641 lines
92 KiB
XML
641 lines
92 KiB
XML
<document id="36C277F672E10BA7C6EA9C606A71B382" ID-DOI="10.1093/zoolinnean/zlad165" ID-ISSN="0024-4082" ID-Zenodo-Dep="11241243" ID-ZooBank="1EEF0D52-180B-4D3D-AB95-91AF3091E272" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1716364298778" checkinUser="plazi" docAuthor="Young, Mark T., Wilberg, Eric W., Johnson, Michela M., Herrera, Yanina, Brandalise, Marco de Andrade, Brignon, Arnaud, Sachs, Sven, Abel, Pascal, Foffa, Davide, Fernández, Marta S., Vignaud, Patrick, Cowgill, Thomas & Brusatte, Stephen L." docDate="2024" docId="03D08506FF9B731179EB0B850D85F848" docLanguage="en" docName="zlad165.pdf" docOrigin="Zoological Journal of the Linnean Society 200 (2)" docSource="http://dx.doi.org/10.1093/zoolinnean/zlad165" docStyle="DocumentStyle:4F230B9370E98E256D973D6DFB57F36C.5:ZoolJLinnSoc.2023-.journal_article" docStyleId="4F230B9370E98E256D973D6DFB57F36C" docStyleName="ZoolJLinnSoc.2023-.journal_article" docStyleVersion="5" docTitle="Teleosauroidea Geoffroy Saint-Hilaire 1831" docType="treatment" docVersion="1" lastPageNumber="593" masterDocId="FFE9FD7EFFB0733F796008100D3AFFFF" masterDocTitle="The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha)" masterLastPageNumber="617" masterPageNumber="547" pageNumber="590" updateTime="1716820941663" updateUser="juliana">
|
||
<mods:mods id="85EEE874FDF9D878E08D762BE739B799" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="38912B52ADF1D5A9ACFBCE3762F690D5">
|
||
<mods:title id="37E9DA7395B93943E4B7FE12E52ECC43">The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="48E55D72F75FB15E61FA3E556BA2593A" type="personal">
|
||
<mods:role id="84E96BAC5835A8081FC84970FD9E78A8">
|
||
<mods:roleTerm id="D82814D9CE51D6C3AE3D7FA488A3654D">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="92C0A70C91B4A04E91F22F29BA45A4B1">Young, Mark T.</mods:namePart>
|
||
<mods:affiliation id="D505033028890BA8337141C7EC5158D7">School of GeoSciences, Grant Institute, The King’s Buildings, University of Edinburgh, James Hutton Road, Edinburgh, EH 9 3 FE, United Kingdom & LWL-Museum für Naturkunde, Sentruper Strasse 285, 48161 Münster, Germany</mods:affiliation>
|
||
<mods:nameIdentifier id="829933773B2DE775DB1BB75F01A31B20" type="email">marktyoung1984@gmail.com</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name id="735061553A109154B4E8032BC339E0D1" type="personal">
|
||
<mods:role id="CBF69AE4849218B1FEED97B4E65D2A1A">
|
||
<mods:roleTerm id="C46931D7AD67F928549CABBE660B44B7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A3B2E067CE337493800F88B242E5BB70">Wilberg, Eric W.</mods:namePart>
|
||
<mods:affiliation id="660A4AC79958A94359EB81440E4C2414">Department of Anatomical Sciences, Stony Brook University, Stony Brook, NY 11794, USA</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="53E8E61CFE36DD10079B0556DE815DBA" type="personal">
|
||
<mods:role id="CC93C7FDC1511A315700265D6AE3F95C">
|
||
<mods:roleTerm id="A9D663F85F1D27120DC345C418A0F725">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="EDC4C6B0ED5E1699C1F6951FB17D3AFC">Johnson, Michela M.</mods:namePart>
|
||
<mods:affiliation id="DEB6707774692A8838AB861757861B16">Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="50EC8B8566AB85F256A9466409C233BE" type="personal">
|
||
<mods:role id="A080DC38724439F86676EA59B74B3B80">
|
||
<mods:roleTerm id="C98A73F0E5A18B37D642F6AE6F5AEB68">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="1FAEE1764551EAA0978B90A4B62B2F0F">Herrera, Yanina</mods:namePart>
|
||
<mods:affiliation id="616A54067E9A00A9F971B9B567CBB89E">Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), División Paleontología Vertebrados, Museo de La Plata, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, B 1900 La Plata, Buenos Aires, Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET) & Departamento de Diversidade e Ecologia, Faculdade de Biociências, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga 6681, Prédio 12, Porto Alegre, 90619 - 900, Brazil & Setor de Paleontologia, Museu de Ciências e Tecnologia, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga 6681, Prédio 40, Porto Alegre, 90619 - 900, Brazil</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="46E9E866577C560D2E0DA25BDB30C506" type="personal">
|
||
<mods:role id="372A21AA5A1F9987AA5C4D9F292E550C">
|
||
<mods:roleTerm id="81B3617E7EED3E3F377D7593942D5FF2">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="563E0E2745C532D8DB49DED10EAF8CD5">Brandalise, Marco de Andrade</mods:namePart>
|
||
<mods:affiliation id="B6FBF695C40BBD50B5A20F9DC809D012">Departamento de Diversidade e Ecologia, Faculdade de Biociências, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga 6681, Prédio 12, Porto Alegre, 90619 - 900, Brazil & Setor de Paleontologia, Museu de Ciências e Tecnologia, Pontifícia Universidade Católica do Rio Grande do Sul, Av. Ipiranga 6681, Prédio 40, Porto Alegre, 90619 - 900, Brazil</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="CD606F246A931C2D8EEFAD6777E6E23F" type="personal">
|
||
<mods:role id="5CAB93F7EB8D699A1B9699A8DA67E68E">
|
||
<mods:roleTerm id="887B1806B6086172E786455B7007BEAA">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="C9300A4CF1A91768740D989F1D977A68">Brignon, Arnaud</mods:namePart>
|
||
<mods:affiliation id="B30104ACC2AB5EBA518A3A3DB9E209CB">5, Villa Jeanne-d’Arc, 92340 Bourg-la-Reine, France</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="27E54A72E34EA127E5D0F911EFE7D840" type="personal">
|
||
<mods:role id="A7C176984D3427131F2A979F1DB34022">
|
||
<mods:roleTerm id="B5867C2CC2765F178B6A4050013CD5C7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B324E5768348875E2972771E15D361C1">Sachs, Sven</mods:namePart>
|
||
<mods:affiliation id="298570C6628F378ED30E73AB7E0B020C">Naturkunde-Museum Bielefeld, Abteilung Geowissenschaften, Adenauerplatz 2, 33602 Bielefeld, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="2CE9640BB8C6AE2B4C2B6D0F372E524F" type="personal">
|
||
<mods:role id="7162328E69F370ABDBA986CE1C98AB7A">
|
||
<mods:roleTerm id="AE4C5A8250B0C199D165BCD0F296834E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="22EDB238E574AC9E842D6245499AB4A9">Abel, Pascal</mods:namePart>
|
||
<mods:affiliation id="69ABB727E69FEC3821C185C48BEA01A9">Eberhard-Karls-University Tübingen, Fachbereich Geowissenschaften, Sigwartstrasse 10, 72074 Tübingen, Germany</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="4A80D3271B01B96E2213D328F12070EA" type="personal">
|
||
<mods:role id="E893F67B05D0A9EF4E6659E5640C3F86">
|
||
<mods:roleTerm id="7A2A579688581D3E1407F4E1A0DE0198">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="B54CA3B3FAC45DEF49AFEF06F2AE3359">Foffa, Davide</mods:namePart>
|
||
<mods:affiliation id="3ED56DA93AD5881EA5D6E62100BAED31">Department of Geosciences, Virginia Tech, 4044 Derring Hall 926 West Campus Drive, Blacksburg 24061, Virginia, USA & School of Geography, Earth and Environmental Sciences, University of Birmingham, Edgbaston, Birmingham B 15 2 TT, United Kingdom</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="B4D2329A8C717A98E389DBB80CF4A53D" type="personal">
|
||
<mods:role id="6058DE5FED4A945DFA81D9F3DEBEE931">
|
||
<mods:roleTerm id="364553D5AA10C477DE8B90C955E9C391">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="91821F6898F67E5AF190D5F34B540FAC">Fernández, Marta S.</mods:namePart>
|
||
<mods:affiliation id="FCCC248A5A727933B2BA0075F5C657D6">Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), División Paleontología Vertebrados, Museo de La Plata, Facultad de Ciencias Naturales y Museo, Universidad Nacional de La Plata, B 1900 La Plata, Buenos Aires, Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET)</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="7514C8E26687BCE43C92AEAB82D8F9B8" type="personal">
|
||
<mods:role id="C8EEDA7179F66394D30AA430F51134D7">
|
||
<mods:roleTerm id="1CB72FF72C36A31319D71310A182778C">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="265ACD45140625788A8BD218D88CB712">Vignaud, Patrick</mods:namePart>
|
||
<mods:affiliation id="66DE2760E5F374AC6953B82D5E9CDF70">Laboratoire de Paléontologie, Evolution, Paléoécosystèmes et Paléoprimatologie, CNRS UMR 7262, Department of Geosciences, University of Poitiers, 86073 Poitiers Cedex 9, France</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="832AFE93653540DDC64B930E5AFB705F" type="personal">
|
||
<mods:role id="E0A126CED4AB960410C4F5E34C152C69">
|
||
<mods:roleTerm id="D59280FCD2E9A8E860F76A6E61BE57E8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="108A5A303197C06BE40165DB576365AB">Cowgill, Thomas</mods:namePart>
|
||
<mods:affiliation id="550D5FC96597E3EEC51EC06D89381757">School of GeoSciences, Grant Institute, The King’s Buildings, University of Edinburgh, James Hutton Road, Edinburgh, EH 9 3 FE, United Kingdom</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="4F5359EAA2AB641C867E559388B807D2" type="personal">
|
||
<mods:role id="0BF2E7F4929D0784FB35E01BE6194E53">
|
||
<mods:roleTerm id="93A38CDFDD14A0AAFB0D1C933D8D68DB">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="468A738862A51795784EBA0AD86AE53B">Brusatte, Stephen L.</mods:namePart>
|
||
<mods:affiliation id="D5AF757022C57CF9D6C3A7C55B854FE3">School of GeoSciences, Grant Institute, The King’s Buildings, University of Edinburgh, James Hutton Road, Edinburgh, EH 9 3 FE, United Kingdom</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="248B186D41A39179427BDC970D7BC825">text</mods:typeOfResource>
|
||
<mods:relatedItem id="EFDC6A51882247B44C23CAF1389939A8" type="host">
|
||
<mods:titleInfo id="D21ADB9005662C0C3FEF52D90DA0A327">
|
||
<mods:title id="214F7C4E2EB949518370295D35A502D2">Zoological Journal of the Linnean Society</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="2D2A497DF7B815D5C5CB5B33F2BE78DA">
|
||
<mods:date id="977589E8E20B308BF536C693483EF0FD">2024</mods:date>
|
||
<mods:detail id="DC67954BF1183A24BBA5268924B32C2A" type="pubDate">
|
||
<mods:number id="A375C1E0C374E26BBC0E9043F0E65385">2024-01-09</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="1C819EFAD108563BE8BD111B388E5666" type="volume">
|
||
<mods:number id="80A715A0F21F96D5128972595A87351E">200</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="C8F88230E0E8E77B3FED06034F9C0296" type="issue">
|
||
<mods:number id="D0ED5C8C5C0D6A8E725BC622A054D723">2</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="65548E57EE6F3B913F47FE569349270F" unit="page">
|
||
<mods:start id="340D29342536CFD409B72DA976EBEFB8">547</mods:start>
|
||
<mods:end id="2BD8C9252AAF2BACB86ACCEEAAD1F653">617</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="9EFB916B51B886717BD62C2AB5D9E3A3">
|
||
<mods:url id="66A15C851AB14BB3968AA7CC47D16034">http://dx.doi.org/10.1093/zoolinnean/zlad165</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="58126A41A022045421C00CAF7E6A74F4">journal article</mods:classification>
|
||
<mods:identifier id="984090E326A79682772C2030269C5C33" type="DOI">10.1093/zoolinnean/zlad165</mods:identifier>
|
||
<mods:identifier id="BD47B8E5C0C677B24043E7D3FCD19F64" type="ISSN">0024-4082</mods:identifier>
|
||
<mods:identifier id="38C834DB94B0E0272A997A431F6FD62B" type="Zenodo-Dep">11241243</mods:identifier>
|
||
<mods:identifier id="39525965572FCBA1E814D9A100A29B88" type="ZooBank">1EEF0D52-180B-4D3D-AB95-91AF3091E272</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03D08506FF9B731179EB0B850D85F848" LSID="urn:lsid:plazi:treatment:03D08506FF9B731179EB0B850D85F848" httpUri="http://treatment.plazi.org/id/03D08506FF9B731179EB0B850D85F848" lastPageId="46" lastPageNumber="593" pageId="43" pageNumber="590">
|
||
<subSubSection id="C363679BFF9B731479EB0B850CE2FC31" pageId="43" pageNumber="590" type="nomenclature">
|
||
<paragraph id="8BC63410FF9B731479EB0B850CE2FC31" blockId="43.[139,737,917,974]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B731479EB0B850CE2FC31" bold="true" pageId="43" pageNumber="590">
|
||
<taxonomicName id="4C794F93FF9B731479EB0B850F64FC50" authority="Geoffroy Saint-Hilaire, 1831" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[139,606,917,943]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="582" phylum="Chordata" rank="superFamily" superFamily="Teleosauroidea">
|
||
Teleosauroidea
|
||
<bibRefCitation id="EFE849E1FF9B731478520B850F64FC50" author="Geoffroy Saint-Hilaire E" box="[306,606,917,943]" pageId="43" pageNumber="590" refId="ref52429" refString="Geoffroy Saint-Hilaire E. Recherches sur de Grands Sauriens Trouves a l'etat Fossile aux Confins Maritimes de la Basse-Normandie, Attribues d'abord au Crocodile, puis Determines sous les Noms de Teleosaurus et Steneosaurus. Paris: Firmin Didot, 1831, 138." type="book" year="1831">Geoffroy Saint-Hilaire, 1831</bibRefCitation>
|
||
</taxonomicName>
|
||
(Zoological Code)
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C363679BFF9B731479140BEC0FCDFBE9" box="[116,759,1020,1047]" pageId="43" pageNumber="590" type="reference_group">
|
||
<paragraph id="8BC63410FF9B731479140BEC0FCDFBE9" blockId="43.[113,759,1020,1114]" box="[116,759,1020,1047]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B731479140BEC0FCDFBE9" bold="true" box="[116,759,1020,1047]" pageId="43" pageNumber="590">
|
||
<taxonomicName id="4C794F93FF9B731479140BEC0C33FBE8" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[116,265,1020,1047]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="582" phylum="Chordata" rank="superFamily" superFamily="Teleosauroidea">
|
||
<emphasis id="B90DE802FF9B731479140BEC0C33FBE8" bold="true" box="[116,265,1020,1047]" italics="true" pageId="43" pageNumber="590">Teleosauroidea</emphasis>
|
||
</taxonomicName>
|
||
<bibRefCitation id="EFE849E1FF9B7314786E0BEC0F5FFBE9" author="Delfino M & Dal Sasso C" box="[270,613,1020,1046]" pageId="43" pageNumber="590" pagination="346 - 54" refId="ref50569" refString="Delfino M, Dal Sasso C. Marine reptiles (Thalattosuchia) from the Early Jurassic of Lombardy (northern Italy). Geobios 2006; 39: 346 - 54. https: // doi. org / 10.1016 / j. geobios. 2005.01.001" type="journal article" year="2006">Delfino and Dal Sasso 2006: 250</bibRefCitation>
|
||
(PhyloCode)
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C363679BFF9B731479110C330DA5FBA5" pageId="43" pageNumber="590" type="description">
|
||
<paragraph id="8BC63410FF9B731479110C330DA5FBA5" blockId="43.[113,759,1020,1114]" pageId="43" pageNumber="590">RegNum registration number 801.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C363679BFF9B731479110C6C0FBCFB6C" box="[113,646,1147,1171]" pageId="43" pageNumber="590" type="type_taxon">
|
||
<paragraph id="8BC63410FF9B731479110C6C0FBCFB6C" blockId="43.[113,646,1147,1171]" box="[113,646,1147,1171]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B731479110C6C0DD9FB6C" box="[113,227,1148,1171]" italics="true" pageId="43" pageNumber="590">
|
||
<typeStatus id="54C28AB2FF9B731479110C6C0D9BFB6C" box="[113,161,1148,1171]" pageId="43" pageNumber="590">Type</typeStatus>
|
||
genus:
|
||
</emphasis>
|
||
<taxonomicName id="4C794F93FF9B731479880C6B0FB8FB6C" authority="Geoffroy Saint-Hilaire, 1825" authorityName="Geoffroy Saint-Hilaire" authorityYear="1825" box="[232,642,1147,1171]" class="Reptilia" family="Teleosauridae" genus="Teleosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="590" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9B731479880C6B0C63FB6C" box="[232,345,1147,1171]" italics="true" pageId="43" pageNumber="590">Teleosaurus</emphasis>
|
||
Geoffroy Saint-Hilaire, 1825
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C363679BFF9B731478E30CA30FFAFAB0" pageId="43" pageNumber="590" type="etymology">
|
||
<paragraph id="8BC63410FF9B731478E30CA30CD3FB32" blockId="43.[113,763,1203,1359]" box="[387,489,1203,1229]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B731478E30CA30CD3FB32" box="[387,489,1203,1229]" italics="true" pageId="43" pageNumber="590">Etymology</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B731479110CC90FFAFAB0" blockId="43.[113,763,1203,1359]" pageId="43" pageNumber="590">
|
||
‘Those resembling
|
||
<taxonomicName id="4C794F93FF9B731478550CC90C9CFB0E" authorityName="Geoffroy Saint-Hilaire" authorityYear="1825" box="[309,422,1241,1265]" class="Reptilia" family="Teleosauridae" genus="Teleosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="590" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9B731478550CC90C9CFB0E" box="[309,422,1241,1265]" italics="true" pageId="43" pageNumber="590">Teleosaurus</emphasis>
|
||
</taxonomicName>
|
||
’. The stem
|
||
<emphasis id="B90DE802FF9B73147B7B0CC90F4DFB0E" box="[539,631,1241,1265]" italics="true" pageId="43" pageNumber="590">Teleosaur</emphasis>
|
||
- is based on the genus
|
||
<taxonomicName id="4C794F93FF9B731479BB0CE90C76FAEE" authorityName="Geoffroy Saint-Hilaire" authorityYear="1825" box="[219,332,1273,1297]" class="Reptilia" family="Teleosauridae" genus="Teleosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="590" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9B731479BB0CE90C76FAEE" box="[219,332,1273,1297]" italics="true" pageId="43" pageNumber="590">Teleosaurus</emphasis>
|
||
</taxonomicName>
|
||
. -
|
||
<emphasis id="B90DE802FF9B7314783F0CE90CA9FAEE" box="[351,403,1273,1297]" italics="true" pageId="43" pageNumber="590">oidea</emphasis>
|
||
, is a Neo-Latin plural suffix for ‘resembling’. Under Article 29.2 of the Zoological Code, the suffix -
|
||
<emphasis id="B90DE802FF9B731479190D270D97FAB0" box="[121,173,1335,1359]" italics="true" pageId="43" pageNumber="590">oidea</emphasis>
|
||
denotes a superfamily rank within the family-group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C363679BFF9B731478090D7F0E52FF31" pageId="43" pageNumber="590" type="description">
|
||
<paragraph id="8BC63410FF9B731478090D7F0F39FA76" blockId="43.[113,762,1391,1516]" box="[361,515,1391,1417]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B731478090D7F0F39FA76" box="[361,515,1391,1417]" italics="true" pageId="43" pageNumber="590">Geological range</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B731479110D850FADFA13" blockId="43.[113,762,1391,1516]" pageId="43" pageNumber="590">
|
||
Early Jurassic (Hettangian–Sinemurian) to Early Cretaceous (late Barremian) (
|
||
<bibRefCitation id="EFE849E1FF9B7314784A0DA50CF0FA32" author="Fanti F & Miyashita T & Cantelli L" box="[298,458,1461,1485]" pageId="43" pageNumber="590" pagination="263 - 74" refId="ref51216" refString="Fanti F, Miyashita T, Cantelli L et al. The largest thalattosuchian (Crocodylomorpha) supports teleosaurid survival across the Jurassic-Cretaceous boundary. Cretaceous Research 2016; 61: 263 - 74. https: // doi. org / 10.1016 / j. cretres. 2015.11.011" type="journal article" year="2016">
|
||
Fanti
|
||
<emphasis id="B90DE802FF9B731478030DA50CB1FA32" box="[355,395,1461,1485]" italics="true" pageId="43" pageNumber="590">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE849E1FF9B731478B40DA50FBFFA32" author="Cortes D & Larsson HCE & Maxwell EE" box="[468,645,1461,1485]" pageId="43" pageNumber="590" pagination="365 - 79" refId="ref50271" refString="Cortes D, Larsson HCE, Maxwell EE et al. An Early Cretaceous teleosauroid (Crocodylomorpha: Thalattosuchia) from Colombia. Ameghiniana 2019; 56: 365 - 79." type="journal article" year="2019">
|
||
Cortés
|
||
<emphasis id="B90DE802FF9B73147B7E0DA50F7CFA32" box="[542,582,1461,1485]" italics="true" pageId="43" pageNumber="590">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE849E1FF9B73147BEF0DA50DF7FA13" author="Johnson MM & Young MT & Brusatte SL" pageId="43" pageNumber="590" pagination="9808" refId="ref54662" refString="Johnson MM, Young MT, Brusatte SL. The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution. PeerJ 2020 a; 8: e 9808. https: // doi. org / 10.7717 / peerj. 9808" type="journal article" year="2020">
|
||
Johnson
|
||
<emphasis id="B90DE802FF9B73147B890DA50DB0FA13" italics="true" pageId="43" pageNumber="590">et al.</emphasis>
|
||
2020
|
||
<bibRefCitation id="EFE849E1FF9B731479A20DC40DF7FA13" author="Young MT & Brignon A & Sachs S" box="[194,205,1492,1516]" pageId="43" pageNumber="590" pagination="510 - 53" refId="ref61294" refString="Young MT, Brignon A, Sachs S et al. Cutting the Gordian knot: a historical and taxonomic revision of the Jurassic crocodylomorph Metriorhynchus. Zoological Journal of the Linnean Society 2021; 192: 510 - 53." type="journal article" year="2021">a</bibRefCitation>
|
||
</bibRefCitation>
|
||
, Young and Sachs 2021,
|
||
<bibRefCitation id="EFE849E1FF9B731478A70DC40FBDFA13" author="Hicham B & Nehili A & Ouzzaouit LA" box="[455,647,1492,1516]" pageId="43" pageNumber="590" pagination="104804" refId="ref53643" refString="Hicham B, Nehili A, Ouzzaouit LA et al. Discovery of the teleosauroid crocodylomorph from the early Jurassic of Chaara cave, Middle Atlas of Morocco. Journal of African Earth Sciences 2023; 198: 104804." type="journal article" year="2023">
|
||
Hicham
|
||
<emphasis id="B90DE802FF9B73147B7F0DC50F7DFA13" box="[543,583,1492,1516]" italics="true" pageId="43" pageNumber="590">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B731478790E1B0F69F9DA" blockId="43.[281,595,1547,1573]" box="[281,595,1547,1573]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B731478790E1B0F69F9DA" box="[281,595,1547,1573]" italics="true" pageId="43" pageNumber="590">PhyloCode phylogenetic definition</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B731479110E220C61F958" blockId="43.[113,762,1586,1704]" pageId="43" pageNumber="590">
|
||
The largest clade within
|
||
<taxonomicName id="4C794F93FF9B7314781A0E220F33F9B5" authorityName="sensu Fraas" authorityYear="1901" box="[378,521,1586,1610]" class="Reptilia" genus="Thalattosuchia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="590" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9B7314781A0E220F33F9B5" box="[378,521,1586,1610]" italics="true" pageId="43" pageNumber="590">Thalattosuchia</emphasis>
|
||
</taxonomicName>
|
||
containing
|
||
<taxonomicName id="4C794F93FF9B73147BE90E220CE8F996" authority="(Lamouroux 1820)" baseAuthorityName="Lamouroux" baseAuthorityYear="1820" class="Reptilia" family="Teleosauridae" genus="Teleosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="590" phylum="Chordata" rank="species" species="cadomensis">
|
||
<emphasis id="B90DE802FF9B73147BE90E220DE4F996" italics="true" pageId="43" pageNumber="590">Teleosaurus cadomensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE849E1FF9B731478620E420CFDF996" author="Lamouroux JVF" box="[258,455,1617,1641]" pageId="43" pageNumber="590" pagination="160 - 4" refId="ref55568" refString="Lamouroux JVF. Sur le crocodile fossile trouve dans les carrieres du bourg d'Allemagne, a un quart de lieue de Cean. Annales Generales des Sciences Physiques 1820; 3: 160 - 4." type="journal article" year="1820">Lamouroux 1820</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
but not
|
||
<taxonomicName id="4C794F93FF9B73147B010E410F7EF976" authority="(Holl 1829) (Metriorhynchoidea)" authorityName="Metriorhynchoidea" baseAuthorityName="Holl" baseAuthorityYear="1829" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="590" phylum="Chordata" rank="species" species="brevirostris">
|
||
<emphasis id="B90DE802FF9B73147B010E410DE6F976" italics="true" pageId="43" pageNumber="590">Metriorhynchus brevirostris</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE849E1FF9B731479920E610C65F976" author="Holl F" box="[242,351,1649,1673]" pageId="43" pageNumber="590" refId="ref53683" refString="Holl F. Handbuch der Petrefactenkunde Teil 1. Dresden: Hilscher, 1829, 416." type="book" year="1829">Holl 1829</bibRefCitation>
|
||
) (
|
||
<emphasis id="B90DE802FF9B731478E00E610F02F976" box="[384,568,1649,1673]" italics="true" pageId="43" pageNumber="590">Metriorhynchoidea</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
. Definition from
|
||
<bibRefCitation id="EFE849E1FF9B731479110E800C6FF957" author="Johnson MM & Young MT & Brignon A" box="[113,341,1680,1704]" pageId="43" pageNumber="590" pagination="1 - 7" refId="ref54877" refString="Johnson MM, Young MT, Brignon A et al. Addition to ' The phylogenetics of Teleosauroidea (Crocodylomorpha; Thalattosuchia) and implications for their ecology and evolution'. Bulletin of Phylogenetic Nomenclature 2022 b; 1: 1 - 7. https: // doi. org / 10.11646 / bpn. 1.1.1" type="journal article" year="2022">
|
||
Johnson
|
||
<emphasis id="B90DE802FF9B731479AB0E810DCEF957" box="[203,244,1680,1704]" italics="true" pageId="43" pageNumber="590">et al</emphasis>
|
||
. (2022b)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B73147D72088009F6FF55" blockId="43.[1042,1228,145,169]" box="[1042,1228,144,170]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B73147D72088009F6FF55" box="[1042,1228,144,170]" italics="true" pageId="43" pageNumber="590">Reference phylogeny</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B73147A4A08A60E52FF31" blockId="43.[810,872,183,206]" box="[810,872,182,206]" pageId="43" pageNumber="590">
|
||
<figureCitation id="13422895FF9B73147A4A08A60E58FF31" box="[810,866,182,206]" captionStart="Figure 3" captionStartId="19.[113,178,1185,1209]" captionTargetBox="[117,1455,147,1154]" captionTargetId="figure-283@19.[114,1458,144,1157]" captionTargetPageId="19" captionText="Figure 3. The equal weights, no constraints strict consensus topology (C0). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S1 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/." figureDoi="http://doi.org/10.5281/zenodo.11241251" httpUri="https://zenodo.org/record/11241251/files/figure.png" pageId="43" pageNumber="590">Fig. 3</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C363679BFF9B73147A5B08FE09A4FC41" pageId="43" pageNumber="590" type="diagnosis">
|
||
<paragraph id="8BC63410FF9B73147A5B08FE0898FEF7" blockId="43.[810,1459,238,958]" box="[827,1442,238,264]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B73147A5B08FE0898FEF7" box="[827,1442,238,264]" italics="true" pageId="43" pageNumber="590">Zoological Code diagnosis and PhyloCode diagnostic apomorphies</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B73147A4A090409A4FC41" blockId="43.[810,1459,238,958]" pageId="43" pageNumber="590">
|
||
Thalattosuchian crocodylomorphs with the following unique combination of characters (13): external and internal antorbital fenestrae are subequal or not distinguishable (169.0); supratemporal fenestrae longer than the orbits (198.2); squamosal does not project further posteriorly than the occipital condyle (reversal in the unnamed Chinese teleosauroid) (249.0); orbits have a dorsal inclination (271.1); in palatal view, the maxilla projects anteriorly with straightened margins creating a sub-rectangular shape (353.2); consistent presence of occipital tuberosities (410.1); within the trigeminal fossa, the foramina are fully divided into two openings by a bridge formed by proötic, dorsal opening interpreted as rostral middle cerebral vein and ventral opening as exit for trigeminal branches (441.1); paired ridges present on the medial ventral surface of the basisphenoid (449.1); main body of the quadrates are strongly inclined (464.2); first and second premaxillary alveoli form a couplet (reversals present within
|
||
<taxonomicName id="4C794F93FF9B73147DE30B19082EFCDE" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[1155,1300,777,801]" class="Reptilia" family="Teleosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="590" phylum="Chordata" rank="family">Teleosauridae</taxonomicName>
|
||
) (584.1); third dentary tooth occludes against the premaxilla–maxilla suture (635.0); coracoid with a fan-shaped distal end and a triangularshaped proximal end (741.1); scapular blade is less than 1.5 times the width of the scapular shaft (shared with the unnamed Toarcian metriorhynchoid) (744.1).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C363679BFF9B73117D520BCE0D85F848" lastPageId="46" lastPageNumber="593" pageId="43" pageNumber="590" type="discussion">
|
||
<paragraph id="8BC63410FF9B73147D520BCE0991FC07" blockId="43.[810,1459,990,1209]" box="[1074,1195,990,1016]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B73147D520BCE0991FC07" box="[1074,1195,990,1016]" italics="true" pageId="43" pageNumber="590">Composition</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B73147A4A0C1408ADFB46" blockId="43.[810,1459,990,1209]" pageId="43" pageNumber="590">
|
||
<taxonomicName id="4C794F93FF9B73147A4A0C140EE3FBE3" box="[810,985,1028,1052]" family="Machimosauridae" pageId="43" pageNumber="582" rank="family">
|
||
<emphasis id="B90DE802FF9B73147A4A0C140EE3FBE3" box="[810,985,1028,1052]" italics="true" pageId="43" pageNumber="590">Machimosauridae</emphasis>
|
||
</taxonomicName>
|
||
(composed of
|
||
<taxonomicName id="4C794F93FF9B73147DAC0C140850FBE3" box="[1228,1386,1028,1052]" genus="Macrospondylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="590" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9B73147DAC0C140850FBE3" box="[1228,1386,1028,1052]" italics="true" pageId="43" pageNumber="590">Macrospondylus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C794F93FF9B73147A4A0C340EE3FBC3" box="[810,985,1060,1084]" family="Machimosauridae" pageId="43" pageNumber="582" rank="subFamily" subFamily="Machimosaurinae">
|
||
<emphasis id="B90DE802FF9B73147A4A0C340EE3FBC3" box="[810,985,1060,1084]" italics="true" pageId="43" pageNumber="590">Machimosaurinae</emphasis>
|
||
</taxonomicName>
|
||
, and based on our phylogenetic analyses possibly also the ‘Hettangian–Sinemurian’ teleosauroid from
|
||
<collectingCountry id="F36E7480FF9B73147A4A0C730EB2FB85" box="[810,904,1123,1146]" name="Morocco" pageId="43" pageNumber="590">Morocco</collectingCountry>
|
||
—
|
||
<bibRefCitation id="EFE849E1FF9B73147AC30C7209B2FB85" author="Hicham B & Nehili A & Ouzzaouit LA" box="[931,1160,1122,1146]" pageId="43" pageNumber="590" pagination="104804" refId="ref53643" refString="Hicham B, Nehili A, Ouzzaouit LA et al. Discovery of the teleosauroid crocodylomorph from the early Jurassic of Chaara cave, Middle Atlas of Morocco. Journal of African Earth Sciences 2023; 198: 104804." type="journal article" year="2023">
|
||
Hicham
|
||
<emphasis id="B90DE802FF9B73147D670C730907FB85" box="[1031,1085,1122,1146]" italics="true" pageId="43" pageNumber="590">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C794F93FF9B73147DBE0C720859FB85" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[1246,1379,1122,1146]" class="Reptilia" family="Teleosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="590" phylum="Chordata" rank="family">
|
||
<emphasis id="B90DE802FF9B73147DBE0C720859FB85" box="[1246,1379,1122,1146]" italics="true" pageId="43" pageNumber="590">Teleosauridae</emphasis>
|
||
</taxonomicName>
|
||
(composed of the unnamed Chinese teleosauroid,
|
||
<taxonomicName id="4C794F93FF9B73147C7B0C920897FB65" box="[1307,1453,1154,1178]" family="Teleosauridae" genus="Indosinosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="590" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9B73147C7B0C920897FB65" box="[1307,1453,1154,1178]" italics="true" pageId="43" pageNumber="590">Indosinosuchus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794F93FF9B73147A4A0CB20E8EFB46" authorityName="Fitzinger" authorityYear="1843" box="[810,948,1186,1209]" family="Teleosauridae" genus="Mystriosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="590" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9B73147A4A0CB20E8EFB46" box="[810,948,1186,1209]" italics="true" pageId="43" pageNumber="590">Mystriosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794F93FF9B73147ADF0CB108B6FB46" authority=", Teleosaurinae, and Aeolodontini" authorityName="Teleosaurinae, and Aeolodontini" box="[959,1420,1185,1209]" genus="Seldsienean" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="43" pageNumber="590" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9B73147ADF0CB10915FB46" box="[959,1071,1185,1209]" italics="true" pageId="43" pageNumber="590">Seldsienean</emphasis>
|
||
, Teleosaurinae, and Aeolodontini
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B73147D5B0CC9099BFB0C" blockId="43.[810,1460,1241,1554]" box="[1083,1185,1241,1267]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B73147D5B0CC9099BFB0C" box="[1083,1185,1241,1267]" italics="true" pageId="43" pageNumber="590">Comments</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B73147A4A0CEF091EF9ED" blockId="43.[810,1460,1241,1554]" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B73147A4A0CEF0EA4FAE8" box="[810,926,1279,1303]" italics="true" pageId="43" pageNumber="590">Authorship:</emphasis>
|
||
While the nomen
|
||
<taxonomicName id="4C794F93FF9B73147D170CEF0828FAE8" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[1143,1298,1279,1303]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="582" phylum="Chordata" rank="superFamily" superFamily="Teleosauroidea">Teleosauroidea</taxonomicName>
|
||
was first used by
|
||
<bibRefCitation id="EFE849E1FF9B73147A2D0D0E09FDFAC9" author="Delfino M & Dal Sasso C" box="[845,1223,1310,1334]" pageId="43" pageNumber="590" pagination="346 - 54" refId="ref50569" refString="Delfino M, Dal Sasso C. Marine reptiles (Thalattosuchia) from the Early Jurassic of Lombardy (northern Italy). Geobios 2006; 39: 346 - 54. https: // doi. org / 10.1016 / j. geobios. 2005.01.001" type="journal article" year="2006">Delfino and Dal Sasso (2006: 250)</bibRefCitation>
|
||
, under the Zoological Code
|
||
<bibRefCitation id="EFE849E1FF9B73147A0F0D2E09A3FAA9" author="Geoffroy Saint-Hilaire E" box="[879,1177,1342,1366]" pageId="43" pageNumber="590" refId="ref52429" refString="Geoffroy Saint-Hilaire E. Recherches sur de Grands Sauriens Trouves a l'etat Fossile aux Confins Maritimes de la Basse-Normandie, Attribues d'abord au Crocodile, puis Determines sous les Noms de Teleosaurus et Steneosaurus. Paris: Firmin Didot, 1831, 138." type="book" year="1831">Geoffroy Saint-Hilaire 1831</bibRefCitation>
|
||
is the nominal authority. The nominal author of a family-group is the author who first erected a family-group taxon that is valid (in fulfilment of Article 11), and in accordance with the Zoological Code Principle of Coordination applied to family-group names (Article 36.1). However, under the PhyloCode,
|
||
<bibRefCitation id="EFE849E1FF9B73147D1E0DCA0889FA0D" author="Delfino M & Dal Sasso C" box="[1150,1459,1498,1522]" pageId="43" pageNumber="590" pagination="346 - 54" refId="ref50569" refString="Delfino M, Dal Sasso C. Marine reptiles (Thalattosuchia) from the Early Jurassic of Lombardy (northern Italy). Geobios 2006; 39: 346 - 54. https: // doi. org / 10.1016 / j. geobios. 2005.01.001" type="journal article" year="2006">Delfino and Dal Sasso (2006)</bibRefCitation>
|
||
is the nominal authority.
|
||
</paragraph>
|
||
<paragraph id="8BC63410FF9B73117A4A0E230C0DFA9B" blockId="43.[810,1459,1587,1705]" lastBlockId="46.[129,778,1231,1975]" lastPageId="46" lastPageNumber="593" pageId="43" pageNumber="590">
|
||
<emphasis id="B90DE802FF9B73147A4A0E230EA0F9B5" box="[810,922,1587,1610]" italics="true" pageId="43" pageNumber="590">Comments:</emphasis>
|
||
As demonstrated by
|
||
<bibRefCitation id="EFE849E1FF9B73147DE30E230865F9B4" author="Johnson MM & Young MT & Steel L" box="[1155,1375,1587,1611]" pageId="43" pageNumber="590" pagination="385 - 418" refId="ref54602" refString="Johnson MM, Young MT, Steel L et al. Re-description of ' Steneosaurus ' obtusidens Andrews, 1909, an unusual macrophagous teleosaurid crocodylomorph from the Middle Jurassic of England. Zoological Journal of the Linnean Society 2018; 182: 385 - 418. https: // doi. org / 10.1093 / zoolinnean / zlx 035" type="journal article" year="2018">
|
||
Johnson
|
||
<emphasis id="B90DE802FF9B73147D820E240835F9B4" box="[1250,1295,1587,1611]" italics="true" pageId="43" pageNumber="590">et al</emphasis>
|
||
. (2018
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE849E1FF9B73147C0E0E230897F9B4" author="Johnson MM & Young MT & Brusatte SL" box="[1390,1453,1587,1611]" pageId="43" pageNumber="590" pagination="9808" refId="ref54662" refString="Johnson MM, Young MT, Brusatte SL. The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution. PeerJ 2020 a; 8: e 9808. https: // doi. org / 10.7717 / peerj. 9808" type="journal article" year="2020">2020a</bibRefCitation>
|
||
, b, c) teleosauroids were far more morphologically diverse than previously thought. The previous use of
|
||
<taxonomicName id="4C794F93FF9B73147C660E6208BEF976" box="[1286,1412,1650,1673]" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="43" pageNumber="590" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9B73147C660E6208BEF976" box="[1286,1412,1650,1673]" italics="true" pageId="43" pageNumber="590">Steneosaurus</emphasis>
|
||
</taxonomicName>
|
||
as a ‘wastebasket’ to place the majority of teleosauroid species has greatly held back our understanding of the clade. For example, the diversity shown in
|
||
<figureCitation id="13422895FF9D7312783B0DCE0C87FA09" box="[347,445,1502,1526]" captionStart="Figure 21" captionStartId="46.[129,194,1019,1043]" captionTargetBox="[132,1470,145,989]" captionTargetId="figure-352@46.[129,1473,142,991]" captionTargetPageId="46" captionText="Figure 21. Comparative plate of teleosauroid crania in dorsal view.A, the teleosaurid Mystriosaurus laurillardi, referred specimen NHMUK PV OR 14781 (holotype of Steneosaurus brevior); B, the aeolodontin teleosaurid Mycterosuchus nasutus, holotype NHMUK PV R 2617; C, the machimosaurid Macrospondylus bollensis, referred specimen NHMUK PV R 5703; D, the machimosaurid Charitomenosuchus leedsi, referred specimen NHMUK PV R 3806; E, the machimosaurin machimosaurid Lemmysuchus obtusidens, holotype NHMUK PV R 3168.All scale bars equal 20 cm. Note, all of these species were previously referred to the genus Steneosaurus, highlighting that taxonomic lumping and paraphyletic genera obscure morphological variation." figureDoi="http://doi.org/10.5281/zenodo.11241287" httpUri="https://zenodo.org/record/11241287/files/figure.png" pageId="45" pageNumber="592">Figure 21</figureCitation>
|
||
would have been placed in the genus
|
||
<taxonomicName id="4C794F93FF9D731279D60DED0C0EF9EB" box="[182,308,1533,1556]" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731279D60DED0C0EF9EB" box="[182,308,1533,1556]" italics="true" pageId="45" pageNumber="592">Steneosaurus</emphasis>
|
||
</taxonomicName>
|
||
prior to the work of
|
||
<bibRefCitation id="EFE849E1FF9D73127B7C0DED0FCFF9EA" author="Johnson MM & Young MT & Steel L" box="[540,757,1533,1557]" pageId="45" pageNumber="592" pagination="385 - 418" refId="ref54602" refString="Johnson MM, Young MT, Steel L et al. Re-description of ' Steneosaurus ' obtusidens Andrews, 1909, an unusual macrophagous teleosaurid crocodylomorph from the Middle Jurassic of England. Zoological Journal of the Linnean Society 2018; 182: 385 - 418. https: // doi. org / 10.1093 / zoolinnean / zlx 035" type="journal article" year="2018">
|
||
Johnson
|
||
<emphasis id="B90DE802FF9D73127B1A0DEE0F9CF9EA" box="[634,678,1533,1557]" italics="true" pageId="45" pageNumber="592">et al</emphasis>
|
||
. (2018
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE849E1FF9D731279110E0D0D8AF9CA" author="Johnson MM & Young MT & Brusatte SL" box="[113,176,1565,1589]" pageId="45" pageNumber="592" pagination="9808" refId="ref54662" refString="Johnson MM, Young MT, Brusatte SL. The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution. PeerJ 2020 a; 8: e 9808. https: // doi. org / 10.7717 / peerj. 9808" type="journal article" year="2020">2020a</bibRefCitation>
|
||
). Even though some species are more closely related to the genera
|
||
<taxonomicName id="4C794F93FF9D7312786B0E2C0C46F9AB" authorityName="Geoffroy Saint-Hilaire" authorityYear="1825" box="[267,380,1596,1620]" class="Reptilia" family="Teleosauridae" genus="Teleosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D7312786B0E2C0C46F9AB" box="[267,380,1596,1620]" italics="true" pageId="45" pageNumber="592">Teleosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C794F93FF9D731278D80E2C0F10F9AB" authorityName="Westphal" authorityYear="1961" box="[440,554,1596,1620]" class="Reptilia" family="Teleosauridae" genus="Platysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731278D80E2C0F10F9AB" box="[440,554,1596,1620]" italics="true" pageId="45" pageNumber="592">Platysuchus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C794F93FF9D73127B200E2C0FF0F9AC" authorityName="Fitzinger" authorityYear="1843" box="[576,714,1596,1619]" family="Teleosauridae" genus="Mystriosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D73127B200E2C0FF0F9AC" box="[576,714,1596,1619]" italics="true" pageId="45" pageNumber="592">Mystriosaurus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C794F93FF9D731279110E4B0DC4F98C" authorityName="Andrews" authorityYear="1913" box="[113,254,1627,1651]" family="Teleosauridae" genus="Mycterosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731279110E4B0DC4F98C" box="[113,254,1627,1651]" italics="true" pageId="45" pageNumber="592">Mycterosuchus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="13422895FF9D7312786D0E4B0CBAF98C" box="[269,384,1627,1651]" captionStart="Figure 21" captionStartId="46.[129,194,1019,1043]" captionTargetBox="[132,1470,145,989]" captionTargetId="figure-352@46.[129,1473,142,991]" captionTargetPageId="46" captionText="Figure 21. Comparative plate of teleosauroid crania in dorsal view.A, the teleosaurid Mystriosaurus laurillardi, referred specimen NHMUK PV OR 14781 (holotype of Steneosaurus brevior); B, the aeolodontin teleosaurid Mycterosuchus nasutus, holotype NHMUK PV R 2617; C, the machimosaurid Macrospondylus bollensis, referred specimen NHMUK PV R 5703; D, the machimosaurid Charitomenosuchus leedsi, referred specimen NHMUK PV R 3806; E, the machimosaurin machimosaurid Lemmysuchus obtusidens, holotype NHMUK PV R 3168.All scale bars equal 20 cm. Note, all of these species were previously referred to the genus Steneosaurus, highlighting that taxonomic lumping and paraphyletic genera obscure morphological variation." figureDoi="http://doi.org/10.5281/zenodo.11241287" httpUri="https://zenodo.org/record/11241287/files/figure.png" pageId="45" pageNumber="592">Fig. 21A, B</figureCitation>
|
||
) and others are more closely related to
|
||
<taxonomicName id="4C794F93FF9D731279EB0E6A0C1CF96D" box="[139,294,1658,1682]" class="Reptilia" family="Teleosauridae" genus="Machimosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731279EB0E6A0C1CF96D" box="[139,294,1658,1682]" italics="true" pageId="45" pageNumber="592">Machimosaurus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C794F93FF9D731278560E6A0CEEF96D" box="[310,468,1658,1682]" genus="Macrospondylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731278560E6A0CEEF96D" box="[310,468,1658,1682]" italics="true" pageId="45" pageNumber="592">Macrospondylus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794F93FF9D731278BE0E6A0FA7F96D" authorityName="Johnson, Young & Brusatte" authorityYear="2020" box="[478,669,1658,1682]" genus="Charitomenosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731278BE0E6A0FA7F96D" box="[478,669,1658,1682]" italics="true" pageId="45" pageNumber="592">Charitomenosuchus</emphasis>
|
||
</taxonomicName>
|
||
, and particularly
|
||
<taxonomicName id="4C794F93FF9D731279AB0E8A0C69F94D" authorityName="Johnson, Young, Steel, Foffa, Smith, Hua, Havlik, Howlett & Dyke" authorityYear="2017" box="[203,339,1690,1714]" genus="Lemmysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731279AB0E8A0C69F94D" box="[203,339,1690,1714]" italics="true" pageId="45" pageNumber="592">Lemmysuchus</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<figureCitation id="13422895FF9D731278000E8A0CEFF94D" box="[352,469,1690,1714]" captionStart="Figure 21" captionStartId="46.[129,194,1019,1043]" captionTargetBox="[132,1470,145,989]" captionTargetId="figure-352@46.[129,1473,142,991]" captionTargetPageId="46" captionText="Figure 21. Comparative plate of teleosauroid crania in dorsal view.A, the teleosaurid Mystriosaurus laurillardi, referred specimen NHMUK PV OR 14781 (holotype of Steneosaurus brevior); B, the aeolodontin teleosaurid Mycterosuchus nasutus, holotype NHMUK PV R 2617; C, the machimosaurid Macrospondylus bollensis, referred specimen NHMUK PV R 5703; D, the machimosaurid Charitomenosuchus leedsi, referred specimen NHMUK PV R 3806; E, the machimosaurin machimosaurid Lemmysuchus obtusidens, holotype NHMUK PV R 3168.All scale bars equal 20 cm. Note, all of these species were previously referred to the genus Steneosaurus, highlighting that taxonomic lumping and paraphyletic genera obscure morphological variation." figureDoi="http://doi.org/10.5281/zenodo.11241287" httpUri="https://zenodo.org/record/11241287/files/figure.png" pageId="45" pageNumber="592">Fig. 21C–E</figureCitation>
|
||
). Based on the phylogenetic analyses presented herein (
|
||
<figureCitation id="13422895FF9D731278ED0EA90CC9F92E" box="[397,499,1721,1745]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStart-4="Figure 7" captionStart-5="Figure 8" captionStart-6="Figure 9" captionStart-7="Figure 10" captionStart-8="Figure 11" captionStart-9="Figure 12" captionStartId-0="19.[113,178,1185,1209]" captionStartId-1="21.[113,178,1219,1243]" captionStartId-2="22.[128,193,1219,1243]" captionStartId-3="23.[113,178,1244,1268]" captionStartId-4="24.[129,194,1442,1466]" captionStartId-5="25.[113,178,1707,1731]" captionStartId-6="27.[113,178,1246,1270]" captionStartId-7="28.[129,194,1301,1325]" captionStartId-8="29.[113,178,1287,1311]" captionStartId-9="30.[128,193,1319,1343]" captionTargetBox-0="[117,1455,147,1154]" captionTargetBox-1="[117,1455,147,1190]" captionTargetBox-2="[132,1470,147,1188]" captionTargetBox-3="[117,1455,147,1213]" captionTargetBox-4="[132,1470,147,1411]" captionTargetBox-5="[117,1455,553,1676]" captionTargetBox-6="[117,1455,147,1215]" captionTargetBox-7="[132,1470,147,1270]" captionTargetBox-8="[117,1455,147,1256]" captionTargetBox-9="[132,1470,147,1289]" captionTargetId-0="figure-283@19.[114,1458,144,1157]" captionTargetId-1="figure-205@21.[114,1458,144,1192]" captionTargetId-2="figure-220@22.[129,1473,144,1191]" captionTargetId-3="figure-242@23.[114,1458,144,1216]" captionTargetId-4="figure-150@24.[129,1473,144,1414]" captionTargetId-5="figure-191@25.[114,1458,550,1679]" captionTargetId-6="figure-185@27.[114,1458,144,1218]" captionTargetId-7="figure-212@28.[129,1473,144,1273]" captionTargetId-8="figure-196@29.[114,1458,144,1259]" captionTargetId-9="figure-177@30.[129,1473,144,1291]" captionTargetPageId-0="19" captionTargetPageId-1="21" captionTargetPageId-2="22" captionTargetPageId-3="23" captionTargetPageId-4="24" captionTargetPageId-5="25" captionTargetPageId-6="27" captionTargetPageId-7="28" captionTargetPageId-8="29" captionTargetPageId-9="30" captionText-0="Figure 3. The equal weights, no constraints strict consensus topology (C0). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S1 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/." captionText-1="Figure 4. The equal weights, crocodyliform constraint strict consensus topology (C1). Length = 4177, CI = 0.302, RI = 0.830, RC = 0.201, HI = 0.698, number of most parsimonious cladograms hit the hold limit. Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S2 for the entire topology). A, Crocodylomorpha; B, Thalattosuchia; C, Neothalattosuchia; D, Teleosauridae; E, Teleosaurinae; F, Aeolodontini; G, Machimosauridae; H, Machimosaurinae; I, Machimosaurini; J, Euthalattosuchia; K, Metriorhynchidae; L, Metriorhynchinae; M, Rhacheosaurini; N, Geosaurinae; O, Geosaurini.Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians. Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/." captionText-2="Figure 5. The equal weights, mesoeucrocodylian constraint strict consensus topology (C2). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit. Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig.S3 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians. Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T.Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/ by/3.0/." captionText-3="Figure 6. The equal weights, neosuchian constraint strict consensus topology (C3). Length = 4174, CI = 0.302, RI = 0.830, RC = 0.251, HI = 0.698, number of most parsimonious cladograms hit the hold limit. Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S4 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Metasuchia; D, Neosuchia; E, Thalattosuchia; F, Neothalattosuchia; G, Teleosauridae; H, Teleosaurinae; I, Aeolodontini; J, Machimosauridae; K, Machimosaurinae; L, Machimosaurini; M, Euthalattosuchia; N, Metriorhynchidae; O, Metriorhynchinae; P, Rhacheosaurini; Q, Geosaurinae; R, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians. Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons. org/licenses/by/3.0/." captionText-4="Figure 7. The equal weights, tethysuchian constraint strict consensus topology (C4). Length = 4172, CI = 0.302, RI = 0.830, RC = 0.251, HI = 0.698, number of most parsimonious cladograms hit the hold limit. Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S5 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Metasuchia; D, Neosuchia; E, Thalattosuchia; F, Neothalattosuchia; G, Teleosaurinae; H, Aeolodontini; I, Machimosaurinae; J, Machimosaurini; K, Metriorhynchidae; L, Metriorhynchinae; M, Rhacheosaurini; N, Geosaurinae; O, Geosaurini.Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians. Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T.Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/." captionText-5="Figure 8. The extended implied weighting (k = 15), no constraints strict consensus topology (C0). Length = 4178, fit = 111.578, CI = 0.302, RI = 0.830, RC = 0.250, HI = 0.698, number of most parsimonious cladograms = 4500. Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig.S6 for the entire topology, Appendix Two).A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians. Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons. org/licenses/by/3.0/." captionText-6="Figure 9. The extended implied weighting (k = 15), crocodyliform constraint strict consensus topology (C1). Length = 4194, fit = 111.226, CI = 0.300, RI = 0.829, RC = 0.249, HI = 0.700, number of most parsimonious cladograms = 4500.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S7 for the entire topology). A, Crocodylomorpha; B, Thalattosuchia; C, Neothalattosuchia; D, Teleosauridae; E, Teleosaurinae; F, Aeolodontini; G, Machimosauridae; H, Machimosaurinae; I, Machimosaurini; J, Euthalattosuchia; K, Metriorhynchidae; L, Metriorhynchinae; M, Rhacheosaurini; N, Geosaurinae; O, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians. Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T.Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/ by/3.0/." captionText-7="Figure 10. The extended implied weighting (k = 15), mesoeucrocodylian constraint strict consensus topology (C2). Length = 4178, fit = 111.578, CI = 0.302, RI = 0.830, RC = 0.250, HI = 0.698, number of most parsimonious cladograms = 4500.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig.S8 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https:// creativecommons.org/licenses/by/3.0/." captionText-8="Figure 11. The extended implied weighting (k = 15), neosuchian constraint strict consensus topology (C3). Length = 4192, fit = 112.001, CI = 0.301, RI = 0.829, RC = 0.249, HI = 0.699, number of most parsimonious cladograms = 4500.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S9 for the entire topology, Appendix Two). A, Crocodylomorpha; B, Crocodyliformes; C, Metasuchia; D, Neosuchia; E, Thalattosuchia; F, Neothalattosuchia; G, Teleosauridae; H, Teleosaurinae; I, Aeolodontini; J, Machimosauridae; K, Machimosaurinae; L, Machimosaurini; M, Euthalattosuchia; N, Metriorhynchidae; O, Metriorhynchinae; P, Rhacheosaurini; Q, Geosaurinae; R, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians. Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/." captionText-9="Figure 12. The extended implied weighting (k = 15), tethysuchian constraint strict consensus topology (C4). Length = 4189, fit = 112.271, CI = 0.301, RI = 0.829, RC = 0.249, HI = 0.699, number of most parsimonious cladograms = 4500.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S10 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Metasuchia; D, Neosuchia; E, Thalattosuchia; F, Neothalattosuchia; G, Teleosauridae; H, Teleosaurinae; I, Aeolodontini; J, Machimosauridae; K, Machimosaurinae; L, Machimosaurini; M, Euthalattosuchia; N, Metriorhynchidae; O, Metriorhynchinae; P, Rhacheosaurini; Q, Geosaurinae; R, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians. Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T.Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/." figureDoi-0="http://doi.org/10.5281/zenodo.11241251" figureDoi-1="http://doi.org/10.5281/zenodo.11241253" figureDoi-2="http://doi.org/10.5281/zenodo.11241255" figureDoi-3="http://doi.org/10.5281/zenodo.11241257" figureDoi-4="http://doi.org/10.5281/zenodo.11241259" figureDoi-5="http://doi.org/10.5281/zenodo.11241261" figureDoi-6="http://doi.org/10.5281/zenodo.11241263" figureDoi-7="http://doi.org/10.5281/zenodo.11241265" figureDoi-8="http://doi.org/10.5281/zenodo.11241267" figureDoi-9="http://doi.org/10.5281/zenodo.11241269" httpUri-0="https://zenodo.org/record/11241251/files/figure.png" httpUri-1="https://zenodo.org/record/11241253/files/figure.png" httpUri-2="https://zenodo.org/record/11241255/files/figure.png" httpUri-3="https://zenodo.org/record/11241257/files/figure.png" httpUri-4="https://zenodo.org/record/11241259/files/figure.png" httpUri-5="https://zenodo.org/record/11241261/files/figure.png" httpUri-6="https://zenodo.org/record/11241263/files/figure.png" httpUri-7="https://zenodo.org/record/11241265/files/figure.png" httpUri-8="https://zenodo.org/record/11241267/files/figure.png" httpUri-9="https://zenodo.org/record/11241269/files/figure.png" pageId="45" pageNumber="592">Figs 3–12</figureCitation>
|
||
), the only way to render
|
||
<taxonomicName id="4C794F93FF9D731279110EC90DD5F90F" box="[113,239,1753,1776]" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731279110EC90DD5F90F" box="[113,239,1753,1776]" italics="true" pageId="45" pageNumber="592">Steneosaurus</emphasis>
|
||
</taxonomicName>
|
||
monophyletic would be to place all thalattosuchians more derived than
|
||
<taxonomicName id="4C794F93FF9D731278560EE80C82F8EF" authorityName="Wilberg, Godoy, Griffiths, Turner & Benson" authorityYear="2023" box="[310,440,1784,1808]" family="Teleosauridae" genus="Turnersuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731278560EE80C82F8EF" box="[310,440,1784,1808]" italics="true" pageId="45" pageNumber="592">Turnersuchus</emphasis>
|
||
</taxonomicName>
|
||
into the genus
|
||
<taxonomicName id="4C794F93FF9D73127B380EE80FECF8F0" box="[600,726,1784,1807]" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D73127B380EE80FECF8F0" box="[600,726,1784,1807]" italics="true" pageId="45" pageNumber="592">Steneosaurus</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="4C794F93FF9D731279110F070CFCF8D0" box="[113,454,1815,1839]" genus="Plagiophthalmosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="592" phylum="Chordata" rank="species" species="gracilirostris">
|
||
<emphasis id="B90DE802FF9D731279110F070CFCF8D0" box="[113,454,1815,1839]" italics="true" pageId="45" pageNumber="592">Plagiophthalmosuchus gracilirostris</emphasis>
|
||
</taxonomicName>
|
||
, originally named
|
||
<taxonomicName id="4C794F93FF9D73127B1D0F070DD2F8B1" baseAuthorityName="Westphal" baseAuthorityYear="1961" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="species" species="gracilirostris">
|
||
<emphasis id="B90DE802FF9D73127B1D0F070DD2F8B1" italics="true" pageId="45" pageNumber="592">Steneosaurus gracilirostris</emphasis>
|
||
</taxonomicName>
|
||
, is recovered outside of
|
||
<taxonomicName id="4C794F93FF9D731278B40F260FAFF8B1" box="[468,661,1846,1870]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="582" parvOrder="Neothalattosuchia" phylum="Chordata" rank="parvOrder">Neothalattosuchia</taxonomicName>
|
||
).The only other way to use the genus
|
||
<taxonomicName id="4C794F93FF9D731278E20F460F3AF892" box="[386,512,1878,1901]" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D731278E20F460F3AF892" box="[386,512,1878,1901]" italics="true" pageId="45" pageNumber="592">Steneosaurus</emphasis>
|
||
</taxonomicName>
|
||
as ‘traditionally’ defined would be to accept it is as a paraphyletic taxon (and accept the usage of paraphyletic taxa), one in which the other traditionally accepted teleosauroid genera (
|
||
<taxonomicName id="4C794F93FF9D73127D1F0DAE09CAFA29" authorityName="Geoffroy Saint-Hilaire" authorityYear="1825" box="[1151,1264,1470,1494]" class="Reptilia" family="Teleosauridae" genus="Teleosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D73127D1F0DAE09CAFA29" box="[1151,1264,1470,1494]" italics="true" pageId="45" pageNumber="592">Teleosaurus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794F93FF9D73127C650DAE084DFA29" authorityName="Westphal" authorityYear="1961" box="[1285,1399,1470,1494]" class="Reptilia" family="Teleosauridae" genus="Platysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D73127C650DAE084DFA29" box="[1285,1399,1470,1494]" italics="true" pageId="45" pageNumber="592">Platysuchus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C794F93FF9D73127A4A0DCE0EFFFA09" box="[810,965,1502,1526]" class="Reptilia" family="Teleosauridae" genus="Machimosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D73127A4A0DCE0EFFFA09" box="[810,965,1502,1526]" italics="true" pageId="45" pageNumber="592">Machimosaurus</emphasis>
|
||
</taxonomicName>
|
||
) and
|
||
<taxonomicName id="4C794F93FF9D73127D6E0DCE09E2FA09" authorityName="Fitzinger" authorityYear="1843" box="[1038,1240,1502,1526]" pageId="45" pageNumber="582" rank="superFamily" superFamily="Metriorhynchoidea">Metriorhynchoidea</taxonomicName>
|
||
, evolved from. That would place
|
||
<taxonomicName id="4C794F93FF9D73127ACF0DED0956F9EA" box="[943,1132,1533,1557]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="582" parvOrder="Neothalattosuchia" phylum="Chordata" rank="parvOrder">Neothalattosuchia</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794F93FF9D73127D170DED082EF9EA" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[1143,1300,1533,1557]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="582" phylum="Chordata" rank="superFamily" superFamily="Teleosauroidea">Teleosauroidea</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794F93FF9D73127C400DED0895F9EA" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[1312,1455,1533,1557]" class="Reptilia" family="Teleosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="family">Teleosauridae</taxonomicName>
|
||
, Aeolodontini,
|
||
<taxonomicName id="4C794F93FF9D73127AB70E0C09AAF9CB" box="[983,1168,1564,1588]" family="Machimosauridae" pageId="45" pageNumber="582" rank="family">Machimosauridae</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C794F93FF9D73127DD20E0C0851F9CB" box="[1202,1387,1564,1588]" family="Machimosauridae" pageId="45" pageNumber="582" rank="subFamily" subFamily="Machimosaurinae">Machimosaurinae</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C794F93FF9D73127A4A0E2C0EEEF9AB" box="[810,980,1596,1620]" pageId="45" pageNumber="582" rank="tribe" tribe="Machimosaurini">Machimosaurini</taxonomicName>
|
||
, as subclades within
|
||
<taxonomicName id="4C794F93FF9D73127DD00E2C0814F9AC" box="[1200,1326,1596,1619]" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9D73127DD00E2C0814F9AC" box="[1200,1326,1596,1619]" italics="true" pageId="45" pageNumber="592">Steneosaurus</emphasis>
|
||
</taxonomicName>
|
||
. Such an action would subsume a remarkable level of morphological variation into a paraphyletic genus, creating a genus that could never be morphologically defined, only defined by what it is not. Having such a genus would allow for fragmentary specimens to be a generic assignment, as argued for by
|
||
<bibRefCitation id="EFE849E1FF9D73127DB20EC908B9F90F" author="Hua S & Pennetier E & Pennetier G" box="[1234,1411,1752,1776]" pageId="45" pageNumber="592" pagination="1 - 8" refId="ref53818" refString="Hua S, Pennetier E, Pennetier G. A juvenile Steneosaurus in the Callovian of Normandy (France): a genus too hastily consigned to the wastebasket? Carnets natures 2021; 8: 1 - 8." type="journal article" year="2021">
|
||
Hua
|
||
<emphasis id="B90DE802FF9D73127C640EC90817F90F" box="[1284,1325,1752,1776]" italics="true" pageId="45" pageNumber="592">et al</emphasis>
|
||
. (2021)
|
||
</bibRefCitation>
|
||
. But that begs the question, what does the designation
|
||
<taxonomicName id="4C794F93FF9D73127C550EE80E7CF8D1" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="592" phylum="Chordata" rank="species" species="undetermined">
|
||
<emphasis id="B90DE802FF9D73127C550EE80889F8F0" box="[1333,1459,1784,1807]" italics="true" pageId="45" pageNumber="592">Steneosaurus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
mean when the genus is paraphyletic, and to such a degree? Although
|
||
<bibRefCitation id="EFE849E1FF9D73127AF40F270977F8B1" author="Hua S & Pennetier E & Pennetier G" box="[916,1101,1846,1870]" pageId="45" pageNumber="592" pagination="1 - 8" refId="ref53818" refString="Hua S, Pennetier E, Pennetier G. A juvenile Steneosaurus in the Callovian of Normandy (France): a genus too hastily consigned to the wastebasket? Carnets natures 2021; 8: 1 - 8." type="journal article" year="2021">
|
||
Hua
|
||
<emphasis id="B90DE802FF9D73127AA90F270ECFF8B1" box="[969,1013,1846,1870]" italics="true" pageId="45" pageNumber="592">et al</emphasis>
|
||
. (2021)
|
||
</bibRefCitation>
|
||
consider the lack of a generic assignment to be a weakness of the
|
||
<bibRefCitation id="EFE849E1FF9D73127DF60F4508BEF891" author="Johnson MM & Young MT & Brusatte SL" box="[1174,1412,1877,1902]" pageId="45" pageNumber="592" pagination="9808" refId="ref54662" refString="Johnson MM, Young MT, Brusatte SL. The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution. PeerJ 2020 a; 8: e 9808. https: // doi. org / 10.7717 / peerj. 9808" type="journal article" year="2020">
|
||
Johnson
|
||
<emphasis id="B90DE802FF9D73127D940F46081AF892" box="[1268,1312,1877,1901]" italics="true" pageId="45" pageNumber="592">et al</emphasis>
|
||
. (2020a)
|
||
</bibRefCitation>
|
||
systematization, we argue it is in fact a strength. Assignments such as
|
||
<taxonomicName id="4C794F93FF9D73127A2A0F840EDFF853" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[842,997,1940,1964]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="45" pageNumber="582" phylum="Chordata" rank="superFamily" superFamily="Teleosauroidea">Teleosauroidea</taxonomicName>
|
||
indeterminate, or
|
||
<taxonomicName id="4C794F93FF9D73127DD00F840852F853" box="[1200,1384,1940,1964]" family="Machimosauridae" pageId="45" pageNumber="582" rank="family">Machimosauridae</taxonomicName>
|
||
indet., show the limits of our current knowledge and gives a clear indication where in the teleosauroid tree we think a specimen belongs. Using a paraphyletic
|
||
<taxonomicName id="4C794F93FF9E731178F40D1E0F28FADA" box="[404,530,1294,1317]" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="46" pageNumber="593" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9E731178F40D1E0F28FADA" box="[404,530,1294,1317]" italics="true" pageId="46" pageNumber="593">Steneosaurus</emphasis>
|
||
</taxonomicName>
|
||
(or any other genus, like
|
||
<taxonomicName id="4C794F93FF9E731179E10D3D0C20FABA" box="[129,282,1325,1349]" class="Reptilia" family="Metriorhynchidae" genus="Metriorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="46" pageNumber="593" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9E731179E10D3D0C20FABA" box="[129,282,1325,1349]" italics="true" pageId="46" pageNumber="593">Metriorhynchus</emphasis>
|
||
</taxonomicName>
|
||
—
|
||
<bibRefCitation id="EFE849E1FF9E731178540D3E0CC6FABA" author="Le Mort J & Martin JE & Picot L" box="[308,508,1325,1349]" pageId="46" pageNumber="593" pagination="102539" refId="ref55995" refString="Le Mort J, Martin JE, Picot L et al. First description of the most complete Metriorhynchus aff. superciliosus (Thalattosuchia) specimen from the Callovian of the Vaches-Noires cliffs (Normandy, France) and limitations in the classification of Metriorhynchidae. Annales de Paleontologie 2022; 108: 102539. https: // doi. org / 10.1016 / j. annpal. 2022.102539" type="journal article" year="2022">
|
||
Le Mort
|
||
<emphasis id="B90DE802FF9E731178F20D3E0C81FABA" box="[402,443,1325,1349]" italics="true" pageId="46" pageNumber="593">et al</emphasis>
|
||
. 2022
|
||
</bibRefCitation>
|
||
) merely provides the illusion of specificity.
|
||
</paragraph>
|
||
<footnote id="E862281EFF9B731479110EEB0866F844" pageId="43" pageNumber="590">
|
||
<paragraph id="8BC63410FF9B731479110EEB0866F844" blockId="43.[113,1452,1787,1979]" pageId="43" pageNumber="590">
|
||
of braincase pneumaticity in thalattosuchians, and the enlargement of the internal vascular canals. For details on the computed tomography scans and internal descriptions, see:
|
||
<bibRefCitation id="EFE849E1FF9B731478A70F070F40F8D0" author="Brusatte SL & Muir A & Young MT" box="[455,634,1815,1839]" pageId="43" pageNumber="590" pagination="1511 - 30" refId="ref49329" refString="Brusatte SL, Muir A, Young MT, et al. The braincase and neurosensory anatomy of an Early Jurassic marine crocodylomorph: implications for crocodilian sinus evolution and sensory transitions. The Anatomical Record 2016; 299: 1511 - 30." type="journal article" year="2016">
|
||
Brusatte
|
||
<emphasis id="B90DE802FF9B73147B7A0F070F7AF8D0" box="[538,576,1815,1839]" italics="true" pageId="43" pageNumber="590">et al</emphasis>
|
||
. 2016
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE849E1FF9B73147BE50F070E0EF8D0" author="Herrera Y & Leardi JM & Fernandez MS" box="[645,820,1815,1839]" pageId="43" pageNumber="590" pagination="5686" refId="ref53484" refString="Herrera Y, Leardi JM, Fernandez MS. Braincase and endocranial anatomy of two thalattosuchian crocodylomorphs and their relevance in understanding their adaptations to the marine environment. PeerJ 2018; 6: e 5686. https: // doi. org / 10.7717 / peerj. 5686" type="journal article" year="2018">
|
||
Herrera
|
||
<emphasis id="B90DE802FF9B73147BB40F070FC0F8D0" box="[724,762,1815,1839]" italics="true" pageId="43" pageNumber="590">et al</emphasis>
|
||
. 2018
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFE849E1FF9B73147A200F070ED5F8D0" author="Cowgill T & Young MT & Schwab JA" box="[832,1007,1815,1839]" pageId="43" pageNumber="590" pagination="812 - 35" refId="ref50391" refString="Cowgill T, Young MT, Schwab JA et al. Cephalic salt gland evolution in Mesozoic pelagic crocodylomorphs. Zoological Journal of the Linnean Society 2023; 197: 812 - 35." type="journal article" year="2023">
|
||
Cowgill
|
||
<emphasis id="B90DE802FF9B73147AEF0F070E8FF8D0" box="[911,949,1815,1839]" italics="true" pageId="43" pageNumber="590">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
. Abbreviations: II, cranial nerve II; IV, cranial nerve IV; V, cranial nerve V; IX, cranial nerve IX; X–XI, cranial nerves X–XI; XII, cranial nerve XII; bsd, basisphenoid diverticulum of the media pharyngeal sinus; cc, cerebral carotid canal; dvs, transverse dural venous sinus; en, dural envelope endocast; pts, paratympanic sinus; cqc, cranioquadrate canal; ifd, infundibular diverticulum of the paratympanic sinus; qud, quadrate diverticulum of the paratympanic sinus; itd, intertympanic diverticulum of the paratympanic sinus; el, endosseous labyrinth; mpt, median pharyngeal sinus; oa, orbital artery canals; otd, otoccipital diverticulum of the paratympanic sinus; pf, pituitary fossa cavity; ptt, paratympanic tube; tov, temporoorbital vessel canals.
|
||
</paragraph>
|
||
</footnote>
|
||
<caption id="DF066498FF9C731379E10E630821F877" ID-DOI="http://doi.org/10.5281/zenodo.11241283" ID-Zenodo-Dep="11241283" httpUri="https://zenodo.org/record/11241283/files/figure.png" pageId="44" pageNumber="591" startId="44.[129,194,1651,1675]" targetBox="[131,1469,156,1621]" targetPageId="44" targetType="figure">
|
||
<paragraph id="8BC63410FF9C731379E10E630821F877" blockId="44.[129,1466,1651,1928]" pageId="44" pageNumber="591">
|
||
<emphasis id="B90DE802FF9C731379E10E630DDFF973" bold="true" box="[129,229,1651,1676]" pageId="44" pageNumber="591">Figure 19.</emphasis>
|
||
Comparative plate of extant crocodylian and thalattosuchian internal rostral anatomy (A–G in dorsal view and H–I in lateral view). A, extant gavialid
|
||
<taxonomicName id="4C794F93FF9C731378080E800F29F958" baseAuthorityName="Gmelin" baseAuthorityYear="1789" box="[360,531,1680,1704]" class="Reptilia" family="Gavialidae" genus="Gavialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="44" pageNumber="591" phylum="Chordata" rank="species" species="gangeticus">
|
||
<emphasis id="B90DE802FF9C731378080E800F29F958" box="[360,531,1680,1704]" italics="true" pageId="44" pageNumber="591">Gavialis gangeticus</emphasis>
|
||
</taxonomicName>
|
||
, referred specimen UF-herp-118998; B, non-neothalattosuchian
|
||
<taxonomicName id="4C794F93FF9C73137D170E800DD5F93B" genus="Plagiophthalmosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="44" pageNumber="591" phylum="Chordata" rank="species" species="gracilirostris">
|
||
<emphasis id="B90DE802FF9C73137D170E800DD5F93B" italics="true" pageId="44" pageNumber="591">Plagiophthalmosuchus gracilirostris</emphasis>
|
||
</taxonomicName>
|
||
, paratype NHMUK PV OR 15500; C, extant gavialid
|
||
<taxonomicName id="4C794F93FF9C73137B8C0EBC0E99F93B" baseAuthorityName="Muller" baseAuthorityYear="1838" box="[748,931,1708,1732]" class="Crocodylia" family="Crocodylidae" genus="Tomistoma" kingdom="Animalia" order="Eusuchia" pageId="44" pageNumber="591" phylum="Chordata" rank="species" species="schlegelii">
|
||
<emphasis id="B90DE802FF9C73137B8C0EBC0E99F93B" box="[748,931,1708,1732]" italics="true" pageId="44" pageNumber="591">Tomistoma schlegelii</emphasis>
|
||
</taxonomicName>
|
||
, referred specimen USNM 211322; D, early diverging metriorhynchoid
|
||
<emphasis id="B90DE802FF9C731378480ED80CE9F920" box="[296,467,1736,1760]" italics="true" pageId="44" pageNumber="591">
|
||
<taxonomicName id="4C794F93FF9C731378480ED80CA7F91F" authorityName="Haeckel" authorityYear="1895" box="[296,413,1736,1760]" class="Reptilia" family="Metriorhynchidae" genus="Pelagosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="44" pageNumber="591" phylum="Chordata" rank="genus">Pelagosaurus</taxonomicName>
|
||
typus
|
||
</emphasis>
|
||
, referred specimen NHMUK PV OR 32599; E, extant crocodylid
|
||
<taxonomicName id="4C794F93FF9C73137D210ED8083BF91F" baseAuthorityName="Cuvier" baseAuthorityYear="1807" box="[1089,1281,1736,1760]" class="Reptilia" family="Crocodylidae" genus="Crocodylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="44" pageNumber="591" phylum="Chordata" rank="species" species="rhombifer">
|
||
<emphasis id="B90DE802FF9C73137D210ED8083BF91F" box="[1089,1281,1736,1760]" italics="true" pageId="44" pageNumber="591">Crocodylus rhombifer</emphasis>
|
||
</taxonomicName>
|
||
, referred specimen MNB AB50.0171; F, H, rhacheosaurin metriorhynchid
|
||
<taxonomicName id="4C794F93FF9C73137BEC0EF40E55F904" baseAuthorityName="Gasparini & Dellape" baseAuthorityYear="1976" box="[652,879,1764,1787]" class="Reptilia" family="Metriorhynchidae" genus="Cricosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="44" pageNumber="591" phylum="Chordata" rank="species" species="araucanensis">
|
||
<emphasis id="B90DE802FF9C73137BEC0EF40E55F904" box="[652,879,1764,1787]" italics="true" pageId="44" pageNumber="591">Cricosaurus araucanensis</emphasis>
|
||
</taxonomicName>
|
||
, holotype MLP 72-IV-7-1; G, I, geosaurin metriorhynchid
|
||
<taxonomicName id="4C794F93FF9C731379E10F100C64F8E7" authorityName="Young, Andrade, Etches & Beatty" authorityYear="2013" box="[129,350,1792,1816]" class="Reptilia" family="Metriorhynchidae" genus="Torvoneustes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="44" pageNumber="591" phylum="Chordata" rank="species" species="coryphaeus">
|
||
<emphasis id="B90DE802FF9C731379E10F100C64F8E7" box="[129,350,1792,1816]" italics="true" pageId="44" pageNumber="591">Torvoneustes coryphaeus</emphasis>
|
||
</taxonomicName>
|
||
, holotype MJML K1863. All scale bars equal 5 cm. Highlighting how apneumatic thalattosuchian rostra were, that the preorbital fenestrae were decoupled from the paranasal sinus system, and how the enlarged salt glands expanded the posterior nasal cavity in metriorhynchoids. For details on the computed tomography scans and internal descriptions, see:
|
||
<bibRefCitation id="EFE849E1FF9C73137D730F2809F8F8AF" author="Cowgill T & Young MT & Schwab JA" box="[1043,1218,1848,1872]" pageId="44" pageNumber="591" pagination="812 - 35" refId="ref50391" refString="Cowgill T, Young MT, Schwab JA et al. Cephalic salt gland evolution in Mesozoic pelagic crocodylomorphs. Zoological Journal of the Linnean Society 2023; 197: 812 - 35." type="journal article" year="2023">
|
||
Cowgill
|
||
<emphasis id="B90DE802FF9C73137D020F2809B2F8AF" box="[1122,1160,1848,1872]" italics="true" pageId="44" pageNumber="591">et al</emphasis>
|
||
. 2023
|
||
</bibRefCitation>
|
||
. Abbreviations: ant fen, antorbital fenestra; antorb, antorbital sinus of the paranasal sinus system; exp, expansion of the nasal cavity; max, maxillary diverticulum of the paranasal sinus system; max cec, maxillary cecal recess; nc, nasal cavity; pre fen, preorbital fenestra; post, postvestibular sinus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF066498FF9D731279110D2E0C9AFA71" ID-DOI="http://doi.org/10.5281/zenodo.11241285" ID-Zenodo-Dep="11241285" httpUri="https://zenodo.org/record/11241285/files/figure.png" pageId="45" pageNumber="592" startId="45.[113,178,1342,1366]" targetBox="[117,1455,147,1313]" targetPageId="45" targetType="figure">
|
||
<paragraph id="8BC63410FF9D731279110D2E0C9AFA71" blockId="45.[113,1457,1342,1423]" pageId="45" pageNumber="592">
|
||
<emphasis id="B90DE802FF9D731279110D2E0DECFAA9" bold="true" box="[113,214,1342,1366]" pageId="45" pageNumber="592">Figure 20.</emphasis>
|
||
The type specimens of the non-neothalattosuchian thalattosuchian
|
||
<taxonomicName id="4C794F93FF9D73127A310D2F09B4FAA8" box="[849,1166,1343,1367]" genus="Plagiophthalmosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="45" pageNumber="592" phylum="Chordata" rank="species" species="gracilirostris">
|
||
<emphasis id="B90DE802FF9D73127A310D2F09B4FAA8" box="[849,1166,1343,1367]" italics="true" pageId="45" pageNumber="592">Plagiophthalmosuchus gracilirostris</emphasis>
|
||
</taxonomicName>
|
||
. A, holotype NHMUK PV OR 14792, nearly complete skeleton in dorsal view; B, paratype NHMUK PV OR 15500, skull in lateral view. Scale bar for A equals 30 cm, and the scale bar for B equals 10 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF066498FF9E731179E10BEB0F4DFB60" ID-DOI="http://doi.org/10.5281/zenodo.11241287" ID-Zenodo-Dep="11241287" httpUri="https://zenodo.org/record/11241287/files/figure.png" pageId="46" pageNumber="593" startId="46.[129,194,1019,1043]" targetBox="[132,1470,145,989]" targetPageId="46" targetType="figure">
|
||
<paragraph id="8BC63410FF9E731179E10BEB0F4DFB60" blockId="46.[129,1449,1019,1184]" pageId="46" pageNumber="593">
|
||
<emphasis id="B90DE802FF9E731179E10BEB0DDFFBEC" bold="true" box="[129,229,1019,1043]" pageId="46" pageNumber="593">Figure 21.</emphasis>
|
||
Comparative plate of teleosauroid crania in dorsal view. A, the teleosaurid
|
||
<taxonomicName id="4C794F93FF9E73117AC10BEC09BBFBEC" authorityName="Kaup" authorityYear="1837" box="[929,1153,1019,1043]" family="Teleosauridae" genus="Mystriosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="593" phylum="Chordata" rank="species" species="laurillardi">
|
||
<emphasis id="B90DE802FF9E73117AC10BEC09BBFBEC" box="[929,1153,1019,1043]" italics="true" pageId="46" pageNumber="593">Mystriosaurus laurillardi</emphasis>
|
||
</taxonomicName>
|
||
, referred specimen NHMUK PV OR 14781 (holotype of
|
||
<taxonomicName id="4C794F93FF9E731178E90C080F7BFBD0" authorityName="Tate & Blake" authorityYear="1876" box="[393,577,1047,1071]" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="46" pageNumber="593" phylum="Chordata" rank="species" species="brevior">
|
||
<emphasis id="B90DE802FF9E731178E90C080F7BFBD0" box="[393,577,1047,1071]" italics="true" pageId="46" pageNumber="593">Steneosaurus brevior</emphasis>
|
||
</taxonomicName>
|
||
); B, the aeolodontin teleosaurid
|
||
<taxonomicName id="4C794F93FF9E73117A180C07097EFBD0" baseAuthorityName="Andrews" baseAuthorityYear="1909" box="[888,1092,1047,1071]" family="Teleosauridae" genus="Mycterosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="593" phylum="Chordata" rank="species" species="nasutus">
|
||
<emphasis id="B90DE802FF9E73117A180C07097EFBD0" box="[888,1092,1047,1071]" italics="true" pageId="46" pageNumber="593">Mycterosuchus nasutus</emphasis>
|
||
</taxonomicName>
|
||
, holotype NHMUK PV R 2617; C, the machimosaurid
|
||
<taxonomicName id="4C794F93FF9E731178370C230F03FBB4" baseAuthorityName="Jager" baseAuthorityYear="1828" box="[343,569,1075,1099]" genus="Macrospondylus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="593" phylum="Chordata" rank="species" species="bollensis">
|
||
<emphasis id="B90DE802FF9E731178370C230F03FBB4" box="[343,569,1075,1099]" italics="true" pageId="46" pageNumber="593">Macrospondylus bollensis</emphasis>
|
||
</taxonomicName>
|
||
, referred specimen NHMUK PV R 5703; D, the machimosaurid
|
||
<taxonomicName id="4C794F93FF9E73117DFE0C2308BFFBB4" baseAuthorityName="Andrews" baseAuthorityYear="1909" box="[1182,1413,1075,1099]" genus="Charitomenosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="593" phylum="Chordata" rank="species" species="leedsi">
|
||
<emphasis id="B90DE802FF9E73117DFE0C2308BFFBB4" box="[1182,1413,1075,1099]" italics="true" pageId="46" pageNumber="593">Charitomenosuchus leedsi</emphasis>
|
||
</taxonomicName>
|
||
, referred specimen NHMUK PV R 3806; E, the machimosaurin machimosaurid
|
||
<taxonomicName id="4C794F93FF9E73117A110C5F096BFB98" baseAuthorityName="Andrews" baseAuthorityYear="1909" box="[881,1105,1103,1127]" genus="Lemmysuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="593" phylum="Chordata" rank="species" species="obtusidens">
|
||
<emphasis id="B90DE802FF9E73117A110C5F096BFB98" box="[881,1105,1103,1127]" italics="true" pageId="46" pageNumber="593">Lemmysuchus obtusidens</emphasis>
|
||
</taxonomicName>
|
||
, holotype NHMUK PV R 3168. All scale bars equal 20 cm. Note, all of these species were previously referred to the genus
|
||
<taxonomicName id="4C794F93FF9E73117AC70C7C0921FB7C" box="[935,1051,1132,1155]" class="Reptilia" family="Teleosauridae" genus="Steneosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="46" pageNumber="593" phylum="Chordata" rank="genus">
|
||
<emphasis id="B90DE802FF9E73117AC70C7C0921FB7C" box="[935,1051,1132,1155]" italics="true" pageId="46" pageNumber="593">Steneosaurus</emphasis>
|
||
</taxonomicName>
|
||
, highlighting that taxonomic lumping and paraphyletic genera obscure morphological variation.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC63410FF9E731179FC0D7C0D85F848" blockId="46.[129,778,1231,1975]" pageId="46" pageNumber="593">
|
||
<bibRefCitation id="EFE849E1FF9E731179FC0D7C0CAEFA7B" author="Hicham B & Nehili A & Ouzzaouit LA" box="[156,404,1388,1412]" pageId="46" pageNumber="593" pagination="104804" refId="ref53643" refString="Hicham B, Nehili A, Ouzzaouit LA et al. Discovery of the teleosauroid crocodylomorph from the early Jurassic of Chaara cave, Middle Atlas of Morocco. Journal of African Earth Sciences 2023; 198: 104804." type="journal article" year="2023">
|
||
Hicham
|
||
<emphasis id="B90DE802FF9E731178600D7C0C08FA7B" box="[256,306,1388,1412]" italics="true" pageId="46" pageNumber="593">et al</emphasis>
|
||
. (2023)
|
||
</bibRefCitation>
|
||
described a specimen from the Hettangian or Sinemurian of
|
||
<collectingCountry id="F36E7480FF9E731178860D9C0F7DFA5C" box="[486,583,1420,1443]" name="Morocco" pageId="46" pageNumber="593">Morocco</collectingCountry>
|
||
that is remarkably similar to early Toarcian teleosauroids (
|
||
<bibRefCitation id="EFE849E1FF9E73117B130DBA0DF8FA1D" author="Johnson MM & Young MT & Brusatte SL" pageId="46" pageNumber="593" pagination="9808" refId="ref54662" refString="Johnson MM, Young MT, Brusatte SL. The phylogenetics of Teleosauroidea (Crocodylomorpha, Thalattosuchia) and implications for their ecology and evolution. PeerJ 2020 a; 8: e 9808. https: // doi. org / 10.7717 / peerj. 9808" type="journal article" year="2020">
|
||
Johnson
|
||
<emphasis id="B90DE802FF9E73117BB50DBB0E39FA3D" box="[725,771,1450,1474]" italics="true" pageId="46" pageNumber="593">et al</emphasis>
|
||
. 2020a
|
||
</bibRefCitation>
|
||
). Two characters from our diagnosis support their placement of the specimen within
|
||
<taxonomicName id="4C794F93FF9E731178920DF90FA1F9FE" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[498,667,1513,1537]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="46" pageNumber="582" phylum="Chordata" rank="superFamily" superFamily="Teleosauroidea">Teleosauroidea</taxonomicName>
|
||
: (i) in palatal view, the maxilla projects anteriorly with straightened margins creating a sub-rectangular shape, and (ii) first and second premaxillary alveoli form a couplet. Moreover, in our phylogenetic analyses the specimen is recovered within
|
||
<taxonomicName id="4C794F93FF9E731179E10E960C1DF961" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[129,295,1670,1694]" class="Reptilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="46" pageNumber="582" phylum="Chordata" rank="superFamily" superFamily="Teleosauroidea">Teleosauroidea</taxonomicName>
|
||
, specifically in
|
||
<taxonomicName id="4C794F93FF9E731178AE0E960FA8F961" box="[462,658,1670,1694]" family="Machimosauridae" pageId="46" pageNumber="582" rank="family">Machimosauridae</taxonomicName>
|
||
. We regard the referral of the specimen to
|
||
<taxonomicName id="4C794F93FF9E731178B50EB50FADF942" box="[469,663,1701,1725]" family="Machimosauridae" pageId="46" pageNumber="582" rank="family">Machimosauridae</taxonomicName>
|
||
to be premature, however, as many of the diagnostic characters for
|
||
<taxonomicName id="4C794F93FF9E731179E10EF40C2DF903" authorityName="Geoffroy Saint-Hilaire" authorityYear="1831" box="[129,279,1764,1788]" class="Reptilia" family="Teleosauridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="46" pageNumber="593" phylum="Chordata" rank="family">Teleosauridae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C794F93FF9E731178340EF40F2CF903" box="[340,534,1764,1788]" family="Machimosauridae" pageId="46" pageNumber="582" rank="family">Machimosauridae</taxonomicName>
|
||
cannot be scored for that specimen. While our understanding of thalattosuchian character distribution prior to the Toarcian is currently limited, at present we concur with
|
||
<bibRefCitation id="EFE849E1FF9E731178960F520FE2F8A5" author="Hicham B & Nehili A & Ouzzaouit LA" box="[502,728,1858,1882]" pageId="46" pageNumber="593" pagination="104804" refId="ref53643" refString="Hicham B, Nehili A, Ouzzaouit LA et al. Discovery of the teleosauroid crocodylomorph from the early Jurassic of Chaara cave, Middle Atlas of Morocco. Journal of African Earth Sciences 2023; 198: 104804." type="journal article" year="2023">
|
||
Hicham
|
||
<emphasis id="B90DE802FF9E73117B330F520F44F8A5" box="[595,638,1858,1882]" italics="true" pageId="46" pageNumber="593">et al</emphasis>
|
||
. (2023)
|
||
</bibRefCitation>
|
||
that the
|
||
<collectingCountry id="F36E7480FF9E731179C80F710C33F887" box="[168,265,1889,1912]" name="Morocco" pageId="46" pageNumber="593">Morocco</collectingCountry>
|
||
specimen is a teleosauroid. Thus, extending the geological range of the clade by possibly 15 million years or more.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |