treatments-xml/data/03/EE/48/03EE4833FFD5082FFF6FFC7F84643ED7.xml
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<document id="A853A86E1C57F264C46F8A947596D770" ID-DOI="10.11646/zootaxa.5023.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5226026" ID-ZooBank="CABE11FD-4FAE-4716-80F8-390376AF1B73" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629427582629" checkinUser="plazi" docAuthor="Johansson, Niklas" docDate="2021" docId="03EE4833FFD5082FFF6FFC7F84643ED7" docLanguage="en" docName="zootaxa.5023.3.1.pdf" docOrigin="Zootaxa 5023 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enicospilus erebus Johansson 2021, sp. nov." docType="treatment" docUuid="6A077FE4-BE6B-4966-9157-476DEC7B29E7" docUuidSource="ZooBank" docVersion="3" lastPageNumber="312" masterDocId="FFD7304BFFDD0824FFF8FF8981453F22" masterDocTitle="Contribution to the subfamily Ophioninae (Hymenoptera: Ichneumonidae) of Southern Europe with the description of one new genus and 15 new species" masterLastPageNumber="334" masterPageNumber="301" pageNumber="309" updateTime="1699072774709" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="A9EAAD63C3B6B6B558961F67AB5DF913">Contribution to the subfamily Ophioninae (Hymenoptera: Ichneumonidae) of Southern Europe with the description of one new genus and 15 new species</mods:title>
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<mods:namePart id="F98B11EB23A1ABA93267DF04AA02E3C5">Johansson, Niklas</mods:namePart>
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<mods:date id="4CC93520436A26F7254E4E81C5C25D17">2021</mods:date>
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<mods:number id="771F399B05AD48ADC5A0EECD9D50CC63">2021-08-19</mods:number>
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<paragraph id="8BF8F925FFD5082CFF6FFC7F80A43B32" blockId="8.[151,481,1014,1076]" box="[151,481,1014,1041]" pageId="8" pageNumber="309">
<heading id="D0B04E49FFD5082CFF6FFC7F80A43B32" bold="true" box="[151,481,1014,1041]" fontSize="11" level="1" pageId="8" pageNumber="309" reason="1">
<emphasis id="B9332537FFD5082CFF6FFC7F80A43B32" bold="true" box="[151,481,1014,1041]" pageId="8" pageNumber="309">
<taxonomicName id="4C4782A6FFD5082CFF6FFC7F80393B33" authority="Johansson, 2021" authorityName="Johansson" authorityYear="2021" box="[151,380,1014,1041]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
<emphasis id="B9332537FFD5082CFF6FFC7F80393B33" bold="true" box="[151,380,1014,1041]" italics="true" pageId="8" pageNumber="309">Enicospilus erebus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A200984CFFD5082CFE7CFC7F80A43B32" box="[388,481,1014,1040]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="137CE5A0FFD5082CFF6FFB93804A3B16" box="[151,271,1050,1076]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,453,1824]" captionTargetId="figure-133@9.[151,1436,453,1824]" captionTargetPageId="9" captionText="FIGURE 6. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="8" pageNumber="309">Figs 6AF</figureCitation>
,
<figureCitation id="137CE5A0FFD5082CFEE2FB9380183B16" box="[282,349,1050,1076]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[164,1403,181,1919]" captionTargetId="figure-21@10.[161,1426,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5226040" httpUri="https://zenodo.org/record/5226040/files/figure.png" pageId="8" pageNumber="309">7AB</figureCitation>
</paragraph>
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urn:lsid:zoobank.org:act:
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<paragraph id="8BF8F925FFD5082CFF6FFB2C83173BC7" blockId="8.[151,1437,1188,2043]" pageId="8" pageNumber="309">
<materialsCitation id="3B2FF378FFD5082CFF6FFB2C83173BC7" collectingDate="1993-06-08" collectionCode="OOLM" collectorName="R. Rausch" country="Greece" county="Iraklion" elevation="1075" location="Weide Fd Skinakas" municipality="Ida-Geb" pageId="8" pageNumber="309" specimenCount="1" specimenCount-female="1" stateProvince="Kriti" typeStatus="holotype">
<emphasis id="B9332537FFD5082CFF6FFB2C80413B9D" bold="true" box="[151,260,1189,1215]" pageId="8" pageNumber="309">
<typeStatus id="54FC4787FFD5082CFF6FFB2C80413B9D" box="[151,260,1189,1215]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
</emphasis>
:
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<collectingCountry id="F350B9B5FFD5082CFEEDFB2C80CE3B9D" box="[277,395,1189,1215]" name="Greece" pageId="8" pageNumber="309">GREECE</collectingCountry>
</emphasis>
:
<specimenCount id="9D4132ACFFD5082CFE63FB2C80F53B9D" box="[411,432,1189,1215]" pageId="8" pageNumber="309" type="female"></specimenCount>
,
<collectingRegion id="498337C7FFD5082CFE38FB2C80BA3B9D" box="[448,511,1189,1215]" country="Greece" name="Kriti" pageId="8" pageNumber="309">Crete</collectingRegion>
,
<collectingCounty id="629981A9FFD5082CFDF5FB2C832C3B9D" box="[525,617,1189,1215]" pageId="8" pageNumber="309">Iraklion</collectingCounty>
,
<collectingMunicipality id="6B9C635FFFD5082CFD8DFB2C83913B9D" box="[629,724,1189,1215]" pageId="8" pageNumber="309">Ida-Geb</collectingMunicipality>
,
<location id="8E98AFFEFFD5082CFD18FB2C82FB3B9D" LSID="urn:lsid:plazi:treatment:03EE4833FFD5082FFF6FFC7F84643ED7:8E98AFFEFFD5082CFD18FB2C82FB3B9D" box="[736,958,1189,1215]" country="Greece" county="Iraklion" municipality="Ida-Geb" name="Weide Fd Skinakas" pageId="8" pageNumber="309" stateProvince="Kriti">Weide Fd Skinakas</location>
, 35˚13N, 24˚56E,
<quantity id="4CBF54C0FFD5082CFB5DFB2C84013B9D" box="[1189,1348,1189,1216]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="8" pageNumber="309" unit="m" value="1075.0" valueMax="1100.0" valueMin="1050.0">
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</quantity>
. a. s.,
<date id="FFF9DFE5FFD5082CFA76FB2C80423BC1" pageId="8" pageNumber="309" value="1993-06-08">
<collectingDate id="EFBD260DFFD5082CFA76FB2C80423BC1" pageId="8" pageNumber="309" value="1993-06-08">8 Jun. 1993</collectingDate>
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, H.u.
<collectorName id="26B29CF3FFD5082CFEBAFB4080F03BC1" box="[322,437,1225,1251]" pageId="8" pageNumber="309">R. Rausch</collectorName>
leg. (
<collectionCode id="ED5661E0FFD5082CFE0CFB43830F3BC7" box="[500,586,1226,1253]" pageId="8" pageNumber="309">OÖLM</collectionCode>
)
</materialsCitation>
</paragraph>
<paragraph id="8BF8F925FFD5082CFF3FFB64837C3ABB" blockId="8.[151,1437,1188,2043]" pageId="8" pageNumber="309">
<materialsCitation id="3B2FF378FFD5082CFF3FFB6483C93A0F" collectingDate="1993-06-08" collectionCode="OOLM" collectorName="R. Rausch" country="Greece" county="Iraklion" elevation="1075" location="Weide Fd Skinakas" municipality="Ida-Geb" pageId="8" pageNumber="309" specimenCount="2" specimenCount-female="2" stateProvince="Kriti" typeStatus="paratype">
<emphasis id="B9332537FFD5082CFF3FFB64807A3A25" bold="true" box="[199,319,1261,1287]" pageId="8" pageNumber="309">
<typeStatus id="54FC4787FFD5082CFF3FFB64807A3A25" box="[199,319,1261,1287]" pageId="8" pageNumber="309" type="paratype">Paratypes</typeStatus>
</emphasis>
:
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<collectingCountry id="F350B9B5FFD5082CFEB5FB6480863A25" box="[333,451,1261,1287]" name="Greece" pageId="8" pageNumber="309">GREECE</collectingCountry>
</emphasis>
:
<specimenCount id="9D4132ACFFD5082CFE29FB64834F3A25" box="[465,522,1261,1287]" pageId="8" pageNumber="309" type="female">2♀♀</specimenCount>
,
<collectingRegion id="498337C7FFD5082CFDEDFB6483113A25" box="[533,596,1261,1287]" country="Greece" name="Kriti" pageId="8" pageNumber="309">Crete</collectingRegion>
,
<collectingCounty id="629981A9FFD5082CFDA7FB6483FE3A25" box="[607,699,1261,1287]" pageId="8" pageNumber="309">Iraklion</collectingCounty>
,
<collectingMunicipality id="6B9C635FFFD5082CFD3DFB6482613A25" box="[709,804,1261,1287]" pageId="8" pageNumber="309">Ida-Geb</collectingMunicipality>
,
<location id="8E98AFFEFFD5082CFCD5FB6485403A25" LSID="urn:lsid:plazi:treatment:03EE4833FFD5082FFF6FFC7F84643ED7:8E98AFFEFFD5082CFCD5FB6485403A25" box="[813,1029,1261,1287]" country="Greece" county="Iraklion" municipality="Ida-Geb" name="Weide Fd Skinakas" pageId="8" pageNumber="309" stateProvince="Kriti">Weide Fd Skinakas</location>
, 35˚13N, 24˚56E,
<quantity id="4CBF54C0FFD5082CFB1BFB64843A3A25" box="[1251,1407,1261,1288]" metricMagnitude="3" metricUnit="m" metricValue="1.075" metricValueMax="1.1" metricValueMin="1.05" pageId="8" pageNumber="309" unit="m" value="1075.0" valueMax="1100.0" valueMin="1050.0">
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. a. s.,
<date id="FFF9DFE5FFD5082CFF4FFA9880783A09" box="[183,317,1297,1323]" pageId="8" pageNumber="309" value="1993-06-08">
<collectingDate id="EFBD260DFFD5082CFF4FFA9880783A09" box="[183,317,1297,1323]" pageId="8" pageNumber="309" value="1993-06-08">8 Jun. 1993</collectingDate>
</date>
, H.u.
<collectorName id="26B29CF3FFD5082CFE8FFA9880AE3A09" box="[375,491,1297,1323]" pageId="8" pageNumber="309">R. Rausch</collectorName>
leg. (
<collectionCode id="ED5661E0FFD5082CFDD3FA9B83C13A0F" box="[555,644,1298,1325]" pageId="8" pageNumber="309">OÖLM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2FF378FFD5082CFD6FFA98843B3A09" box="[663,1406,1296,1323]" collectorName="Arkadien S. Paparis" country="Greece" elevation="800" location="Peloponnesos" pageId="8" pageNumber="309" specimenCount="1" specimenCount-female="1" stateProvince="Peloponnisos" typeStatus="paratype">
<specimenCount id="9D4132ACFFD5082CFD6FFA9883FF3A09" box="[663,698,1297,1323]" pageId="8" pageNumber="309" type="female">1♀</specimenCount>
,
<collectingRegion id="498337C7FFD5082CFD30FA9882203A09" box="[712,869,1297,1323]" country="Greece" name="Peloponnisos" pageId="8" pageNumber="309">Peloponnesos</collectingRegion>
,
<collectorName id="26B29CF3FFD5082CFC97FA9885143A09" box="[879,1105,1297,1323]" pageId="8" pageNumber="309">Arkadien S. Paparis</collectorName>
, 37˚22N, 22˚14E,
<quantity id="4CBF54C0FFD5082CFACCFA98843B3A09" box="[1332,1406,1297,1323]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="309" unit="m" value="800.0">
<specimenCount id="9D4132ACFFD5082CFACCFA98843B3A09" box="[1332,1406,1297,1323]" pageId="8" pageNumber="309" type="generic">
<elevation id="006A1E16FFD5082CFACCFA98843B3A09" box="[1332,1406,1297,1323]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="309" unit="m" value="800.0">800 m</elevation>
</specimenCount>
</quantity>
</materialsCitation>
<materialsCitation id="3B2FF378FFD5082CFA86FA9883D73A73" collectingDate="1992-05-18" collectionCode="OOLM" collectorName="R Rausch" country="Greece" pageId="8" pageNumber="309" specimenCount="1" typeStatus="paratype">
. a. s.,
<date id="FFF9DFE5FFD5082CFF4EFABC80083A6D" box="[182,333,1333,1359]" pageId="8" pageNumber="309" value="1992-05-18">
<collectingDate id="EFBD260DFFD5082CFF4EFABC80083A6D" box="[182,333,1333,1359]" pageId="8" pageNumber="309" value="1992-05-18">18 May 1992</collectingDate>
</date>
, H.u.
<collectorName id="26B29CF3FFD5082CFE7DFABC80B63A6D" box="[389,499,1333,1359]" pageId="8" pageNumber="309">R Rausch</collectorName>
leg. (
<collectionCode id="ED5661E0FFD5082CFDC9FABF83CF3A73" box="[561,650,1334,1361]" pageId="8" pageNumber="309">OÖLM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2FF378FFD5082CFD64FABC84DC3A6D" box="[668,1433,1333,1359]" collectingDate="1960-05-22" collectionCode="NHMW" collectorName="K. Kusdas" country="Greece" location="Zachlorou" pageId="8" pageNumber="309" specimenCount="1" specimenCount-male="1" stateProvince="Peloponnisos" typeStatus="paratype">
<specimenCount id="9D4132ACFFD5082CFD64FABC83843A6D" box="[668,705,1333,1359]" pageId="8" pageNumber="309" type="male">1♂</specimenCount>
,
<collectingRegion id="498337C7FFD5082CFD35FABC82143A6D" box="[717,849,1333,1359]" country="Greece" name="Peloponnisos" pageId="8" pageNumber="309">Peloponnes</collectingRegion>
,
<location id="8E98AFFEFFD5082CFCA3FABC82943A6D" LSID="urn:lsid:plazi:treatment:03EE4833FFD5082FFF6FFC7F84643ED7:8E98AFFEFFD5082CFCA3FABC82943A6D" box="[859,977,1333,1359]" country="Greece" name="Zachlorou" pageId="8" pageNumber="309" stateProvince="Peloponnisos">Zachlorou</location>
,
<date id="FFF9DFE5FFD5082CFC24FABC85363A6D" box="[988,1139,1333,1359]" pageId="8" pageNumber="309" value="1960-05-22">
<collectingDate id="EFBD260DFFD5082CFC24FABC85363A6D" box="[988,1139,1333,1359]" pageId="8" pageNumber="309" value="1960-05-22">22 May 1960</collectingDate>
</date>
,
<collectorName id="26B29CF3FFD5082CFB85FABC85B53A6D" box="[1149,1264,1333,1359]" pageId="8" pageNumber="309">K. Kusdas</collectorName>
leg. (
<collectionCode id="ED5661E0FFD5082CFAD6FABC84D53A6D" box="[1326,1424,1333,1359]" country="Austria" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="309" type="Museum">NHMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2FF378FFD5082CFF6FFAD082923A57" box="[151,983,1369,1397]" collectingDate="1998-05-30" collectionCode="OOLM" collectorName="Ma. Halada" country="Turkey" location="Gaziantep" pageId="8" pageNumber="309" specimenCount="1" specimenCount-female="1" stateProvince="Gaziantep" typeStatus="paratype">
<emphasis id="B9332537FFD5082CFF6FFAD0804A3A51" bold="true" box="[151,271,1369,1395]" pageId="8" pageNumber="309">
<collectingCountry id="F350B9B5FFD5082CFF6FFAD0804A3A51" box="[151,271,1369,1395]" name="Turkey" pageId="8" pageNumber="309">TURKEY</collectingCountry>
</emphasis>
:
<specimenCount id="9D4132ACFFD5082CFEE6FAD080053A51" box="[286,320,1369,1395]" pageId="8" pageNumber="309" type="female">1♀</specimenCount>
,
<quantity id="4CBF54C0FFD5082CFEB6FAD080D03A51" box="[334,405,1369,1395]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="309" unit="km" value="30.0">30 km</quantity>
W
<collectingRegion id="498337C7FFD5082CFE44FAD083753A51" box="[444,560,1369,1395]" country="Turkey" name="Gaziantep" pageId="8" pageNumber="309">Gaziantep</collectingRegion>
,
<date id="FFF9DFE5FFD5082CFDC3FAD083963A51" box="[571,723,1369,1395]" pageId="8" pageNumber="309" value="1998-05-30">
<collectingDate id="EFBD260DFFD5082CFDC3FAD083963A51" box="[571,723,1369,1395]" pageId="8" pageNumber="309" value="1998-05-30">30 May 1998</collectingDate>
</date>
,
<collectorName id="26B29CF3FFD5082CFD26FAD082263A51" box="[734,867,1369,1395]" pageId="8" pageNumber="309">Ma. Halada</collectorName>
, (
<collectionCode id="ED5661E0FFD5082CFC8EFAD3828A3A57" box="[886,975,1370,1397]" pageId="8" pageNumber="309">OÖLM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B2FF378FFD5082CFC1AFAD083703ABB" collectingDate="1998-06-10" collectionCode="OOLM" collectorName="Ma. Halada" country="Turkey" location="Hasanbeyli" pageId="8" pageNumber="309" specimenCount="2" specimenCount-female="2" stateProvince="Osmaniye" typeStatus="paratype">
<specimenCount id="9D4132ACFFD5082CFC1AFAD0855E3A51" box="[994,1051,1369,1395]" pageId="8" pageNumber="309" type="female">2♀♀</specimenCount>
,
<locationDeviation id="9A4AE007FFD5082CFBD0FAD085AC3A51" box="[1064,1257,1369,1395]" location="Osmaniye" pageId="8" pageNumber="309">
<quantity id="4CBF54C0FFD5082CFBD0FAD0852A3A51" box="[1064,1135,1369,1395]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="8" pageNumber="309" unit="km" value="30.0">30 km</quantity>
<collectingRegion id="498337C7FFD5082CFB8DFAD085AC3A51" box="[1141,1257,1369,1395]" country="Turkey" name="Osmaniye" pageId="8" pageNumber="309">Osmaniye</collectingRegion>
</locationDeviation>
,
<location id="8E98AFFEFFD5082CFB0CFAD084303A51" LSID="urn:lsid:plazi:treatment:03EE4833FFD5082FFF6FFC7F84643ED7:8E98AFFEFFD5082CFB0CFAD084303A51" box="[1268,1397,1369,1395]" country="Turkey" name="Hasanbeyli" pageId="8" pageNumber="309" stateProvince="Osmaniye">Hasanbeyli</location>
,
<date id="FFF9DFE5FFD5082CFA78FAD080423AB5" pageId="8" pageNumber="309" value="1998-06-10">
<collectingDate id="EFBD260DFFD5082CFA78FAD080423AB5" pageId="8" pageNumber="309" value="1998-06-10">10 Jun. 1998</collectingDate>
</date>
,
<collectorName id="26B29CF3FFD5082CFEEAFAF480D13AB5" box="[274,404,1405,1431]" pageId="8" pageNumber="309">Ma. Halada</collectorName>
leg. (
<collectionCode id="ED5661E0FFD5082CFE2CFAF783683ABB" box="[468,557,1406,1433]" pageId="8" pageNumber="309">OÖLM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35DAAAEFFD5082CFF3FFA2882373AFD" pageId="8" pageNumber="309" type="etymology">
<paragraph id="8BF8F925FFD5082CFF3FFA2882373AFD" blockId="8.[151,1437,1188,2043]" pageId="8" pageNumber="309">
<emphasis id="B9332537FFD5082CFF3FFA28800D3A99" bold="true" box="[199,328,1441,1467]" pageId="8" pageNumber="309">Etymology</emphasis>
: The name refers to Erebus, who is the personification of darkness in greek mythology, both referring to the dark coloration and nocturnal behavior of this species.
</paragraph>
</subSubSection>
<subSubSection id="C35DAAAEFFD5082CFF3FFA60853939F9" pageId="8" pageNumber="309" type="diagnosis">
<paragraph id="8BF8F925FFD5082CFF3FFA60853939F9" blockId="8.[151,1437,1188,2043]" pageId="8" pageNumber="309">
<emphasis id="B9332537FFD5082CFF3FFA60807F3921" bold="true" box="[199,314,1513,1539]" pageId="8" pageNumber="309">Diagnosis</emphasis>
: Usually a very distinctive species, easily identified by the dark head and metasoma in combination with the number and shape of the alar sclerites. The species also occurs in varieties with more or less testaceous head and mesosoma.
<taxonomicName id="4C4782A6FFD5082CFE76F9B883213969" authority="Johansson, 2021" authorityName="Johansson" authorityYear="2021" box="[398,612,1585,1611]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
<emphasis id="B9332537FFD5082CFE76F9B883213969" box="[398,612,1585,1611]" italics="true" pageId="8" pageNumber="309">Enicospilus erebus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A200984CFFD5082CFD95F9B883833969" box="[621,710,1585,1611]" pageId="8" pageNumber="309" rank="species">sp. nov.</taxonomicNameLabel>
is distinguished from similar species by the constitution of the alar sclerites, the polished, almost parallel sided and scarcely punctate scutellum and the quite dark colour of the body. The light form resembles
<taxonomicName id="4C4782A6FFD5082CFE04F9F0828739B1" authority="Johansson, 2018" authorityName="Johansson" authorityYear="2018" box="[508,962,1657,1683]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="cederbergi">
<emphasis id="B9332537FFD5082CFE04F9F083BB39B1" box="[508,766,1657,1683]" italics="true" pageId="8" pageNumber="309">Enicospilus cederbergi</emphasis>
<bibRefCitation id="EFD684D4FFD5082CFCFEF9F0828739B1" author="Johansson, N." box="[774,962,1657,1683]" pageId="8" pageNumber="309" pagination="1 - 21" refId="ref17214" refString="Johansson, N. (2018) Review of the Swedish Enicospilus (Hymenoptera; Ichneumonidae; Ophioninae) with description of three new species and an illustrated key to species. European Journal of Taxonomy, 483, 1 - 21. https: // doi. org / 10.5852 / ejt. 2018.483" type="journal article" year="2018">Johansson, 2018</bibRefCitation>
</taxonomicName>
, but differs in the number of alar sclerites, the polished scutellum and face and the the less numerous flagellomeres. Notably specimens from
<collectingCountry id="F350B9B5FFD5082CFB10F91484723995" box="[1256,1335,1693,1719]" name="Greece" pageId="8" pageNumber="309">Greece</collectingCountry>
have the mesosoma and head mostly dark while the Turkish specimens are less strongly infuscate.
</paragraph>
</subSubSection>
<subSubSection id="C35DAAAEFFD5082FFF3FF96C83563E44" lastPageId="11" lastPageNumber="312" pageId="8" pageNumber="309" type="description">
<paragraph id="8BF8F925FFD5082DFF3FF96C821D3E8C" blockId="8.[151,1437,1188,2043]" lastBlockId="9.[151,1437,151,430]" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
<emphasis id="B9332537FFD5082CFF3FF96C801439DD" bold="true" box="[199,337,1765,1791]" pageId="8" pageNumber="309">Description</emphasis>
: Fore wing length
<quantity id="4CBF54C0FFD5082CFDD0F96C83E039DD" box="[552,677,1765,1792]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="8" pageNumber="309" unit="mm" value="14.5" valueMax="15.0" valueMin="14.0">1415 mm</quantity>
. Body length
<quantity id="4CBF54C0FFD5082CFCB9F96C82F839DD" box="[833,957,1765,1791]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="8" pageNumber="309" unit="mm" value="19.0" valueMax="20.0" valueMin="18.0">1820 mm</quantity>
. Number of flagellomeres 4851. Temples slightly buccate, in lateral view about 0.40.5 × the width of compound eye (
<figureCitation id="137CE5A0FFD5082CFBFDF880851F3806" box="[1029,1114,1801,1828]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,453,1824]" captionTargetId="figure-133@9.[151,1436,453,1824]" captionTargetPageId="9" captionText="FIGURE 6. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="8" pageNumber="309">Fig 6D</figureCitation>
). In male wider, about 0.9 × the width of the compound eye. Face moderately wide (
<figureCitation id="137CE5A0FFD5082CFCF2F8A482CC386A" box="[778,905,1837,1864]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,453,1824]" captionTargetId="figure-133@9.[151,1436,453,1824]" captionTargetPageId="9" captionText="FIGURE 6. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="8" pageNumber="309">Figs 6AB</figureCitation>
), distinctly wider in the male. Malar space narrow, about 0.1 × the width of the mandibular base. Gap between lateral ocelli and inner margin of compound eye about 0.1 × the diameter of ocellus or virtually absent. Clypeus flat, anterior margin weakly convex in anterior view, centrally straight. Mandible strongly twisted with upper tooth about 2.0 × as long as lower tooth. First flagellomere stout, about 3.54.0 × as long as apically wide. Mid- and preapical flagellomeres about 1.11.3 × as long as wide. Mesopleuron polished, densely punctate. Interstices between punctures about equal to their diameter, punctures centrally sometimes merging. Pleurosternal angles distinct, almost right angled. Mesoscutum mostly polished with notauli weakly indicated anteriorly, punctate. Scutellum long, almost parallel sided, polished with a few scattered punctures and lateral carinae along its entire length (
<figureCitation id="137CE5A0FFD4082DFD13FF5682003FD8" box="[747,837,223,250]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,453,1824]" captionTargetId="figure-133@9.[151,1436,453,1824]" captionTargetPageId="9" captionText="FIGURE 6. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="9" pageNumber="310">Fig. 6C</figureCitation>
). Propodeum with anterior transverse carina distinct. Posterior to carina rugulose, anterior to carinae smooth, slightly polished. Proximal sclerite entirely pigmented, strongly sclerotised, triangular, central sclerite entirely absent. Distal sclerite crescent-shaped, weakly sclerotised (
<figureCitation id="137CE5A0FFD4082DFF58FEC281B23E44" box="[160,247,331,358]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,453,1824]" captionTargetId="figure-133@9.[151,1436,453,1824]" captionTargetPageId="9" captionText="FIGURE 6. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="9" pageNumber="310">Fig. 6E</figureCitation>
). Radius weakly sinuate before junction with pterostigma. Hind femur quite stout, about 7.08.0 × longer than wide. Inner spur of hind tibia about 0.5 × as long as hind metatarsus. Hind tarsal claws in female pectinate throughout their entire length, dorsally evenly curved (
<figureCitation id="137CE5A0FFD4082DFD04FE1A820E3E8C" box="[764,843,403,430]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,453,1824]" captionTargetId="figure-133@9.[151,1436,453,1824]" captionTargetPageId="9" captionText="FIGURE 6. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="9" pageNumber="310">Fig 6F</figureCitation>
).
</paragraph>
<caption id="DF38A9ADFFD4082DFF6FF8BE83943856" ID-DOI="http://doi.org/10.5281/zenodo.5226038" ID-Zenodo-Dep="5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="9" pageNumber="310" startId="9.[151,250,1847,1871]" targetBox="[151,1436,453,1824]" targetPageId="9">
<paragraph id="8BF8F925FFD4082DFF6FF8BE83943856" blockId="9.[151,1436,1847,1908]" pageId="9" pageNumber="310">
<emphasis id="B9332537FFD4082DFF6FF8BE8051386D" bold="true" box="[151,276,1847,1871]" pageId="9" pageNumber="310">FIGURE 6.</emphasis>
<taxonomicName id="4C4782A6FFD4082DFEE4F8B180983872" authority="Johansson, 2021" authorityName="Johansson" authorityYear="2021" box="[284,477,1848,1872]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
<emphasis id="B9332537FFD4082DFEE4F8B180983872" box="[284,477,1848,1872]" italics="true" pageId="9" pageNumber="310">Enicospilus erebus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A200984CFFD4082DFE1CF8B18371386D" box="[484,564,1848,1871]" pageId="9" pageNumber="310" rank="species">sp. nov.</taxonomicNameLabel>
paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws.
</paragraph>
</caption>
<caption id="DF38A9ADFFD7082EFF6FF81E85263892" ID-DOI="http://doi.org/10.5281/zenodo.5226040" ID-Zenodo-Dep="5226040" box="[151,1123,1943,1970]" httpUri="https://zenodo.org/record/5226040/files/figure.png" pageId="10" pageNumber="311" startId="10.[151,250,1943,1967]" targetBox="[164,1403,181,1919]" targetPageId="10">
<paragraph id="8BF8F925FFD7082EFF6FF81E85263892" blockId="10.[151,1123,1943,1970]" box="[151,1123,1943,1970]" pageId="10" pageNumber="311">
<emphasis id="B9332537FFD7082EFF6FF81E80563892" bold="true" box="[151,275,1943,1968]" pageId="10" pageNumber="311">FIGURE 7.</emphasis>
<taxonomicName id="4C4782A6FFD7082EFEE1F811809C3892" authority="Johansson, 2021" authorityName="Johansson" authorityYear="2021" box="[281,473,1944,1968]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
<emphasis id="B9332537FFD7082EFEE1F811809C3892" box="[281,473,1944,1968]" italics="true" pageId="10" pageNumber="311">Enicospilus erebus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A200984CFFD7082EFE27F811836B388D" box="[479,558,1944,1967]" pageId="10" pageNumber="311" rank="species">sp. nov.</taxonomicNameLabel>
paratype females (OÖLM). AB) Habitus, lateral view.
</paragraph>
</caption>
<paragraph id="8BF8F925FFD6082FFF3FFF1E83563E44" blockId="11.[151,1437,151,501]" pageId="11" pageNumber="312">
<emphasis id="B9332537FFD6082FFF3FFF1E80153F93" box="[199,336,151,177]" italics="true" pageId="11" pageNumber="312">Colouration</emphasis>
: The species is very variable in colour. The dark form (
<figureCitation id="137CE5A0FFD6082FFC38FF1E850B3F93" box="[960,1102,151,177]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,453,1824]" captionTargetId="figure-133@9.[151,1436,453,1824]" captionTargetPageId="9" captionText="FIGURE 6. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="11" pageNumber="312">Figs 6A, 6D</figureCitation>
,
<figureCitation id="137CE5A0FFD6082FFBA2FF1E853B3F93" box="[1114,1150,151,177]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[164,1403,181,1919]" captionTargetId="figure-21@10.[161,1426,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5226040" httpUri="https://zenodo.org/record/5226040/files/figure.png" pageId="11" pageNumber="312">7A</figureCitation>
) has the head, mesosoma and most of metasoma infuscate, while the legs (excluding coxae), antennae (including scapus), mandibles (excluding teeth) and the first and second tergite are testaceous. The pale form (
<figureCitation id="137CE5A0FFD6082FFC21FF5685123FDB" box="[985,1111,223,249]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,453,1824]" captionTargetId="figure-133@9.[151,1436,453,1824]" captionTargetPageId="9" captionText="FIGURE 6. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="11" pageNumber="312">Figs 6BC</figureCitation>
,
<figureCitation id="137CE5A0FFD6082FFB9DFF5685C33FDB" box="[1125,1158,223,249]" captionStart="FIGURE 7" captionStartId="10.[151,250,1943,1967]" captionTargetBox="[164,1403,181,1919]" captionTargetId="figure-21@10.[161,1426,181,1919]" captionTargetPageId="10" captionText="FIGURE 7. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) Habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.5226040" httpUri="https://zenodo.org/record/5226040/files/figure.png" pageId="11" pageNumber="312">7B</figureCitation>
) is testaceous with only the posterior segments of metasoma, the ovipositor, the mandibular teeth and claws black. There are intermediate specimens regarding colouration. Wing membrane slightly yellowish. Fore wing veins infuscate. Alar sclerites and pterostigma pale brown (
<figureCitation id="137CE5A0FFD6082FFE57FEC283433E44" box="[431,518,331,358]" captionStart="FIGURE 6" captionStartId="9.[151,250,1847,1871]" captionTargetBox="[151,1436,453,1824]" captionTargetId="figure-133@9.[151,1436,453,1824]" captionTargetPageId="9" captionText="FIGURE 6. Enicospilus erebus sp. nov. paratype females (OÖLM). AB) head, anterior view; C) scutellum, dorsal view; D) head, lateral view; E) alar sclerites; F) hind tarsal claws." figureDoi="http://doi.org/10.5281/zenodo.5226038" httpUri="https://zenodo.org/record/5226038/files/figure.png" pageId="11" pageNumber="312">Fig. 6E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35DAAAEFFD6082FFF3FFEE684643ED7" pageId="11" pageNumber="312" type="discussion">
<paragraph id="8BF8F925FFD6082FFF3FFEE683063E8F" blockId="11.[151,1437,151,501]" pageId="11" pageNumber="312">
<emphasis id="B9332537FFD6082FFF3FFEE680783EAB" bold="true" box="[199,317,367,393]" pageId="11" pageNumber="312">Remarks:</emphasis>
One barcoded pale female specimen from
<collectingCountry id="F350B9B5FFD6082FFCEFFEE682223EAB" box="[791,871,367,393]" name="Turkey" pageId="11" pageNumber="312">Turkey</collectingCountry>
displays about 0.55% sequence divergence in relation to two dark females from
<collectingCountry id="F350B9B5FFD6082FFE14FE1A837A3E8F" box="[492,575,403,429]" name="Greece" pageId="11" pageNumber="312">Greece</collectingCountry>
.
</paragraph>
<paragraph id="8BF8F925FFD6082FFF3FFE3F84643ED7" blockId="11.[151,1437,151,501]" pageId="11" pageNumber="312">
<emphasis id="B9332537FFD6082FFF3FFE3F80293EF3" bold="true" box="[199,364,438,465]" pageId="11" pageNumber="312">DNA Barcode</emphasis>
: The DNA barcode sequences of
<specimenCount id="9D4132ACFFD6082FFD1CFE3E82DD3EF3" box="[740,920,439,465]" pageId="11" pageNumber="312" type="generic">three specimens</specimenCount>
of
<taxonomicName id="4C4782A6FFD6082FFC45FE3E85D53EF3" authority="Johansson, 2021" authorityName="Johansson" authorityYear="2021" box="[957,1168,439,465]" class="Insecta" family="Ichneumonidae" genus="Enicospilus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="erebus" status="sp. nov.">
<emphasis id="B9332537FFD6082FFC45FE3E85D53EF3" box="[957,1168,439,465]" italics="true" pageId="11" pageNumber="312">Enicospilus erebus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A200984CFFD6082FFB6EFE3E85A83EF3" box="[1174,1261,439,465]" pageId="11" pageNumber="312" rank="species">sp. nov.</taxonomicNameLabel>
are available at the BOLD systems database: sample id. NJP727/728/729, sequence id. LNU2563-21/2564-21/2565-21.
</paragraph>
</subSubSection>
</treatment>
</document>