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<document id="DB8D1DF058B2E943E06A78BF920982F7" ID-CLB-Dataset="21232" ID-DOI="10.11646/zootaxa.4608.1.3" ID-GBIF-Dataset="c4d067fa-4c56-4248-8084-caf4dd1eea21" ID-ISSN="1175-5326" ID-Zenodo-Dep="3993611" ID-ZooBank="3540218C-34FA-4D91-925A-3E89C6C3A7C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1597950731833" checkinUser="felipe" docAuthor="Venugopal, Akhil S. &amp; Thomas, Sabu K." docDate="2019" docId="03FB8782FFF277524AFEF9A4301DFCFC" docLanguage="en" docName="zootaxa.4608.1.3.pdf" docOrigin="Zootaxa 4608 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pheropsophus lissoderus Chaudoir 1850" docType="treatment" docVersion="8" lastPageNumber="77" masterDocId="FFC2FFFAFFF8775E4A69FFAD3561FE32" masterDocTitle="Bombardier beetles of the genus Pheropsophus Solier 1833 (Carabidae: Brachininae: Brachinini) from Indian subcontinent" masterLastPageNumber="89" masterPageNumber="65" pageNumber="75" updateTime="1698845024281" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="239DDF48C755CE5A3DC2800B49CBDE0D">Bombardier beetles of the genus Pheropsophus Solier 1833 (Carabidae: Brachininae: Brachinini) from Indian subcontinent</mods:title>
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<mods:namePart id="8BA3FBB3B75B9E5D556CD02F45B283EE">Venugopal, Akhil S.</mods:namePart>
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<mods:namePart id="8DDCCE607B14AE13007B04751908EDBE">Thomas, Sabu K.</mods:namePart>
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<mods:date id="BA77BC591FCE013C827A830DD9024F69">2019</mods:date>
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<mods:number id="9BB186A1C0AD1B83F1DB6DCEA57740AB">2019-05-17</mods:number>
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<treatment id="03FB8782FFF277524AFEF9A4301DFCFC" ID-DOI="http://doi.org/10.5281/zenodo.4335113" ID-GBIF-Taxon="167284785" ID-Zenodo-Dep="4335113" LSID="urn:lsid:plazi:treatment:03FB8782FFF277524AFEF9A4301DFCFC" httpUri="http://treatment.plazi.org/id/03FB8782FFF277524AFEF9A4301DFCFC" lastPageId="12" lastPageNumber="77" pageId="10" pageNumber="75">
<subSubSection id="C348651FFFF277544AFEF9A437D9F816" box="[151,696,1545,1572]" pageId="10" pageNumber="75" type="nomenclature">
<paragraph id="8BED3694FFF277544AFEF9A437D9F816" blockId="10.[151,696,1545,1608]" box="[151,696,1545,1572]" pageId="10" pageNumber="75">
<heading id="D0A581F8FFF277544AFEF9A437D9F816" bold="true" box="[151,696,1545,1572]" fontSize="11" level="1" pageId="10" pageNumber="75" reason="1">
<taxonomicName id="4C524D17FFF277544AFEF9A437D9F816" ID-CoL="6V8J8" authority="CHAUDOIR, 1850" authorityName="Chaudoir" authorityYear="1850" box="[151,696,1545,1572]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="lissoderus">
<emphasis id="B926EA86FFF277544AFEF9A437D9F816" bold="true" box="[151,696,1545,1572]" pageId="10" pageNumber="75">
<emphasis id="B926EA86FFF277544AFEF9A434A5F816" bold="true" box="[151,452,1545,1572]" italics="true" pageId="10" pageNumber="75">Pheropsophus lissoderus</emphasis>
<bibRefCitation id="EFC34B65FFF277544BA2F9A737D9F816" author="Chaudoir, M." box="[459,696,1546,1572]" pageId="10" pageNumber="75" pagination="62 - 206" refId="ref18853" refString="Chaudoir, M. (1850) Supplement a la faune des carabiques de la Russie. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 23 (3), 62 - 206." type="journal article" year="1850">CHAUDOIR, 1850</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFF277544AFEF983345DF87A" box="[151,316,1581,1608]" pageId="10" pageNumber="75" type="description">
<paragraph id="8BED3694FFF277544AFEF983345DF87A" blockId="10.[151,696,1545,1608]" box="[151,316,1581,1608]" pageId="10" pageNumber="75">
(
<figureCitation id="13692A11FFF277544AC9F983359FF87A" box="[160,254,1582,1608]" captionStart="FIGURE 2" captionStartId="11.[151,250,1867,1892]" captionTargetBox="[220,1413,191,1818]" captionTargetId="figure@11.[204,1425,179,1827]" captionTargetPageId="11" captionText="FIGURE 2. Dorsal habitus of Pheropsophus from India (cont.)A) Pheropsophus lissoderus (Holotype, Tibet, MNHN), B) P. nanodes (Verified specimen, S.E.Tibet, BMNH), C) P. picicollis (Syntype, Siam (now Thailand) MNHN), D) P. nigricollis (Holotype, Bangalore, BMNH), E) P. occipitalis (Verified specimen, China, BMNH), F) P. prophylax (Holotype, Manipur, BMNH), G) P. scythropus (Holotype, Gopaldhara, BMNH), H) P. scythropus dissolutus (Laos, BMNH)" figureDoi="http://doi.org/10.5281/zenodo.3993615" httpUri="https://zenodo.org/record/3993615/files/figure.png" pageId="10" pageNumber="75">Fig. 2 A</figureCitation>
,
<figureCitation id="13692A11FFF277544B63F9803455F87A" box="[266,308,1581,1608]" captionStart="FIGURE 5" captionStartId="20.[151,250,1882,1907]" captionTargetBox="[174,1420,189,1789]" captionTargetId="figure@20.[153,1438,176,1822]" captionTargetPageId="20" captionText="FIGURE 5. Pheropsophus male genitalia, Right lateral aspectA) Pheropsophus devagiriensis sp. nov. (Holotype, India: Tamilnadu: Courtallam), B) P. indicus sp. nov. (Holotype, India: Kerala: Palakkad), C) P. bimaculatus (India: Kerala: Nilambur), D) P. lissoderus (India: Kerala: Kalpetta), E) P. occipitalis (India: Kerala: Kalpetta), F) P. picicollis (India: Kerala: Chelari), G) P. sobrinus (India: Kerala: Palakkad), H) P. tripustulatus (India: Kerala: Nilambur). Scale Bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3993621" httpUri="https://zenodo.org/record/3993621/files/figure.png" pageId="10" pageNumber="75">5 D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFF277544AFEF9DF3429F906" pageId="10" pageNumber="75" type="reference_group">
<paragraph id="8BED3694FFF277544AFEF9DF37CAF8B8" blockId="10.[151,1265,1649,1736]" box="[151,683,1649,1674]" pageId="10" pageNumber="75">
<emphasis id="B926EA86FFF277544AFEF9DF35C4F8B8" box="[151,165,1650,1674]" italics="true" pageId="10" pageNumber="75">=</emphasis>
<taxonomicName id="4C524D17FFF277544ACCF9DF3741F8B8" ID-CoL="4G38V" authority="BATES, 1886" authorityName="Bates" authorityYear="1886" box="[165,544,1649,1674]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="catoirei">
<emphasis id="B926EA86FFF277544ACCF9DF34ECF8B8" box="[165,397,1650,1674]" italics="true" pageId="10" pageNumber="75">Pheropsophus catoirei</emphasis>
<bibRefCitation id="EFC34B65FFF277544BFAF9DF3741F8B8" author="Bates, H. W." box="[403,544,1649,1674]" pageId="10" pageNumber="75" pagination="199 - 212" refId="ref18382" refString="Bates, H. W. (1886) On the Geodephagous Coleoptera collected by George Lewis in Ceylon. The Annals and Magazine of Natural History including Zoology, Botany and Geology, 5 (17), 199 - 212. https: // doi. org / 10.1080 / 00222938609460134" type="journal article" year="1886">BATES, 1886</bibRefCitation>
</taxonomicName>
(non Dejean)
</paragraph>
<paragraph id="8BED3694FFF277544AFEF93C3190F89B" blockId="10.[151,1265,1649,1736]" box="[151,1265,1680,1705]" pageId="10" pageNumber="75">
<treatmentCitationGroup id="AB4211BAFFF277544AFEF93C3190F89B" box="[151,1265,1680,1705]" pageId="10" pageNumber="75">
<treatmentCitation id="0AF31085FFF277544AFEF93C3700F89B" author="Chaudoir, M." box="[151,609,1680,1705]" page="79" pageId="10" pageNumber="75" year="1850">
<taxonomicName id="4C524D17FFF277544AFEF93C3700F89B" ID-CoL="6V8J8" authority="Chaudoir, 1850: 79" authorityName="Chaudoir" authorityPageNumber="79" authorityYear="1850" box="[151,609,1680,1705]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="lissoderus">
<emphasis id="B926EA86FFF277544AFEF93C34F4F89B" box="[151,405,1681,1705]" italics="true" pageId="10" pageNumber="75">Pheropsophus lissoderus</emphasis>
<bibRefCitation id="EFC34B65FFF277544BF2F93D3700F89B" author="Chaudoir, M." box="[411,609,1680,1705]" pageId="10" pageNumber="75" pagination="62 - 206" refId="ref18853" refString="Chaudoir, M. (1850) Supplement a la faune des carabiques de la Russie. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 23 (3), 62 - 206." type="journal article" year="1850">Chaudoir, 1850: 79</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF277544802F93D37A6F89B" author="Chaudoir, M." box="[619,711,1680,1705]" page="24" pageId="10" pageNumber="75" year="1876">
<bibRefCitation id="EFC34B65FFF277544802F93D37A6F89B" author="Chaudoir, M." box="[619,711,1680,1705]" pageId="10" pageNumber="75" pagination="11 - 104" refId="ref18920" refString="Chaudoir, M. (1876) Monographie de Brachynides. Annales de la Societe entomologique de Belgique, 19, 11 - 104." type="journal article" year="1876">1876: 24</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF2775448B8F93D36C1F89B" author="Andrewes, H. E." box="[721,928,1680,1705]" page="21" pageId="10" pageNumber="75" year="1929">
<bibRefCitation id="EFC34B65FFF2775448B8F93D36C1F89B" author="Andrewes, H. E." box="[721,928,1680,1705]" pageId="10" pageNumber="75" refId="ref18155" refString="Andrewes, H. E. (1929) The fauna of British India, including Ceylon and Burma. Coleoptera. Carabidae. Vol. I. Carabinae. Taylor &amp; Francis, London, xviii + 431 pp., 9 pls." type="book" year="1929">Andrewes, 1929: 21</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF2775449C2F93C3175F89B" author="Andrewes, H. E." box="[939,1044,1681,1705]" page="274" pageId="10" pageNumber="75" year="1930">
<bibRefCitation id="EFC34B65FFF2775449C2F93C3175F89B" author="Andrewes, H. E." box="[939,1044,1681,1705]" pageId="10" pageNumber="75" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">1930: 274</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF277544E77F93D318CF89B" author="Hrdlicka, J." box="[1054,1261,1680,1705]" page="480" pageId="10" pageNumber="75" year="2017">
<bibRefCitation id="EFC34B65FFF277544E77F93D318CF89B" author="Hrdlicka, J." box="[1054,1261,1680,1705]" pageId="10" pageNumber="75" pagination="471 - 480" refId="ref19780" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden / Boston, 471 - 480." type="book chapter" year="2017">Hrdlička, 2017: 480</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BED3694FFF277544AFEF90236C4F8F5" blockId="10.[151,1265,1649,1736]" box="[151,933,1711,1736]" pageId="10" pageNumber="75">
<treatmentCitationGroup id="AB4211BAFFF277544AFEF90236C4F8F5" box="[151,933,1711,1736]" pageId="10" pageNumber="75">
<taxonomicName id="4C524D17FFF277544AFEF90234DBF8F5" ID-CoL="4G38V" authority="Bates" authorityName="Bates" authorityYear="1886" box="[151,442,1711,1735]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="75" phylum="Arthropoda" rank="species" species="catoirei">
<emphasis id="B926EA86FFF277544AFEF902341DF8F5" box="[151,380,1711,1735]" italics="true" pageId="10" pageNumber="75">Pheropsophus catoirei</emphasis>
Bates
</taxonomicName>
(non Dejean), 1886: 201;
<treatmentCitation id="0AF31085FFF2775448ADF90236C1F8F5" author="Andrewes, H. E." box="[708,928,1711,1736]" page="274" pageId="10" pageNumber="75" year="1930">
<bibRefCitation id="EFC34B65FFF2775448ADF90236C1F8F5" author="Andrewes, H. E." box="[708,928,1711,1736]" pageId="10" pageNumber="75" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes, 1930: 274</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BED3694FFF277544AFEF9583429F906" blockId="10.[151,1436,1781,1988]" pageId="10" pageNumber="75">
<emphasis id="B926EA86FFF277544AFEF958348BF922" bold="true" box="[151,490,1781,1808]" pageId="10" pageNumber="75">Specimens examined (n=12)</emphasis>
:
<materialsCitation id="3B3A3CC9FFF277544B95F95B3429F906" ID-GBIF-Occurrence="2846550332" collectionCode="MNHN" collectorName="M. Melly" location="Tibet" pageId="10" pageNumber="75" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus id="54E98836FFF277544B95F95B3705F922" box="[508,612,1782,1808]" pageId="10" pageNumber="75" type="holotype">Holotype</typeStatus>
(by monotypy), labelled “ExMusaeo Chaudoir”,
<collectingRegion id="4996F876FFF277544ECAF95B3180F922" box="[1187,1249,1782,1808]" country="China" name="Xizang" pageId="10" pageNumber="75">Tibet</collectingRegion>
, coll.
<collectorName id="26A75342FFF277544F43F95B30F9F922" box="[1322,1432,1782,1808]" pageId="10" pageNumber="75">M. Melly</collectorName>
, type in
<collectionCode id="ED43AE51FFF277544A82F8B73425F906" box="[235,324,1818,1844]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="75">MNHN</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFF277524AAEF89337D6FCB5" lastPageId="12" lastPageNumber="77" pageId="10" pageNumber="75" type="description">
<paragraph id="8BED3694FFF277544AAEF89336C9F992" blockId="10.[151,1436,1781,1988]" pageId="10" pageNumber="75">
Other specimens examined:
<materialsCitation id="3B3A3CC9FFF277544860F89331DCF96A" ID-GBIF-Occurrence="2846550343" box="[521,1213,1854,1880]" collectorName="Dupuis" location="Orientalis. Indes" pageId="10" pageNumber="75" specimenCount="1">
<specimenCount id="9D54FD1DFFF277544860F893375BF96A" box="[521,570,1854,1880]" pageId="10" pageNumber="75" type="generic">1ex.</specimenCount>
, “
<location id="8E8D604FFFF27754483BF8933673F96A" LSID="urn:lsid:plazi:treatment:03FB8782FFF277524AFEF9A4301DFCFC:8E8D604FFFF27754483BF8933673F96A" box="[594,786,1854,1880]" name="Orientalis. Indes" pageId="10" pageNumber="75">Orientalis. Indes</location>
, Ex Musaeo Mniszech”, coll.
<collectorName id="26A75342FFF277544E00F89331DCF96A" box="[1129,1213,1854,1880]" pageId="10" pageNumber="75">Dupuis</collectorName>
</materialsCitation>
,
<materialsCitation id="3B3A3CC9FFF277544EA3F89330E3F96A" ID-GBIF-Occurrence="2846550348" box="[1226,1410,1854,1880]" collectionCode="MNHN" collectorName="Paris" pageId="10" pageNumber="75" specimenCount="1">
in
<collectionCode id="ED43AE51FFF277544E81F893305FF96A" box="[1256,1342,1854,1880]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="75">MNHN</collectionCode>
<collectorName id="26A75342FFF277544F2FF89330E3F96A" box="[1350,1410,1854,1880]" pageId="10" pageNumber="75">Paris</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3A3CC9FFF277544FE7F893317FF94E" ID-GBIF-Occurrence="2846550301" collectingDate="2017-09-13" collectorName="K. Ashly" country="India" location="Kalpetta" pageId="10" pageNumber="75" specimenCount="2" stateProvince="Kerala">
<specimenCount id="9D54FD1DFFF277544FE7F89335A7F94E" pageId="10" pageNumber="75" type="generic">2 exs.</specimenCount>
,
<collectingCountry id="F3457604FFF277544AB9F8CF346FF94E" box="[208,270,1890,1916]" name="India" pageId="10" pageNumber="75">India</collectingCountry>
:
<collectingRegion id="4996F876FFF277544B71F8CF3406F94E" box="[280,359,1890,1916]" country="India" name="Kerala" pageId="10" pageNumber="75">Kerala</collectingRegion>
:
<location id="8E8D604FFFF277544B19F8CF34B3F94E" LSID="urn:lsid:plazi:treatment:03FB8782FFF277524AFEF9A4301DFCFC:8E8D604FFFF277544B19F8CF34B3F94E" box="[368,466,1890,1916]" country="India" name="Kalpetta" pageId="10" pageNumber="75" stateProvince="Kerala">Kalpetta</location>
,
<date id="FFEC1054FFF277544BB5F8CF373FF94E" box="[476,606,1890,1916]" pageId="10" pageNumber="75" value="2017-09-13">
<collectingDate id="EFA8E9BCFFF277544BB5F8CF373FF94E" box="[476,606,1890,1916]" pageId="10" pageNumber="75" value="2017-09-13">13.IX.2017</collectingDate>
</date>
,
<collectingMethod id="52134E83FFF277544801F8CF37BEF94E" box="[616,735,1890,1916]" pageId="10" pageNumber="75">Pitfall trap</collectingMethod>
(paddy field), coll.
<collectorName id="26A75342FFF2775449D1F8CF317FF94E" box="[952,1054,1890,1916]" pageId="10" pageNumber="75">K. Ashly</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3A3CC9FFF277544E40F8CF3470F992" ID-GBIF-Occurrence="2846550326" collectingDate="2017-10-16" country="India" location="Coimbatore" pageId="10" pageNumber="75" specimenCount="7">
<specimenCount id="9D54FD1DFFF277544E40F8CF3108F94E" box="[1065,1129,1890,1916]" pageId="10" pageNumber="75" type="generic">7 exs.</specimenCount>
<collectingCountry id="F3457604FFF277544E06F8CF31CCF94E" box="[1135,1197,1890,1916]" name="India" pageId="10" pageNumber="75">India</collectingCountry>
:
<location id="8E8D604FFFF277544EDFF8CF305EF94E" LSID="urn:lsid:plazi:treatment:03FB8782FFF277524AFEF9A4301DFCFC:8E8D604FFFF277544EDFF8CF305EF94E" box="[1206,1343,1890,1916]" country="India" name="Coimbatore" pageId="10" pageNumber="75">Coimbatore</location>
: TNAU,
<date id="FFEC1054FFF277544AFEF82B3470F992" box="[151,273,1926,1952]" pageId="10" pageNumber="75" value="2017-10-16">
<collectingDate id="EFA8E9BCFFF277544AFEF82B3470F992" box="[151,273,1926,1952]" pageId="10" pageNumber="75" value="2017-10-16">16.X.2017</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3A3CC9FFF277544B75F82B36C2F992" ID-GBIF-Occurrence="2846550363" box="[284,931,1926,1952]" collectorName="B. Horne" country="India" determinerName="S. V. Akhil" location="Kutch" municipality="Gujrat" pageId="10" pageNumber="75" specimenCount="1">
<specimenCount id="9D54FD1DFFF277544B75F82B342DF992" box="[284,332,1926,1952]" pageId="10" pageNumber="75" type="generic">1ex.</specimenCount>
<collectingCountry id="F3457604FFF277544B34F82B34FCF992" box="[349,413,1926,1952]" name="India" pageId="10" pageNumber="75">India</collectingCountry>
:
<collectingMunicipality id="6B89ACEEFFF277544BCEF82B3492F992" box="[423,499,1926,1952]" pageId="10" pageNumber="75">Gujrat</collectingMunicipality>
:
<location id="8E8D604FFFF277544B97F82B3724F992" LSID="urn:lsid:plazi:treatment:03FB8782FFF277524AFEF9A4301DFCFC:8E8D604FFFF277544B97F82B3724F992" box="[510,581,1926,1952]" country="India" municipality="Gujrat" name="Kutch" pageId="10" pageNumber="75">Kutch</location>
, coll.
<collectorName id="26A75342FFF2775448E1F82B378AF992" box="[648,747,1926,1952]" pageId="10" pageNumber="75">B.Horne</collectorName>
, det.
<determinerName id="7E31DA50FFF27754494FF82B36F8F992" box="[806,921,1926,1952]" pageId="10" pageNumber="75">S.V.Akhil</determinerName>
</materialsCitation>
.
</paragraph>
<paragraph id="8BED3694FFF277544AAEF8043700F9F6" blockId="10.[151,1436,1781,1988]" box="[199,609,1961,1988]" pageId="10" pageNumber="75">
<emphasis id="B926EA86FFF277544AAEF8043438F9F6" bold="true" box="[199,345,1961,1988]" pageId="10" pageNumber="75">Description.</emphasis>
Length: 15.516.0 mm
</paragraph>
<caption id="DF2D661CFFF377554AFEF8E631B3F9E2" ID-DOI="http://doi.org/10.5281/zenodo.3993615" ID-Zenodo-Dep="3993615" httpUri="https://zenodo.org/record/3993615/files/figure.png" pageId="11" pageNumber="76" startId="11.[151,250,1867,1892]" targetBox="[220,1413,191,1818]" targetPageId="11">
<paragraph id="8BED3694FFF377554AFEF8E631B3F9E2" blockId="11.[151,1437,1867,2000]" pageId="11" pageNumber="76">
<emphasis id="B926EA86FFF377554AFEF8E63473F956" bold="true" box="[151,274,1867,1892]" pageId="11" pageNumber="76">FIGURE 2.</emphasis>
Dorsal habitus of
<taxonomicName id="4C524D17FFF377554BA2F8E1373AF956" authorityName="Solier" authorityYear="1833" box="[459,603,1868,1892]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="76" phylum="Arthropoda" rank="genus">
<emphasis id="B926EA86FFF377554BA2F8E1373AF956" box="[459,603,1868,1892]" italics="true" pageId="11" pageNumber="76">Pheropsophus</emphasis>
</taxonomicName>
from India (cont.)A)
<taxonomicName id="4C524D17FFF37755492BF8E1315EF956" authorityName="Chaudoir" authorityYear="1850" box="[834,1087,1868,1892]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="lissoderus">
<emphasis id="B926EA86FFF37755492BF8E1315EF956" box="[834,1087,1868,1892]" italics="true" pageId="11" pageNumber="76">Pheropsophus lissoderus</emphasis>
</taxonomicName>
(Holotype, Tibet, MNHN), B)
<taxonomicName id="4C524D17FFF377554FE3F8E1358AF9BA" authorityName="Bates" authorityYear="1892" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="nanodes">
<emphasis id="B926EA86FFF377554FE3F8E1358AF9BA" italics="true" pageId="11" pageNumber="76">P. nanodes</emphasis>
</taxonomicName>
(Verified specimen, S.E.Tibet, BMNH), C)
<taxonomicName id="4C524D17FFF3775548A9F8DD3656F9BA" authorityName="Chaudoir" authorityYear="1876" box="[704,823,1904,1928]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="picicollis">
<emphasis id="B926EA86FFF3775548A9F8DD3656F9BA" box="[704,823,1904,1928]" italics="true" pageId="11" pageNumber="76">P. picicollis</emphasis>
</taxonomicName>
(Syntype, Siam (now Thailand) MNHN), D)
<taxonomicName id="4C524D17FFF377554F73F8DD30FDF9BA" authorityName="Arrow" authorityYear="1901" box="[1306,1436,1904,1928]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="nigricollis">
<emphasis id="B926EA86FFF377554F73F8DD30FDF9BA" box="[1306,1436,1904,1928]" italics="true" pageId="11" pageNumber="76">P. nigricollis</emphasis>
</taxonomicName>
(Holotype, Bangalore, BMNH), E)
<taxonomicName id="4C524D17FFF377554864F83937EEF99E" baseAuthorityName="Macleay" baseAuthorityYear="1825" box="[525,655,1940,1964]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="occipitalis">
<emphasis id="B926EA86FFF377554864F83937EEF99E" box="[525,655,1940,1964]" italics="true" pageId="11" pageNumber="76">P. occipitalis</emphasis>
</taxonomicName>
(Verified specimen, China, BMNH), F)
<taxonomicName id="4C524D17FFF377554E5FF83931D4F99E" authorityName="Heller H. E. Andrewes" authorityYear="1903" box="[1078,1205,1940,1964]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="prophylax">
<emphasis id="B926EA86FFF377554E5FF83931D4F99E" box="[1078,1205,1940,1964]" italics="true" pageId="11" pageNumber="76">P. prophylax</emphasis>
</taxonomicName>
(Holotype, Manipur, BMNH), G)
<taxonomicName id="4C524D17FFF377554B70F81534FEF9E2" authorityName="Andrewes" authorityYear="1923" box="[281,415,1976,2000]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="76" phylum="Arthropoda" rank="species" species="scythropus">
<emphasis id="B926EA86FFF377554B70F81534FEF9E2" box="[281,415,1976,2000]" italics="true" pageId="11" pageNumber="76">P. scythropus</emphasis>
</taxonomicName>
(Holotype, Gopaldhara, BMNH), H)
<taxonomicName id="4C524D17FFF377554946F8153141F9E2" authorityName="ANDREWES" authorityYear="1923" box="[815,1056,1976,2000]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="76" phylum="Arthropoda" rank="subSpecies" species="scythropus" subSpecies="dissolutus">
<emphasis id="B926EA86FFF377554946F8153141F9E2" box="[815,1056,1976,2000]" italics="true" pageId="11" pageNumber="76">P. scythropus dissolutus</emphasis>
</taxonomicName>
(Laos, BMNH)
</paragraph>
</caption>
<paragraph id="8BED3694FFF477524AAEFF3531C3FEC8" blockId="12.[151,1437,152,719]" pageId="12" pageNumber="77">
<emphasis id="B926EA86FFF477524AAEFF353445FE81" bold="true" box="[199,292,152,179]" pageId="12" pageNumber="77">Colour:</emphasis>
Head and pronotum reddish brown; head with labrum and edges of clypeus reddish yellow coloured, rest reddish brown; pronotum with lateral margin black; antennal segments 14 light reddish yellow, segments 511 dark brown; elytra black; legs reddish yellow with large black spot on the apex of the femur.
</paragraph>
<paragraph id="8BED3694FFF477524AAEFEA836B6FF2C" blockId="12.[151,1437,152,719]" box="[199,983,260,287]" pageId="12" pageNumber="77">
<emphasis id="B926EA86FFF477524AAEFEA83471FF2D" bold="true" box="[199,272,261,287]" pageId="12" pageNumber="77">Head:</emphasis>
Narrower than pronotum; disc smooth; eyes highly protruding.
</paragraph>
<paragraph id="8BED3694FFF477524AAEFE8434D4FF54" blockId="12.[151,1437,152,719]" pageId="12" pageNumber="77">
<emphasis id="B926EA86FFF477524AAEFE843428FF71" bold="true" box="[199,329,297,323]" pageId="12" pageNumber="77">Pronotum:</emphasis>
Wider than the head, as long as wide, square shaped; anterior margin truncated squarely; disc smooth without wrinkles.
</paragraph>
<paragraph id="8BED3694FFF477524AAEFEDC3038FFE0" blockId="12.[151,1437,152,719]" pageId="12" pageNumber="77">
<emphasis id="B926EA86FFF477524AAEFEDC347CFFB9" bold="true" box="[199,285,369,395]" pageId="12" pageNumber="77">Elytra:</emphasis>
Subparallel, narrowed towards the base, elytral base and apex squared; humerus straight, prominent; the median transverse band dagger shaped, transverse, margins strongly serrated, dilated near the outer edge that it does not reach, and truncated squarely by the 2
<superScript id="7C279BDCFFF4775248CFFE1A37D7FFF7" attach="left" box="[678,694,439,453]" fontSize="6" pageId="12" pageNumber="77">nd</superScript>
striation; apical yellow band broader covering the intervals.
</paragraph>
<paragraph id="8BED3694FFF477524AAEFE7137D6FCB5" blockId="12.[151,1437,152,719]" pageId="12" pageNumber="77">
<emphasis id="B926EA86FFF477524AAEFE713764FFC5" bold="true" box="[199,517,476,503]" pageId="12" pageNumber="77">Geographical distribution.</emphasis>
<collectingCountry id="F3457604FFF477524864FE703700FFC5" box="[525,609,477,503]" name="India" pageId="12" pageNumber="77">INDIA</collectingCountry>
:
<collectingRegion id="4996F876FFF477524803FE713621FFC4" box="[618,832,476,502]" country="India" name="Arunachal Pradesh" pageId="12" pageNumber="77">Arunachal Pradesh</collectingRegion>
(
<bibRefCitation id="EFC34B65FFF477524926FE713696FFC5" author="Hrdlicka, J." box="[847,1015,476,503]" pageId="12" pageNumber="77" pagination="471 - 480" refId="ref19780" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden / Boston, 471 - 480." type="book chapter" year="2017">Hrdlička 2017</bibRefCitation>
);
<collectingRegion id="4996F876FFF477524E62FE7131B8FFC4" box="[1035,1241,476,502]" country="India" name="Himachal Pradesh" pageId="12" pageNumber="77">Himachal Pradesh</collectingRegion>
(
<bibRefCitation id="EFC34B65FFF477524E8EFE7130F1FFC5" author="Hrdlicka, J." box="[1255,1424,476,503]" pageId="12" pageNumber="77" pagination="471 - 480" refId="ref19780" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden / Boston, 471 - 480." type="book chapter" year="2017">Hrdlička 2017</bibRefCitation>
); Kashmir (
<bibRefCitation id="EFC34B65FFF477524B6CFDAD34CDFC29" author="Hrdlicka, J." box="[261,428,512,539]" pageId="12" pageNumber="77" pagination="471 - 480" refId="ref19780" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden / Boston, 471 - 480." type="book chapter" year="2017">Hrdlička 2017</bibRefCitation>
); Bengal: Darjiling (
<bibRefCitation id="EFC34B65FFF4775248E7FDAD3654FC29" author="Hrdlicka, J." box="[654,821,512,539]" pageId="12" pageNumber="77" pagination="471 - 480" refId="ref19780" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden / Boston, 471 - 480." type="book chapter" year="2017">Hrdlička 2017</bibRefCitation>
);
<collectingRegion id="4996F876FFF477524921FDAD36B3FC28" box="[840,978,512,538]" country="India" name="Uttarakhand" pageId="12" pageNumber="77">Uttarakhand</collectingRegion>
(
<bibRefCitation id="EFC34B65FFF4775249B6FDAD31E6FC29" author="Hrdlicka, J." box="[991,1159,512,539]" pageId="12" pageNumber="77" pagination="471 - 480" refId="ref19780" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden / Boston, 471 - 480." type="book chapter" year="2017">Hrdlička 2017</bibRefCitation>
); Gujrat: Kutch (New re- cord);
<collectingRegion id="4996F876FFF477524A89FD893409FC0C" box="[224,360,548,574]" country="India" name="Tamil Nadu" pageId="12" pageNumber="77">Tamil Nadu</collectingRegion>
: Coimbatore (New record);
<collectingRegion id="4996F876FFF4775248F7FD89378CFC0C" box="[670,749,548,574]" country="India" name="Kerala" pageId="12" pageNumber="77">Kerala</collectingRegion>
: Kalpetta (New record);
<collectingCountry id="F3457604FFF477524996FD8831F6FC0D" box="[1023,1175,549,575]" name="Sri Lanka" pageId="12" pageNumber="77">SRI LANKA</collectingCountry>
: Kandy and Peradeniya (
<bibRefCitation id="EFC34B65FFF477524AF6FDE53439FC51" author="Andrewes, H. E." box="[159,344,584,611]" pageId="12" pageNumber="77" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes 1930</bibRefCitation>
);
<collectingCountry id="F3457604FFF477524B06FDE43480FC51" box="[367,481,585,611]" name="Bhutan" pageId="12" pageNumber="77">BHUTAN</collectingCountry>
(
<bibRefCitation id="EFC34B65FFF477524B9BFDE537FCFC51" author="Hrdlicka, J." box="[498,669,584,611]" pageId="12" pageNumber="77" pagination="471 - 480" refId="ref19780" refString="Hrdlicka, J. (2017) Brachininae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata-Myxophaga-Adephaga. Revised and Updated Edition. Brill, Leiden / Boston, 471 - 480." type="book chapter" year="2017">Hrdlička 2017</bibRefCitation>
).
<collectingCountry id="F3457604FFF4775248DBFDE4366EFC51" box="[690,783,585,611]" name="China" pageId="12" pageNumber="77">CHINA</collectingCountry>
:
<collectingRegion id="4996F876FFF477524972FDE53634FC50" box="[795,853,584,610]" country="China" name="Xizang" pageId="12" pageNumber="77">Tibet</collectingRegion>
(
<bibRefCitation id="EFC34B65FFF47752490EFDE53177FC51" author="Chaudoir, M." box="[871,1046,584,611]" pageId="12" pageNumber="77" pagination="62 - 206" refId="ref18853" refString="Chaudoir, M. (1850) Supplement a la faune des carabiques de la Russie. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 23 (3), 62 - 206." type="journal article" year="1850">Chaudoir 1850</bibRefCitation>
);
<collectingCountry id="F3457604FFF477524E44FDE431D6FC51" box="[1069,1207,585,611]" name="Pakistan" pageId="12" pageNumber="77">PAKISTAN</collectingCountry>
: Islamabad.
<collectingRegion id="4996F876FFF477524F21FDE53452FCB4" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="12" pageNumber="77">Khyber Pakhtunkhwa</collectingRegion>
. Muzaffarabad (
<bibRefCitation id="EFC34B65FFF477524B80FDC137CDFCB5" author="Ullah, M. &amp; Naeem, M. &amp; Mahmood, K. &amp; Rafi, M. A." box="[489,684,620,647]" pageId="12" pageNumber="77" pagination="173 - 184" refId="ref20904" refString="Ullah, M., Naeem, M., Mahmood, K. &amp; Rafi, M. A. (2017) Faunistic studies of the tribe Brachinini (Carabidae: Coleoptera) from northern Pakistan. Zootaxa, 4232 (2), 173 - 184. https: // doi. org / 10.11646 / zootaxa. 4232.2.2" type="journal article" year="2017">
Ullah
<emphasis id="B926EA86FFF477524859FDC03709FCB4" box="[560,616,620,646]" italics="true" pageId="12" pageNumber="77">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFF477524AAEFD3C301DFCFC" pageId="12" pageNumber="77" type="discussion">
<paragraph id="8BED3694FFF477524AAEFD3C301DFCFC" blockId="12.[151,1437,152,719]" pageId="12" pageNumber="77">
<emphasis id="B926EA86FFF477524AAEFD3C345AFC99" bold="true" box="[199,315,657,683]" pageId="12" pageNumber="77">Remarks.</emphasis>
Very similar to
<taxonomicName id="4C524D17FFF477524B9EFD3C370AFC98" authorityName="Bates" authorityYear="1886" box="[503,619,657,682]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="77" phylum="Arthropoda" rank="species" species="catoirei">
<emphasis id="B926EA86FFF477524B9EFD3C370AFC98" box="[503,619,657,682]" italics="true" pageId="12" pageNumber="77">P. catoirei</emphasis>
</taxonomicName>
, differs by the shape of pronotum which is as wide as long and square shaped, not narrowing posteriorly; elytral median transverse band is more transverse, more serrated at its edges.
</paragraph>
</subSubSection>
</treatment>
</document>