for D." httpUri="https://zenodo.org/record/280468/files/figure.png" pageId="13" pageNumber="48">Fig. 11</figureCitation>
A, B) reaching distal 0.20.3 of antennal scale. First segment distinctly longer than distal two segments combined, slightly overreaching midlength of antennal scale, dorsodistal margin armed with 13 small teeth; stylocerite reaching or slightly overreaching distolateral angle of first peduncular segment, sharply pointed, mesial margin sinuous. Second segment 0.4 length of first segment, with l large dorsolateral distal tooth. Third segment less than half as long as second segment, bearing 1 small dorsodistal tooth. Lateral flagellum with thickened aesthetasc-bearing portion about 0.3 times as long as carapace.
</paragraph>
<caption id="DF2D663C7D45DC3BF483F8C3FAB7F831" httpUri="https://zenodo.org/record/280468/files/figure.png" pageId="13" pageNumber="48" targetBox="[175,1407,554,1873]" targetPageId="13">
<paragraph id="8BED36B47D45DC3BF483F8C3FAB7F831" blockId="13.[151,1436,1896,2011]" pageId="13" pageNumber="48">
<emphasis id="B926EAA67D45DC3BF483F8C3FED7F894" bold="true" box="[151,287,1896,1919]" pageId="13" pageNumber="48">FIGURE 11.</emphasis>
<taxonomicName id="4C524D377D45DC3BF533F8C2FE2CF895" box="[295,484,1897,1919]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="shinkaiae" status="sp. nov.">
<emphasis id="B926EAA67D45DC3BF533F8C2FE2CF895" box="[295,484,1897,1919]" italics="true" pageId="13" pageNumber="48">Lebbeus shinkaiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557DD7D45DC3BF5FFF8C2FDF5F894" box="[491,573,1897,1918]" pageId="13" pageNumber="48" rank="species">
<emphasis id="B926EAA67D45DC3BF5FFF8C2FDF5F894" bold="true" box="[491,573,1897,1918]" pageId="13" pageNumber="48">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, female (cl 18.2 mm), DS
<taxonomicName id="4C524D377D45DC3BF7AFF8C2FB89F895" box="[955,1089,1897,1919]" class="Polychaeta" family="Chrysopetalidae" genus="Shinkai" kingdom="Animalia" order="Phyllodocida" pageId="13" pageNumber="48" phylum="Annelida" rank="genus">
<emphasis id="B926EAA67D45DC3BF7AFF8C2FB89F895" box="[955,1089,1897,1919]" italics="true" pageId="13" pageNumber="48">Shinkai 2000</emphasis>
</taxonomicName>
, dive #1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for AC, E, F; 1 mm for D.
</paragraph>
</caption>
<caption id="DF2D663C7D46DC38F483F897FA4AF827" httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="14" pageNumber="49" targetBox="[166,1404,219,1795]" targetPageId="14">
<paragraph id="8BED36B47D46DC38F483F897FA4AF827" blockId="14.[151,1436,1852,1997]" pageId="14" pageNumber="49">
<emphasis id="B926EAA67D46DC38F483F897FEE8F8BB" bold="true" box="[151,288,1852,1875]" pageId="14" pageNumber="49">FIGURE 12.</emphasis>
<taxonomicName id="4C524D377D46DC38F53CF897FE2DF8B8" box="[296,485,1852,1874]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="49" phylum="Arthropoda" rank="species" species="shinkaiae" status="sp. nov.">
<emphasis id="B926EAA67D46DC38F53CF897FE2DF8B8" box="[296,485,1852,1874]" italics="true" pageId="14" pageNumber="49">Lebbeus shinkaiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557DD7D46DC38F5F8F896FDF6F8B8" box="[492,574,1853,1874]" pageId="14" pageNumber="49" rank="species">
<emphasis id="B926EAA67D46DC38F5F8F896FDF6F8B8" bold="true" box="[492,574,1853,1874]" pageId="14" pageNumber="49">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, female (cl 18.2 mm), DS
<taxonomicName id="4C524D377D46DC38F7AFF897FB89F8B8" box="[955,1089,1852,1874]" class="Polychaeta" family="Chrysopetalidae" genus="Shinkai" kingdom="Animalia" order="Phyllodocida" pageId="14" pageNumber="49" phylum="Annelida" rank="genus">
<emphasis id="B926EAA67D46DC38F7AFF897FB89F8B8" box="[955,1089,1852,1874]" italics="true" pageId="14" pageNumber="49">Shinkai 2000</emphasis>
</taxonomicName>
, dive #1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, DF, G, I, J; 1 mm for B, C, H.
</paragraph>
</caption>
<caption id="DF2D663C7D47DC39F483FCE0FC91FC74" httpUri="https://zenodo.org/record/280470/files/figure.png" pageId="15" pageNumber="50" targetBox="[179,1410,214,802]" targetPageId="15">
<paragraph id="8BED36B47D47DC39F483FCE0FC91FC74" blockId="15.[151,1436,843,927]" pageId="15" pageNumber="50">
<emphasis id="B926EAA67D47DC39F483FCE0FEE8FC8B" bold="true" box="[151,288,843,866]" pageId="15" pageNumber="50">FIGURE 13.</emphasis>
<taxonomicName id="4C524D377D47DC39F53CFCE7FE2DFC88" box="[296,485,844,866]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="15" pageNumber="50" phylum="Arthropoda" rank="species" species="shinkaiae" status="sp. nov.">
<emphasis id="B926EAA67D47DC39F53CFCE7FE2DFC88" box="[296,485,844,866]" italics="true" pageId="15" pageNumber="50">Lebbeus shinkaiae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557DD7D47DC39F5F8FCE7FDF6FC8B" box="[492,574,844,865]" pageId="15" pageNumber="50" rank="species">
<emphasis id="B926EAA67D47DC39F5F8FCE7FDF6FC8B" bold="true" box="[492,574,844,865]" pageId="15" pageNumber="50">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype, male (cl 9.0 mm), JAMSTEC 072424072432. A, carapace and cephalic appendages, lateral view; B, endopod of left first pleopod, dorsal (anterior) view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 2 mm for A; 1 mm for B, C.
</paragraph>
</caption>
<paragraph id="8BED36B47D47DC39F4DDFC67FB50FBC1" blockId="15.[151,1437,971,2003]" pageId="15" pageNumber="50">
Antenna (
<figureCitation id="13692A317D47DC39F52EFC60FE43FC09" box="[314,395,971,995]" captionStart="FIGURE 11" captionStartId="13.[151,250,1896,1919]" captionTargetBox="[175,1407,554,1873]" captionTargetId="figure@13.[151,1436,527,1887]" captionTargetPageId="13" captionText="FIGURE 11. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for A C, E, F; 1 mm for D." httpUri="https://zenodo.org/record/280468/files/figure.png" pageId="15" pageNumber="50">Fig. 11</figureCitation>
A, B, E) with basicerite bearing moderately small ventrolateral tooth; carpocerite reaching midlength of antennal scale. Antennal scale about 0.5 times as long as carapace and 2.4 times longer than wide; lateral margin nearly straight; distolateral tooth not reaching rounded distal margin of lamella.
</paragraph>
<paragraph id="8BED36B47D47DC39F4DDFB9CFC3CFB35" blockId="15.[151,1437,971,2003]" pageId="15" pageNumber="50">
Mouthparts similar to those of other species of the genus. Third maxilliped (
<figureCitation id="13692A317D47DC39F038FB9CFBB4FBA5" box="[1068,1148,1079,1103]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">Fig. 12</figureCitation>
A) overreaching antennal scale by 0.60.7 length of ultimate segment; ultimate segment 3.63.7 times longer than penultimate segment, tapering distally, with short row of darkly pigmented corneous spines distomesially (
<figureCitation id="13692A317D47DC39F07DFBD4FB76FB7D" box="[1129,1214,1151,1175]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">Fig. 12</figureCitation>
B); antepenultimate segment shorter than distal two segments combined, armed with 1 small tooth on distolateral margin and 1 minute spinule at ventrolateral distal angle (
<figureCitation id="13692A317D47DC39F63AFB6CFD48FB35" box="[558,640,1223,1247]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">Fig. 12</figureCitation>
C); lateral surface bluntly ridged.
</paragraph>
<paragraph id="8BED36B47D47DC39F4DDFB40FD3BFACD" blockId="15.[151,1437,971,2003]" pageId="15" pageNumber="50">
Strap-like, terminally hooked epipods present on third maxilliped to third pereopod, corresponding setobranchs present on first to fourth pereopod (
<figureCitation id="13692A317D47DC39F697FAA4FD1CFACD" box="[643,724,1295,1319]" captionStart="FIGURE 11" captionStartId="13.[151,250,1896,1919]" captionTargetBox="[175,1407,554,1873]" captionTargetId="figure@13.[151,1436,527,1887]" captionTargetPageId="13" captionText="FIGURE 11. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. A, anterior part of carapace and cephalic appendages, lateral view; B, same, dorsal view; C, telson, dorsal view; D, posterior part of telson, dorsal view; E, left antennal peduncle and scale, ventral view; F, coxae of second to fourth pereopods, showing presence of epipod on third pereopod. Scale bars: 5 mm for A C, E, F; 1 mm for D." httpUri="https://zenodo.org/record/280468/files/figure.png" pageId="15" pageNumber="50">Fig. 11</figureCitation>
F).
</paragraph>
<paragraph id="8BED36B47D47DC39F4DDFA98FD80F932" blockId="15.[151,1437,971,2003]" pageId="15" pageNumber="50">
First pereopod (
<figureCitation id="13692A317D47DC39F56EFA98FE02FAA1" box="[378,458,1331,1355]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">Fig. 12</figureCitation>
D) moderately stout, reaching distal margin of antennal scale; dactylus (
<figureCitation id="13692A317D47DC39F0F9FA98FAF5FAA1" box="[1261,1341,1331,1355]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">Fig. 12</figureCitation>
E) about 0.7 times as long as palm, terminating in 2 darkly pigmented corneous claws; fixed finger terminating in single corneous claw. Second pereopod (
<figureCitation id="13692A317D47DC39F5E7FAD0FD8CFA79" box="[499,580,1403,1427]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">Fig. 12</figureCitation>
F) overreaching antennal scale by 0.4 length of carpus; dactylus terminating in 2 small corneous claws; carpus divided into 7 articles. Third to fifth pereopods moderately long and slender, similar and slightly decreasing in length posteriorly. Third pereopod (
<figureCitation id="13692A317D47DC39F760FA68FC0CFA31" box="[884,964,1475,1499]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">Fig. 12</figureCitation>
G) overreaching antennal scale by 0.60.7 length of propodus; dactylus (
<figureCitation id="13692A317D47DC39F5F3FA43FDFFF9EA" box="[487,567,1512,1536]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">Fig. 12</figureCitation>
H) 0.150.18 times as long as propodus, moderately stout (3.03.2 times longer than deep), terminating in acute, darkly pigmented unguis, armed with 4 or 5 darkly pigmented accessory spinules on flexor margin, distalmost accessory spinule subterminal, distinctly larger than others, making tip of dactylus appearing biunguiculate; carpus about 0.5 times as long as propodus; merus armed with 05 lateral spines. Fourth pereopod (
<figureCitation id="13692A317D47DC39F570F9D3FE7FF97A" box="[356,439,1656,1680]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">
Fig.
<date id="FFEC10747D47DC39F58FF9D3FE7FF97A" box="[411,439,1656,1680]" pageId="15" pageNumber="50" value="1912-01">12</date>
</figureCitation>
I) overreaching antennal scale by 0.3 length of propodus; merus with 05 ventrolateral spines. Fifth pereopod (
<figureCitation id="13692A317D47DC39F5A6F930FDC1F959" box="[434,521,1691,1715]" captionStart="FIGURE 12" captionStartId="14.[151,250,1852,1875]" captionTargetBox="[166,1404,219,1795]" captionTargetId="figure@14.[151,1436,193,1831]" captionTargetPageId="14" captionText="FIGURE 12. Lebbeus shinkaiae sp. nov., holotype, female (cl 18.2 mm), DS Shinkai 2000, dive # 1182, Hatoma Knoll, Okinawa Trough, Japan, NSMT-Cr 21982. Left thoracic appendages. A, third maxilliped, lateral view; B, same, distal part of ultimate segment, dorsal (extensor) view; C, same, distal part of antepenultimate segment, lateral view; D, first pereopod, lateral view; E, same, chela, extensor view; F, second pereopod, lateral view; G, third pereopod, lateral view; H, same, dactylus, lateral view; I, fourth pereopod, lateral view; J, fifth pereopod, lateral view. Scale bars: 2 mm for A, D F, G, I, J; 1 mm for B, C, H." httpUri="https://zenodo.org/record/280469/files/figure.png" pageId="15" pageNumber="50">Fig. 12</figureCitation>
J) overreaching distal margin of antennal scale by length of dactylus; merus unarmed or armed with 1 lateral spine.
</paragraph>
<paragraph id="8BED36B47D47DC39F4DDF94FFEDBF839" blockId="15.[151,1437,971,2003]" pageId="15" pageNumber="50">
<emphasis id="B926EAA67D47DC39F4DDF94FFEC5F916" box="[201,269,1764,1788]" italics="true" pageId="15" pageNumber="50">Males</emphasis>
. Carapace (
<figureCitation id="13692A317D47DC39F589F948FE39F911" box="[413,497,1763,1787]" captionStart="FIGURE 13" captionStartId="15.[151,250,843,866]" captionTargetBox="[179,1410,214,802]" captionTargetId="figure@15.[151,1436,193,822]" captionTargetPageId="15" captionText="FIGURE 13. Lebbeus shinkaiae sp. nov., paratype, male (cl 9.0 mm), JAMSTEC 072424 072432. A, carapace and cephalic appendages, lateral view; B, endopod of left first pleopod, dorsal (anterior) view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 2 mm for A; 1 mm for B, C." httpUri="https://zenodo.org/record/280470/files/figure.png" pageId="15" pageNumber="50">Fig. 13</figureCitation>
A) with nearly straight dorsal margin in lateral view; postrostral median carina extending beyond midlength of carapace. Fourth abdominal pleuron bearing small but distinct posteroventral tooth. Antennular peduncle (
<figureCitation id="13692A317D47DC39F584F880FE17F8A9" box="[400,479,1835,1859]" captionStart="FIGURE 13" captionStartId="15.[151,250,843,866]" captionTargetBox="[179,1410,214,802]" captionTargetId="figure@15.[151,1436,193,822]" captionTargetPageId="15" captionText="FIGURE 13. Lebbeus shinkaiae sp. nov., paratype, male (cl 9.0 mm), JAMSTEC 072424 072432. A, carapace and cephalic appendages, lateral view; B, endopod of left first pleopod, dorsal (anterior) view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 2 mm for A; 1 mm for B, C." httpUri="https://zenodo.org/record/280470/files/figure.png" pageId="15" pageNumber="50">Fig. 13</figureCitation>
A) reaching nearly to distal margin of antennal scale; outer antennular flagellum with thickened aesthetasc-bearing portion about 0.4 times as long as carapace; inner flagellum thin, subequal in length to carapace. Endopod of first pleopod (
<figureCitation id="13692A317D47DC39F643F8D8FD61F861" box="[599,681,1907,1931]" captionStart="FIGURE 13" captionStartId="15.[151,250,843,866]" captionTargetBox="[179,1410,214,802]" captionTargetId="figure@15.[151,1436,193,822]" captionTargetPageId="15" captionText="FIGURE 13. Lebbeus shinkaiae sp. nov., paratype, male (cl 9.0 mm), JAMSTEC 072424 072432. A, carapace and cephalic appendages, lateral view; B, endopod of left first pleopod, dorsal (anterior) view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 2 mm for A; 1 mm for B, C." httpUri="https://zenodo.org/record/280470/files/figure.png" pageId="15" pageNumber="50">Fig. 13</figureCitation>
B) without lobe just lateral to base of terminally located appendix interna. Appendix masculina on second pleopod shorter than appendix interna, bearing about 15 setae in distal 0.4 (
<figureCitation id="13692A317D47DC39F48BF810FF39F839" box="[159,241,1979,2003]" captionStart="FIGURE 13" captionStartId="15.[151,250,843,866]" captionTargetBox="[179,1410,214,802]" captionTargetId="figure@15.[151,1436,193,822]" captionTargetPageId="15" captionText="FIGURE 13. Lebbeus shinkaiae sp. nov., paratype, male (cl 9.0 mm), JAMSTEC 072424 072432. A, carapace and cephalic appendages, lateral view; B, endopod of left first pleopod, dorsal (anterior) view; C, appendices interna and masculina of left second pleopod, mesial view. Scale bars: 2 mm for A; 1 mm for B, C." httpUri="https://zenodo.org/record/280470/files/figure.png" pageId="15" pageNumber="50">Fig. 13</figureCitation>
C).
</paragraph>
<paragraph id="8BED36B47D58DC26F4DDFF3CFB23FF45" blockId="16.[151,1436,151,535]" box="[201,1259,151,176]" pageId="16" pageNumber="51">
<emphasis id="B926EAA67D58DC26F4DDFF3CFE54FF5A" bold="true" box="[201,412,151,176]" pageId="16" pageNumber="51">Coloration in life.</emphasis>
Entirely reddish or light yellowish brown or tan. Cornea darkly pigmented.
</paragraph>
</subSubSection>
<subSubSection id="C348653F7D58DC26F4DDFF10FEE3FE25" pageId="16" pageNumber="51" type="distribution">
<paragraph id="8BED36B47D58DC26F4DDFF10FCDEFF3E" blockId="16.[151,1436,151,535]" box="[201,790,187,212]" pageId="16" pageNumber="51">
<emphasis id="B926EAA67D58DC26F4DDFF10FEAAFF3E" bold="true" box="[201,354,187,212]" pageId="16" pageNumber="51">Distribution.</emphasis>
Okinawa Trough,
<collectingCountry id="F34576247D58DC26F621FF16FDB0FF3E" box="[565,632,189,212]" name="Japan" pageId="16" pageNumber="51">Japan</collectingCountry>
,
<quantity id="4CAA9B517D58DC26F696FF17FCDAFF3E" box="[642,786,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.091" metricValueMax="1.491" metricValueMin="0.691" pageId="16" pageNumber="51" unit="m" value="1091.0" valueMax="1491.0" valueMin="691.0">6911491 m</quantity>
.
</paragraph>
<paragraph id="8BED36B47D58DC26F4DDFF4BFEE3FE25" blockId="16.[151,1436,151,535]" pageId="16" pageNumber="51">
<emphasis id="B926EAA67D58DC26F4DDFF4BFEE5FF12" bold="true" box="[201,301,224,248]" pageId="16" pageNumber="51">Ecology.</emphasis>
In the vent fields of the Hatoma Knoll, this species was abundant in deep-sea mussel beds,
<taxonomicName id="4C524D377D58DC26F13BFF4BFD5AFEF6" authority="Hashimoto &amp; Okutani, 1994" authorityName="Hashimoto &amp; Okutani" authorityYear="1994" class="Bivalvia" family="Mytilidae" genus="Bathymodiolus" kingdom="Animalia" order="Mytiloida" pageId="16" pageNumber="51" phylum="Mollusca" rank="species" species="platifrons">
<emphasis id="B926EAA67D58DC26F13BFF4BFE86FEF6" italics="true" pageId="16" pageNumber="51">Bathymodiolus platifrons</emphasis>
Hashimoto &amp; Okutani, 1994
</taxonomicName>
. Other decapod crustaceans seen in the vent fields include the alvinocaridid shrimps
<taxonomicName id="4C524D377D58DC26F574FE83FC69FED5" authority="Kikuchi &amp; Ohta, 1995" authorityName="Kikuchi &amp; Ohta" authorityYear="1995" box="[352,929,295,320]" class="Malacostraca" family="Alvinocarididae" genus="Alvinocaris" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="longirostris">
<emphasis id="B926EAA67D58DC26F574FE83FDBDFEAA" box="[352,629,296,320]" italics="true" pageId="16" pageNumber="51">Alvinocaris longirostris</emphasis>
<bibRefCitation id="EFC34B457D58DC26F692FE8CFC69FED5" author="Kikuchi" box="[646,929,295,319]" pageId="16" pageNumber="51" refString="Kikuchi, T. &amp; Ohta, S. (1995) Two caridean shrimps of the families Bresiliidae and Hippolytidae from a hydrothermal field on the Iheya Ridge, off the Ryukyu Islands, Japan. Journal of Crustacean Biology, 15, 771 - 785." type="journal article" year="1995">Kikuchi &amp; Ohta, 1995</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C524D377D58DC26F7F8FE83FEAEFE8E" authority="Kikuchi &amp; Hashimoto, 2000" authorityName="Kikuchi &amp; Hashimoto" authorityYear="2000" class="Malacostraca" family="Alvinocarididae" genus="Shinkaicaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="leurokolos">
<emphasis id="B926EAA67D58DC26F7F8FE83FB37FEAA" box="[1004,1279,296,320]" italics="true" pageId="16" pageNumber="51">Shinkaicaris leurokolos</emphasis>
(Kikuchi &amp; Hashimoto, 2000)
</taxonomicName>
and the galatheoid squat lobster
<taxonomicName id="4C524D377D58DC26F6FAFEE7FB2AFE8E" authority="Baba &amp; Williams, 1998" authorityName="Baba &amp; Williams" authorityYear="1998" box="[750,1250,332,356]" class="Malacostraca" family="Galatheidae" genus="Shinkaia" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="crosnieri">
<emphasis id="B926EAA67D58DC26F6FAFEE7FC77FE8E" box="[750,959,332,356]" italics="true" pageId="16" pageNumber="51">Shinkaia crosnieri</emphasis>
Baba &amp; Williams, 1998
</taxonomicName>
. Observation of video recordings showed that individuals of
<taxonomicName id="4C524D377D58DC26F68FFEDBFCA6FE62" box="[667,878,368,392]" class="Malacostraca" family="Hippolytidae" genus="Lebbeus" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="51" phylum="Arthropoda" rank="species" species="shinkaiae">
<emphasis id="B926EAA67D58DC26F68FFEDBFCA6FE62" box="[667,878,368,392]" italics="true" pageId="16" pageNumber="51">Lebbeus shinkaiae</emphasis>
</taxonomicName>
pick some food items (probably bacterial mats) from the substrate by using the chelae of the first pereopods. It was found that they aggregate on the broken mussels for food.
</paragraph>
</subSubSection>
<subSubSection id="C348653F7D58DC26F4DDFE77FD49FDFD" pageId="16" pageNumber="51" type="etymology">
<paragraph id="8BED36B47D58DC26F4DDFE77FD49FDFD" blockId="16.[151,1436,151,535]" pageId="16" pageNumber="51">
<emphasis id="B926EAA67D58DC26F4DDFE77FE87FE1E" bold="true" box="[201,335,476,500]" pageId="16" pageNumber="51">Etymology.</emphasis>
From the name of the deep submersible
<taxonomicName id="4C524D377D58DC26F708FE77FC78FE1E" box="[796,944,476,500]" class="Polychaeta" family="Chrysopetalidae" genus="Shinkai" kingdom="Animalia" order="Phyllodocida" pageId="16" pageNumber="51" phylum="Annelida" rank="genus">
<emphasis id="B926EAA67D58DC26F708FE77FC78FE1E" box="[796,944,476,500]" italics="true" pageId="16" pageNumber="51">Shinkai 2000</emphasis>
</taxonomicName>
, which contributed substantially to biological studies of chemosynthetic communities.
</paragraph>
</subSubSection>
</treatment>
</document>