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<document id="983F2A08BE4E692EE7874C93E54176FC" ID-DOI="10.11646/zootaxa.5563.1.17" ID-ISSN="1175-5326" ID-Zenodo-Dep="14595935" ID-ZooBank="08458DF2-248E-421D-9D14-5937E53DD55B" IM.bibliography_approvedBy="jonas" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="jonas" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="jonas" checkinTime="1735932756273" checkinUser="plazi" docAuthor="Ríos-Tamayo, Duniesky &amp; Goloboff, Pablo A." docDate="2025" docId="03FB65461A1E7A18FF1ADCF8CDB47685" docLanguage="en" docName="zootaxa.5563.1.17.pdf" docOrigin="Zootaxa 5563 (1)" docSource="https://doi.org/10.11646/zootaxa.5563.1.17" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Actinopus chilikuti Ríos-Tamayo &amp; Goloboff, 2025, sp. nov." docType="treatment" docVersion="4" lastPageNumber="260" masterDocId="FFC21D3E1A1F7A1DFF8DDA5BCE0B7224" masterDocTitle="A new trapdoor spider of the genus Actinopus (Araneae: Mygalomorphae: Actinopodidae) from Argentina" masterLastPageNumber="263" masterPageNumber="255" pageNumber="256" updateTime="1737662141028" updateUser="jonas" zenodo-license-document="CLOSED">
<mods:mods id="C81B7E659CA5BB1A7DB6FA163E5143AD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="31FAFD8DFA4D4C056D9AF24BBE22B727">
<mods:title id="723F6A1C71304F76CD0A4319339DB39C">A new trapdoor spider of the genus Actinopus (Araneae: Mygalomorphae: Actinopodidae) from Argentina</mods:title>
</mods:titleInfo>
<mods:name id="1E37E5F7693B52FD244B672E6B39F58F" type="personal">
<mods:role id="75C8E7EAA2A6825FA35243AB636323FE">
<mods:roleTerm id="6A1AEEA41A35A1BD153DD1B8CB159D53">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="77F051546573FCC5019AAB9E557ECC3F">Ríos-Tamayo, Duniesky</mods:namePart>
<mods:affiliation id="251EDC394CA192602AF1DF9D1A5A89DD">Instituto de Invertebrados, Fundación Miguel Lillo. Miguel Lillo 251, CP 4000. San Miguel de Tucumán, Argentina &amp; Unidad Ejecutora Lillo (Fundación Miguel Lillo-CONICET). Miguel Lillo 251, CP 4000. San Miguel de Tucumán, Argentina</mods:affiliation>
</mods:name>
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<mods:role id="570B983F56320ED3F4AA51F07496FE25">
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</mods:role>
<mods:namePart id="47609ECC0C5973BEBC061569E430322D">Goloboff, Pablo A.</mods:namePart>
<mods:affiliation id="43E5B0DAC2EA70A799361239BC466E7F">Unidad Ejecutora Lillo (Fundación Miguel Lillo-CONICET). Miguel Lillo 251, CP 4000. San Miguel de Tucumán, Argentina</mods:affiliation>
</mods:name>
<mods:typeOfResource id="45CD39F443FC874E03083F379B464811">text</mods:typeOfResource>
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<mods:title id="ECA3D943CEC136607FA657DB5737BD12">Zootaxa</mods:title>
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<mods:part id="649B3AEAA25BF7A1159ABE469A2B851D">
<mods:date id="4EB7FC5BF5C192E241B4CEF344558503">2025</mods:date>
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<mods:number id="64629B3D020902D5733D9952FCCE0CA2">2025-01-03</mods:number>
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<mods:number id="2D7BCD9E80B63DC4307FA0A050430710">5563</mods:number>
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<mods:url id="0648A7CAE0E30E2340DE044358F56627">https://doi.org/10.11646/zootaxa.5563.1.17</mods:url>
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<mods:classification id="25587A01FAAE066C54754294DDE9AC38">journal article</mods:classification>
<mods:identifier id="FEFC2B1B927B514D95CDBEC088509403" type="DOI">10.11646/zootaxa.5563.1.17</mods:identifier>
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<treatment id="03FB65461A1E7A18FF1ADCF8CDB47685" ID-DOI="http://doi.org/10.5281/zenodo.14596679" ID-Zenodo-Dep="14596679" LSID="urn:lsid:plazi:treatment:03FB65461A1E7A18FF1ADCF8CDB47685" httpUri="http://treatment.plazi.org/id/03FB65461A1E7A18FF1ADCF8CDB47685" lastPageId="5" lastPageNumber="260" pageId="1" pageNumber="256">
<subSubSection id="C34887DB1A1E7A1CFF1ADCF8CFE87499" box="[151,483,1698,1725]" pageId="1" pageNumber="256" type="nomenclature">
<paragraph id="8BEDD4501A1E7A1CFF1ADCF8CFE87499" blockId="1.[151,483,1698,1761]" box="[151,483,1698,1725]" pageId="1" pageNumber="256">
<heading id="D0A5633C1A1E7A1CFF1ADCF8CFE87499" bold="true" box="[151,483,1698,1725]" fontSize="11" level="1" pageId="1" pageNumber="256" reason="1">
<emphasis id="B92608421A1E7A1CFF1ADCF8CFE87499" bold="true" box="[151,483,1698,1725]" pageId="1" pageNumber="256">
<taxonomicName id="4C52AFD31A1E7A1CFF1ADCF8CF757499" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2025" box="[151,382,1698,1725]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="chilikuti" status="sp. nov.">
<emphasis id="B92608421A1E7A1CFF1ADCF8CF757499" bold="true" box="[151,382,1698,1725]" italics="true" pageId="1" pageNumber="256">Actinopus chilikuti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A215B5391A1E7A1CFE08DCF8CFE87499" box="[389,483,1699,1725]" pageId="1" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34887DB1A1E7A1CFF1ADC9CCF0074C5" box="[151,267,1734,1761]" pageId="1" pageNumber="256" type="description">
<paragraph id="8BEDD4501A1E7A1CFF1ADC9CCF0074C5" blockId="1.[151,483,1698,1761]" box="[151,267,1734,1761]" pageId="1" pageNumber="256">
(
<figureCitation id="1369C8D51A1E7A1CFF12DC9CCF0974C5" box="[159,258,1734,1761]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,250,1709,1733]" captionStartId-1="3.[151,250,1011,1035]" captionStartId-2="4.[151,250,1746,1770]" captionStartId-3="5.[151,250,1898,1922]" captionStartId-4="6.[151,250,914,938]" captionTargetBox-0="[151,1436,181,1685]" captionTargetBox-1="[152,1435,181,987]" captionTargetBox-2="[151,1436,181,1723]" captionTargetBox-3="[189,1399,1242,1872]" captionTargetBox-4="[166,1419,185,882]" captionTargetId-0="figure-21@2.[151,1436,181,1685]" captionTargetId-1="figure-21@3.[152,1435,181,987]" captionTargetId-2="figure-21@4.[151,1436,181,1723]" captionTargetId-3="figure-542@5.[176,1412,1242,1872]" captionTargetId-4="figure-21@6.[162,1425,181,890]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Actinopus chilikuti sp. nov., male holotype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, ocular pattern, dorsal view; D, right tibia II, retrolateral view; E–F, abdomen; E, dorsal view; F, ventral view. Scale bars: 1 mm." captionText-1="FIGURE 2. Actinopus chilikuti sp. nov., male holotype. A, right palp, retrolateral view; B, right palp, prolateral view; C–F, copulatory bulb; C, dorsal view; D, ventral view; E, prolateral view; F, retrolateral view. Scale bars: 1 mm." captionText-2="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." captionText-3="FIGURE 4. Copulatory bulbs with type I morphology inArgentina, dorsal views. A, Actinopus clavero Ríos-Tamayo &amp; Goloboff 2018; B, Actinopus reycali Ríos-Tamayo &amp; Goloboff 2018; C, Actinopus chilikuti sp. nov.; D, Actinopus septemtrionalis Ríos- Tamayo &amp; Goloboff 2018. Arrows indicate the paraembolic apophysis (PA). Scale bars: 1 mm." captionText-4="FIGURE 5. Known distribution of the species with Type I morphology in Argentina." figureDoi-0="http://doi.org/10.5281/zenodo.14595937" figureDoi-1="http://doi.org/10.5281/zenodo.14595939" figureDoi-2="http://doi.org/10.5281/zenodo.14595941" figureDoi-3="http://doi.org/10.5281/zenodo.14595943" figureDoi-4="http://doi.org/10.5281/zenodo.14595945" httpUri-0="https://zenodo.org/record/14595937/files/figure.png" httpUri-1="https://zenodo.org/record/14595939/files/figure.png" httpUri-2="https://zenodo.org/record/14595941/files/figure.png" httpUri-3="https://zenodo.org/record/14595943/files/figure.png" httpUri-4="https://zenodo.org/record/14595945/files/figure.png" pageId="1" pageNumber="256">Figs 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34887DB1A1E7A1CFF1ADD50CB5C7507" box="[151,1367,1803,1827]" pageId="1" pageNumber="256" type="reference_group">
<paragraph id="8BEDD4501A1E7A1CFF1ADD50CB5C7507" blockId="1.[151,1367,1803,1827]" box="[151,1367,1803,1827]" pageId="1" pageNumber="256">
<treatmentCitationGroup id="AB42F37E1A1E7A1CFF1ADD50CB5C7507" box="[151,1367,1803,1827]" pageId="1" pageNumber="256">
<treatmentCitation id="0AF3F2411A1E7A1CFF1ADD50CD1B7507" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[151,784,1803,1827]" httpUri="http://treatment.plazi.org/id/C22D980AFFE6F37572DFFF35D68FFA76" page="26" pageId="1" pageNumber="256" year="2018">
<taxonomicName id="4C52AFD31A1E7A1CFF1ADD50CD1B7507" ID-CoL="64PLS" authority="Rios-Tamayo &amp; Goloboff, 2018: 26" authorityName="Rios-Tamayo &amp; Goloboff" authorityPageNumber="26" authorityYear="2018" box="[151,784,1803,1827]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="256" phylum="Arthropoda" rank="species" species="septemtrionalis">
<emphasis id="B92608421A1E7A1CFF1ADD50CF947507" box="[151,415,1803,1827]" italics="true" pageId="1" pageNumber="256">Actinopus septemtrionalis</emphasis>
<bibRefCitation id="EFC3A9A11A1E7A1CFE28DD50CD1B7507" author="Rios-Tamayo, D. &amp; Goloboff, P. A." box="[421,784,1803,1827]" pageId="1" pageNumber="256" pagination="1 - 83" refId="ref6268" refString="Rios-Tamayo, D. &amp; Goloboff, P. A. (2018) Taxonomic revision and morphology of the trapdoor spider genus Actinopus (Mygalomorphae: Actinopodidae) in Argentina. Bulletin of the American Museum of Natural History, 419, 1 - 83." type="journal article" year="2018">Ríos-Tamayo &amp; Goloboff, 2018: 26</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(only female from
<collectingRegion id="49961AB21A1E7A1CFC58DD57CA857507" box="[981,1166,1803,1827]" country="Argentina" name="Formosa" pageId="1" pageNumber="256">Formosa province</collectingRegion>
). Misidentification.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34887DB1A1E7A1FFF1ADD0ACD0675E5" lastPageId="2" lastPageNumber="257" pageId="1" pageNumber="256" type="materials_examined">
<paragraph id="8BEDD4501A1E7A1CFF1ADD0ACB3975F3" blockId="1.[151,1437,1873,2008]" pageId="1" pageNumber="256">
<materialsCitation id="3B3ADE0D1A1E7A1CFF1ADD0ACB9675AB" collectingDate="2023-11-23" collectorName="Rios-Tamayo. D &amp; P. Goloboff" country="Argentina" county="Prov" latitude="-25.972542" location="Parque Nacional &quot; Copo &quot;" longLatPrecision="1" longitude="-61.958523" municipality="Parque Nacional" pageId="1" pageNumber="256" specimenCode="FML-ARAN 02273" specimenCount="1" specimenCount-male="1" stateProvince="Santiago del Estero" typeStatus="holotype">
<emphasis id="B92608421A1E7A1CFF1ADD0ACF00754F" bold="true" box="[151,267,1873,1899]" pageId="1" pageNumber="256">
<typeStatus id="54E96AF21A1E7A1CFF1ADD0ACF0D754F" box="[151,262,1873,1899]" pageId="1" pageNumber="256" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F34594C01A1E7A1CFE99DD0ACFB7754F" box="[276,444,1873,1899]" name="Argentina" pageId="1" pageNumber="256">ARGENTINA</collectingCountry>
:
<collectingRegion id="49961AB21A1E7A1CFE47DD0ACCA77548" box="[458,684,1873,1900]" country="Argentina" name="Santiago del Estero" pageId="1" pageNumber="256">Santiago del Estero</collectingRegion>
Prov:
<specimenCount id="9D541FD91A1E7A1CFD70DD0ACD207548" box="[765,811,1873,1900]" count="1" pageId="1" pageNumber="256" type="male">1 ♂</specimenCount>
(
<specimenCode id="DBF47C2B1A1E7A1CFCB0DD0ACA2F754F" box="[829,1060,1873,1899]" pageId="1" pageNumber="256">FML-ARAN 02273</specimenCode>
) from
<location id="8E8D828B1A1E7A1CFBFADD09CB93754F" LSID="urn:lsid:plazi:treatment:03FB65461A1E7A18FF1ADCF8CDB47685:8E8D828B1A1E7A1CFBFADD09CB93754F" box="[1143,1432,1873,1900]" country="Argentina" county="Prov" latitude="-25.972542" longLatPrecision="1" longitude="-61.958523" municipality="Parque Nacional" name="Parque Nacional &quot; Copo &quot;" pageId="1" pageNumber="256" stateProvince="Santiago del Estero">Parque Nacional “Copo”</location>
, seccional
<location id="8E8D828B1A1E7A1CFE84DD2DCF8D75AB" LSID="urn:lsid:plazi:treatment:03FB65461A1E7A18FF1ADCF8CDB47685:8E8D828B1A1E7A1CFE84DD2DCF8D75AB" box="[265,390,1909,1936]" country="Argentina" county="Prov" latitude="-25.972542" longLatPrecision="1" longitude="-61.958523" municipality="Parque Nacional" name="La Portada" pageId="1" pageNumber="256" stateProvince="Santiago del Estero">La Portada</location>
(
<geoCoordinate id="EE66B2971A1E7A1CFE1ADD2ECC4B75B4" box="[407,576,1909,1936]" degrees="25" direction="south" minutes="58" orientation="latitude" pageId="1" pageNumber="256" precision="1" seconds="21.15" value="-25.972542">25°58’21.15”S</geoCoordinate>
<geoCoordinate id="EE66B2971A1E7A1CFDC4DD2ECCF475B4" box="[585,767,1909,1936]" degrees="61" direction="west" minutes="57" orientation="longitude" pageId="1" pageNumber="256" precision="1" seconds="30.68" value="-61.958523">61°57’30.68”W</geoCoordinate>
),
<date id="FFECF2901A1E7A1CFC98DD2ECDFE75AB" box="[789,1013,1909,1935]" pageId="1" pageNumber="256" value="2023-11-23">
<collectingDate id="EFA80B781A1E7A1CFC98DD2ECDFE75AB" box="[789,1013,1909,1935]" pageId="1" pageNumber="256" value="2023-11-23">23 November 2023</collectingDate>
</date>
,
<collectorName id="26A7B1861A1E7A1CFB8EDD2ECAB575B4" box="[1027,1214,1909,1936]" pageId="1" pageNumber="256">Ríos-Tamayo. D</collectorName>
&amp;
<collectorName id="26A7B1861A1E7A1CFB6BDD2DCB6275AB" box="[1254,1385,1909,1936]" pageId="1" pageNumber="256">P. Goloboff</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B3ADE0D1A1E7A1CFF1ADDC1CEC875F3" collectingDate="2023-11-24" collectorName="Rios-Tamayo. D &amp; J. C Stazonelli" country="Argentina" county="Prov" latitude="-25.972542" location="La Portada" longLatPrecision="1" longitude="-61.958523" municipality="Parque Nacional" pageId="1" pageNumber="256" specimenCode="FML-ARAN 02274" specimenCount="1" specimenCount-female="1" stateProvince="Santiago del Estero" typeStatus="paratype">
<typeStatus id="54E96AF21A1E7A1CFF1ADDC1CF037590" box="[151,264,1946,1972]" pageId="1" pageNumber="256" type="paratype">Paratypes</typeStatus>
. Same data as holotype,
<specimenCount id="9D541FD91A1E7A1CFD9ADDC2CC4B7590" box="[535,576,1945,1972]" count="1" pageId="1" pageNumber="256" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBF47C2B1A1E7A1CFDC0DDC2CD397597" box="[589,818,1945,1971]" pageId="1" pageNumber="256">FML-ARAN 02274</specimenCode>
),
<date id="FFECF2901A1E7A1CFCC9DDC2CA157597" box="[836,1054,1945,1971]" pageId="1" pageNumber="256" value="2023-11-24">
<collectingDate id="EFA80B781A1E7A1CFCC9DDC2CA157597" box="[836,1054,1945,1971]" pageId="1" pageNumber="256" value="2023-11-24">24 November 2023</collectingDate>
</date>
,
<collectorName id="26A7B1861A1E7A1CFBA5DDC2CAD47590" box="[1064,1247,1945,1972]" pageId="1" pageNumber="256">Ríos-Tamayo. D</collectorName>
&amp;
<collectorName id="26A7B1861A1E7A1CFA8FDDC2CB977597" box="[1282,1436,1945,1971]" pageId="1" pageNumber="256">J.C Stazonelli</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B3ADE0D1A1E7A1CFF42DDE6CB3975F3" box="[207,1330,1981,2008]" collectingDate="2023-11-23" collectorName="Rios-Tamayo. D &amp; J. C Stazonelli &amp; P. Goloboff" country="Argentina" county="Prov" isTailEnumeration="true" latitude="-25.972542" location="La Portada" longLatPrecision="1" longitude="-61.958523" municipality="Parque Nacional" pageId="1" pageNumber="256" specimenCode="FML-ARAN 02275" specimenCount="3" specimenCount-male="3" stateProvince="Santiago del Estero" typeStatus="paratype">
<specimenCount id="9D541FD91A1E7A1CFF42DDE6CF0575FC" box="[207,270,1981,2008]" count="3" pageId="1" pageNumber="256" type="male">3 ♂♂</specimenCount>
(
<specimenCode id="DBF47C2B1A1E7A1CFE91DDE6CC0975FC" box="[284,514,1981,2008]" pageId="1" pageNumber="256">FML-ARAN 02275</specimenCode>
),
<date id="FFECF2901A1E7A1CFD98DDE6CCFB75F3" box="[533,752,1981,2007]" pageId="1" pageNumber="256" value="2023-11-23">
<collectingDate id="EFA80B781A1E7A1CFD98DDE6CCFB75F3" box="[533,752,1981,2007]" pageId="1" pageNumber="256" value="2023-11-23">23 November 2023</collectingDate>
</date>
,
<collectorName id="26A7B1861A1E7A1CFD76DDE6CDBC75FC" box="[763,951,1981,2008]" pageId="1" pageNumber="256">Ríos-Tamayo. D</collectorName>
,
<collectorName id="26A7B1861A1E7A1CFC4FDDE6CA5675F3" box="[962,1117,1981,2007]" pageId="1" pageNumber="256">J.C Stazonelli</collectorName>
&amp;
<collectorName id="26A7B1861A1E7A1CFB0CDDE5CB0975F3" box="[1153,1282,1981,2008]" pageId="1" pageNumber="256">P. Goloboff</collectorName>
leg.
</materialsCitation>
</paragraph>
<caption id="DF2D84D81A1D7A1FFF1ADCF6CFDC7529" ID-DOI="http://doi.org/10.5281/zenodo.14595937" ID-Zenodo-Dep="14595937" httpUri="https://zenodo.org/record/14595937/files/figure.png" pageId="2" pageNumber="257" startId="2.[151,250,1709,1733]" targetBox="[151,1436,181,1685]" targetPageId="2" targetType="figure">
<paragraph id="8BEDD4501A1D7A1FFF1ADCF6CFDC7529" blockId="2.[151,1436,1709,1805]" pageId="2" pageNumber="257">
<emphasis id="B92608421A1D7A1FFF1ADCF6CF1F74E1" bold="true" box="[151,276,1709,1733]" pageId="2" pageNumber="257">FIGURE 1.</emphasis>
<taxonomicName id="4C52AFD31A1D7A1FFE97DCF5CFD174E1" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2025" box="[282,474,1709,1733]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="chilikuti" status="sp. nov.">
<emphasis id="B92608421A1D7A1FFE97DCF5CFD174E1" box="[282,474,1709,1733]" italics="true" pageId="2" pageNumber="257">Actinopus chilikuti</emphasis>
</taxonomicName>
<emphasis id="B92608421A1D7A1FFE6DDCF6CC3374E1" bold="true" box="[480,568,1709,1733]" pageId="2" pageNumber="257">
<taxonomicNameLabel id="A215B5391A1D7A1FFE6DDCF6CC3F74E1" box="[480,564,1709,1733]" pageId="2" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male holotype.
<emphasis id="B92608421A1D7A1FFD50DCF6CCFE74E1" bold="true" box="[733,757,1709,1733]" pageId="2" pageNumber="257">A,</emphasis>
cephalothorax, dorsal view, arrows indicate the presence of mites;
<emphasis id="B92608421A1D7A1FFF1ADC8ACEA574CD" bold="true" box="[151,174,1745,1769]" pageId="2" pageNumber="257">B,</emphasis>
sternum, ventral view;
<emphasis id="B92608421A1D7A1FFE2FDC8ACFB074CD" bold="true" box="[418,443,1745,1769]" pageId="2" pageNumber="257">C,</emphasis>
ocular pattern, dorsal view;
<emphasis id="B92608421A1D7A1FFD6FDC8ACCF174CD" bold="true" box="[738,762,1745,1769]" pageId="2" pageNumber="257">D,</emphasis>
right tibia II, retrolateral view;
<emphasis id="B92608421A1D7A1FFBCFDC8ACA7874CD" bold="true" box="[1090,1139,1745,1769]" pageId="2" pageNumber="257">E–F,</emphasis>
abdomen;
<emphasis id="B92608421A1D7A1FFB68DC8ACAF774CD" bold="true" box="[1253,1276,1745,1769]" pageId="2" pageNumber="257">E,</emphasis>
dorsal view;
<emphasis id="B92608421A1D7A1FFA04DC8ACB9774CD" bold="true" box="[1417,1436,1745,1769]" pageId="2" pageNumber="257">F,</emphasis>
ventral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BEDD4501A1D7A1FFF4ADD04CD0675E5" blockId="2.[151,1437,1887,1986]" pageId="2" pageNumber="257">
<emphasis id="B92608421A1D7A1FFF4ADD04CFF7755D" bold="true" box="[199,508,1887,1913]" pageId="2" pageNumber="257">Other material examined.</emphasis>
<materialsCitation id="3B3ADE0D1A1D7A1FFD8CDD3BCD0675E5" collectingDate="2010-06-27" collectingDateMax="2010-07-05" collectingDateMin="2010-06-27" collectorName="Izquierdo. M &amp; Kopuchian. C &amp; Calderon. L." country="Argentina" latitude="-24.300068" location="Reserva Natural Formosa" longLatPrecision="1" longitude="-61.801502" municipality="Seccional Rio Teuco" pageId="2" pageNumber="257" specimenCode="MACN-Ar 27307, MACN-Ar 27307-t1" specimenCount="1" specimenCount-female="1" stateProvince="Formosa">
<collectingCountry id="F34594C01A1D7A1FFD8CDD3BCCA2755E" box="[513,681,1888,1914]" name="Argentina" pageId="2" pageNumber="257">ARGENTINA</collectingCountry>
:
<collectingRegion id="49961AB21A1D7A1FFD39DD3BCD1D755E" box="[692,790,1888,1914]" country="Argentina" name="Formosa" pageId="2" pageNumber="257">Formosa</collectingRegion>
Prov:
<specimenCount id="9D541FD91A1D7A1FFCEDDD3BCD80755E" box="[864,907,1888,1914]" count="1" pageId="2" pageNumber="257" type="female">1 ♀</specimenCount>
(
<specimenCode id="DBF47C2B1A1D7A1FFC14DD3BCA6C755E" box="[921,1127,1888,1914]" pageId="2" pageNumber="257">MACN-Ar 27307</specimenCode>
), tissue sample (
<specimenCode id="DBF47C2B1A1D7A1FFA92DD3BCEF675BA" pageId="2" pageNumber="257">MACN-Ar 27307-t1</specimenCode>
) from
<location id="8E8D828B1A1D7A1FFEDDDDDFCC7375BA" LSID="urn:lsid:plazi:treatment:03FB65461A1E7A18FF1ADCF8CDB47685:8E8D828B1A1D7A1FFEDDDDDFCC7375BA" box="[336,632,1923,1950]" country="Argentina" latitude="-24.300068" longLatPrecision="1" longitude="-61.801502" municipality="Seccional Rio Teuco" name="Reserva Natural Formosa" pageId="2" pageNumber="257" stateProvince="Formosa">Reserva Natural Formosa</location>
:
<collectingMunicipality id="6B894E2A1A1D7A1FFD0BDDD8CD7E75BA" box="[646,885,1923,1950]" pageId="2" pageNumber="257">Seccional Rio Teuco</collectingMunicipality>
, (
<geoCoordinate id="EE66B2971A1D7A1FFC01DDD8CA2D75BA" box="[908,1062,1923,1950]" degrees="24" direction="south" minutes="18" orientation="latitude" pageId="2" pageNumber="257" precision="1" seconds="0.25" value="-24.300068">24°18’0.25”S</geoCoordinate>
<geoCoordinate id="EE66B2971A1D7A1FFBBDDDD8CAD375BA" box="[1072,1240,1923,1950]" degrees="61" direction="west" minutes="48" orientation="longitude" pageId="2" pageNumber="257" precision="1" seconds="5.41" value="-61.801502">61°48’5.41”W</geoCoordinate>
),
<date id="FFECF2901A1D7A1FFB63DDDFCED975E6" pageId="2" pageNumber="257" value="2010-06-27" valueMax="2010-07-05" valueMin="2010-06-27">
<collectingDate id="EFA80B781A1D7A1FFB63DDDFCED975E6" pageId="2" pageNumber="257" value="2010-06-27" valueMax="2010-07-05" valueMin="2010-06-27">27 June–5 July 2010</collectingDate>
</date>
,
<collectorName id="26A7B1861A1D7A1FFF50DDFCCF7A75E6" box="[221,369,1959,1986]" pageId="2" pageNumber="257">Izquierdo. M</collectorName>
,
<collectorName id="26A7B1861A1D7A1FFEF0DDFCCC1275E6" box="[381,537,1959,1986]" pageId="2" pageNumber="257">Kopuchian. C</collectorName>
and
<collectorName id="26A7B1861A1D7A1FFDC2DDFCCCD775E6" box="[591,732,1959,1986]" pageId="2" pageNumber="257">Calderón. L.</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF2D84D81A1C7A1EFF1AD9A8CADD760A" ID-DOI="http://doi.org/10.5281/zenodo.14595939" ID-Zenodo-Dep="14595939" httpUri="https://zenodo.org/record/14595939/files/figure.png" pageId="3" pageNumber="258" startId="3.[151,250,1011,1035]" targetBox="[152,1435,181,987]" targetPageId="3" targetType="figure">
<paragraph id="8BEDD4501A1C7A1EFF1AD9A8CADD760A" blockId="3.[151,1436,1011,1071]" pageId="3" pageNumber="258">
<emphasis id="B92608421A1C7A1EFF1AD9A8CF1F762F" bold="true" box="[151,276,1011,1035]" pageId="3" pageNumber="258">FIGURE 2.</emphasis>
<taxonomicName id="4C52AFD31A1C7A1EFE91D9A8CFD6762F" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2025" box="[284,477,1011,1035]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="chilikuti" status="sp. nov.">
<emphasis id="B92608421A1C7A1EFE91D9A8CFD6762F" box="[284,477,1011,1035]" italics="true" pageId="3" pageNumber="258">Actinopus chilikuti</emphasis>
</taxonomicName>
<emphasis id="B92608421A1C7A1EFE69D9A8CC36762F" bold="true" box="[484,573,1011,1035]" pageId="3" pageNumber="258">
<taxonomicNameLabel id="A215B5391A1C7A1EFE69D9A8CC32762F" box="[484,569,1011,1035]" pageId="3" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
male holotype.
<emphasis id="B92608421A1C7A1EFD68D9A8CCF6762F" bold="true" box="[741,765,1011,1035]" pageId="3" pageNumber="258">A,</emphasis>
right palp, retrolateral view;
<emphasis id="B92608421A1C7A1EFBA3D9A8CA4E762F" bold="true" box="[1070,1093,1011,1035]" pageId="3" pageNumber="258">B,</emphasis>
right palp, prolateral view;
<emphasis id="B92608421A1C7A1EFAEAD9A8CB9E762F" bold="true" box="[1383,1429,1011,1035]" pageId="3" pageNumber="258">C–F</emphasis>
, copulatory bulb;
<emphasis id="B92608421A1C7A1EFEC9DE4CCF56760B" bold="true" box="[324,349,1047,1071]" pageId="3" pageNumber="258">C,</emphasis>
dorsal view;
<emphasis id="B92608421A1C7A1EFE6BDE4CCFF5760B" bold="true" box="[486,510,1047,1071]" pageId="3" pageNumber="258">D,</emphasis>
ventral view;
<emphasis id="B92608421A1C7A1EFD1DDE4CCCAC760B" bold="true" box="[656,679,1047,1071]" pageId="3" pageNumber="258">E,</emphasis>
prolateral view;
<emphasis id="B92608421A1C7A1EFCD9DE4CCD63760B" bold="true" box="[852,872,1047,1071]" pageId="3" pageNumber="258">F,</emphasis>
retrolateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<subSubSection id="C34887DB1A1C7A1EFF4ADEDACB80769B" pageId="3" pageNumber="258" type="etymology">
<paragraph id="8BEDD4501A1C7A1EFF4ADEDACB80769B" blockId="3.[151,1437,1153,2044]" pageId="3" pageNumber="258">
<emphasis id="B92608421A1C7A1EFF4ADEDACF4576BF" bold="true" box="[199,334,1153,1179]" pageId="3" pageNumber="258">Etymology.</emphasis>
The specific epithet works as a noun in apposition referring to the small size of the specimens of this species. In Quechua Aymara (very commonly spoken in
<collectingRegion id="49961AB21A1C7A1EFCB2DEFECA15769B" box="[831,1054,1189,1215]" country="Argentina" name="Santiago del Estero" pageId="3" pageNumber="258">Santiago del Estero</collectingRegion>
), ch’ilikuti means “little, small”.
</paragraph>
</subSubSection>
<subSubSection id="C34887DB1A1C7A1EFF4ADE92CA6C77BC" pageId="3" pageNumber="258" type="diagnosis">
<paragraph id="8BEDD4501A1C7A1EFF4ADE92CA6C77BC" blockId="3.[151,1437,1153,2044]" pageId="3" pageNumber="258">
<emphasis id="B92608421A1C7A1EFF4ADE92CF4976C7" bold="true" box="[199,322,1225,1251]" pageId="3" pageNumber="258">Diagnosis.</emphasis>
Members of
<taxonomicName id="4C52AFD31A1C7A1EFE59DE92CCAD76C7" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2025" box="[468,678,1225,1251]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="chilikuti" status="sp. nov.">
<emphasis id="B92608421A1C7A1EFE59DE92CCAD76C7" box="[468,678,1225,1251]" italics="true" pageId="3" pageNumber="258">Actinopus chilikuti</emphasis>
</taxonomicName>
<emphasis id="B92608421A1C7A1EFD21DE92CD0F76C7" bold="true" box="[684,772,1225,1251]" pageId="3" pageNumber="258">
<taxonomicNameLabel id="A215B5391A1C7A1EFD21DE92CD0F76C7" box="[684,772,1225,1251]" pageId="3" pageNumber="258" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are small spiders. Males can be distinguished from all other species with
<typeStatus id="54E96AF21A1C7A1EFEAADEB5CF54772C" box="[295,351,1262,1288]" pageId="3" pageNumber="258">Type</typeStatus>
I morphology in
<collectingCountry id="F34594C01A1C7A1EFD92DEB6CC857723" box="[543,654,1261,1287]" name="Argentina" pageId="3" pageNumber="258">Argentina</collectingCountry>
by having a copulatory bulb with a more curved embolus (
<figureCitation id="1369C8D51A1C7A1EFAACDEB6CB937723" box="[1313,1432,1261,1287]" captionStart="FIGURE 2" captionStartId="3.[151,250,1011,1035]" captionTargetBox="[152,1435,181,987]" captionTargetId="figure-21@3.[152,1435,181,987]" captionTargetPageId="3" captionText="FIGURE 2. Actinopus chilikuti sp. nov., male holotype. A, right palp, retrolateral view; B, right palp, prolateral view; C–F, copulatory bulb; C, dorsal view; D, ventral view; E, prolateral view; F, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595939" httpUri="https://zenodo.org/record/14595939/files/figure.png" pageId="3" pageNumber="258">Figs 2C–F</figureCitation>
,
<figureCitation id="1369C8D51A1C7A1EFF1ADF4ACEB0770F" box="[151,187,1297,1323]" captionStart="FIGURE 4" captionStartId="5.[151,250,1898,1922]" captionTargetBox="[189,1399,1242,1872]" captionTargetId="figure-542@5.[176,1412,1242,1872]" captionTargetPageId="5" captionText="FIGURE 4. Copulatory bulbs with type I morphology inArgentina, dorsal views. A, Actinopus clavero Ríos-Tamayo &amp; Goloboff 2018; B, Actinopus reycali Ríos-Tamayo &amp; Goloboff 2018; C, Actinopus chilikuti sp. nov.; D, Actinopus septemtrionalis Ríos- Tamayo &amp; Goloboff 2018. Arrows indicate the paraembolic apophysis (PA). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595943" httpUri="https://zenodo.org/record/14595943/files/figure.png" pageId="3" pageNumber="258">4C</figureCitation>
), with keels well developed (similar to
<taxonomicName id="4C52AFD31A1C7A1EFDF0DF4ACCE1770F" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[637,746,1297,1323]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="reycali">
<emphasis id="B92608421A1C7A1EFDF0DF4ACCE1770F" box="[637,746,1297,1323]" italics="true" pageId="3" pageNumber="258">A. reycali</emphasis>
</taxonomicName>
) and PA developed (smaller in
<taxonomicName id="4C52AFD31A1C7A1EFBDDDF4ACAB5770F" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[1104,1214,1297,1323]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="reycali">
<emphasis id="B92608421A1C7A1EFBDDDF4ACAB5770F" box="[1104,1214,1297,1323]" italics="true" pageId="3" pageNumber="258">A. reycali</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C52AFD31A1C7A1EFB70DF4ACB7B770F" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[1277,1392,1297,1323]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="clavero">
<emphasis id="B92608421A1C7A1EFB70DF4ACB7B770F" box="[1277,1392,1297,1323]" italics="true" pageId="3" pageNumber="258">A. clavero</emphasis>
</taxonomicName>
but bigger in
<taxonomicName id="4C52AFD31A1C7A1EFE8ADF6ECFDE776B" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[263,469,1333,1359]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="septemtrionalis">
<emphasis id="B92608421A1C7A1EFE8ADF6ECFDE776B" box="[263,469,1333,1359]" italics="true" pageId="3" pageNumber="258">A. septemtrionalis</emphasis>
</taxonomicName>
,
<figureCitation id="1369C8D51A1C7A1EFE68DF6ECC2E776B" box="[485,549,1333,1359]" captionStart="FIGURE 4" captionStartId="5.[151,250,1898,1922]" captionTargetBox="[189,1399,1242,1872]" captionTargetId="figure-542@5.[176,1412,1242,1872]" captionTargetPageId="5" captionText="FIGURE 4. Copulatory bulbs with type I morphology inArgentina, dorsal views. A, Actinopus clavero Ríos-Tamayo &amp; Goloboff 2018; B, Actinopus reycali Ríos-Tamayo &amp; Goloboff 2018; C, Actinopus chilikuti sp. nov.; D, Actinopus septemtrionalis Ríos- Tamayo &amp; Goloboff 2018. Arrows indicate the paraembolic apophysis (PA). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595943" httpUri="https://zenodo.org/record/14595943/files/figure.png" pageId="3" pageNumber="258">Fig 4</figureCitation>
). Females can be differentiated from
<taxonomicName id="4C52AFD31A1C7A1EFC58DF6ECA48776B" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[981,1091,1333,1359]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="reycali">
<emphasis id="B92608421A1C7A1EFC58DF6ECA48776B" box="[981,1091,1333,1359]" italics="true" pageId="3" pageNumber="258">A. reycali</emphasis>
</taxonomicName>
and
<taxonomicName id="4C52AFD31A1C7A1EFBF3DF6ECB47776B" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[1150,1356,1333,1359]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="septemtrionalis">
<emphasis id="B92608421A1C7A1EFBF3DF6ECB47776B" box="[1150,1356,1333,1359]" italics="true" pageId="3" pageNumber="258">A. septemtrionalis</emphasis>
</taxonomicName>
by the shorter and more slender spermathecae (
<figureCitation id="1369C8D51A1C7A1EFDDBDF02CCAC7757" box="[598,679,1369,1395]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="3" pageNumber="258">Fig 3D</figureCitation>
, similar to
<taxonomicName id="4C52AFD31A1C7A1EFCACDF02CD997757" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[801,914,1369,1395]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="clavero">
<emphasis id="B92608421A1C7A1EFCACDF02CD997757" box="[801,914,1369,1395]" italics="true" pageId="3" pageNumber="258">A. clavero</emphasis>
</taxonomicName>
). It is distinguished from
<taxonomicName id="4C52AFD31A1C7A1EFB3DDF02CB2A7757" authorityName="Rios-Tamayo &amp; Goloboff" authorityYear="2018" box="[1200,1313,1369,1395]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="258" phylum="Arthropoda" rank="species" species="clavero">
<emphasis id="B92608421A1C7A1EFB3DDF02CB2A7757" box="[1200,1313,1369,1395]" italics="true" pageId="3" pageNumber="258">A. clavero</emphasis>
</taxonomicName>
by a larger number of retrolateral thorns on tibia II (30
<emphasis id="B92608421A1C7A1EFD0EDF25CC9177B3" box="[643,666,1406,1431]" italics="true" pageId="3" pageNumber="258">vs</emphasis>
20,
<figureCitation id="1369C8D51A1C7A1EFD46DF26CD1177B3" box="[715,794,1405,1431]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="3" pageNumber="258">Fig 3G</figureCitation>
) and fewer on tibia I (0
<emphasis id="B92608421A1C7A1EFBA7DF25CA4A77B3" box="[1066,1089,1406,1431]" italics="true" pageId="3" pageNumber="258">vs</emphasis>
5).
</paragraph>
</subSubSection>
<subSubSection id="C34887DB1A1C7A18FF4ADFF9CF077659" lastPageId="5" lastPageNumber="260" pageId="3" pageNumber="258" type="description">
<paragraph id="8BEDD4501A1C7A18FF4ADFF9CF827365" blockId="3.[151,1437,1153,2044]" lastBlockId="5.[151,1437,151,1185]" lastPageId="5" lastPageNumber="260" pageId="3" pageNumber="258">
<emphasis id="B92608421A1C7A1EFF4ADFF9CF58779F" box="[199,339,1442,1467]" italics="true" pageId="3" pageNumber="258">Description.</emphasis>
Male
<typeStatus id="54E96AF21A1C7A1EFE10DFFACFF4779F" box="[413,511,1441,1467]" pageId="3" pageNumber="258" type="holotype">holotype</typeStatus>
(FML-ARAN 02273). Total length: 6.61.
<emphasis id="B92608421A1C7A1EFC6EDFFACA5A779F" box="[995,1105,1441,1467]" italics="true" pageId="3" pageNumber="258">Carapace</emphasis>
(
<figureCitation id="1369C8D51A1C7A1EFBEFDFFACABD779F" box="[1122,1206,1441,1467]" captionStart="FIGURE 1" captionStartId="2.[151,250,1709,1733]" captionTargetBox="[151,1436,181,1685]" captionTargetId="figure-21@2.[151,1436,181,1685]" captionTargetPageId="2" captionText="FIGURE 1. Actinopus chilikuti sp. nov., male holotype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, ocular pattern, dorsal view; D, right tibia II, retrolateral view; E–F, abdomen; E, dorsal view; F, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595937" httpUri="https://zenodo.org/record/14595937/files/figure.png" pageId="3" pageNumber="258">Fig 1A</figureCitation>
): length 3.30, width 2.93; cephalic region: length 1.89, width 2.01; clypeus small, 7 bristles between PME, 4 behind each PME–PLE.
<emphasis id="B92608421A1C7A1EFF1ADFB1CEED7427" box="[151,230,1514,1539]" italics="true" pageId="3" pageNumber="258">Fovea:</emphasis>
length 0.44, width 0.80.
<emphasis id="B92608421A1C7A1EFD8ADFB1CD997427" box="[519,914,1513,1539]" italics="true" pageId="3" pageNumber="258">Eyes, diameters and interdistances</emphasis>
(
<figureCitation id="1369C8D51A1C7A1EFC2FDFB2CDFE7427" box="[930,1013,1513,1539]" captionStart="FIGURE 1" captionStartId="2.[151,250,1709,1733]" captionTargetBox="[151,1436,181,1685]" captionTargetId="figure-21@2.[151,1436,181,1685]" captionTargetPageId="2" captionText="FIGURE 1. Actinopus chilikuti sp. nov., male holotype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, ocular pattern, dorsal view; D, right tibia II, retrolateral view; E–F, abdomen; E, dorsal view; F, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595937" httpUri="https://zenodo.org/record/14595937/files/figure.png" pageId="3" pageNumber="258">Fig 1C</figureCitation>
): AME 0.19 (large), ALE 0.27, PME 0.14, PLE 0.16, ocular group length 0.54, anterior width 1.32, 1.23 posterior width; AME–ALE 0.20, PME–PLE 0.06, AME–PME 0.19, ALE–PLE 0.19, AME–AME 0.09, PME-PME 0,74.
<emphasis id="B92608421A1C7A1EFB8BDC6ACA81746F" box="[1030,1162,1585,1611]" italics="true" pageId="3" pageNumber="258">Chelicerae:</emphasis>
apex with 7 recumbent and elongated bristles over fang base. Rastellum with 4 marginal blunt spines and 2 small dorsal ones. Cheliceral furrow with 4 large promarginal and 5 retromarginal teeth and 4 denticles, near promarginal teeth.
<emphasis id="B92608421A1C7A1EFB57DC22CB3B74B7" box="[1242,1328,1657,1683]" italics="true" pageId="3" pageNumber="258">Labium</emphasis>
(
<figureCitation id="1369C8D51A1C7A1EFAB0DC22CB8674B7" box="[1341,1421,1657,1683]" captionStart="FIGURE 1" captionStartId="2.[151,250,1709,1733]" captionTargetBox="[151,1436,181,1685]" captionTargetId="figure-21@2.[151,1436,181,1685]" captionTargetPageId="2" captionText="FIGURE 1. Actinopus chilikuti sp. nov., male holotype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, ocular pattern, dorsal view; D, right tibia II, retrolateral view; E–F, abdomen; E, dorsal view; F, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595937" httpUri="https://zenodo.org/record/14595937/files/figure.png" pageId="3" pageNumber="258">Fig 1B</figureCitation>
): length 0.52, width 0.55.
<emphasis id="B92608421A1C7A1EFE3CDCC6CC1E7493" box="[433,533,1693,1719]" italics="true" pageId="3" pageNumber="258">Maxillae</emphasis>
(
<figureCitation id="1369C8D51A1C7A1EFDABDCC6CC737493" box="[550,632,1693,1719]" captionStart="FIGURE 1" captionStartId="2.[151,250,1709,1733]" captionTargetBox="[151,1436,181,1685]" captionTargetId="figure-21@2.[151,1436,181,1685]" captionTargetPageId="2" captionText="FIGURE 1. Actinopus chilikuti sp. nov., male holotype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, ocular pattern, dorsal view; D, right tibia II, retrolateral view; E–F, abdomen; E, dorsal view; F, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595937" httpUri="https://zenodo.org/record/14595937/files/figure.png" pageId="3" pageNumber="258">Fig 1B</figureCitation>
): anterior length 0.78, posterior length 1.18, width 0.78.
<emphasis id="B92608421A1C7A1EFA8ADCC6CB6F7493" box="[1287,1380,1693,1719]" italics="true" pageId="3" pageNumber="258">Sternum</emphasis>
(
<figureCitation id="1369C8D51A1C7A1EFAF8DCC6CEB174FF" captionStart="FIGURE 1" captionStartId="2.[151,250,1709,1733]" captionTargetBox="[151,1436,181,1685]" captionTargetId="figure-21@2.[151,1436,181,1685]" captionTargetPageId="2" captionText="FIGURE 1. Actinopus chilikuti sp. nov., male holotype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, ocular pattern, dorsal view; D, right tibia II, retrolateral view; E–F, abdomen; E, dorsal view; F, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595937" httpUri="https://zenodo.org/record/14595937/files/figure.png" pageId="3" pageNumber="258">Fig 1B</figureCitation>
): Center flat, length 2.15, maximum width 1.66, minimum width 0.87. Postlabial sigilla square, posterior sigilla small and well defined.
<emphasis id="B92608421A1C7A1EFE2CDCBECC0074DB" box="[417,523,1765,1791]" italics="true" pageId="3" pageNumber="258">Abdomen</emphasis>
(
<figureCitation id="1369C8D51A1C7A1EFD94DCBECC8E74DB" box="[537,645,1765,1791]" captionStart="FIGURE 1" captionStartId="2.[151,250,1709,1733]" captionTargetBox="[151,1436,181,1685]" captionTargetId="figure-21@2.[151,1436,181,1685]" captionTargetPageId="2" captionText="FIGURE 1. Actinopus chilikuti sp. nov., male holotype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, ocular pattern, dorsal view; D, right tibia II, retrolateral view; E–F, abdomen; E, dorsal view; F, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595937" httpUri="https://zenodo.org/record/14595937/files/figure.png" pageId="3" pageNumber="258">Fig 1E, F</figureCitation>
): length 2.99, covered by small bristles. PMS: length 0.32; PLS: length of basal:medial:apical segments 0.38:0.17:0.13.
<emphasis id="B92608421A1C7A1EFD3ADD52CDC57507" box="[695,974,1801,1827]" italics="true" pageId="3" pageNumber="258">Lengths of legs and palp</emphasis>
: I: 3.92, 1.46, 2.52, 2.83, 1.60, 12.33. II: 3.70, 1.40, 2.40, 2.76, 1.54, 11.80. III: 2.92, 1.42, 1.77, 2.74, 1,59, 10.44. IV: 3.67, 1.51, 2.87, 3.04, 1.85, 12.94. Palp: 4.15, 1.98, 3.50,–, 0.81, 10.44.
<emphasis id="B92608421A1C7A1EFDB5DD0ACCB3754F" box="[568,696,1873,1899]" italics="true" pageId="3" pageNumber="258">Chaetotaxy</emphasis>
: Trochanter: III, p 1; I-II-IV, 0. Femora: I-II, 0; III, d 1-1-2; IV, 0. Patellae: I- II 0; III, d 1-1-1-1, p 16, 14 on the margin (crown of thorns); IV, p 21.Tibiae: I, r 1-1-2, v 1-2-1-2-1-3; II, r 13, v 1-1-2-3; III, v 2-2-2, r 2-2, d 1, with apical crown of 14 thorns; IV, v 1-1-1-1-1-3 (spiniform). Metatarsi: I, r 1-1-1-1-1-1-2, v 2-2-2, p 1-1-2-1-1; II, r 1-1-1-2-2, v 2-1-1-1-4 (apical), p 1-1; III, p 1-1-2-1-1-2, v 1-1-1-1, r 1-1-2-1-2-3; IV, v 2-2, p 1-2-3-2-1-3, r 1-0-2-1-2. Tarsi: I, r 2-1-2-2-1-4, v 2-2-1-1-1, p 1-1-1-1; II, r 18, v 1-1-1-1- 1-1-1, p 1-1-1-1-1; III, r 1-2-1-1, p 1-1-1; IV, r 1, p 1-2-2-2-1. Total number of retrolateral spines on tibiae I, II (
<figureCitation id="1369C8D51A1B7A19FAF8DDDECEB775E7" captionStart="FIGURE 1" captionStartId="2.[151,250,1709,1733]" captionTargetBox="[151,1436,181,1685]" captionTargetId="figure-21@2.[151,1436,181,1685]" captionTargetPageId="2" captionText="FIGURE 1. Actinopus chilikuti sp. nov., male holotype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, ocular pattern, dorsal view; D, right tibia II, retrolateral view; E–F, abdomen; E, dorsal view; F, ventral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595937" httpUri="https://zenodo.org/record/14595937/files/figure.png" pageId="4" pageNumber="259">Fig 1D</figureCitation>
): 4 and 13, respectively. Palp: tibia long, thin (
<figureCitation id="1369C8D51A1B7A19FD4BDDF2CD3275E7" box="[710,825,1961,1987]" captionStart="FIGURE 2" captionStartId="3.[151,250,1011,1035]" captionTargetBox="[152,1435,181,987]" captionTargetId="figure-21@3.[152,1435,181,987]" captionTargetPageId="3" captionText="FIGURE 2. Actinopus chilikuti sp. nov., male holotype. A, right palp, retrolateral view; B, right palp, prolateral view; C–F, copulatory bulb; C, dorsal view; D, ventral view; E, prolateral view; F, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595939" httpUri="https://zenodo.org/record/14595939/files/figure.png" pageId="4" pageNumber="259">Fig 2A, B</figureCitation>
). Bulb (
<figureCitation id="1369C8D51A1B7A19FC1EDDF2CA0A75E7" box="[915,1025,1961,1987]" captionStart="FIGURE 2" captionStartId="3.[151,250,1011,1035]" captionTargetBox="[152,1435,181,987]" captionTargetId="figure-21@3.[152,1435,181,987]" captionTargetPageId="3" captionText="FIGURE 2. Actinopus chilikuti sp. nov., male holotype. A, right palp, retrolateral view; B, right palp, prolateral view; C–F, copulatory bulb; C, dorsal view; D, ventral view; E, prolateral view; F, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595939" httpUri="https://zenodo.org/record/14595939/files/figure.png" pageId="4" pageNumber="259">Fig 2C–F</figureCitation>
) slim and slender. TA developed, PA developed. Embolus without denticles and with PS, PI, and Pac well developed.
<emphasis id="B92608421A1B7A19FB95DD96CAC075C3" box="[1048,1227,1997,2023]" italics="true" pageId="4" pageNumber="259">Pseudoscopula:</emphasis>
Tarsi: I–II, sparse; III-IV, dense, uniform. Metatarsi: I-II absent, III apical and sparse, IV 1:3 apical.
<emphasis id="B92608421A1A7A18FBA5DACCCAC77295" box="[1064,1228,151,177]" italics="true" pageId="5" pageNumber="260">Trichobothria:</emphasis>
Tibiae: I 3-3; II-II 4-3; IV 5-5. Metatarsi: I-II 8; III 6; IV 7. Tarsi: I -II 5; III 4; IV 5.
<emphasis id="B92608421A1A7A18FCFEDAE0CA4872F1" box="[883,1091,187,213]" italics="true" pageId="5" pageNumber="260">Colour in alcohol:</emphasis>
cephalic region and chelicerae reddish brown, carapace lighter. Abdomen brown; book lungs with light markings. Sternum reddish brown with a lighter center mark. Legs like cephalic region, legs III-IV with lighter tarsi–metatarsi; palp with tibia lighter. Spinnerets yellowish.
</paragraph>
<caption id="DF2D84D81A1B7A19FF1ADC89CCAC7516" ID-DOI="http://doi.org/10.5281/zenodo.14595941" ID-Zenodo-Dep="14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="4" pageNumber="259" startId="4.[151,250,1746,1770]" targetBox="[151,1436,181,1723]" targetPageId="4" targetType="figure">
<paragraph id="8BEDD4501A1B7A19FF1ADC89CCAC7516" blockId="4.[151,1436,1746,1843]" pageId="4" pageNumber="259">
<emphasis id="B92608421A1B7A19FF1ADC89CF1974CE" bold="true" box="[151,274,1746,1770]" pageId="4" pageNumber="259">FIGURE 3.</emphasis>
<taxonomicName id="4C52AFD31A1B7A19FE95DC88CFDD74CF" authorityName="Ríos-Tamayo &amp; Goloboff" authorityYear="2025" box="[280,470,1747,1771]" class="Arachnida" family="Actinopodidae" genus="Actinopus" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="chilikuti" status="sp. nov.">
<emphasis id="B92608421A1B7A19FE95DC88CFDD74CF" box="[280,470,1747,1771]" italics="true" pageId="4" pageNumber="259">Actinopus chilikuti</emphasis>
</taxonomicName>
<emphasis id="B92608421A1B7A19FE51DC88CC3874CF" bold="true" box="[476,563,1747,1771]" pageId="4" pageNumber="259">
<taxonomicNameLabel id="A215B5391A1B7A19FE51DC88CC2574CF" box="[476,558,1747,1771]" pageId="4" pageNumber="259" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
female paratype.
<emphasis id="B92608421A1B7A19FD6BDC89CCF474CE" bold="true" box="[742,767,1746,1770]" pageId="4" pageNumber="259">A,</emphasis>
cephalothorax, dorsal view, arrows indicate the presence of mites;
<emphasis id="B92608421A1B7A19FF1ADCACCEA5752B" bold="true" box="[151,174,1783,1807]" pageId="4" pageNumber="259">B,</emphasis>
sternum, ventral view;
<emphasis id="B92608421A1B7A19FE10DCADCFBD752A" bold="true" box="[413,438,1782,1806]" pageId="4" pageNumber="259">C,</emphasis>
abdomen, dorsal view;
<emphasis id="B92608421A1B7A19FD2ADCACCCCB752B" bold="true" box="[679,704,1783,1807]" pageId="4" pageNumber="259">D,</emphasis>
spermathecae;
<emphasis id="B92608421A1B7A19FCD0DCACCD7F752B" bold="true" box="[861,884,1783,1807]" pageId="4" pageNumber="259">E,</emphasis>
abdomen, ventral view;
<emphasis id="B92608421A1B7A19FBE3DCACCA89752B" bold="true" box="[1134,1154,1783,1807]" pageId="4" pageNumber="259">F,</emphasis>
ocular pattern, dorsal view;
<emphasis id="B92608421A1B7A19FF1ADD41CEBA7516" bold="true" box="[151,177,1818,1842]" pageId="4" pageNumber="259">G,</emphasis>
right tibia II, retrolateral view. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="8BEDD4501A1A7A18FF4ADB10CD39711D" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="260">
Female
<typeStatus id="54E96AF21A1A7A18FEADDB10CF8B7341" box="[288,384,331,357]" pageId="5" pageNumber="260" type="paratype">paratype</typeStatus>
(FML-ARAN 02274). Total length: 9.57. Carapace (
<figureCitation id="1369C8D51A1A7A18FC47DB10CA157341" box="[970,1054,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="5" pageNumber="260">Fig 3A</figureCitation>
): length 4.58, width 4.44; cephalic region: length 3.20, width 4.40; clypeus small with 12 bristles; 10 bristles between
<collectionCode id="ED434C951A1A7A18FBD2DB34CA9073AD" box="[1119,1179,367,393]" pageId="5" pageNumber="260">PME</collectionCode>
, 4 behind each
<collectionCode id="ED434C951A1A7A18FAD8DB34CB8573AD" box="[1365,1422,367,393]" pageId="5" pageNumber="260">PME</collectionCode>
–
<collectionCode id="ED434C951A1A7A18FF1ADBC8CEC77389" box="[151,204,403,429]" pageId="5" pageNumber="260">PLE</collectionCode>
, 5 bristles and one larger on both sides of ocular group.
<emphasis id="B92608421A1A7A18FCC1DBC8CD997388" box="[844,914,403,428]" italics="true" pageId="5" pageNumber="260">Fovea</emphasis>
: length 0.47, width 1.08.
<emphasis id="B92608421A1A7A18FB35DBC8CF3873F5" italics="true" pageId="5" pageNumber="260">Eyes, diameters and interdistances</emphasis>
(
<figureCitation id="1369C8D51A1A7A18FECEDBECCF9F73F5" box="[323,404,439,465]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="5" pageNumber="260">Fig 3F</figureCitation>
):
<collectionCode id="ED434C951A1A7A18FE24DBECCFEC73F5" box="[425,487,439,465]" pageId="5" pageNumber="260">AME</collectionCode>
0.11,
<collectionCode id="ED434C951A1A7A18FDA0DBECCC6873F5" box="[557,611,439,465]" pageId="5" pageNumber="260">ALE</collectionCode>
0.27,
<collectionCode id="ED434C951A1A7A18FD20DBECCCEC73F5" box="[685,743,439,465]" pageId="5" pageNumber="260">PME</collectionCode>
0.21,
<collectionCode id="ED434C951A1A7A18FCA2DBECCD6A73F5" box="[815,865,439,465]" pageId="5" pageNumber="260">PLE</collectionCode>
0.17, ocular group 0.80, anterior width 2.39, 2.09 posterior width;
<collectionCode id="ED434C951A1A7A18FEC1DB80CF8173D1" box="[332,394,475,501]" pageId="5" pageNumber="260">AME</collectionCode>
–
<collectionCode id="ED434C951A1A7A18FE14DB80CFC473D1" box="[409,463,475,501]" pageId="5" pageNumber="260">ALE</collectionCode>
0.73,
<collectionCode id="ED434C951A1A7A18FD9EDB80CC4773D1" box="[531,588,475,501]" pageId="5" pageNumber="260">PME</collectionCode>
–
<collectionCode id="ED434C951A1A7A18FDD7DB80CC8773D1" box="[602,652,475,501]" pageId="5" pageNumber="260">PLE</collectionCode>
0.11,
<collectionCode id="ED434C951A1A7A18FD40DB80CD0073D1" box="[717,779,475,501]" pageId="5" pageNumber="260">AME</collectionCode>
–
<collectionCode id="ED434C951A1A7A18FC94DB80CD5873D1" box="[793,851,475,501]" pageId="5" pageNumber="260">PME</collectionCode>
0.47,
<collectionCode id="ED434C951A1A7A18FC18DB80CDC073D1" box="[917,971,475,501]" pageId="5" pageNumber="260">ALE</collectionCode>
–
<collectionCode id="ED434C951A1A7A18FC54DB80CA0073D1" box="[985,1035,475,501]" pageId="5" pageNumber="260">PLE</collectionCode>
0.53,
<collectionCode id="ED434C951A1A7A18FBC0DB80CA8073D1" box="[1101,1163,475,501]" pageId="5" pageNumber="260">AME</collectionCode>
–
<collectionCode id="ED434C951A1A7A18FB14DB80CADC73D1" box="[1177,1239,475,501]" pageId="5" pageNumber="260">AME</collectionCode>
0.54,
<collectionCode id="ED434C951A1A7A18FA96DB80CB5E73D1" box="[1307,1365,475,501]" pageId="5" pageNumber="260">PME</collectionCode>
–
<specimenCode id="DBF47C2B1A1A7A18FAEEDB80CEC0703D" collectionCode="PME" pageId="5" pageNumber="260">PME 1.32</specimenCode>
.
<emphasis id="B92608421A1A7A18FF55DBA4CF58703D" box="[216,339,511,537]" italics="true" pageId="5" pageNumber="260">Chelicerae</emphasis>
: cheliceral apex with 9 recumbent and elongated bristles covering base of fang. Rastellum with 10 marginal, blunt spines and 5 small dorsal. Cheliceral furrow with 6 promarginal and 5 retromarginal teeth and 8 denticles in furrow, near promargin.
<emphasis id="B92608421A1A7A18FDBCD81CCC8C7045" box="[561,647,583,609]" italics="true" pageId="5" pageNumber="260">Labium</emphasis>
(
<figureCitation id="1369C8D51A1A7A18FD1BD81CCCE37045" box="[662,744,583,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="5" pageNumber="260">Fig 3B</figureCitation>
): length 1.13, width 0.97, with 12 anterior cuspules.
<emphasis id="B92608421A1A7A18FAB4D81CCB967045" box="[1337,1437,583,609]" italics="true" pageId="5" pageNumber="260">Maxillae</emphasis>
(
<figureCitation id="1369C8D51A1A7A18FF12D830CEFC70A1" box="[159,247,619,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="5" pageNumber="260">Fig 3B</figureCitation>
): anterior length 1.24, posterior length 1.67, width 1.35, with ~95/100 cuspules 1:3 ventral anterior, expanding to anterior face.
<emphasis id="B92608421A1A7A18FE47D8D4CC2C708D" box="[458,551,655,681]" italics="true" pageId="5" pageNumber="260">Sternum</emphasis>
(
<figureCitation id="1369C8D51A1A7A18FDBBD8D4CC83708D" box="[566,648,655,681]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="5" pageNumber="260">Fig 3B</figureCitation>
): Center flat, length: 3.38, maximum wide 2.67, minimum width 1.63. Postlabial sigilla square, posterior ones shallow, well defined.
<emphasis id="B92608421A1A7A18FCC1D8E8CDBD70E9" box="[844,950,691,717]" italics="true" pageId="5" pageNumber="260">Abdomen</emphasis>
(
<figureCitation id="1369C8D51A1A7A18FC49D8E8CA3E70E9" box="[964,1077,691,717]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="5" pageNumber="260">Fig 3C, E</figureCitation>
): length 5.02, covered by brown bristles.
<collectionCode id="ED434C951A1A7A18FF7BD88CCF3970D5" box="[246,306,727,753]" country="USA" httpUri="http://grbio.org/cool/9hjz-r0xi" name="Peabody Essex Museum" pageId="5" pageNumber="260">PMS</collectionCode>
: length 0.56;
<collectionCode id="ED434C951A1A7A18FE45D88CCFF370D5" box="[456,504,727,753]" pageId="5" pageNumber="260">PLS</collectionCode>
with basal: medial: apical articles 0.62:0.23:0.20.
<emphasis id="B92608421A1A7A18FBA1D88CCB3570D5" box="[1068,1342,727,753]" italics="true" pageId="5" pageNumber="260">Lengths of legs and palp</emphasis>
: I: 2.76, 1.83, 1.46, 1.64, 0.91, 8.60. II: 2.79, 1.86, 1.44, 1.71, 0.85, 8.65. III: 2.67, 2.09, 1.11, 1.73, 0.78, 8.38. IV: 3.33, 1.95, 2.05, 2.04, 0.79, 10.16. Palp: 2.46, 1.66, 1.65, –, 1.81, 7.58.
</paragraph>
<paragraph id="8BEDD4501A1A7A18FF4AD918CF077659" blockId="5.[151,1437,151,1185]" pageId="5" pageNumber="260">
<emphasis id="B92608421A1A7A18FF4AD918CF4C7179" box="[199,327,835,861]" italics="true" pageId="5" pageNumber="260">Chaetotaxy</emphasis>
: All femora: 0. Patellae: I–II, 0; III, p 4 + 9 on margin, r 7 on margin; IV, p-d 38; palp, p 1-1. Tibiae: I, v 1-1-1; II, r 30, v 1-1 spiniform setae; III, d 1, with apical crown of 15 thorns, r 12; IV, 0; palp, p 1-1-1-1-1-1, r 1-1-2-1-1. Metatarsi: I, r 7, v 1-1-2, p 14; II, r 1-2-1-1-1-1, p 1-1-1-1-2, v 1-1-2 near to the r ones; III, r 17 on row, v 2, p 10 on row; IV, p 2-1-2, d 1 anterior. Tarsi: I, r 10, v2, p 6; II, p 11, v 2, r 8; III, r 11, v-p 16; IV, v-p 24; palp, r 16, p 10, v 2 apical. Total number of retrolateral spines on tibiae I and II (
<figureCitation id="1369C8D51A1A7A18FC19D988CDED71C9" box="[916,998,979,1005]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="5" pageNumber="260">Fig 3G</figureCitation>
): 0 and 30, respectively.
<emphasis id="B92608421A1A7A18FB7AD988CB9771C9" box="[1271,1436,979,1005]" italics="true" pageId="5" pageNumber="260">Trichobothria:</emphasis>
Tibiae: I 3-3; II-III 3-4; IV 5-6. Metatarsi: I-II 9; III-IV 7. Tarsi: I 6; II 5; III-IV 4.
<emphasis id="B92608421A1A7A18FBD6D9ACCAB87635" box="[1115,1203,1015,1041]" italics="true" pageId="5" pageNumber="260">Colour:</emphasis>
cephalic region and chelicerae greenish brown, carapace lighter. Abdomen brown; book lungs with light markings. Sternum greenish brown with a lighter center mark (
<figureCitation id="1369C8D51A1A7A18FDA2DE64CC8F767D" box="[559,644,1087,1113]" captionStart="FIGURE 3" captionStartId="4.[151,250,1746,1770]" captionTargetBox="[151,1436,181,1723]" captionTargetId="figure-21@4.[151,1436,181,1723]" captionTargetPageId="4" captionText="FIGURE 3. Actinopus chilikuti sp. nov., female paratype. A, cephalothorax, dorsal view, arrows indicate the presence of mites; B, sternum, ventral view; C, abdomen, dorsal view; D, spermathecae; E, abdomen, ventral view; F, ocular pattern, dorsal view; G, right tibia II, retrolateral view. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.14595941" httpUri="https://zenodo.org/record/14595941/files/figure.png" pageId="5" pageNumber="260">Fig 3B</figureCitation>
), labium and maxillae darker. Legs like cephalic region. Spinnerets yellowish.
</paragraph>
</subSubSection>
<subSubSection id="C34887DB1A1A7A18FF4ADEDCCDB47685" box="[199,959,1158,1185]" pageId="5" pageNumber="260" type="distribution">
<paragraph id="8BEDD4501A1A7A18FF4ADEDCCDB47685" blockId="5.[151,1437,151,1185]" box="[199,959,1158,1185]" pageId="5" pageNumber="260">
<emphasis id="B92608421A1A7A18FF4ADEDCCF5B7685" box="[199,336,1159,1185]" italics="true" pageId="5" pageNumber="260">Distribution</emphasis>
. Provinces of Santiago del Estero and
<collectingCountry id="F34594C01A1A7A18FC8FDEDCCD6F7685" box="[770,868,1159,1185]" name="Taiwan" pageId="5" pageNumber="260">Formosa</collectingCountry>
(
<figureCitation id="1369C8D51A1A7A18FCFEDEDCCDBA7685" box="[883,945,1158,1185]" captionStart="FIGURE 5" captionStartId="6.[151,250,914,938]" captionTargetBox="[166,1419,185,882]" captionTargetId="figure-21@6.[162,1425,181,890]" captionTargetPageId="6" captionText="FIGURE 5. Known distribution of the species with Type I morphology in Argentina." figureDoi="http://doi.org/10.5281/zenodo.14595945" httpUri="https://zenodo.org/record/14595945/files/figure.png" pageId="5" pageNumber="260">Fig 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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