treatments-xml/data/03/D6/71/03D67155FFAAFFACFF25FC83A42228C9.xml

<document id="82CBA3A08AA6EE51D911558BD4357FAE" ID-CLB-Dataset="7679" ID-DOI="10.11646/zootaxa.4942.1.2" ID-GBIF-Dataset="25d73144-c90c-45c3-a9a2-2454f87a9aef" ID-ISSN="1175-5326" ID-Zenodo-Dep="4596097" ID-ZooBank="79353662-7653-4F41-8B39-40E6E4B2E005" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615448142785" checkinUser="plazi" docAuthor="Haddad, Charles R., Jin, Chi, Platnick, Norman I. &amp; Booysen, Ruan" docDate="2021" docId="03D67155FFAAFFACFF25FC83A42228C9" docLanguage="en" docName="zootaxa.4942.1.2.pdf" docOrigin="Zootaxa 4942 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Capobula Haddad &amp; Jin &amp; Platnick &amp; Booysen 2021, gen. nov." docType="treatment" docVersion="6" lastPageNumber="51" masterDocId="FFEF092DFFAFFFA6FFB2FFBAA0202918" masterDocTitle="Capobula gen. nov., a new Afrotropical dark sac spider genus related to Orthobula Simon, 1897 (Araneae: Trachelidae)" masterLastPageNumber="71" masterPageNumber="41" pageNumber="46" updateTime="1698933034801" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="82C40B3DA8B3A8000580DA0AE416C29E">Capobula gen. nov., a new Afrotropical dark sac spider genus related to Orthobula Simon, 1897 (Araneae: Trachelidae)</mods:title>
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<mods:namePart id="472027AA4AB6233328EDF0531FF5291D">Haddad, Charles R.</mods:namePart>
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<mods:affiliation id="A30870D7DBEF1BAC193B9D1392376F9F">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa. &amp; haddadcr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 2317 - 7760; &amp; booysenr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 0841 - 9143</mods:affiliation>
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<mods:namePart id="7DDF326845AF1708132B5A7DAA534CA8">Jin, Chi</mods:namePart>
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<mods:affiliation id="0406AC92CD6503777C57078C9EE7197F">College of Landscape and Ecological Engineering, Hebei University of Engineering, Handan, Hebei 056038, P. R. China. &amp; jinchi _ spider @ 163. com; https: // orcid. org / 0000 - 0002 - 4694 - 4870</mods:affiliation>
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<mods:namePart id="E13ED1D4A8DEACC982C6F9AE6CF63E13">Platnick, Norman I.</mods:namePart>
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<mods:affiliation id="F9A921EADA7C65E51E9AC1059D8091AB">Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 th Street, New York, NY 10024, U. S. A. &amp; https: // orcid. org / 0000 - 0003 - 4868 - 7529; deceased</mods:affiliation>
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<mods:namePart id="B5F73F22074CD48E9953A4DEFBB7E115">Booysen, Ruan</mods:namePart>
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<mods:affiliation id="01104908FDD59E78364D30C78085946D">Department of Zoology &amp; Entomology, University of the Free State, P. O. Box 339, Bloemfontein 9300, South Africa. &amp; haddadcr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 2317 - 7760; &amp; booysenr @ ufs. ac. za; https: // orcid. org / 0000 - 0002 - 0841 - 9143</mods:affiliation>
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<paragraph id="8BC0C043FFAAFFA3FF25FC83A1F92A4B" blockId="5.[151,473,825,852]" box="[151,473,825,852]" pageId="5" pageNumber="46">
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Genus 
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<emphasis id="B90B1C51FFAAFFA3FF5CFC83A1432A4C" bold="true" box="[238,355,825,852]" italics="true" pageId="5" pageNumber="46">Capobula</emphasis>
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<taxonomicNameLabel id="A238A12AFFAAFFA3FED9FC83A1F92A4B" box="[363,473,825,851]" pageId="5" pageNumber="46" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<paragraph id="8BC0C043FFAAFFA3FF25FC3AA29A2A82" blockId="5.[151,698,896,922]" box="[151,698,896,922]" pageId="5" pageNumber="46">
<emphasis id="B90B1C51FFAAFFA3FF25FC3AA11C2A82" bold="true" box="[151,316,896,922]" pageId="5" pageNumber="46">
<typeStatus id="54C47EE1FFAAFFA3FF25FC3AA0F42A82" box="[151,212,896,922]" pageId="5" pageNumber="46">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C7FBBC0FFAAFFA3FEF1FC3AA29A2A82" authority="Simon, 1896 a" authorityName="Simon" authorityYear="1896" box="[323,698,896,922]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="infima">
<emphasis id="B90B1C51FFAAFFA3FEF1FC3AA2302A82" box="[323,528,896,922]" italics="true" pageId="5" pageNumber="46">Orthobula infima</emphasis>
<bibRefCitation id="EFEEBDB2FFAAFFA3FDA5FC3AA29A2A82" author="Simon, E." box="[535,698,896,922]" pageId="5" pageNumber="46" pagination="400 - 422" refId="ref18785" refString="Simon, E. (1896 a) Descriptions d'arachnides nouveaux de la famille des Clubionidae. Annales de la Societe Entomologique de Belgique, 40, 400 - 422." type="journal article" year="1896">Simon, 1896a</bibRefCitation>
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</paragraph>
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<caption id="DF0090CBFFAAFFA3FF25F8F7A2AB2EED" ID-DOI="http://doi.org/10.5281/zenodo.4596103" ID-Zenodo-Dep="4596103" httpUri="https://zenodo.org/record/4596103/files/figure.png" pageId="5" pageNumber="46" startId="5.[151,264,1869,1893]" targetBox="[151,1436,946,1846]" targetPageId="5">
<paragraph id="8BC0C043FFAAFFA3FF25F8F7A2AB2EED" blockId="5.[151,1436,1869,2038]" pageId="5" pageNumber="46">
<emphasis id="B90B1C51FFAAFFA3FF25F8F7A1672E7D" bold="true" box="[151,327,1869,1893]" pageId="5" pageNumber="46">FIGURES 3–10.</emphasis>
<taxonomicName id="4C7FBBC0FFAAFFA3FEFEF8F7A1902E7D" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[332,432,1869,1893]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="46" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90B1C51FFAAFFA3FEFEF8F7A1902E7D" box="[332,432,1869,1893]" italics="true" pageId="5" pageNumber="46">Capobula</emphasis>
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<emphasis id="B90B1C51FFAAFFA3FE07F8F7A2332E7D" bold="true" box="[437,531,1869,1893]" pageId="5" pageNumber="46">
<taxonomicNameLabel id="A238A12AFFAAFFA3FE07F8F7A2332E7D" box="[437,531,1869,1893]" pageId="5" pageNumber="46" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
dorsal habitus photographs: 3 
<taxonomicName id="4C7FBBC0FFAAFFA3FCF5F8F4A39B2E7D" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[839,955,1870,1893]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="capensis" status="sp. nov.">
<emphasis id="B90B1C51FFAAFFA3FCF5F8F4A39B2E7D" box="[839,955,1870,1893]" italics="true" pageId="5" pageNumber="46">C. capensis</emphasis>
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<emphasis id="B90B1C51FFAAFFA3FC73F8F7A4072E7D" bold="true" box="[961,1063,1869,1893]" pageId="5" pageNumber="46">
<taxonomicNameLabel id="A238A12AFFAAFFA3FC73F8F7A4072E7D" box="[961,1063,1869,1893]" pageId="5" pageNumber="46" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, female holotype from Pakhuis Pass; 4 Same, paratype male from Pakhuis Pass; 5 
<taxonomicName id="4C7FBBC0FFAAFFA3FDE8F8C8A36F2E91" authority="(Simon, 1896)" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" baseAuthorityName="Simon" baseAuthorityYear="1896" box="[602,847,1905,1929]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="infima" status="comb. nov.">
<emphasis id="B90B1C51FFAAFFA3FDE8F8C8A2982E91" box="[602,696,1905,1929]" italics="true" pageId="5" pageNumber="46">C. infima</emphasis>
(Simon, 1896)
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<emphasis id="B90B1C51FFAAFFA3FCE6F8CBA3E72E91" bold="true" box="[852,967,1905,1929]" pageId="5" pageNumber="46">
<taxonomicNameLabel id="A238A12AFFAAFFA3FCE6F8CBA3E72E91" box="[852,967,1905,1929]" pageId="5" pageNumber="46" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, female from Fisherhaven; 6 Same, male from De Hoop Nature Reserve; 7 
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<emphasis id="B90B1C51FFAAFFA3FE72F82CA2182EB5" box="[448,568,1942,1965]" italics="true" pageId="5" pageNumber="46">C. montana</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFAAFFA3FDF2F82FA2882EB5" bold="true" box="[576,680,1941,1965]" pageId="5" pageNumber="46">
<taxonomicNameLabel id="A238A12AFFAAFFA3FDF2F82FA2882EB5" box="[576,680,1941,1965]" pageId="5" pageNumber="46" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, female holotype from Mount Coke State Forest; 8 Same, paratype male from Shamwari Game Reserve; 9 
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<emphasis id="B90B1C51FFAAFFA3FE43F800A2542EC9" box="[497,628,1977,2001]" italics="true" pageId="5" pageNumber="46">C. neethlingi</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFAAFFA3FDC9F803A2C22EC9" bold="true" box="[635,738,1977,2001]" pageId="5" pageNumber="46">
<taxonomicNameLabel id="A238A12AFFAAFFA3FDC9F803A2C22EC9" box="[635,738,1977,2001]" pageId="5" pageNumber="46" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, female holotype from Saasveld Pass; 10 
<taxonomicName id="4C7FBBC0FFAAFFA3FB3FF800A5082EC9" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[1165,1320,1977,2001]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="46" phylum="Arthropoda" rank="species" species="ukhahlamba" status="sp. nov.">
<emphasis id="B90B1C51FFAAFFA3FB3FF800A5082EC9" box="[1165,1320,1977,2001]" italics="true" pageId="5" pageNumber="46">C. ukhahlamba</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFAAFFA3FA9CF803A5B52EC9" bold="true" box="[1326,1429,1977,2001]" pageId="5" pageNumber="46">
<taxonomicNameLabel id="A238A12AFFAAFFA3FA9CF803A5B52EC9" box="[1326,1429,1977,2001]" pageId="5" pageNumber="46" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, paratype female from Royal Natal National Park.
</paragraph>
</caption>
<subSubSection id="C36593C8FFA9FFA0FF25FF2CA5B22B9E" pageId="6" pageNumber="47" type="diagnosis">
<paragraph id="8BC0C043FFA9FFA0FF25FF2CA5B22B9E" blockId="6.[151,1437,150,753]" pageId="6" pageNumber="47">
<emphasis id="B90B1C51FFA9FFA0FF25FF2CA13229A8" bold="true" box="[151,274,150,176]" pageId="6" pageNumber="47">Diagnosis.</emphasis>
<taxonomicName id="4C7FBBC0FFA9FFA0FEA5FF2DA1A529A9" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[279,389,151,177]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90B1C51FFA9FFA0FEA5FF2DA1A529A9" box="[279,389,151,177]" italics="true" pageId="6" pageNumber="47">Capobula</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFA9FFA0FE3EFF2DA1D429A9" bold="true" box="[396,500,151,177]" pageId="6" pageNumber="47">
<taxonomicNameLabel id="A238A12AFFA9FFA0FE3EFF2DA1D429A9" box="[396,500,151,177]" pageId="6" pageNumber="47" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be recognized from all other trachelid genera except 
<taxonomicName id="4C7FBBC0FFA9FFA0FBDCFF2DA4C329A9" authorityName="Simon" authorityYear="1897" box="[1134,1251,151,177]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFA9FFA0FBDCFF2DA4C329A9" box="[1134,1251,151,177]" italics="true" pageId="6" pageNumber="47">Orthobula</emphasis>
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by the combination of a large number of paired ventral spines on the anterior tibiae, metatarsi and tarsi, the absence of ventral cusps in both sexes, the deep pits on the carapace, and the presence of a large ventral sclerite on the abdomen of males. Although 
<taxonomicName id="4C7FBBC0FFA9FFA0FF59FEB9A2162805" authority="Haddad, 2006" authorityName="Haddad" authorityYear="2006" box="[235,566,259,285]" class="Arachnida" family="Corinnidae" genus="Spinotrachelas" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFA9FFA0FF59FEB9A1B32805" box="[235,403,259,285]" italics="true" pageId="6" pageNumber="47">Spinotrachelas</emphasis>
<bibRefCitation id="EFEEBDB2FFA9FFA0FE28FEB9A2162805" author="Haddad, C. R." box="[410,566,259,285]" pageId="6" pageNumber="47" pagination="85 - 93" refId="ref16673" refString="Haddad, C. R. (2006) Spinotrachelas, a new genus of tracheline sac spiders from South Africa (Araneae: Corinnidae). African Invertebrates, 47, 85 - 93." type="journal article" year="2006">Haddad, 2006</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C7FBBC0FFA9FFA0FDDEFEB9A2C12805" authorityName="Haddad &amp; Lyle" authorityYear="2008" box="[620,737,259,285]" class="Arachnida" family="Corinnidae" genus="Poachelas" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFA9FFA0FDDEFEB9A2C12805" box="[620,737,259,285]" italics="true" pageId="6" pageNumber="47">Poachelas</emphasis>
</taxonomicName>
also have heavily spined anterior legs, both have ventral cusps on the anterior legs, lack carapace pits, and lack a ventral abdominal scutum (
<bibRefCitation id="EFEEBDB2FFA9FFA0FC48FE9DA4B62859" author="Haddad, C. R." box="[1018,1174,295,321]" pageId="6" pageNumber="47" pagination="85 - 93" refId="ref16673" refString="Haddad, C. R. (2006) Spinotrachelas, a new genus of tracheline sac spiders from South Africa (Araneae: Corinnidae). African Invertebrates, 47, 85 - 93." type="journal article" year="2006">Haddad 2006</bibRefCitation>
; 
<bibRefCitation id="EFEEBDB2FFA9FFA0FB12FE9DA5B12859" author="Haddad, C. R. &amp; Lyle, R." box="[1184,1425,295,321]" pageId="6" pageNumber="47" pagination="37 - 76" refId="ref16709" refString="Haddad, C. R. &amp; Lyle, R. (2008) Three new genera of tracheline sac spiders from southern Africa (Araneae: Corinnidae). African Invertebrates, 49, 37 - 76. https: // doi. org / 10.5733 / afin. 049.0204" type="journal article" year="2008">Haddad &amp; Lyle 2008</bibRefCitation>
). Female 
<taxonomicName id="4C7FBBC0FFA9FFA0FF40FEF1A140287D" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[242,352,331,357]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90B1C51FFA9FFA0FF40FEF1A140287D" box="[242,352,331,357]" italics="true" pageId="6" pageNumber="47">Capobula</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFA9FFA0FEDBFEF1A1F3287D" bold="true" box="[361,467,331,357]" pageId="6" pageNumber="47">
<taxonomicNameLabel id="A238A12AFFA9FFA0FEDBFEF1A1F3287D" box="[361,467,331,357]" pageId="6" pageNumber="47" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be recognized by the laterally positioned, teardrop-shaped primary spermathecae that are in the same transverse plane as the copulatory openings, while in 
<taxonomicName id="4C7FBBC0FFA9FFA0FC7DFED5A4642891" authorityName="Simon" authorityYear="1897" box="[975,1092,367,393]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFA9FFA0FC7DFED5A4642891" box="[975,1092,367,393]" italics="true" pageId="6" pageNumber="47">Orthobula</emphasis>
</taxonomicName>
the spermathecae are oval and positioned posteriorly, close to the midline of the epigyne, and far from the anteriorly positioned copulatory openings (e.g. 
<bibRefCitation id="EFEEBDB2FFA9FFA0FEB4FE0DA1C528C9" author="Marusik, Y. M. &amp; Ozkutuk, R. S. &amp; Kunt, K. B." box="[262,485,439,465]" pageId="6" pageNumber="47" pagination="157 - 162" refId="ref18076" refString="Marusik, Y. M., Ozkutuk, R. S. &amp; Kunt, K. B. (2013) On the identity and distribution of the poorly known spider Orthobula charitonovi (Mikhailov, 1986) (Aranei: Corinnidae). Arthropoda Selecta, 22, 157 - 162." type="journal article" year="2013">
Marusik 
<emphasis id="B90B1C51FFA9FFA0FED8FE0DA18228C9" box="[362,418,439,465]" italics="true" pageId="6" pageNumber="47">et al.</emphasis>
2013
</bibRefCitation>
: figs 19–22). Further, 
<taxonomicName id="4C7FBBC0FFA9FFA0FD6FFE0DA36B28C9" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[733,843,439,465]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90B1C51FFA9FFA0FD6FFE0DA36B28C9" box="[733,843,439,465]" italics="true" pageId="6" pageNumber="47">Capobula</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFA9FFA0FCE1FE0DA39C28C9" bold="true" box="[851,956,439,465]" pageId="6" pageNumber="47">
<taxonomicNameLabel id="A238A12AFFA9FFA0FCE1FE0DA39C28C9" box="[851,956,439,465]" pageId="6" pageNumber="47" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
lack pits along the midline of the carapace (
<figureCitation id="1344DCC6FFA9FFA0FF2DFE61A13228ED" box="[159,274,475,501]" captionStart-0="FIGURES 3–10" captionStart-1="FIGURES 11–19" captionStartId-0="5.[151,264,1869,1893]" captionStartId-1="6.[151,264,1833,1857]" captionTargetBox-0="[151,1436,946,1846]" captionTargetBox-1="[151,1436,777,1810]" captionTargetId-0="figure-265@5.[151,1436,946,1846]" captionTargetId-1="figure-327@6.[151,1436,777,1810]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 3–10. Capobula gen. nov. dorsal habitus photographs: 3 C. capensis spec. nov., female holotype from Pakhuis Pass; 4 Same, paratype male from Pakhuis Pass; 5 C. infima (Simon, 1896) comb. nov., female from Fisherhaven; 6 Same, male from De Hoop Nature Reserve; 7 C. montana spec. nov., female holotype from Mount Coke State Forest; 8 Same, paratype male from Shamwari Game Reserve; 9 C. neethlingi spec. nov., female holotype from Saasveld Pass; 10 C. ukhahlamba spec. nov., paratype female from Royal Natal National Park." captionText-1="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi-0="http://doi.org/10.5281/zenodo.4596103" figureDoi-1="http://doi.org/10.5281/zenodo.4596107" httpUri-0="https://zenodo.org/record/4596103/files/figure.png" httpUri-1="https://zenodo.org/record/4596107/files/figure.png" pageId="6" pageNumber="47">Figs 3–11</figureCitation>
), which are present in 
<taxonomicName id="4C7FBBC0FFA9FFA0FDA2FE61A2A528ED" authorityName="Simon" authorityYear="1897" box="[528,645,475,501]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFA9FFA0FDA2FE61A2A528ED" box="[528,645,475,501]" italics="true" pageId="6" pageNumber="47">Orthobula</emphasis>
</taxonomicName>
(e.g. 
<bibRefCitation id="EFEEBDB2FFA9FFA0FD77FE61A38528ED" author="Marusik, Y. M. &amp; Ozkutuk, R. S. &amp; Kunt, K. B." box="[709,933,475,501]" pageId="6" pageNumber="47" pagination="157 - 162" refId="ref18076" refString="Marusik, Y. M., Ozkutuk, R. S. &amp; Kunt, K. B. (2013) On the identity and distribution of the poorly known spider Orthobula charitonovi (Mikhailov, 1986) (Aranei: Corinnidae). Arthropoda Selecta, 22, 157 - 162." type="journal article" year="2013">
Marusik 
<emphasis id="B90B1C51FFA9FFA0FC9BFE61A34228ED" box="[809,866,475,501]" italics="true" pageId="6" pageNumber="47">et al.</emphasis>
2013
</bibRefCitation>
: figs 1, 4; 
<bibRefCitation id="EFEEBDB2FFA9FFA0FBA8FE61A49C28ED" author="Ramirez, M. J." box="[1050,1212,475,501]" pageId="6" pageNumber="47" pagination="1" refId="ref18592" refString="Ramirez, M. J. (2014) The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1 ‾ 374. https: // doi. org / 10.1206 / 821.1" type="journal article" year="2014">Ramírez 2014</bibRefCitation>
: fig. 6A). 
<taxonomicName id="4C7FBBC0FFA9FFA0FA9CFE61A5BC28ED" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[1326,1436,475,501]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90B1C51FFA9FFA0FA9CFE61A5BC28ED" box="[1326,1436,475,501]" italics="true" pageId="6" pageNumber="47">Capobula</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFA9FFA0FF25FE45A1222B01" bold="true" box="[151,258,511,537]" pageId="6" pageNumber="47">
<taxonomicNameLabel id="A238A12AFFA9FFA0FF25FE45A1222B01" box="[151,258,511,537]" pageId="6" pageNumber="47" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
can be separated from 
<taxonomicName id="4C7FBBC0FFA9FFA0FDA3FE45A2A62B01" authorityName="Simon" authorityYear="1897" box="[529,646,511,537]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFA9FFA0FDA3FE45A2A62B01" box="[529,646,511,537]" italics="true" pageId="6" pageNumber="47">Orthobula</emphasis>
</taxonomicName>
by the male palpal tegulum being pear-shaped and approximately as broad as the cymbium, and the embolus with a broad base, clear bend near its midpoint, and tip directed retrodistally (e.g. 
<figureCitation id="1344DCC6FFA9FFA0FF63FDFDA1062B79" box="[209,294,582,609]" captionStart="FIGURES 47–53" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1436,489,1867]" captionTargetId="figure-157@10.[151,1436,489,1867]" captionTargetPageId="10" captionText="FIGURES 47–53. Capobula infima (Simon, 1896) comb. nov., female (47) and male (48–53) from De Hoop Nature Reserve, SEM photographs: 47 Epigyne, ventral; 48 Left palpal femur and patella, ventral; 49 Left palpal patella, retrolateral (rectangle indicating lyriform organ associated with patellar apophysis); 50 Lyriform organ, enlarged; 51 Right palp, detail of retrolateral tibial apophysis; 52 Left palp, ventral; 53 Embolus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596117" httpUri="https://zenodo.org/record/4596117/files/figure.png" pageId="6" pageNumber="47">Fig. 52</figureCitation>
). In 
<taxonomicName id="4C7FBBC0FFA9FFA0FEE8FDFDA1EF2B79" authorityName="Simon" authorityYear="1897" box="[346,463,583,609]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFA9FFA0FEE8FDFDA1EF2B79" box="[346,463,583,609]" italics="true" pageId="6" pageNumber="47">Orthobula</emphasis>
</taxonomicName>
the tegulum is subtriangular and strongly swollen basally towards the prolateral side, and the embolus usually straight, narrow, finely coiled and directed distally (e.g. 
<bibRefCitation id="EFEEBDB2FFA9FFA0FB90FDD1A5212B9D" author="Marusik, Y. M. &amp; Ozkutuk, R. S. &amp; Kunt, K. B." box="[1058,1281,619,645]" pageId="6" pageNumber="47" pagination="157 - 162" refId="ref18076" refString="Marusik, Y. M., Ozkutuk, R. S. &amp; Kunt, K. B. (2013) On the identity and distribution of the poorly known spider Orthobula charitonovi (Mikhailov, 1986) (Aranei: Corinnidae). Arthropoda Selecta, 22, 157 - 162." type="journal article" year="2013">
Marusik 
<emphasis id="B90B1C51FFA9FFA0FB34FDD1A49E2B9D" box="[1158,1214,619,645]" italics="true" pageId="6" pageNumber="47">et al.</emphasis>
2013
</bibRefCitation>
: figs 11–16).
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFA9FFA0FF25FD35A19A2BE9" pageId="6" pageNumber="47" type="etymology">
<paragraph id="8BC0C043FFA9FFA0FF25FD35A19A2BE9" blockId="6.[151,1437,150,753]" pageId="6" pageNumber="47">
<emphasis id="B90B1C51FFA9FFA0FF25FD35A13E2BB1" bold="true" box="[151,286,655,681]" pageId="6" pageNumber="47">Etymology.</emphasis>
The genus name is a combination of 
<collectingRegion id="49BB0EA1FFA9FFA0FD64FD35A3322BB1" box="[726,786,655,681]" country="South Africa" name="Western Cape" pageId="6" pageNumber="47">Cape</collectingRegion>
, referring to the Western and 
<collectingRegion id="49BB0EA1FFA9FFA0FBDEFD35A5242BB1" box="[1132,1284,655,681]" country="South Africa" name="Eastern Cape" pageId="6" pageNumber="47">Eastern Cape</collectingRegion>
Provinces of 
<collectingCountry id="F36880D3FFA9FFA0FF25FD09A1072BD5" box="[151,295,691,717]" name="South Africa" pageId="6" pageNumber="47">South Africa</collectingCountry>
from where the majority of the species have been recorded, and 
<taxonomicName id="4C7FBBC0FFA9FFA0FBB4FD09A45B2BD5" authorityName="Simon" authorityYear="1897" box="[1030,1147,691,717]" class="Arachnida" family="Corinnidae" genus="Orthobula" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B90B1C51FFA9FFA0FBB4FD09A45B2BD5" box="[1030,1147,691,717]" italics="true" pageId="6" pageNumber="47">Orthobula</emphasis>
</taxonomicName>
, to which it is closely related. Gender is feminine.
</paragraph>
</subSubSection>
<caption id="DF0090CBFFA9FFA0FF25F893A44E2EB5" ID-DOI="http://doi.org/10.5281/zenodo.4596107" ID-Zenodo-Dep="4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="6" pageNumber="47" startId="6.[151,264,1833,1857]" targetBox="[151,1436,777,1810]" targetPageId="6">
<paragraph id="8BC0C043FFA9FFA0FF25F893A44E2EB5" blockId="6.[151,1436,1833,1966]" pageId="6" pageNumber="47">
<emphasis id="B90B1C51FFA9FFA0FF25F893A1732E59" bold="true" box="[151,339,1833,1857]" pageId="6" pageNumber="47">FIGURES 11–19.</emphasis>
<taxonomicName id="4C7FBBC0FFA9FFA0FEE8F893A2802E59" authority="(Simon, 1896)" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" baseAuthorityName="Simon" baseAuthorityYear="1896" box="[346,672,1833,1857]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="47" phylum="Arthropoda" rank="species" species="infima" status="comb. nov.">
<emphasis id="B90B1C51FFA9FFA0FEE8F893A2262E59" box="[346,518,1833,1857]" italics="true" pageId="6" pageNumber="47">Capobula infima</emphasis>
(Simon, 1896)
</taxonomicName>
<emphasis id="B90B1C51FFA9FFA0FD15F890A33C2E5A" bold="true" box="[679,796,1834,1858]" pageId="6" pageNumber="47">
<taxonomicNameLabel id="A238A12AFFA9FFA0FD15F890A33C2E5A" box="[679,796,1834,1858]" pageId="6" pageNumber="47" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral.
</paragraph>
</caption>
<subSubSection id="C36593C8FFA8FFACFF75FF2CA4A728B5" lastPageId="10" lastPageNumber="51" pageId="7" pageNumber="48" type="description">
<paragraph id="8BC0C043FFA8FFACFF75FF2CA404287D" blockId="7.[151,1437,150,467]" lastBlockId="10.[151,1437,151,465]" lastPageId="10" lastPageNumber="51" pageId="7" pageNumber="48">
<emphasis id="B90B1C51FFA8FFA1FF75FF2CA17929A8" bold="true" box="[199,345,150,176]" pageId="7" pageNumber="48">Description.</emphasis>
Small spiders, 
<quantity id="4C876DA6FFA8FFA1FDBDFF2CA29629A9" box="[527,694,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.025" metricValueMax="2.35" metricValueMin="1.7" pageId="7" pageNumber="48" unit="mm" value="2.025" valueMax="2.35" valueMin="1.7">1.70–2.35 mm</quantity>
in total length; carapace bright orange to deep red-brown (
<figureCitation id="1344DCC6FFA8FFA1FAD8FF2DA0F229CD" captionStart="FIGURES 3–10" captionStartId="5.[151,264,1869,1893]" captionTargetBox="[151,1436,946,1846]" captionTargetId="figure-265@5.[151,1436,946,1846]" captionTargetPageId="5" captionText="FIGURES 3–10. Capobula gen. nov. dorsal habitus photographs: 3 C. capensis spec. nov., female holotype from Pakhuis Pass; 4 Same, paratype male from Pakhuis Pass; 5 C. infima (Simon, 1896) comb. nov., female from Fisherhaven; 6 Same, male from De Hoop Nature Reserve; 7 C. montana spec. nov., female holotype from Mount Coke State Forest; 8 Same, paratype male from Shamwari Game Reserve; 9 C. neethlingi spec. nov., female holotype from Saasveld Pass; 10 C. ukhahlamba spec. nov., paratype female from Royal Natal National Park." figureDoi="http://doi.org/10.5281/zenodo.4596103" httpUri="https://zenodo.org/record/4596103/files/figure.png" pageId="7" pageNumber="48">Figs 3–10</figureCitation>
); carapace oval, broadest at middle of coxae II, eye region narrow, fovea indistinct, a short shallow depression (
<figureCitation id="1344DCC6FFA8FFA1FF12FF65A0D029E1" box="[160,240,223,249]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="7" pageNumber="48">Fig. 11</figureCitation>
); posterior margin very slightly concave; carapace surface finely granulose, with series of deep pits forming striae, pits absent from midline of carapace (
<figureCitation id="1344DCC6FFA8FFA1FD3EFEB9A32C2805" box="[652,780,259,285]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="7" pageNumber="48">Figs 11, 12</figureCitation>
), each pit with central pore; surface sparsely covered with scattered fine curved setae (
<figureCitation id="1344DCC6FFA8FFA1FE7BFE9DA2692859" box="[457,585,295,321]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="7" pageNumber="48">Figs 12, 13</figureCitation>
). All eyes surrounded by black rings (
<figureCitation id="1344DCC6FFA8FFA1FC5BFE9DA47C2859" box="[1001,1116,295,321]" captionStart="FIGURES 3–10" captionStartId="5.[151,264,1869,1893]" captionTargetBox="[151,1436,946,1846]" captionTargetId="figure-265@5.[151,1436,946,1846]" captionTargetPageId="5" captionText="FIGURES 3–10. Capobula gen. nov. dorsal habitus photographs: 3 C. capensis spec. nov., female holotype from Pakhuis Pass; 4 Same, paratype male from Pakhuis Pass; 5 C. infima (Simon, 1896) comb. nov., female from Fisherhaven; 6 Same, male from De Hoop Nature Reserve; 7 C. montana spec. nov., female holotype from Mount Coke State Forest; 8 Same, paratype male from Shamwari Game Reserve; 9 C. neethlingi spec. nov., female holotype from Saasveld Pass; 10 C. ukhahlamba spec. nov., paratype female from Royal Natal National Park." figureDoi="http://doi.org/10.5281/zenodo.4596103" httpUri="https://zenodo.org/record/4596103/files/figure.png" pageId="7" pageNumber="48">Figs 3–10</figureCitation>
); anterior eye row procurved in anterior view, recurved in dorsal view, anterior median eyes approximately ¾ anterior lateral eye diameter; anterior median eyes separated by approximately ½ their diameter, nearly touching anterior lateral eyes (
<figureCitation id="1344DCC6FFA8FFA1FAB8FED5A5AE2891" box="[1290,1422,367,393]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="7" pageNumber="48">Figs 13, 14</figureCitation>
); posterior eye row recurved in dorsal view, posterior median eyes oval, posterior lateral eyes round; posterior median eyes approximately 1¼ times posterior lateral eye diameter; MOQ narrower anteriorly than posteriorly, length and posterior width approximately equal. Chilum indistinct, a tiny transverse sclerite; cheliceral promargin and retromargin with two teeth each, cheliceral escort seta present (
<figureCitation id="1344DCC6FFA7FFAEFC94FF01A38829CE" box="[806,936,186,214]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="8" pageNumber="49">Figs 15, 16</figureCitation>
); fang with distinct serrula (
<figureCitation id="1344DCC6FFA7FFAEFB56FF01A51829CE" box="[1252,1336,187,214]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="8" pageNumber="49">Fig. 16</figureCitation>
); endites convergent, not sexually dimorphic, slightly notched laterally (
<figureCitation id="1344DCC6FFA7FFAEFCC3FF65A3E729E1" box="[881,967,223,249]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="8" pageNumber="49">Fig. 17</figureCitation>
), with distinct serrula comprising sharp, ventrally curved denticles (
<figureCitation id="1344DCC6FFA7FFAEFE77FEB9A2372805" box="[453,535,259,285]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="8" pageNumber="49">Fig. 18</figureCitation>
); dense maxillar hair tuft on mesal margins (
<figureCitation id="1344DCC6FFA7FFAEFBB6FEB9A4772805" box="[1028,1111,259,285]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="8" pageNumber="49">Fig. 17</figureCitation>
); labium trapezoidal, slightly wider than long. Pleural bars sclerotised, isolated; sternum shield-shaped, longer than broad, surface smooth with deep pits without pores, sparsely covered in long straight setae (
<figureCitation id="1344DCC6FFA7FFAEFCD4FEF1A39D287D" box="[870,957,331,357]" captionStart="FIGURES 11–19" captionStartId="6.[151,264,1833,1857]" captionTargetBox="[151,1436,777,1810]" captionTargetId="figure-327@6.[151,1436,777,1810]" captionTargetPageId="6" captionText="FIGURES 11–19. Capobula infima (Simon, 1896) comb. nov., female from De Hoop Nature Reserve, SEM photographs: 11 Carapace, dorsal; 12 Same, detail of carapace texture, showing pits and fine tubercles on striae, dorsal; 13 Eye region, dorsal; 14 Eye region, frontal; 15 Chelicerae, ventral (arrow indicating promarginal escort seta); 16 Fang, ventral (arrow indicating fang serrula); 17 Endites and labium, ventral; 18 Detail of endite serrula, ventral; 19 Sternum, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596107" httpUri="https://zenodo.org/record/4596107/files/figure.png" pageId="8" pageNumber="49">Figs 19</figureCitation>
, 
<figureCitation id="1344DCC6FFA7FFAEFC7BFEF1A3C9287D" box="[969,1001,331,357]" captionStart="FIGURES 20–29" captionStartId="7.[151,264,1869,1893]" captionTargetBox="[151,1436,489,1846]" captionTargetId="figure-173@7.[151,1436,489,1846]" captionTargetPageId="7" captionText="FIGURES 20–29. Capobula infima (Simon, 1896) comb. nov., female (20, 23–29) and male (21, 22) from De Hoop Nature Reserve, SEM photographs: 20 Detail of sternum texture; 21 Femur I, ventral; 22 Femur I, detail of fine distal tubercles on ventral surface, ventral; 23 Patella III, prolateral (arrow indicating lyriform organ); 24 Patella IV, retrolateral (arrow indicating lyriform organ at end of patellar indentation: PI); 25 Detail of lyriform organ at end of PI, retrolateral ventral; 26 Tibia I spines, ventral; 27 Metatarsus I spines, ventral; 28 Metatarsus I, detail of spine bases, ventral; 29 Tarsus I spines, ventro-distal." figureDoi="http://doi.org/10.5281/zenodo.4596109" httpUri="https://zenodo.org/record/4596109/files/figure.png" pageId="8" pageNumber="49">20</figureCitation>
); precoxal triangles present, intercoxal sclerites present between all coxal pairs. Leg formula 4132, sparsely covered in long fine setae; femora I with mesal convex curvature, all femora strongly constricted proximally, with sparse ventral tubercles in basal half (
<figureCitation id="1344DCC6FFA7FFAEFA8DFE29A09728C9" captionStart="FIGURES 20–29" captionStartId="7.[151,264,1869,1893]" captionTargetBox="[151,1436,489,1846]" captionTargetId="figure-173@7.[151,1436,489,1846]" captionTargetPageId="7" captionText="FIGURES 20–29. Capobula infima (Simon, 1896) comb. nov., female (20, 23–29) and male (21, 22) from De Hoop Nature Reserve, SEM photographs: 20 Detail of sternum texture; 21 Femur I, ventral; 22 Femur I, detail of fine distal tubercles on ventral surface, ventral; 23 Patella III, prolateral (arrow indicating lyriform organ); 24 Patella IV, retrolateral (arrow indicating lyriform organ at end of patellar indentation: PI); 25 Detail of lyriform organ at end of PI, retrolateral ventral; 26 Tibia I spines, ventral; 27 Metatarsus I spines, ventral; 28 Metatarsus I, detail of spine bases, ventral; 29 Tarsus I spines, ventro-distal." figureDoi="http://doi.org/10.5281/zenodo.4596109" httpUri="https://zenodo.org/record/4596109/files/figure.png" pageId="8" pageNumber="49">Figs 21, 22</figureCitation>
); patellae with small lyriform organ on prolateral side (
<figureCitation id="1344DCC6FFA7FFAEFC97FE0DA35928C9" box="[805,889,439,465]" captionStart="FIGURES 20–29" captionStartId="7.[151,264,1869,1893]" captionTargetBox="[151,1436,489,1846]" captionTargetId="figure-173@7.[151,1436,489,1846]" captionTargetPageId="7" captionText="FIGURES 20–29. Capobula infima (Simon, 1896) comb. nov., female (20, 23–29) and male (21, 22) from De Hoop Nature Reserve, SEM photographs: 20 Detail of sternum texture; 21 Femur I, ventral; 22 Femur I, detail of fine distal tubercles on ventral surface, ventral; 23 Patella III, prolateral (arrow indicating lyriform organ); 24 Patella IV, retrolateral (arrow indicating lyriform organ at end of patellar indentation: PI); 25 Detail of lyriform organ at end of PI, retrolateral ventral; 26 Tibia I spines, ventral; 27 Metatarsus I spines, ventral; 28 Metatarsus I, detail of spine bases, ventral; 29 Tarsus I spines, ventro-distal." figureDoi="http://doi.org/10.5281/zenodo.4596109" httpUri="https://zenodo.org/record/4596109/files/figure.png" pageId="8" pageNumber="49">Fig. 23</figureCitation>
), patellar indentation on retrolateral side narrow, with lyriform organ at proximal end (
<figureCitation id="1344DCC6FFA7FFAEFD86FE61A29428ED" box="[564,692,474,501]" captionStart="FIGURES 20–29" captionStartId="7.[151,264,1869,1893]" captionTargetBox="[151,1436,489,1846]" captionTargetId="figure-173@7.[151,1436,489,1846]" captionTargetPageId="7" captionText="FIGURES 20–29. Capobula infima (Simon, 1896) comb. nov., female (20, 23–29) and male (21, 22) from De Hoop Nature Reserve, SEM photographs: 20 Detail of sternum texture; 21 Femur I, ventral; 22 Femur I, detail of fine distal tubercles on ventral surface, ventral; 23 Patella III, prolateral (arrow indicating lyriform organ); 24 Patella IV, retrolateral (arrow indicating lyriform organ at end of patellar indentation: PI); 25 Detail of lyriform organ at end of PI, retrolateral ventral; 26 Tibia I spines, ventral; 27 Metatarsus I spines, ventral; 28 Metatarsus I, detail of spine bases, ventral; 29 Tarsus I spines, ventro-distal." figureDoi="http://doi.org/10.5281/zenodo.4596109" httpUri="https://zenodo.org/record/4596109/files/figure.png" pageId="8" pageNumber="49">Figs 24, 25</figureCitation>
); anterior legs with strong paired ventral spines on tibiae, metatarsi and tarsi (
<figureCitation id="1344DCC6FFA7FFAEFEB8FE45A1AF2B02" box="[266,399,511,538]" captionStart="FIGURES 20–29" captionStartId="7.[151,264,1869,1893]" captionTargetBox="[151,1436,489,1846]" captionTargetId="figure-173@7.[151,1436,489,1846]" captionTargetPageId="7" captionText="FIGURES 20–29. Capobula infima (Simon, 1896) comb. nov., female (20, 23–29) and male (21, 22) from De Hoop Nature Reserve, SEM photographs: 20 Detail of sternum texture; 21 Femur I, ventral; 22 Femur I, detail of fine distal tubercles on ventral surface, ventral; 23 Patella III, prolateral (arrow indicating lyriform organ); 24 Patella IV, retrolateral (arrow indicating lyriform organ at end of patellar indentation: PI); 25 Detail of lyriform organ at end of PI, retrolateral ventral; 26 Tibia I spines, ventral; 27 Metatarsus I spines, ventral; 28 Metatarsus I, detail of spine bases, ventral; 29 Tarsus I spines, ventro-distal." figureDoi="http://doi.org/10.5281/zenodo.4596109" httpUri="https://zenodo.org/record/4596109/files/figure.png" pageId="8" pageNumber="49">Figs 26–29</figureCitation>
); metatarsi IV with sparse chemosensory setae and trichobothria dorsally, ventral preening brush at distal end (
<figureCitation id="1344DCC6FFA7FFAEFEC5FD99A1EC2B25" box="[375,460,547,573]" captionStart="FIGURES 30–38" captionStartId="8.[151,264,1832,1856]" captionTargetBox="[151,1436,777,1809]" captionTargetId="figure-313@8.[151,1436,777,1809]" captionTargetPageId="8" captionText="FIGURES 30–38. Capobula infima (Simon, 1896) comb. nov. female from De Hoop Nature Reserve, SEM photographs: 30 Metatarsus IV, distal preening brush and metatarsal stopper, retrolateral; 31 Tarsus I, dorsal, indicating chemosensory seta (cs), trichobothria (tr) and tarsal organ (to); 32 Tarsus I, trichobothrium, dorsal; 33 Same, detail of trichobothrium base, dorsal; 34 Tarsus I, detail of tarsal organ, dorsal; 35 Tarsal claws, leg IV, prolateral; 36 Tarsal claws, leg III, distal; 37 Palpal claw, prolateral distal; 38 Dorsal abdominal setae, dorsal, arrow indicating sigillum." figureDoi="http://doi.org/10.5281/zenodo.4596113" httpUri="https://zenodo.org/record/4596113/files/figure.png" pageId="8" pageNumber="49">Fig. 30</figureCitation>
); tarsi with sparse tactile hairs, few dorsal trichobothria and chemosensory setae (
<figureCitation id="1344DCC6FFA7FFAEFAD8FD99A0C12B79" captionStart="FIGURES 30–38" captionStartId="8.[151,264,1832,1856]" captionTargetBox="[151,1436,777,1809]" captionTargetId="figure-313@8.[151,1436,777,1809]" captionTargetPageId="8" captionText="FIGURES 30–38. Capobula infima (Simon, 1896) comb. nov. female from De Hoop Nature Reserve, SEM photographs: 30 Metatarsus IV, distal preening brush and metatarsal stopper, retrolateral; 31 Tarsus I, dorsal, indicating chemosensory seta (cs), trichobothria (tr) and tarsal organ (to); 32 Tarsus I, trichobothrium, dorsal; 33 Same, detail of trichobothrium base, dorsal; 34 Tarsus I, detail of tarsal organ, dorsal; 35 Tarsal claws, leg IV, prolateral; 36 Tarsal claws, leg III, distal; 37 Palpal claw, prolateral distal; 38 Dorsal abdominal setae, dorsal, arrow indicating sigillum." figureDoi="http://doi.org/10.5281/zenodo.4596113" httpUri="https://zenodo.org/record/4596113/files/figure.png" pageId="8" pageNumber="49">Figs 31, 32</figureCitation>
); trichobothria with sunken distal plate, distal margin of hood overlapping plate, hood with four curved ridges, roughly concentric (
<figureCitation id="1344DCC6FFA7FFAEFE2DFDD1A1D22B9D" box="[415,498,619,645]" captionStart="FIGURES 30–38" captionStartId="8.[151,264,1832,1856]" captionTargetBox="[151,1436,777,1809]" captionTargetId="figure-313@8.[151,1436,777,1809]" captionTargetPageId="8" captionText="FIGURES 30–38. Capobula infima (Simon, 1896) comb. nov. female from De Hoop Nature Reserve, SEM photographs: 30 Metatarsus IV, distal preening brush and metatarsal stopper, retrolateral; 31 Tarsus I, dorsal, indicating chemosensory seta (cs), trichobothria (tr) and tarsal organ (to); 32 Tarsus I, trichobothrium, dorsal; 33 Same, detail of trichobothrium base, dorsal; 34 Tarsus I, detail of tarsal organ, dorsal; 35 Tarsal claws, leg IV, prolateral; 36 Tarsal claws, leg III, distal; 37 Palpal claw, prolateral distal; 38 Dorsal abdominal setae, dorsal, arrow indicating sigillum." figureDoi="http://doi.org/10.5281/zenodo.4596113" httpUri="https://zenodo.org/record/4596113/files/figure.png" pageId="8" pageNumber="49">Fig. 33</figureCitation>
); tarsal organ oval, very slightly elevated from integument, surface finely wrinkled, opening oval and distally placed (
<figureCitation id="1344DCC6FFA7FFAEFDBEFD35A2AD2BB1" box="[524,653,655,681]" captionStart="FIGURES 30–38" captionStartId="8.[151,264,1832,1856]" captionTargetBox="[151,1436,777,1809]" captionTargetId="figure-313@8.[151,1436,777,1809]" captionTargetPageId="8" captionText="FIGURES 30–38. Capobula infima (Simon, 1896) comb. nov. female from De Hoop Nature Reserve, SEM photographs: 30 Metatarsus IV, distal preening brush and metatarsal stopper, retrolateral; 31 Tarsus I, dorsal, indicating chemosensory seta (cs), trichobothria (tr) and tarsal organ (to); 32 Tarsus I, trichobothrium, dorsal; 33 Same, detail of trichobothrium base, dorsal; 34 Tarsus I, detail of tarsal organ, dorsal; 35 Tarsal claws, leg IV, prolateral; 36 Tarsal claws, leg III, distal; 37 Palpal claw, prolateral distal; 38 Dorsal abdominal setae, dorsal, arrow indicating sigillum." figureDoi="http://doi.org/10.5281/zenodo.4596113" httpUri="https://zenodo.org/record/4596113/files/figure.png" pageId="8" pageNumber="49">Figs 31, 34</figureCitation>
); paired tarsal claws short, with two teeth and moderately dense tenant setae forming claw tufts in between (
<figureCitation id="1344DCC6FFA7FFAEFD88FD09A29E2BD6" box="[570,702,690,718]" captionStart="FIGURES 30–38" captionStartId="8.[151,264,1832,1856]" captionTargetBox="[151,1436,777,1809]" captionTargetId="figure-313@8.[151,1436,777,1809]" captionTargetPageId="8" captionText="FIGURES 30–38. Capobula infima (Simon, 1896) comb. nov. female from De Hoop Nature Reserve, SEM photographs: 30 Metatarsus IV, distal preening brush and metatarsal stopper, retrolateral; 31 Tarsus I, dorsal, indicating chemosensory seta (cs), trichobothria (tr) and tarsal organ (to); 32 Tarsus I, trichobothrium, dorsal; 33 Same, detail of trichobothrium base, dorsal; 34 Tarsus I, detail of tarsal organ, dorsal; 35 Tarsal claws, leg IV, prolateral; 36 Tarsal claws, leg III, distal; 37 Palpal claw, prolateral distal; 38 Dorsal abdominal setae, dorsal, arrow indicating sigillum." figureDoi="http://doi.org/10.5281/zenodo.4596113" httpUri="https://zenodo.org/record/4596113/files/figure.png" pageId="8" pageNumber="49">Figs 35, 36</figureCitation>
); palpal claw simple, sharply curved distally (
<figureCitation id="1344DCC6FFA7FFAEFB74FD09A53B2BD5" box="[1222,1307,691,717]" captionStart="FIGURES 30–38" captionStartId="8.[151,264,1832,1856]" captionTargetBox="[151,1436,777,1809]" captionTargetId="figure-313@8.[151,1436,777,1809]" captionTargetPageId="8" captionText="FIGURES 30–38. Capobula infima (Simon, 1896) comb. nov. female from De Hoop Nature Reserve, SEM photographs: 30 Metatarsus IV, distal preening brush and metatarsal stopper, retrolateral; 31 Tarsus I, dorsal, indicating chemosensory seta (cs), trichobothria (tr) and tarsal organ (to); 32 Tarsus I, trichobothrium, dorsal; 33 Same, detail of trichobothrium base, dorsal; 34 Tarsus I, detail of tarsal organ, dorsal; 35 Tarsal claws, leg IV, prolateral; 36 Tarsal claws, leg III, distal; 37 Palpal claw, prolateral distal; 38 Dorsal abdominal setae, dorsal, arrow indicating sigillum." figureDoi="http://doi.org/10.5281/zenodo.4596113" httpUri="https://zenodo.org/record/4596113/files/figure.png" pageId="8" pageNumber="49">Fig. 37</figureCitation>
). Abdomen oval, clearly larger in females than males, with dorsal scutum in males only; dorsum with very sparse fine setae and two pairs of sigilla, prominent in females, barely distinguishable on scutum in males (
<figureCitation id="1344DCC6FFA6FFAFFB2CFF2DA53329A9" box="[1182,1299,151,177]" captionStart="FIGURES 3–10" captionStartId="5.[151,264,1869,1893]" captionTargetBox="[151,1436,946,1846]" captionTargetId="figure-265@5.[151,1436,946,1846]" captionTargetPageId="5" captionText="FIGURES 3–10. Capobula gen. nov. dorsal habitus photographs: 3 C. capensis spec. nov., female holotype from Pakhuis Pass; 4 Same, paratype male from Pakhuis Pass; 5 C. infima (Simon, 1896) comb. nov., female from Fisherhaven; 6 Same, male from De Hoop Nature Reserve; 7 C. montana spec. nov., female holotype from Mount Coke State Forest; 8 Same, paratype male from Shamwari Game Reserve; 9 C. neethlingi spec. nov., female holotype from Saasveld Pass; 10 C. ukhahlamba spec. nov., paratype female from Royal Natal National Park." figureDoi="http://doi.org/10.5281/zenodo.4596103" httpUri="https://zenodo.org/record/4596103/files/figure.png" pageId="9" pageNumber="50">Figs 3–10</figureCitation>
, 
<figureCitation id="1344DCC6FFA6FFAFFA92FF2DA51F29A9" box="[1312,1343,151,177]" captionStart="FIGURES 30–38" captionStartId="8.[151,264,1832,1856]" captionTargetBox="[151,1436,777,1809]" captionTargetId="figure-313@8.[151,1436,777,1809]" captionTargetPageId="8" captionText="FIGURES 30–38. Capobula infima (Simon, 1896) comb. nov. female from De Hoop Nature Reserve, SEM photographs: 30 Metatarsus IV, distal preening brush and metatarsal stopper, retrolateral; 31 Tarsus I, dorsal, indicating chemosensory seta (cs), trichobothria (tr) and tarsal organ (to); 32 Tarsus I, trichobothrium, dorsal; 33 Same, detail of trichobothrium base, dorsal; 34 Tarsus I, detail of tarsal organ, dorsal; 35 Tarsal claws, leg IV, prolateral; 36 Tarsal claws, leg III, distal; 37 Palpal claw, prolateral distal; 38 Dorsal abdominal setae, dorsal, arrow indicating sigillum." figureDoi="http://doi.org/10.5281/zenodo.4596113" httpUri="https://zenodo.org/record/4596113/files/figure.png" pageId="9" pageNumber="50">38</figureCitation>
); venter without post-epigastric sclerites, with large ventral sclerite in males that is wider than long, absent in females; small, weakly sclerotized inframamillary sclerite present in females, indistinct in males. Spinnerets (observed in 
<taxonomicName id="4C7FBBC0FFA6FFAFFA86FF65A5BD29E1" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" baseAuthorityName="Simon" baseAuthorityYear="1896" box="[1332,1437,223,249]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="infima" status="comb. nov.">
<emphasis id="B90B1C51FFA6FFAFFA86FF65A5BD29E1" box="[1332,1437,223,249]" italics="true" pageId="9" pageNumber="50">C. infima</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFA6FFAFFF25FEB9A13A2805" bold="true" box="[151,282,259,285]" pageId="9" pageNumber="50">
<taxonomicNameLabel id="A238A12AFFA6FFAFFF25FEB9A13A2805" box="[151,282,259,285]" pageId="9" pageNumber="50" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
only): female (
<figureCitation id="1344DCC6FFA6FFAFFE7CFEB9A2A12805" box="[462,641,259,285]" captionStart="FIGURES 39–46" captionStartId="9.[151,264,1825,1849]" captionTargetBox="[151,1436,597,1801]" captionTargetId="figure-219@9.[151,1436,597,1801]" captionTargetPageId="9" captionText="FIGURES 39–46. Capobula infima (Simon, 1896) comb. nov., female (39, 41–43) and male (40, 44–46) from De Hoop Nature Reserve, SEM photographs: 39–40 Spinnerets, general view; 41, 44 Anterior lateral spinnerets; 42, 45 Posterior median spinnerets; 43, 46 Posterior lateral spinnerets. Abbreviations: Ac—aciniform gland spigot; Cy—cylindrical gland spigot; mAmp—minor ampullate gland spigot; MAmp—major ampullate gland spigot; Nu—nubbin; Pi—piriform gland spigot; Ta—tartipore." figureDoi="http://doi.org/10.5281/zenodo.4596115" httpUri="https://zenodo.org/record/4596115/files/figure.png" pageId="9" pageNumber="50">Figs 39, 41–43</figureCitation>
): anterior lateral spinnerets of female each with one major ampullate gland spigot, one nubbin and approximately ten piriform gland spigots; posterior median spinnerets of female each with one small minor ampullate gland spigot and five large cylindrical gland spigots; posterior lateral spinnerets of female each with two large cylindrical gland spigots and approximately ten aciniform gland spigots; male (
<figureCitation id="1344DCC6FFA6FFAFFAF3FED5A0C328B6" captionStart="FIGURES 39–46" captionStartId="9.[151,264,1825,1849]" captionTargetBox="[151,1436,597,1801]" captionTargetId="figure-219@9.[151,1436,597,1801]" captionTargetPageId="9" captionText="FIGURES 39–46. Capobula infima (Simon, 1896) comb. nov., female (39, 41–43) and male (40, 44–46) from De Hoop Nature Reserve, SEM photographs: 39–40 Spinnerets, general view; 41, 44 Anterior lateral spinnerets; 42, 45 Posterior median spinnerets; 43, 46 Posterior lateral spinnerets. Abbreviations: Ac—aciniform gland spigot; Cy—cylindrical gland spigot; mAmp—minor ampullate gland spigot; MAmp—major ampullate gland spigot; Nu—nubbin; Pi—piriform gland spigot; Ta—tartipore." figureDoi="http://doi.org/10.5281/zenodo.4596115" httpUri="https://zenodo.org/record/4596115/files/figure.png" pageId="9" pageNumber="50">Figs 40, 44–46</figureCitation>
): anterior lateral spinnerets of male each with one major ampullate gland spigot and five piriform gland spigots; posterior median spinnerets of male each with only one minor ampullate gland spigot and one tartipore evident; posterior lateral spinnerets of male each with only a single tartipore discernible. Female epigyne with small paired copulatory openings positioned anteriorly in epigyne (
<figureCitation id="1344DCC6FFA6FFAFFD5DFE45A3612B01" box="[751,833,511,537]" captionStart="FIGURES 47–53" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1436,489,1867]" captionTargetId="figure-157@10.[151,1436,489,1867]" captionTargetPageId="10" captionText="FIGURES 47–53. Capobula infima (Simon, 1896) comb. nov., female (47) and male (48–53) from De Hoop Nature Reserve, SEM photographs: 47 Epigyne, ventral; 48 Left palpal femur and patella, ventral; 49 Left palpal patella, retrolateral (rectangle indicating lyriform organ associated with patellar apophysis); 50 Lyriform organ, enlarged; 51 Right palp, detail of retrolateral tibial apophysis; 52 Left palp, ventral; 53 Embolus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596117" httpUri="https://zenodo.org/record/4596117/files/figure.png" pageId="9" pageNumber="50">Fig. 47</figureCitation>
), in same transverse plane as teardrop-shaped laterally positioned primary spermathecae (e.g. 
<figureCitation id="1344DCC6FFA6FFAFFDEAFD99A2912B25" box="[600,689,546,573]" captionStart="FIGURES 54–58" captionStartId="12.[151,264,1667,1691]" captionTargetBox="[171,1421,571,1638]" captionTargetId="figure-183@12.[164,1423,561,1644]" captionTargetPageId="12" captionText="FIGURES 54–58. Capobula capensis spec. nov., genitalic morphology of female (54–55) and male (56–58) paratypes from Pakhuis Pass: 54 Epigyne, ventral; 55 Vulva, dorsal; 56–58 Left palp (56 prolateral, 57 ventral, 58 retrolateral). Scale bars = 0.25 mm. Abbreviations: b—bursa; cd—copulatory duct; co—copulatory opening; ps—primary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4596119" httpUri="https://zenodo.org/record/4596119/files/figure.png" pageId="9" pageNumber="50">Figs 54</figureCitation>
, 
<figureCitation id="1344DCC6FFA6FFAFFD0FFD98A2FE2B25" box="[701,734,546,573]" captionStart="FIGURES 59–64" captionStartId="14.[151,264,1535,1559]" captionTargetBox="[156,1432,634,1511]" captionTargetId="figure-249@14.[151,1436,630,1516]" captionTargetPageId="14" captionText="FIGURES 59–64. Capobula infima (Simon, 1896) comb. nov., genitalic morphology of female (59, 60) and male (61–64) from De Hoop Nature Reserve: 59 Epigyne, ventral; 60 Vulva, dorsal; 61–63 Left palp (61 prolateral, 62 ventral, 63 retrolateral); 64 Palpal femur, patella and tibia, retro-dorsal. Scale bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4596121" httpUri="https://zenodo.org/record/4596121/files/figure.png" pageId="9" pageNumber="50">59</figureCitation>
); copulatory ducts directed posteriorly along midline, with membranous anterior bursae originating near initial part of ducts; posteriorly, copulatory ducts bend laterally to enter spermathecae along their mesal margin (e.g. 
<figureCitation id="1344DCC6FFA5FFACFD68FF01A31129CD" box="[730,817,186,213]" captionStart="FIGURES 54–58" captionStartId="12.[151,264,1667,1691]" captionTargetBox="[171,1421,571,1638]" captionTargetId="figure-183@12.[164,1423,561,1644]" captionTargetPageId="12" captionText="FIGURES 54–58. Capobula capensis spec. nov., genitalic morphology of female (54–55) and male (56–58) paratypes from Pakhuis Pass: 54 Epigyne, ventral; 55 Vulva, dorsal; 56–58 Left palp (56 prolateral, 57 ventral, 58 retrolateral). Scale bars = 0.25 mm. Abbreviations: b—bursa; cd—copulatory duct; co—copulatory opening; ps—primary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4596119" httpUri="https://zenodo.org/record/4596119/files/figure.png" pageId="10" pageNumber="51">Figs 55</figureCitation>
, 
<figureCitation id="1344DCC6FFA5FFACFC8DFF01A37F29CE" box="[831,863,187,214]" captionStart="FIGURES 59–64" captionStartId="14.[151,264,1535,1559]" captionTargetBox="[156,1432,634,1511]" captionTargetId="figure-249@14.[151,1436,630,1516]" captionTargetPageId="14" captionText="FIGURES 59–64. Capobula infima (Simon, 1896) comb. nov., genitalic morphology of female (59, 60) and male (61–64) from De Hoop Nature Reserve: 59 Epigyne, ventral; 60 Vulva, dorsal; 61–63 Left palp (61 prolateral, 62 ventral, 63 retrolateral); 64 Palpal femur, patella and tibia, retro-dorsal. Scale bars = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4596121" httpUri="https://zenodo.org/record/4596121/files/figure.png" pageId="10" pageNumber="51">60</figureCitation>
); secondary spermathecae not distinguished. Male palpal femur, patella and tibia all with ventral apophyses (
<figureCitation id="1344DCC6FFA5FFACFC91FF65A38629E1" box="[803,934,222,249]" captionStart="FIGURES 47–53" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1436,489,1867]" captionTargetId="figure-157@10.[151,1436,489,1867]" captionTargetPageId="10" captionText="FIGURES 47–53. Capobula infima (Simon, 1896) comb. nov., female (47) and male (48–53) from De Hoop Nature Reserve, SEM photographs: 47 Epigyne, ventral; 48 Left palpal femur and patella, ventral; 49 Left palpal patella, retrolateral (rectangle indicating lyriform organ associated with patellar apophysis); 50 Lyriform organ, enlarged; 51 Right palp, detail of retrolateral tibial apophysis; 52 Left palp, ventral; 53 Embolus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596117" httpUri="https://zenodo.org/record/4596117/files/figure.png" pageId="10" pageNumber="51">Figs 48–52</figureCitation>
), patella with lyriform organ associated with apophysis (
<figureCitation id="1344DCC6FFA5FFACFEAAFEB9A1BE2805" box="[280,414,258,285]" captionStart="FIGURES 47–53" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1436,489,1867]" captionTargetId="figure-157@10.[151,1436,489,1867]" captionTargetPageId="10" captionText="FIGURES 47–53. Capobula infima (Simon, 1896) comb. nov., female (47) and male (48–53) from De Hoop Nature Reserve, SEM photographs: 47 Epigyne, ventral; 48 Left palpal femur and patella, ventral; 49 Left palpal patella, retrolateral (rectangle indicating lyriform organ associated with patellar apophysis); 50 Lyriform organ, enlarged; 51 Right palp, detail of retrolateral tibial apophysis; 52 Left palp, ventral; 53 Embolus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596117" httpUri="https://zenodo.org/record/4596117/files/figure.png" pageId="10" pageNumber="51">Figs 49, 50</figureCitation>
); tegulum pear-shaped, broadest basally, tapering distally, slightly broader than cymbium, with narrow sperm duct running down centre of tegulum in ventral view, with single basal loop (e.g. 
<figureCitation id="1344DCC6FFA5FFACFABAFE9DA57C2859" box="[1288,1372,294,321]" captionStart="FIGURES 54–58" captionStartId="12.[151,264,1667,1691]" captionTargetBox="[171,1421,571,1638]" captionTargetId="figure-183@12.[164,1423,561,1644]" captionTargetPageId="12" captionText="FIGURES 54–58. Capobula capensis spec. nov., genitalic morphology of female (54–55) and male (56–58) paratypes from Pakhuis Pass: 54 Epigyne, ventral; 55 Vulva, dorsal; 56–58 Left palp (56 prolateral, 57 ventral, 58 retrolateral). Scale bars = 0.25 mm. Abbreviations: b—bursa; cd—copulatory duct; co—copulatory opening; ps—primary spermatheca." figureDoi="http://doi.org/10.5281/zenodo.4596119" httpUri="https://zenodo.org/record/4596119/files/figure.png" pageId="10" pageNumber="51">Fig. 57</figureCitation>
); embolus distal, curving retrodistally (
<figureCitation id="1344DCC6FFA5FFACFDABFEF1A24C287D" box="[537,620,330,357]" captionStart="FIGURES 47–53" captionStartId="10.[151,264,1890,1914]" captionTargetBox="[151,1436,489,1867]" captionTargetId="figure-157@10.[151,1436,489,1867]" captionTargetPageId="10" captionText="FIGURES 47–53. Capobula infima (Simon, 1896) comb. nov., female (47) and male (48–53) from De Hoop Nature Reserve, SEM photographs: 47 Epigyne, ventral; 48 Left palpal femur and patella, ventral; 49 Left palpal patella, retrolateral (rectangle indicating lyriform organ associated with patellar apophysis); 50 Lyriform organ, enlarged; 51 Right palp, detail of retrolateral tibial apophysis; 52 Left palp, ventral; 53 Embolus, ventral." figureDoi="http://doi.org/10.5281/zenodo.4596117" httpUri="https://zenodo.org/record/4596117/files/figure.png" pageId="10" pageNumber="51">Fig. 53</figureCitation>
), tegulum without any other structures.
</paragraph>
<caption id="DF0090CBFFA8FFA1FF25F8F7A4DC2EEE" ID-DOI="http://doi.org/10.5281/zenodo.4596109" ID-Zenodo-Dep="4596109" httpUri="https://zenodo.org/record/4596109/files/figure.png" pageId="7" pageNumber="48" startId="7.[151,264,1869,1893]" targetBox="[151,1436,489,1846]" targetPageId="7">
<paragraph id="8BC0C043FFA8FFA1FF25F8F7A4DC2EEE" blockId="7.[151,1437,1869,2038]" pageId="7" pageNumber="48">
<emphasis id="B90B1C51FFA8FFA1FF25F8F7A1732E7D" bold="true" box="[151,339,1869,1893]" pageId="7" pageNumber="48">FIGURES 20–29.</emphasis>
<taxonomicName id="4C7FBBC0FFA8FFA1FEEAF8F4A2B92E7D" authority="(Simon, 1896)" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" baseAuthorityName="Simon" baseAuthorityYear="1896" box="[344,665,1869,1894]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="48" phylum="Arthropoda" rank="species" species="infima" status="comb. nov.">
<emphasis id="B90B1C51FFA8FFA1FEEAF8F4A2222E7E" box="[344,514,1870,1894]" italics="true" pageId="7" pageNumber="48">Capobula infima</emphasis>
(Simon, 1896)
</taxonomicName>
<emphasis id="B90B1C51FFA8FFA1FD2CF8F4A3312E7E" bold="true" box="[670,785,1870,1894]" pageId="7" pageNumber="48">
<taxonomicNameLabel id="A238A12AFFA8FFA1FD2CF8F4A3312E7E" box="[670,785,1870,1894]" pageId="7" pageNumber="48" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, female (20, 23–29) and male (21, 22) from De Hoop Nature Reserve, SEM photographs: 20 Detail of sternum texture; 21 Femur I, ventral; 22 Femur I, detail of fine distal tubercles on ventral surface, ventral; 23 Patella III, prolateral (arrow indicating lyriform organ); 24 Patella IV, retrolateral (arrow indicating lyriform organ at end of patellar indentation: PI); 25 Detail of lyriform organ at end of PI, retrolateral ventral; 26 Tibia I spines, ventral; 27 Metatarsus I spines, ventral; 28 Metatarsus I, detail of spine bases, ventral; 29 Tarsus I spines, ventro-distal.
</paragraph>
</caption>
<caption id="DF0090CBFFA7FFAEFF25F892A35B2EC8" ID-DOI="http://doi.org/10.5281/zenodo.4596113" ID-Zenodo-Dep="4596113" httpUri="https://zenodo.org/record/4596113/files/figure.png" pageId="8" pageNumber="49" startId="8.[151,264,1832,1856]" targetBox="[151,1436,777,1809]" targetPageId="8">
<paragraph id="8BC0C043FFA7FFAEFF25F892A35B2EC8" blockId="8.[151,1437,1832,2000]" pageId="8" pageNumber="49">
<emphasis id="B90B1C51FFA7FFAEFF25F892A1752E58" bold="true" box="[151,341,1832,1856]" pageId="8" pageNumber="49">FIGURES 30–38.</emphasis>
<taxonomicName id="4C7FBBC0FFA7FFAEFEEEF892A2832E58" authority="(Simon, 1896)" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" baseAuthorityName="Simon" baseAuthorityYear="1896" box="[348,675,1832,1856]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="49" phylum="Arthropoda" rank="species" species="infima" status="comb. nov.">
<emphasis id="B90B1C51FFA7FFAEFEEEF892A2282E58" box="[348,520,1832,1856]" italics="true" pageId="8" pageNumber="49">Capobula infima</emphasis>
(Simon, 1896)
</taxonomicName>
<emphasis id="B90B1C51FFA7FFAEFD18F892A33F2E58" bold="true" box="[682,799,1832,1856]" pageId="8" pageNumber="49">
<taxonomicNameLabel id="A238A12AFFA7FFAEFD18F892A33F2E58" box="[682,799,1832,1856]" pageId="8" pageNumber="49" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
female from De Hoop Nature Reserve, SEM photographs: 30 Metatarsus IV, distal preening brush and metatarsal stopper, retrolateral; 31 Tarsus I, dorsal, indicating chemosensory seta (cs), trichobothria (tr) and tarsal organ (to); 32 Tarsus I, trichobothrium, dorsal; 33 Same, detail of trichobothrium base, dorsal; 34 Tarsus I, detail of tarsal organ, dorsal; 35 Tarsal claws, leg IV, prolateral; 36 Tarsal claws, leg III, distal; 37 Palpal claw, prolateral distal; 38 Dorsal abdominal setae, dorsal, arrow indicating sigillum.
</paragraph>
</caption>
<caption id="DF0090CBFFA6FFAFFF25F89BA55A2EBD" ID-DOI="http://doi.org/10.5281/zenodo.4596115" ID-Zenodo-Dep="4596115" httpUri="https://zenodo.org/record/4596115/files/figure.png" pageId="9" pageNumber="50" startId="9.[151,264,1825,1849]" targetBox="[151,1436,597,1801]" targetPageId="9">
<paragraph id="8BC0C043FFA6FFAFFF25F89BA55A2EBD" blockId="9.[151,1437,1825,1957]" pageId="9" pageNumber="50">
<emphasis id="B90B1C51FFA6FFAFFF25F89BA1732E21" bold="true" box="[151,339,1825,1849]" pageId="9" pageNumber="50">FIGURES 39–46.</emphasis>
<taxonomicName id="4C7FBBC0FFA6FFAFFEEBF89BA2BA2E21" authority="(Simon, 1896)" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" baseAuthorityName="Simon" baseAuthorityYear="1896" box="[345,666,1825,1849]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="50" phylum="Arthropoda" rank="species" species="infima" status="comb. nov.">
<emphasis id="B90B1C51FFA6FFAFFEEBF89BA2232E21" box="[345,515,1825,1849]" italics="true" pageId="9" pageNumber="50">Capobula infima</emphasis>
(Simon, 1896)
</taxonomicName>
<emphasis id="B90B1C51FFA6FFAFFD2DF89BA3322E21" bold="true" box="[671,786,1825,1849]" pageId="9" pageNumber="50">
<taxonomicNameLabel id="A238A12AFFA6FFAFFD2DF89BA3322E21" box="[671,786,1825,1849]" pageId="9" pageNumber="50" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
, female (39, 41–43) and male (40, 44–46) from De Hoop Nature Reserve, SEM photographs: 39–40 Spinnerets, general view; 41, 44 Anterior lateral spinnerets; 42, 45 Posterior median spinnerets; 43, 46 Posterior lateral spinnerets. Abbreviations: Ac—aciniform gland spigot; Cy—cylindrical gland spigot; mAmp—mi-nor ampullate gland spigot; MAmp—major ampullate gland spigot; Nu—nubbin; Pi—piriform gland spigot; Ta—tartipore.
</paragraph>
</caption>
<paragraph id="8BC0C043FFA5FFACFF75FED4A4A728B5" blockId="10.[151,1437,151,465]" pageId="10" pageNumber="51">
<emphasis id="B90B1C51FFA5FFACFF75FED4A1472890" bold="true" box="[199,359,366,392]" pageId="10" pageNumber="51">Composition.</emphasis>
<taxonomicName id="4C7FBBC0FFA5FFACFEDDFED5A2C52892" authority="(Simon, 1896 a)" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" baseAuthorityName="Simon" baseAuthorityYear="1896" box="[367,741,367,394]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="infima" status="comb. nov.">
<emphasis id="B90B1C51FFA5FFACFEDDFED5A20E2891" box="[367,558,367,393]" italics="true" pageId="10" pageNumber="51">Capobula infima</emphasis>
(
<bibRefCitation id="EFEEBDB2FFA5FFACFD8FFED5A2FD2892" author="Simon, E." box="[573,733,367,394]" pageId="10" pageNumber="51" pagination="400 - 422" refId="ref18785" refString="Simon, E. (1896 a) Descriptions d'arachnides nouveaux de la famille des Clubionidae. Annales de la Societe Entomologique de Belgique, 40, 400 - 422." type="journal article" year="1896">Simon, 1896a</bibRefCitation>
)
</taxonomicName>
<emphasis id="B90B1C51FFA5FFACFD5FFED5A34F2891" bold="true" box="[749,879,367,393]" pageId="10" pageNumber="51">
<taxonomicNameLabel id="A238A12AFFA5FFACFD5FFED5A34F2891" box="[749,879,367,393]" pageId="10" pageNumber="51" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
(
<typeStatus id="54C47EE1FFA5FFACFCCDFED5A3912891" box="[895,945,367,393]" pageId="10" pageNumber="51">type</typeStatus>
species) and four new species, 
<taxonomicName id="4C7FBBC0FFA5FFACFAA8FED5A5BC2890" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[1306,1436,367,393]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="capensis" status="sp. nov.">
<emphasis id="B90B1C51FFA5FFACFAA8FED5A5BC2890" box="[1306,1436,367,393]" italics="true" pageId="10" pageNumber="51">C. capensis</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFA5FFACFF25FE29A12A28B5" bold="true" box="[151,266,403,429]" pageId="10" pageNumber="51">
<taxonomicNameLabel id="A238A12AFFA5FFACFF25FE29A12A28B5" box="[151,266,403,429]" pageId="10" pageNumber="51" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C7FBBC0FFA5FFACFEAAFE29A1BA28B4" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[280,410,403,429]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="montana" status="sp. nov.">
<emphasis id="B90B1C51FFA5FFACFEAAFE29A1BA28B4" box="[280,410,403,429]" italics="true" pageId="10" pageNumber="51">C. montana</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFA5FFACFE13FE29A23428B5" bold="true" box="[417,532,403,429]" pageId="10" pageNumber="51">
<taxonomicNameLabel id="A238A12AFFA5FFACFE13FE29A23428B5" box="[417,532,403,429]" pageId="10" pageNumber="51" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C7FBBC0FFA5FFACFD90FE29A29228B5" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[546,690,403,429]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="neethlingi" status="sp. nov.">
<emphasis id="B90B1C51FFA5FFACFD90FE29A29228B5" box="[546,690,403,429]" italics="true" pageId="10" pageNumber="51">C. neethlingi</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFA5FFACFD0BFE29A30C28B5" bold="true" box="[697,812,403,429]" pageId="10" pageNumber="51">
<taxonomicNameLabel id="A238A12AFFA5FFACFD0BFE29A30C28B5" box="[697,812,403,429]" pageId="10" pageNumber="51" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="4C7FBBC0FFA5FFACFCD0FE29A42D28B5" authority="Haddad &amp; Jin &amp; Platnick &amp; Booysen, 2021" authorityName="Haddad &amp; Jin &amp; Platnick &amp; Booysen" authorityYear="2021" box="[866,1037,403,429]" class="Arachnida" family="Trachelidae" genus="Capobula" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="51" phylum="Arthropoda" rank="species" species="ukhahlamba" status="sp. nov.">
<emphasis id="B90B1C51FFA5FFACFCD0FE29A42D28B5" box="[866,1037,403,429]" italics="true" pageId="10" pageNumber="51">C. ukhahlamba</emphasis>
</taxonomicName>
<emphasis id="B90B1C51FFA5FFACFBA6FE29A4A728B5" bold="true" box="[1044,1159,403,429]" pageId="10" pageNumber="51">
<taxonomicNameLabel id="A238A12AFFA5FFACFBA6FE29A4A728B5" box="[1044,1159,403,429]" pageId="10" pageNumber="51" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C36593C8FFA5FFACFF75FE0CA42228C9" box="[199,1026,438,465]" pageId="10" pageNumber="51" type="distribution">
<paragraph id="8BC0C043FFA5FFACFF75FE0CA42228C9" blockId="10.[151,1437,151,465]" box="[199,1026,438,465]" pageId="10" pageNumber="51">
<emphasis id="B90B1C51FFA5FFACFF75FE0CA14128C8" bold="true" box="[199,353,438,464]" pageId="10" pageNumber="51">Distribution.</emphasis>
Only known from 
<collectingCountry id="F36880D3FFA5FFACFD85FE0DA2E628C9" box="[567,710,439,465]" name="South Africa" pageId="10" pageNumber="51">South Africa</collectingCountry>
and the enclave of 
<collectingCountry id="F36880D3FFA5FFACFC12FE0DA3DE28C9" box="[928,1022,439,465]" name="Lesotho" pageId="10" pageNumber="51">Lesotho</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>