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<document id="1CA45AB56201CC31C19FFEB5562FAB9B" ID-DOI="10.11646/zootaxa.5569.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="14703528" ID-ZooBank="96194340-054D-49C0-BD89-D47899BCAF14" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1737371250141" checkinUser="plazi" docAuthor="Yamana, Yusuke, Hirashima, Kentarou, Sato, Masanori &amp; Yamamori, Luna" docDate="2025" docId="03CC879FF9449D57FF63217B8936AB73" docLanguage="en" docName="zootaxa.5569.1.1.pdf" docOrigin="Zootaxa 5569 (1)" docSource="https://doi.org/10.11646/zootaxa.5569.1.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patinapta deformis Yamana, Hirashima, Sato &amp; Yamamori, 2025, sp. nov." docType="treatment" docVersion="1" lastPageNumber="28" masterDocId="FFF5FFE7F9529D4CFFF4243A8C2BAC76" masterDocTitle="First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan" masterLastPageNumber="54" masterPageNumber="1" pageNumber="23" updateTime="1737473723657" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title id="C50E1BE568B5336A589C12FBF3A65E5C">First revision of the apodid holothurian genus Patinapta Heding, 1928 (Synaptidae) in Japanese waters, with establishment of four new species from Kyushu Island and Ryukyu Islands, southwestern Japan</mods:title>
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<mods:namePart id="D597D23B9FD20910C78CBF1FB8408BF9">Yamana, Yusuke</mods:namePart>
<mods:affiliation id="29B2CDBC777955C63C648CB7D23C6D6B">Wakayama Prefectural Museum of Natural History (WMNH), Invertebrate Specimen House, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan</mods:affiliation>
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<mods:namePart id="D76283256B5835F4F0DDBDD48329E0A8">Hirashima, Kentarou</mods:namePart>
<mods:affiliation id="09BF366CC8063E93E87D638277DE0DB3">Wakayama Prefectural Museum of Natural History (WMNH), 1 Room Curators Head Office, Funo 370 - 1, Kainan, Wakayama 642 - 0001, Japan</mods:affiliation>
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<mods:namePart id="649486317B7B1BDD3CB58040453A9D06">Sato, Masanori</mods:namePart>
<mods:affiliation id="36F8647692FC434978A70D11E7A2D6D4">Graduate School of Science and Engineering, Kagoshima University, Kagoshima 890 - 0065, Japan &amp; Present address: 5233 Hiramatsu, Aira, Kagoshima 899 - 5652, Japan</mods:affiliation>
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<mods:namePart id="068ED170173D29C7D7F88BC5D18A5D6C">Yamamori, Luna</mods:namePart>
<mods:affiliation id="DF3DC6AB70D3092532BEA225F2C58AF3">Seto Marine biological laboratory (SMBL) of Kyoto University, 459, Shirahama, Wakayama 649 - 2211, Japan</mods:affiliation>
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<paragraph id="8BDA3689F9449D5AFF63217B8DC8A92D" blockId="22.[151,705,1344,1443]" box="[151,483,1344,1371]" pageId="22" pageNumber="23">
<heading id="D09281E5F9449D5AFF63217B8DC8A92D" bold="true" box="[151,483,1344,1371]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<emphasis id="B911EA9BF9449D5AFF63217B8DC8A92D" bold="true" box="[151,483,1344,1371]" pageId="22" pageNumber="23">
<taxonomicName id="4C654D0AF9449D5AFF63217B8D55A92D" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[151,382,1344,1371]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="22" pageNumber="23" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF9449D5AFF63217B8D55A92D" bold="true" box="[151,382,1344,1371]" italics="true" pageId="22" pageNumber="23">Patinapta deformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257E0F9449D5AFE71217B8DC8A92D" box="[389,483,1345,1371]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C37F6502F9449D5AFF63215C8EEAA909" box="[151,705,1381,1408]" pageId="22" pageNumber="23" type="vernacular_names">
<paragraph id="8BDA3689F9449D5AFF63215C8EEAA909" blockId="22.[151,705,1344,1443]" box="[151,705,1381,1408]" pageId="22" pageNumber="23">[New Japanese name: Yoron-himo-ikari-namako]</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5AFF6321B38DA4A98B" pageId="22" pageNumber="23" type="description">
<paragraph id="8BDA3689F9449D5AFF6321B38DE0A9D5" blockId="22.[151,705,1344,1443]" box="[151,459,1417,1443]" pageId="22" pageNumber="23">
(
<figureCitation id="135E2A0CF9449D5AFF6B21B38D0AA9D5" box="[159,289,1417,1443]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStartId-0="23.[151,250,1446,1470]" captionStartId-1="25.[151,250,1699,1723]" captionStartId-2="26.[151,250,516,540]" captionTargetBox-0="[151,1436,309,1423]" captionTargetBox-1="[151,1436,181,1667]" captionTargetBox-2="[164,1423,188,485]" captionTargetId-0="figure-69@23.[151,1436,309,1423]" captionTargetId-1="figure-20@25.[151,1436,181,1675]" captionTargetId-2="figure-29@26.[151,1435,181,493]" captionTargetPageId-0="23" captionTargetPageId-1="25" captionTargetPageId-2="26" captionText-0="FIGURE 12. External view of holotype specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (specimen WMNH-2015-INV-106): A, lateral view, anterior left, dorsal top; B, oral-side view of the tentacles; C, out-side view of the tentacles." captionText-1="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." captionText-2="FIGURE 14. Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima (holotype specimen, WMNH-2015-INV-106), described as Patinapta deformis sp. nov., in the present study. The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stand for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi-0="http://doi.org/10.5281/zenodo.14703556" figureDoi-1="http://doi.org/10.5281/zenodo.14703560" figureDoi-2="http://doi.org/10.5281/zenodo.14703562" httpUri-0="https://zenodo.org/record/14703556/files/figure.png" httpUri-1="https://zenodo.org/record/14703560/files/figure.png" httpUri-2="https://zenodo.org/record/14703562/files/figure.png" pageId="22" pageNumber="23">Figs 1214</figureCitation>
;
<tableCitation id="C6E70332F9449D5AFED821B38DA1A9D5" box="[300,394,1417,1443]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="22" pageNumber="23">Tables 1</tableCitation>
,
<tableCitation id="C6E70332F9449D5AFE6C21B38D83A9D5" box="[408,424,1417,1443]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="22" pageNumber="23">6</tableCitation>
,
<tableCitation id="C6E70332F9449D5AFE4021B38DE9A9D5" box="[436,450,1417,1443]" captionStart="TABLE 8" captionStartId="31.[152,244,152,176]" captionTargetPageId="31" captionText="TABLE 8. List of WMNH-INV specimens of Patinapta deformis sp. nov. as of 2024." pageId="22" pageNumber="23">8</tableCitation>
)
</paragraph>
<paragraph id="8BDA3689F9449D5AFF6321FD8DA4A98B" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">
<specimenCount id="9D63FD00F9449D5AFF6321FD8D68A9A9" box="[151,323,1479,1503]" count="3" pageId="22" pageNumber="23" type="generic">Three specimens</specimenCount>
from
<location id="8EBA6052F9449D5AFE7521FD8E12A9A9" LSID="urn:lsid:plazi:treatment:03CC879FF9449D57FF63217B8936AB73:8EBA6052F9449D5AFE7521FD8E12A9A9" box="[385,569,1479,1503]" municipality="We" name="Yoron-jima Island" pageId="22" pageNumber="23">Yoron-jima Island</location>
were identified as new to science.
<collectingMunicipality id="6BBEACF3F9449D5AFC6B21FD8FEBA9A8" box="[927,960,1479,1502]" pageId="22" pageNumber="23">We</collectingMunicipality>
herein described the morphological characters of this new species.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5AFF63223D8DC3AAF8" pageId="22" pageNumber="23" type="diagnosis">
<paragraph id="8BDA3689F9449D5AFF63223D8DC3AAF8" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">
<emphasis id="B911EA9BF9449D5AFF63223D8D39AA57" bold="true" box="[151,274,1543,1569]" pageId="22" pageNumber="23">Diagnosis.</emphasis>
Total shapes of anchor plates strongly distorted, their large perforations (mostly 23) fewer than those of the two congener nominal species (mostly 47), around these few perforations, marginal area broad and flat. Basal T-shaped end of anchor ossicles large, semicircular-shape. Ciliated funnels attached at inner side body wall of right dorso-lateral interradial (
<emphasis id="B911EA9BF9449D5AFE5B224E8DF3AAF8" box="[431,472,1652,1678]" italics="true" pageId="22" pageNumber="23">IR4</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5AFF3322A288F2AAC4" box="[199,1241,1687,1714]" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8BDA3689F9449D5AFF3322A288F2AAC4" blockId="22.[151,1437,1479,2038]" box="[199,1241,1687,1714]" pageId="22" pageNumber="23">
<emphasis id="B911EA9BF9449D5AFF3322A28D78AAC7" bold="true" box="[199,339,1687,1714]" pageId="22" pageNumber="23">Type series.</emphasis>
<materialsCitation id="3B0D3CD4F9449D5AFEAE22A28EF7AAC4" box="[346,732,1687,1714]" pageId="22" pageNumber="23">
<typeStatus id="54DE882BF9449D5AFEAE22A28DEEAAC4" box="[346,453,1688,1714]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
, WMNH-2015-INV-106
</materialsCitation>
;
<materialsCitation id="3B0D3CD4F9449D5AFD1322A288FEAAC4" box="[743,1237,1687,1714]" pageId="22" pageNumber="23">
<specimenCount id="9D63FD00F9449D5AFD1322A28FADAAC4" box="[743,902,1688,1714]" count="2" pageId="22" pageNumber="23" type="generic" typeStatus="paratype">two paratypes</specimenCount>
, WMNH-2015-INV-105, 108
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9449D5BFF3322818D56AC8F" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="8BDA3689F9449D5AFF3322818E71ABD8" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">
<emphasis id="B911EA9BF9449D5AFF3322818880AAA3" bold="true" box="[199,1195,1723,1750]" pageId="22" pageNumber="23">
Description of
<typeStatus id="54DE882BF9449D5AFE8322868DF5AAA0" box="[375,478,1724,1750]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
(of WMNH-2015-INV-106: middle-size complete specimen).
</emphasis>
External and internal morphologies as follows (
<tableCitation id="C6E70332F9449D5AFE4122DA8E22AA8C" box="[437,521,1760,1786]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="22" pageNumber="23">Table 1</tableCitation>
): 30 years-preserved specimen pale peanut color (
<figureCitation id="135E2A0CF9449D5AFBC122DA88ACAA8C" box="[1077,1159,1760,1786]" captionStart="FIGURE 12" captionStartId="23.[151,250,1446,1470]" captionTargetBox="[151,1436,309,1423]" captionTargetId="figure-69@23.[151,1436,309,1423]" captionTargetPageId="23" captionText="FIGURE 12. External view of holotype specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (specimen WMNH-2015-INV-106): A, lateral view, anterior left, dorsal top; B, oral-side view of the tentacles; C, out-side view of the tentacles." figureDoi="http://doi.org/10.5281/zenodo.14703556" httpUri="https://zenodo.org/record/14703556/files/figure.png" pageId="22" pageNumber="23">Fig. 12</figureCitation>
). Tentacles 12, each with 10 digits of 5 pairs. On the oral-side of tentacle stem, about 1020 sensory cups sporadically arranged in U- shape. Polian vesicle in medio-ventral position, fusiform, double (
<tableCitation id="C6E70332F9449D5AFCB323128FB5AB34" box="[839,926,1832,1858]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="22" pageNumber="23">Table 1</tableCitation>
), stone canal single thread-like, attached to narrow space between basal parts of tentacle ampullae at mediodorsal (
<emphasis id="B911EA9BF9449D5AFC4323768FF4AB10" box="[951,991,1868,1894]" italics="true" pageId="22" pageNumber="23">IR5</emphasis>
), suddenly turned upward and forward, resulted in vesicle facing forward, distally with mold-like madreporite. Ciliated funnels in one crowded band along midline of right dorsal interradius (
<emphasis id="B911EA9BF9449D5AFDD523AE8E61ABD8" box="[545,586,1940,1966]" italics="true" pageId="22" pageNumber="23">IR4</emphasis>
).
</paragraph>
<paragraph id="8BDA3689F9449D5AFF332382889CAB80" blockId="22.[151,1437,1479,2038]" pageId="22" pageNumber="23">Inside body, two tufts of gonad tubules attached with both sides of anterior dorsal mesentery, also two whitish tubule organs adhered to upper and under sides of intestine canal. Intestine canal lacking loop.</paragraph>
<paragraph id="8BDA3689F9459D5BFF3324AD8D56AC8F" blockId="23.[151,1436,151,285]" pageId="23" pageNumber="24">Body wall coarse surface, hard and thick, sparsely scattered with ossicles, arranging latitudinal layout of all as sets of anchor and plate, their directions somewhat ordered but without symmetrical design against each axial of longitudinal muscle.</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9459D56FF3325398824A9B3" lastPageId="26" lastPageNumber="27" pageId="23" pageNumber="24" type="etymology">
<paragraph id="8BDA3689F9459D5BFF33253988B3AD6B" blockId="23.[151,1436,151,285]" box="[199,1176,259,285]" pageId="23" pageNumber="24">
<emphasis id="B911EA9BF9459D5BFF3325398D65AD6B" bold="true" box="[199,334,259,285]" pageId="23" pageNumber="24">Etymology.</emphasis>
The species name
<taxonomicName id="4C654D0AF9459D5BFDD625398EAFAD6B" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[546,644,259,285]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="deformis">
<emphasis id="B911EA9BF9459D5BFDD625398EAFAD6B" box="[546,644,259,285]" italics="true" pageId="23" pageNumber="24">deformis</emphasis>
</taxonomicName>
, alluding to the strongly distorted anchor plate.
</paragraph>
<caption id="DF1A6601F9459D5BFF63219C8EE8AA70" ID-DOI="http://doi.org/10.5281/zenodo.14703556" ID-Zenodo-Dep="14703556" httpUri="https://zenodo.org/record/14703556/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1446,1470]" targetBox="[151,1436,309,1423]" targetPageId="23" targetType="figure">
<paragraph id="8BDA3689F9459D5BFF63219C8EE8AA70" blockId="23.[151,1437,1446,1543]" pageId="23" pageNumber="24">
<emphasis id="B911EA9BF9459D5BFF63219C8D0FA9C8" bold="true" box="[151,292,1446,1470]" pageId="23" pageNumber="24">FIGURE 12.</emphasis>
External view of holotype specimen collected from Yoron-jima Island, Kagoshima, described as
<taxonomicName id="4C654D0AF9459D5BFACC219D8CDBA994" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="23" pageNumber="24" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF9459D5BFACC219D8CDBA994" italics="true" pageId="23" pageNumber="24">Patinapta deformis</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9459D5BFF0221F18D6DA995" bold="true" box="[246,326,1483,1507]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="A22257E0F9459D5BFF0221F18D6DA995" box="[246,326,1483,1507]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study (specimen WMNH-2015-INV-106): A, lateral view, anterior left, dorsal top; B, oral-side view of the tentacles; C, out-side view of the tentacles.
</paragraph>
</caption>
<paragraph id="8BDA3689F9459D5BFF6322638E86AA04" blockId="23.[151,685,1624,1651]" box="[151,685,1624,1651]" pageId="23" pageNumber="24">
<heading id="D09281E5F9459D5BFF6322638E86AA04" bold="true" box="[151,685,1624,1651]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="B911EA9BF9459D5BFF6322638E86AA04" bold="true" box="[151,685,1624,1651]" pageId="23" pageNumber="24">The present results of ossicle morphologies</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9459D5BFF63229B8D4BAB19" blockId="23.[151,1437,1697,1903]" pageId="23" pageNumber="24">
Body wall ossicles anchors and anchor plates (
<figureCitation id="135E2A0CF9459D5BFD59229B8F13AACD" box="[685,824,1697,1723]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="23" pageNumber="24">Fig. 13B, D</figureCitation>
;
<tableCitation id="C6E70332F9459D5BFCB1229B8FB7AACD" box="[837,924,1697,1723]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="23" pageNumber="24">Table 6</tableCitation>
). Anchor plates ranged within 98128 μm in length. Size of perforations noticeably variable between central area and marginal area. Largest perforation situated noticeably biased from the center, surrounded by scattering smaller perforations. Few numbers of largest perforations and several small perforations equipped with 418 spaced sharp-teeth arraign along with inner rim. Two calculated indicators for anchor plates and a parameter were variated within
<emphasis id="B911EA9BF9459D5BFC82230B8F8DAB3D" box="[886,934,1841,1867]" italics="true" pageId="23" pageNumber="24">APS</emphasis>
=0.622.4;
<emphasis id="B911EA9BF9459D5BFBD3230B8864AB3C" box="[1063,1103,1841,1866]" italics="true" pageId="23" pageNumber="24">Ntp</emphasis>
=27;
<emphasis id="B911EA9BF9459D5BFB6C230B88E1AB3D" box="[1176,1226,1841,1867]" italics="true" pageId="23" pageNumber="24">APT</emphasis>
=11.820.8% (
<figureCitation id="135E2A0CF9459D5BFA9B230B8CD9AB19" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="23" pageNumber="24">Fig. 13 B, D</figureCitation>
;
<tableCitation id="C6E70332F9459D5BFF09236F8D79AB19" box="[253,338,1877,1903]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="23" pageNumber="24">Table 6</tableCitation>
).
</paragraph>
<caption id="DF1A6601F94A9D54FF6C24AD8942ACC6" box="[152,1385,151,176]" pageId="24" pageNumber="25" startId="24.[152,244,151,175]" targetBox="[163,1425,192,1866]" targetIsTable="true" targetPageId="24" targetType="table">
<paragraph id="8BDA3689F94A9D54FF6C24AD8942ACC6" blockId="24.[152,1385,151,176]" box="[152,1385,151,176]" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF6C24AD8D25ACD9" bold="true" box="[152,270,151,175]" pageId="24" pageNumber="25">TABLE 6.</emphasis>
Measurements (mean±SD μm) and counts of ossicles of three specimens of
<taxonomicName id="4C654D0AF94A9D54FBB424A28926ACD9" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[1088,1293,151,175]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="24" pageNumber="25" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF94A9D54FBB424A28926ACD9" box="[1088,1293,151,175]" italics="true" pageId="24" pageNumber="25">Patinapta deformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257E0F94A9D54FAE024A28942ACC6" box="[1300,1385,152,176]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BDA3689F94A9D54FF5724FA89BAAB3C" pageId="24" pageNumber="25">
<table id="F965C429F94A62B3FF5724FA89BAAB3C" box="[163,1425,192,1866]" gridcols="8" gridrows="42" pageId="24" pageNumber="25">
<tr id="355534CBF94A62B3FF5724FA89BAACAE" box="[163,1425,192,216]" gridrow="0" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5724FA8D32ACAE" box="[163,281,192,216]" gridcol="0" gridrow="0" pageId="24" pageNumber="25">Body parts</th>
<th id="76845DB7F94A62B3FEB824FA8DD6ACAE" box="[332,509,192,216]" gridcol="1" gridrow="0" pageId="24" pageNumber="25">(Ossicle-type)</th>
<th id="76845DB7F94A62B3FDEE24FA8F0BACAE" box="[538,800,192,216]" colspan="2" colspanRight="1" gridcol="2" gridrow="0" pageId="24" pageNumber="25">2015-INV-105</th>
<th id="76845DB7F94A62B3FCC824FA887FACAE" box="[828,1108,192,216]" colspan="2" colspanRight="1" gridcol="4" gridrow="0" pageId="24" pageNumber="25">2015-INV-106</th>
<th id="76845DB7F94A62B3FB8F24FA89BAACAE" box="[1147,1425,192,216]" colspan="2" colspanRight="1" gridcol="6" gridrow="0" pageId="24" pageNumber="25">2015-INV-108</th>
</tr>
<tr id="355534CBF94A62B3FF5724D289BAAD76" box="[163,1425,232,256]" gridrow="1" pageId="24" pageNumber="25" rowspan-7="1">
<th id="76845DB7F94A62B3FF5724D28D32AD76" box="[163,281,232,256]" gridcol="0" gridrow="1" pageId="24" pageNumber="25">Tentacle</th>
<td id="76845DB7F94A62B3FEB824D28DD6AD76" box="[332,509,232,256]" gridcol="1" gridrow="1" pageId="24" pageNumber="25">
Rods (
<emphasis id="B911EA9BF94A9D54FE4F24D38DE3AD76" box="[443,456,233,256]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE24D28F0BAD76" box="[538,800,232,256]" colspan="2" colspanRight="1" gridcol="2" gridrow="1" pageId="24" pageNumber="25">13</td>
<td id="76845DB7F94A62B3FCC824D2887FAD76" box="[828,1108,232,256]" colspan="2" colspanRight="1" gridcol="4" gridrow="1" pageId="24" pageNumber="25">11</td>
<td id="76845DB7F94A62B3FB8F24D28922AD76" box="[1147,1289,232,256]" gridcol="6" gridrow="1" pageId="24" pageNumber="25">5</td>
</tr>
<tr id="355534CBF94A62B3FF57252A89BAAD5E" box="[163,1425,272,296]" gridrow="2" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57252A8D32AD5E" box="[163,281,272,296]" gridcol="0" gridrow="2" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB8252A8DD6AD5E" box="[332,509,272,296]" gridcol="1" gridrow="2" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE252A8EA5AD5E" box="[538,654,272,296]" gridcol="2" gridrow="2" pageId="24" pageNumber="25">53.7±14.3</td>
<td id="76845DB7F94A62B3FD5E252A8F0BAD5E" box="[682,800,272,296]" gridcol="3" gridrow="2" pageId="24" pageNumber="25">(3577)</td>
<td id="76845DB7F94A62B3FCC8252A8F99AD5E" box="[828,946,272,296]" gridcol="4" gridrow="2" pageId="24" pageNumber="25">63.8± 11.9</td>
<td id="76845DB7F94A62B3FC30252A887FAD5E" box="[964,1108,272,296]" gridcol="5" gridrow="2" pageId="24" pageNumber="25">(5285)</td>
<td id="76845DB7F94A62B3FB8F252A8922AD5E" box="[1147,1289,272,296]" gridcol="6" gridrow="2" pageId="24" pageNumber="25">67.4± 7.6</td>
<td id="76845DB7F94A62B3FAEF252A89BAAD5E" box="[1307,1425,272,296]" gridcol="7" gridrow="2" pageId="24" pageNumber="25">(6080)</td>
</tr>
<tr id="355534CBF94A62B3FF57250289BAAD27" box="[163,1425,312,337]" gridrow="3" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5725028D32AD27" box="[163,281,312,337]" gridcol="0" gridrow="3" pageId="24" pageNumber="25">Breadth</th>
<td id="76845DB7F94A62B3FEB825028DD6AD27" box="[332,509,312,337]" gridcol="1" gridrow="3" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE25028EA5AD27" box="[538,654,312,337]" gridcol="2" gridrow="3" pageId="24" pageNumber="25">8.5± 2.5</td>
<td id="76845DB7F94A62B3FD5E25028F0BAD27" box="[682,800,312,337]" gridcol="3" gridrow="3" pageId="24" pageNumber="25">(514)</td>
<td id="76845DB7F94A62B3FCC825028F99AD27" box="[828,946,312,337]" gridcol="4" gridrow="3" pageId="24" pageNumber="25">9.4± 2.7</td>
<td id="76845DB7F94A62B3FC302502887FAD27" box="[964,1108,312,337]" gridcol="5" gridrow="3" pageId="24" pageNumber="25">(715)</td>
<td id="76845DB7F94A62B3FB8F25028922AD27" box="[1147,1289,312,337]" gridcol="6" gridrow="3" pageId="24" pageNumber="25">10.6± 1.1</td>
<td id="76845DB7F94A62B3FAEF250289BAAD27" box="[1307,1425,312,337]" gridcol="7" gridrow="3" pageId="24" pageNumber="25">(912)</td>
</tr>
<tr id="355534CBF94A62B3FF57255A89BAAD0F" box="[163,1425,352,377]" gridrow="4" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57255A8D32AD0F" box="[163,281,352,377]" gridcol="0" gridrow="4" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4425588CE4AD0F" box="[176,207,354,377]" italics="true" pageId="24" pageNumber="25">RP</emphasis>
a (%)
</th>
<td id="76845DB7F94A62B3FEB8255A8DD6AD0F" box="[332,509,352,377]" gridcol="1" gridrow="4" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE255A8EA5AD0F" box="[538,654,352,377]" gridcol="2" gridrow="4" pageId="24" pageNumber="25">16.3± 5.0</td>
<td id="76845DB7F94A62B3FD5E255A8F0BAD0F" box="[682,800,352,377]" gridcol="3" gridrow="4" pageId="24" pageNumber="25">(10.126.3)</td>
<td id="76845DB7F94A62B3FCC8255A8F99AD0F" box="[828,946,352,377]" gridcol="4" gridrow="4" pageId="24" pageNumber="25">14.7± 3.0</td>
<td id="76845DB7F94A62B3FC30255A887FAD0F" box="[964,1108,352,377]" gridcol="5" gridrow="4" pageId="24" pageNumber="25">(10.720.3)</td>
<td id="76845DB7F94A62B3FB8F255A8922AD0F" box="[1147,1289,352,377]" gridcol="6" gridrow="4" pageId="24" pageNumber="25">15.8± 2.1</td>
<td id="76845DB7F94A62B3FAEF255A89BAAD0F" box="[1307,1425,352,377]" gridcol="7" gridrow="4" pageId="24" pageNumber="25">(13.818.3)</td>
</tr>
<tr id="355534CBF94A62B3FF5725B389BAADD7" box="[163,1425,393,417]" gridrow="5" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5725B38D32ADD7" box="[163,281,393,417]" gridcol="0" gridrow="5" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4425B38CFBADD6" box="[176,208,393,416]" italics="true" pageId="24" pageNumber="25">RC</emphasis>
b
</th>
<td id="76845DB7F94A62B3FEB825B38DD6ADD7" box="[332,509,393,417]" gridcol="1" gridrow="5" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE25B38EA5ADD7" box="[538,654,393,417]" gridcol="2" gridrow="5" pageId="24" pageNumber="25">11.1± 1.7</td>
<td id="76845DB7F94A62B3FD5E25B38F0BADD7" box="[682,800,393,417]" gridcol="3" gridrow="5" pageId="24" pageNumber="25">(5.317.3)</td>
<td id="76845DB7F94A62B3FCC825B38F99ADD7" box="[828,946,393,417]" gridcol="4" gridrow="5" pageId="24" pageNumber="25">7.6± 1.7</td>
<td id="76845DB7F94A62B3FC3025B3887FADD7" box="[964,1108,393,417]" gridcol="5" gridrow="5" pageId="24" pageNumber="25">(5.412.2)</td>
<td id="76845DB7F94A62B3FB8F25B38922ADD7" box="[1147,1289,393,417]" gridcol="6" gridrow="5" pageId="24" pageNumber="25">7.9± 2.7</td>
<td id="76845DB7F94A62B3FAEF25B389BAADD7" box="[1307,1425,393,417]" gridcol="7" gridrow="5" pageId="24" pageNumber="25">(5.011.7)</td>
</tr>
<tr id="355534CBF94A62B3FF57258B89BAADBF" box="[163,1425,433,457]" gridrow="6" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57258B89BAADBF" box="[163,1425,433,457]" colspan="8" colspanRight="7" gridcol="0" gridrow="6" pageId="24" pageNumber="25">Ventral side</th>
</tr>
<tr id="355534CBF94A62B3FF5725E389BAAD84" box="[163,1425,473,498]" gridrow="7" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF5725E38D32AD84" box="[163,281,473,498]" gridcol="0" gridrow="7" pageId="24" pageNumber="25">L-Muscle</th>
<td id="76845DB7F94A62B3FEB825E38DD6AD84" box="[332,509,473,498]" gridcol="1" gridrow="7" pageId="24" pageNumber="25">
Granules (
<emphasis id="B911EA9BF94A9D54FE3B25E08DF7AD87" box="[463,476,474,497]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE25E38F0BAD84" box="[538,800,473,498]" colspan="2" colspanRight="1" gridcol="2" gridrow="7" pageId="24" pageNumber="25">7</td>
<td id="76845DB7F94A62B3FC3025E3887FAD84" box="[964,1108,473,498]" gridcol="5" gridrow="7" pageId="24" pageNumber="25">9</td>
<td id="76845DB7F94A62B3FB8F25E38922AD84" box="[1147,1289,473,498]" gridcol="6" gridrow="7" pageId="24" pageNumber="25">9</td>
</tr>
<tr id="355534CBF94A62B3FF57263889BAAE6C" box="[163,1425,514,538]" gridrow="8" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726388D32AE6C" box="[163,281,514,538]" gridcol="0" gridrow="8" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB826388DD6AE6C" box="[332,509,514,538]" gridcol="1" gridrow="8" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE26388EA5AE6C" box="[538,654,514,538]" gridcol="2" gridrow="8" pageId="24" pageNumber="25">30.3 ± 3.9</td>
<td id="76845DB7F94A62B3FD5E26388F0BAE6C" box="[682,800,514,538]" gridcol="3" gridrow="8" pageId="24" pageNumber="25">(2436)</td>
<td id="76845DB7F94A62B3FCC826388F99AE6C" box="[828,946,514,538]" gridcol="4" gridrow="8" pageId="24" pageNumber="25">29.3 ± 4.4</td>
<td id="76845DB7F94A62B3FC302638887FAE6C" box="[964,1108,514,538]" gridcol="5" gridrow="8" pageId="24" pageNumber="25">(2336)</td>
<td id="76845DB7F94A62B3FB8F26388922AE6C" box="[1147,1289,514,538]" gridcol="6" gridrow="8" pageId="24" pageNumber="25">28.8 ± 3.9</td>
<td id="76845DB7F94A62B3FAEF263889BAAE6C" box="[1307,1425,514,538]" gridcol="7" gridrow="8" pageId="24" pageNumber="25">(2434)</td>
</tr>
<tr id="355534CBF94A62B3FF57261089BAAE34" box="[163,1425,554,578]" gridrow="9" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726108D32AE34" box="[163,281,554,578]" gridcol="0" gridrow="9" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB826108DD6AE34" box="[332,509,554,578]" gridcol="1" gridrow="9" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE26108EA5AE34" box="[538,654,554,578]" gridcol="2" gridrow="9" pageId="24" pageNumber="25">14.3 ±1.5</td>
<td id="76845DB7F94A62B3FD5E26108F0BAE34" box="[682,800,554,578]" gridcol="3" gridrow="9" pageId="24" pageNumber="25">(1216)</td>
<td id="76845DB7F94A62B3FCC826108F99AE34" box="[828,946,554,578]" gridcol="4" gridrow="9" pageId="24" pageNumber="25">16.8 ± 2.3</td>
<td id="76845DB7F94A62B3FC302610887FAE34" box="[964,1108,554,578]" gridcol="5" gridrow="9" pageId="24" pageNumber="25">(1321)</td>
<td id="76845DB7F94A62B3FB8F26108922AE34" box="[1147,1289,554,578]" gridcol="6" gridrow="9" pageId="24" pageNumber="25">13.8 ± 1.9</td>
<td id="76845DB7F94A62B3FAEF261089BAAE34" box="[1307,1425,554,578]" gridcol="7" gridrow="9" pageId="24" pageNumber="25">(1017)</td>
</tr>
<tr id="355534CBF94A62B3FF57266B89BAAE1C" box="[163,1425,593,618]" gridrow="10" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57266B8D32AE1C" box="[163,281,593,618]" gridcol="0" gridrow="10" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4426698CF9AE1C" box="[176,210,595,618]" italics="true" pageId="24" pageNumber="25">GP</emphasis>
c
</th>
<td id="76845DB7F94A62B3FEB8266B8DD6AE1C" box="[332,509,593,618]" gridcol="1" gridrow="10" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE266B8EA5AE1C" box="[538,654,593,618]" gridcol="2" gridrow="10" pageId="24" pageNumber="25">48.4 ± 11.2</td>
<td id="76845DB7F94A62B3FD5E266B8F0BAE1C" box="[682,800,593,618]" gridcol="3" gridrow="10" pageId="24" pageNumber="25">(36.166.7)</td>
<td id="76845DB7F94A62B3FCC8266B887FAE1C" box="[828,1108,593,618]" colspan="2" colspanRight="1" gridcol="4" gridrow="10" pageId="24" pageNumber="25">59.5 ± 15.2 (36.182.6)</td>
<td id="76845DB7F94A62B3FB8F266B8922AE1C" box="[1147,1289,593,618]" gridcol="6" gridrow="10" pageId="24" pageNumber="25">48.2 ± 5.9</td>
<td id="76845DB7F94A62B3FAEF266B89BAAE1C" box="[1307,1425,593,618]" gridcol="7" gridrow="10" pageId="24" pageNumber="25">(39.455.6)</td>
</tr>
<tr id="355534CBF94A62B3FF57264089BAAEE5" box="[163,1425,634,659]" gridrow="11" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726408D32AEE5" box="[163,281,634,659]" gridcol="0" gridrow="11" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4426418CCEAEE4" box="[176,229,635,658]" italics="true" pageId="24" pageNumber="25">GCO</emphasis>
d (%)
</th>
<td id="76845DB7F94A62B3FEB826408DD6AEE5" box="[332,509,634,659]" gridcol="1" gridrow="11" pageId="24" pageNumber="25">(Ratio)</td>
<td id="76845DB7F94A62B3FDEE26408EA5AEE5" box="[538,654,634,659]" gridcol="2" gridrow="11" pageId="24" pageNumber="25">14.3</td>
<td id="76845DB7F94A62B3FD5E26408F0BAEE5" box="[682,800,634,659]" gridcol="3" gridrow="11" pageId="24" pageNumber="25">(O1: C6)</td>
<td id="76845DB7F94A62B3FCC826408F99AEE5" box="[828,946,634,659]" gridcol="4" gridrow="11" pageId="24" pageNumber="25">44.4</td>
<td id="76845DB7F94A62B3FC302640887FAEE5" box="[964,1108,634,659]" gridcol="5" gridrow="11" pageId="24" pageNumber="25">(O4: C5)</td>
<td id="76845DB7F94A62B3FB8F26408922AEE5" box="[1147,1289,634,659]" gridcol="6" gridrow="11" pageId="24" pageNumber="25">22.2</td>
<td id="76845DB7F94A62B3FAEF264089BAAEE5" box="[1307,1425,634,659]" gridcol="7" gridrow="11" pageId="24" pageNumber="25">(O2: C7)</td>
</tr>
<tr id="355534CBF94A62B3FF57269989BAAECD" box="[163,1425,675,699]" gridrow="12" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF5726998D32AECD" box="[163,281,675,699]" gridcol="0" gridrow="12" pageId="24" pageNumber="25">Body wall</th>
<td id="76845DB7F94A62B3FEB826998DD6AECD" box="[332,509,675,699]" gridcol="1" gridrow="12" pageId="24" pageNumber="25">
Anchor plates (
<emphasis id="B911EA9BF94A9D54FE1C269E8DDEAECD" box="[488,501,676,699]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE26998F0BAECD" box="[538,800,675,699]" colspan="2" colspanRight="1" gridcol="2" gridrow="12" pageId="24" pageNumber="25">7</td>
<td id="76845DB7F94A62B3FC302699887FAECD" box="[964,1108,675,699]" gridcol="5" gridrow="12" pageId="24" pageNumber="25">4</td>
<td id="76845DB7F94A62B3FB8F26998922AECD" box="[1147,1289,675,699]" gridcol="6" gridrow="12" pageId="24" pageNumber="25">7</td>
</tr>
<tr id="355534CBF94A62B3FF5726F189BAAE95" box="[163,1425,715,739]" gridrow="13" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726F18D32AE95" box="[163,281,715,739]" gridcol="0" gridrow="13" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB826F18DD6AE95" box="[332,509,715,739]" gridcol="1" gridrow="13" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE26F18EA5AE95" box="[538,654,715,739]" gridcol="2" gridrow="13" pageId="24" pageNumber="25">127.7 ± 6.3</td>
<td id="76845DB7F94A62B3FD5E26F18F0BAE95" box="[682,800,715,739]" gridcol="3" gridrow="13" pageId="24" pageNumber="25">(119138)</td>
<td id="76845DB7F94A62B3FCC826F1887FAE95" box="[828,1108,715,739]" colspan="2" colspanRight="1" gridcol="4" gridrow="13" pageId="24" pageNumber="25">110.5 ± 6.2 (104119)</td>
<td id="76845DB7F94A62B3FB8F26F18922AE95" box="[1147,1289,715,739]" gridcol="6" gridrow="13" pageId="24" pageNumber="25">95.7 ± 8.5</td>
<td id="76845DB7F94A62B3FAEF26F189BAAE95" box="[1307,1425,715,739]" gridcol="7" gridrow="13" pageId="24" pageNumber="25">(83108)</td>
</tr>
<tr id="355534CBF94A62B3FF5726C989BAAF7D" box="[163,1425,755,779]" gridrow="14" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5726C98D32AF7D" box="[163,281,755,779]" gridcol="0" gridrow="14" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB826C98DD6AF7D" box="[332,509,755,779]" gridcol="1" gridrow="14" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE26C98EA5AF7D" box="[538,654,755,779]" gridcol="2" gridrow="14" pageId="24" pageNumber="25">99.0 ± 5.1</td>
<td id="76845DB7F94A62B3FD5E26C98F0BAF7D" box="[682,800,755,779]" gridcol="3" gridrow="14" pageId="24" pageNumber="25">(94107)</td>
<td id="76845DB7F94A62B3FCC826C98F99AF7D" box="[828,946,755,779]" gridcol="4" gridrow="14" pageId="24" pageNumber="25">70.5 ± 3.9</td>
<td id="76845DB7F94A62B3FC3026C9887FAF7D" box="[964,1108,755,779]" gridcol="5" gridrow="14" pageId="24" pageNumber="25">(6776)</td>
<td id="76845DB7F94A62B3FB8F26C98922AF7D" box="[1147,1289,755,779]" gridcol="6" gridrow="14" pageId="24" pageNumber="25">64.1 ± 6.4</td>
<td id="76845DB7F94A62B3FAEF26C989BAAF7D" box="[1307,1425,755,779]" gridcol="7" gridrow="14" pageId="24" pageNumber="25">(5469)</td>
</tr>
<tr id="355534CBF94A62B3FF57272189BAAF42" box="[163,1425,795,820]" gridrow="15" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5727218D32AF42" box="[163,281,795,820]" gridcol="0" gridrow="15" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4427268CF7AF45" box="[176,220,796,819]" italics="true" pageId="24" pageNumber="25">APS</emphasis>
e
</th>
<td id="76845DB7F94A62B3FEB827218DD6AF42" box="[332,509,795,820]" gridcol="1" gridrow="15" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE27218EA5AF42" box="[538,654,795,820]" gridcol="2" gridrow="15" pageId="24" pageNumber="25">9.9 ± 5.7</td>
<td id="76845DB7F94A62B3FD5E27218F0BAF42" box="[682,800,795,820]" gridcol="3" gridrow="15" pageId="24" pageNumber="25">(2.313.5)</td>
<td id="76845DB7F94A62B3FCC827218F99AF42" box="[828,946,795,820]" gridcol="4" gridrow="15" pageId="24" pageNumber="25">5.1 ± 4.8</td>
<td id="76845DB7F94A62B3FC302721887FAF42" box="[964,1108,795,820]" gridcol="5" gridrow="15" pageId="24" pageNumber="25">(0.611.7)</td>
<td id="76845DB7F94A62B3FB8F27218922AF42" box="[1147,1289,795,820]" gridcol="6" gridrow="15" pageId="24" pageNumber="25">2.5 ± 2.5</td>
<td id="76845DB7F94A62B3FAEF272189BAAF42" box="[1307,1425,795,820]" gridcol="7" gridrow="15" pageId="24" pageNumber="25">(0.66.1)</td>
</tr>
<tr id="355534CBF94A62B3FF57277E89BAAF2A" box="[163,1425,836,860]" gridrow="16" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57277E8D32AF2A" box="[163,281,836,860]" gridcol="0" gridrow="16" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44277F8CFEAF2A" box="[176,213,837,860]" italics="true" pageId="24" pageNumber="25">Ntp</emphasis>
f
</th>
<td id="76845DB7F94A62B3FEB8277E8DD6AF2A" box="[332,509,836,860]" gridcol="1" gridrow="16" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE277E8EA5AF2A" box="[538,654,836,860]" gridcol="2" gridrow="16" pageId="24" pageNumber="25">3.6 ± 1.4</td>
<td id="76845DB7F94A62B3FD5E277E8F0BAF2A" box="[682,800,836,860]" gridcol="3" gridrow="16" pageId="24" pageNumber="25">(15)</td>
<td id="76845DB7F94A62B3FCC8277E8F99AF2A" box="[828,946,836,860]" gridcol="4" gridrow="16" pageId="24" pageNumber="25">4.5 ± 1.7</td>
<td id="76845DB7F94A62B3FC30277E887FAF2A" box="[964,1108,836,860]" gridcol="5" gridrow="16" pageId="24" pageNumber="25">(37)</td>
<td id="76845DB7F94A62B3FB8F277E8922AF2A" box="[1147,1289,836,860]" gridcol="6" gridrow="16" pageId="24" pageNumber="25">2.3 ± 0.8</td>
<td id="76845DB7F94A62B3FAEF277E89BAAF2A" box="[1307,1425,836,860]" gridcol="7" gridrow="16" pageId="24" pageNumber="25">(13)</td>
</tr>
<tr id="355534CBF94A62B3FF57275189BAAFF2" box="[163,1425,875,900]" gridrow="17" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5727518D32AFF2" box="[163,281,875,900]" gridcol="0" gridrow="17" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4427568CF6AFF5" box="[176,221,876,899]" italics="true" pageId="24" pageNumber="25">APT</emphasis>
g (%)
</th>
<td id="76845DB7F94A62B3FEB827518DD6AFF2" box="[332,509,875,900]" gridcol="1" gridrow="17" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE27518EA5AFF2" box="[538,654,875,900]" gridcol="2" gridrow="17" pageId="24" pageNumber="25">9.9 ± 3.7</td>
<td id="76845DB7F94A62B3FD5E27518F0BAFF2" box="[682,800,875,900]" gridcol="3" gridrow="17" pageId="24" pageNumber="25">(2.313.5)</td>
<td id="76845DB7F94A62B3FCC827518F99AFF2" box="[828,946,875,900]" gridcol="4" gridrow="17" pageId="24" pageNumber="25">14.6 ± 2.1</td>
<td id="76845DB7F94A62B3FC302751887FAFF2" box="[964,1108,875,900]" gridcol="5" gridrow="17" pageId="24" pageNumber="25">(11.816.7)</td>
<td id="76845DB7F94A62B3FB8F27518922AFF2" box="[1147,1289,875,900]" gridcol="6" gridrow="17" pageId="24" pageNumber="25">11.8 ± 3.8</td>
<td id="76845DB7F94A62B3FAEF275189BAAFF2" box="[1307,1425,875,900]" gridcol="7" gridrow="17" pageId="24" pageNumber="25">(8.018.8)</td>
</tr>
<tr id="355534CBF94A62B3FF5727AE89BAAFDB" box="[163,1425,916,941]" gridrow="18" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF5727AE8D32AFDB" box="[163,281,916,941]" gridcol="0" gridrow="18" pageId="24" pageNumber="25">Body wall</th>
<td id="76845DB7F94A62B3FEB827AE8DD6AFDB" box="[332,509,916,941]" gridcol="1" gridrow="18" pageId="24" pageNumber="25">
Anchor (
<emphasis id="B911EA9BF94A9D54FE3327AF8DFFAFDA" box="[455,468,917,940]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE27AE8F0BAFDB" box="[538,800,916,941]" colspan="2" colspanRight="1" gridcol="2" gridrow="18" pageId="24" pageNumber="25">7</td>
<td id="76845DB7F94A62B3FC3027AE887FAFDB" box="[964,1108,916,941]" gridcol="5" gridrow="18" pageId="24" pageNumber="25">4</td>
<td id="76845DB7F94A62B3FB8F27AE8922AFDB" box="[1147,1289,916,941]" gridcol="6" gridrow="18" pageId="24" pageNumber="25">8</td>
</tr>
<tr id="355534CBF94A62B3FF57278689BAAFA3" box="[163,1425,956,981]" gridrow="19" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5727868D32AFA3" box="[163,281,956,981]" gridcol="0" gridrow="19" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB827868DD6AFA3" box="[332,509,956,981]" gridcol="1" gridrow="19" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE27868EA5AFA3" box="[538,654,956,981]" gridcol="2" gridrow="19" pageId="24" pageNumber="25">166.3±9.5</td>
<td id="76845DB7F94A62B3FD5E27868F0BAFA3" box="[682,800,956,981]" gridcol="3" gridrow="19" pageId="24" pageNumber="25">(157181)</td>
<td id="76845DB7F94A62B3FCC82786887FAFA3" box="[828,1108,956,981]" colspan="2" colspanRight="1" gridcol="4" gridrow="19" pageId="24" pageNumber="25">157.8 ± 12.5 (144174)</td>
<td id="76845DB7F94A62B3FB8F27868922AFA3" box="[1147,1289,956,981]" gridcol="6" gridrow="19" pageId="24" pageNumber="25">158.1 ± 7.1</td>
<td id="76845DB7F94A62B3FAEF278689BAAFA3" box="[1307,1425,956,981]" gridcol="7" gridrow="19" pageId="24" pageNumber="25">(147169)</td>
</tr>
<tr id="355534CBF94A62B3FF5727DF89BAAF8B" box="[163,1425,997,1021]" gridrow="20" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5727DF8D32AF8B" box="[163,281,997,1021]" gridcol="0" gridrow="20" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB827DF8DD6AF8B" box="[332,509,997,1021]" gridcol="1" gridrow="20" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE27DF8EA5AF8B" box="[538,654,997,1021]" gridcol="2" gridrow="20" pageId="24" pageNumber="25">85.0 ± 5.9</td>
<td id="76845DB7F94A62B3FD5E27DF8F0BAF8B" box="[682,800,997,1021]" gridcol="3" gridrow="20" pageId="24" pageNumber="25">(7693)</td>
<td id="76845DB7F94A62B3FCC827DF8F99AF8B" box="[828,946,997,1021]" gridcol="4" gridrow="20" pageId="24" pageNumber="25">85.3 ± 5.2</td>
<td id="76845DB7F94A62B3FC3027DF887FAF8B" box="[964,1108,997,1021]" gridcol="5" gridrow="20" pageId="24" pageNumber="25">(7990)</td>
<td id="76845DB7F94A62B3FB8F27DF8922AF8B" box="[1147,1289,997,1021]" gridcol="6" gridrow="20" pageId="24" pageNumber="25">68.5 ± 9.4</td>
<td id="76845DB7F94A62B3FAEF27DF89BAAF8B" box="[1307,1425,997,1021]" gridcol="7" gridrow="20" pageId="24" pageNumber="25">(5884)</td>
</tr>
<tr id="355534CBF94A62B3FF57203789BAA853" box="[163,1425,1037,1061]" gridrow="21" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720378D32A853" box="[163,281,1037,1061]" gridcol="0" gridrow="21" pageId="24" pageNumber="25">Teeth</th>
<td id="76845DB7F94A62B3FEB820378DD6A853" box="[332,509,1037,1061]" gridcol="1" gridrow="21" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE20378EA5A853" box="[538,654,1037,1061]" gridcol="2" gridrow="21" pageId="24" pageNumber="25">1.6 ± 0.9</td>
<td id="76845DB7F94A62B3FD5E20378F0BA853" box="[682,800,1037,1061]" gridcol="3" gridrow="21" pageId="24" pageNumber="25">(03)</td>
<td id="76845DB7F94A62B3FCC820378F99A853" box="[828,946,1037,1061]" gridcol="4" gridrow="21" pageId="24" pageNumber="25">3.3 ± 1.5</td>
<td id="76845DB7F94A62B3FC302037887FA853" box="[964,1108,1037,1061]" gridcol="5" gridrow="21" pageId="24" pageNumber="25">(25)</td>
<td id="76845DB7F94A62B3FB8F20378922A853" box="[1147,1289,1037,1061]" gridcol="6" gridrow="21" pageId="24" pageNumber="25">3.7 ± 0.7</td>
<td id="76845DB7F94A62B3FAEF203789BAA853" box="[1307,1425,1037,1061]" gridcol="7" gridrow="21" pageId="24" pageNumber="25">(35)</td>
</tr>
<tr id="355534CBF94A62B3FF57200F89BAA838" box="[163,1425,1077,1102]" gridrow="22" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57200F8D32A838" box="[163,281,1077,1102]" gridcol="0" gridrow="22" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44200C8CCAA83B" box="[176,225,1078,1101]" italics="true" pageId="24" pageNumber="25">ASW</emphasis>
h(%)
</th>
<td id="76845DB7F94A62B3FEB8200F8DD6A838" box="[332,509,1077,1102]" gridcol="1" gridrow="22" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE200F8EA5A838" box="[538,654,1077,1102]" gridcol="2" gridrow="22" pageId="24" pageNumber="25">23.1 ± 1.2</td>
<td id="76845DB7F94A62B3FD5E200F8F0BA838" box="[682,800,1077,1102]" gridcol="3" gridrow="22" pageId="24" pageNumber="25">(21.525.0)</td>
<td id="76845DB7F94A62B3FCC8200F8F99A838" box="[828,946,1077,1102]" gridcol="4" gridrow="22" pageId="24" pageNumber="25">20.1 ± 0.7</td>
<td id="76845DB7F94A62B3FC30200F887FA838" box="[964,1108,1077,1102]" gridcol="5" gridrow="22" pageId="24" pageNumber="25">(19.421.1)</td>
<td id="76845DB7F94A62B3FB8F200F8922A838" box="[1147,1289,1077,1102]" gridcol="6" gridrow="22" pageId="24" pageNumber="25">25.0 ± 2.1</td>
<td id="76845DB7F94A62B3FAEF200F89BAA838" box="[1307,1425,1077,1102]" gridcol="7" gridrow="22" pageId="24" pageNumber="25">(21.428.4)</td>
</tr>
<tr id="355534CBF94A62B3FF57206789BAA800" box="[163,1425,1117,1142]" gridrow="23" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720678D32A800" box="[163,281,1117,1142]" gridcol="0" gridrow="23" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4420648CCFA803" box="[176,228,1118,1141]" italics="true" pageId="24" pageNumber="25">AEW</emphasis>
i (%)
</th>
<td id="76845DB7F94A62B3FEB820678DD6A800" box="[332,509,1117,1142]" gridcol="1" gridrow="23" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE20678EA5A800" box="[538,654,1117,1142]" gridcol="2" gridrow="23" pageId="24" pageNumber="25">43.0 ± 3.0</td>
<td id="76845DB7F94A62B3FD5E20678F0BA800" box="[682,800,1117,1142]" gridcol="3" gridrow="23" pageId="24" pageNumber="25">(40.247.4)</td>
<td id="76845DB7F94A62B3FCC820678F99A800" box="[828,946,1117,1142]" gridcol="4" gridrow="23" pageId="24" pageNumber="25">48.1 ± 3.1</td>
<td id="76845DB7F94A62B3FC302067887FA800" box="[964,1108,1117,1142]" gridcol="5" gridrow="23" pageId="24" pageNumber="25">(43.851.1)</td>
<td id="76845DB7F94A62B3FB8F20678922A800" box="[1147,1289,1117,1142]" gridcol="6" gridrow="23" pageId="24" pageNumber="25">52.6 ± 5.6</td>
<td id="76845DB7F94A62B3FAEF206789BAA800" box="[1307,1425,1117,1142]" gridcol="7" gridrow="23" pageId="24" pageNumber="25">(45.262.1)</td>
</tr>
<tr id="355534CBF94A62B3FF5720BC89BAA8E8" box="[163,1425,1158,1182]" gridrow="24" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720BC89BAA8E8" box="[163,1425,1158,1182]" colspan="8" colspanRight="7" gridcol="0" gridrow="24" pageId="24" pageNumber="25">Dorsal side</th>
</tr>
<tr id="355534CBF94A62B3FF57209489BAA8B0" box="[163,1425,1198,1222]" gridrow="25" pageId="24" pageNumber="25" rowspan-4="1">
<th id="76845DB7F94A62B3FF5720948D32A8B0" box="[163,281,1198,1222]" gridcol="0" gridrow="25" pageId="24" pageNumber="25">L-Muscle</th>
<td id="76845DB7F94A62B3FEB820948DD6A8B0" box="[332,509,1198,1222]" gridcol="1" gridrow="25" pageId="24" pageNumber="25">
Granules (
<emphasis id="B911EA9BF94A9D54FE3B20958DF7A8B0" box="[463,476,1199,1222]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE20948F0BA8B0" box="[538,800,1198,1222]" colspan="2" colspanRight="1" gridcol="2" gridrow="25" pageId="24" pageNumber="25">9</td>
<td id="76845DB7F94A62B3FC302094887FA8B0" box="[964,1108,1198,1222]" gridcol="5" gridrow="25" pageId="24" pageNumber="25">9</td>
<td id="76845DB7F94A62B3FB8F209489BAA8B0" box="[1147,1425,1198,1222]" colspan="2" colspanRight="1" gridcol="6" gridrow="25" pageId="24" pageNumber="25">12</td>
</tr>
<tr id="355534CBF94A62B3FF5720EC89BAA899" box="[163,1425,1238,1263]" gridrow="26" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720EC8D32A899" box="[163,281,1238,1263]" gridcol="0" gridrow="26" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB820EC8DD6A899" box="[332,509,1238,1263]" gridcol="1" gridrow="26" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE20EC8EA5A899" box="[538,654,1238,1263]" gridcol="2" gridrow="26" pageId="24" pageNumber="25">30.6 ± 2.5</td>
<td id="76845DB7F94A62B3FD5E20EC8F0BA899" box="[682,800,1238,1263]" gridcol="3" gridrow="26" pageId="24" pageNumber="25">(2834)</td>
<td id="76845DB7F94A62B3FCC820EC8F99A899" box="[828,946,1238,1263]" gridcol="4" gridrow="26" pageId="24" pageNumber="25">29.3 ± 2.5</td>
<td id="76845DB7F94A62B3FC3020EC887FA899" box="[964,1108,1238,1263]" gridcol="5" gridrow="26" pageId="24" pageNumber="25">(2533)</td>
<td id="76845DB7F94A62B3FB8F20EC8922A899" box="[1147,1289,1238,1263]" gridcol="6" gridrow="26" pageId="24" pageNumber="25">32.8 ± 2.1</td>
<td id="76845DB7F94A62B3FAEF20EC89BAA899" box="[1307,1425,1238,1263]" gridcol="7" gridrow="26" pageId="24" pageNumber="25">(2936)</td>
</tr>
<tr id="355534CBF94A62B3FF5720C489BAA961" box="[163,1425,1278,1303]" gridrow="27" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5720C48D32A961" box="[163,281,1278,1303]" gridcol="0" gridrow="27" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB820C48DD6A961" box="[332,509,1278,1303]" gridcol="1" gridrow="27" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE20C48EA5A961" box="[538,654,1278,1303]" gridcol="2" gridrow="27" pageId="24" pageNumber="25">13.4 ± 3.0</td>
<td id="76845DB7F94A62B3FD5E20C48F0BA961" box="[682,800,1278,1303]" gridcol="3" gridrow="27" pageId="24" pageNumber="25">(718)</td>
<td id="76845DB7F94A62B3FCC820C48F99A961" box="[828,946,1278,1303]" gridcol="4" gridrow="27" pageId="24" pageNumber="25">15.4 ± 3.5</td>
<td id="76845DB7F94A62B3FC3020C4887FA961" box="[964,1108,1278,1303]" gridcol="5" gridrow="27" pageId="24" pageNumber="25">(1019)</td>
<td id="76845DB7F94A62B3FB8F20C48922A961" box="[1147,1289,1278,1303]" gridcol="6" gridrow="27" pageId="24" pageNumber="25">15.0 ± 1.5</td>
<td id="76845DB7F94A62B3FAEF20C489BAA961" box="[1307,1425,1278,1303]" gridcol="7" gridrow="27" pageId="24" pageNumber="25">(1318)</td>
</tr>
<tr id="355534CBF94A62B3FF57211C89BAA949" box="[163,1425,1318,1343]" gridrow="28" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57211C8D32A949" box="[163,281,1318,1343]" gridcol="0" gridrow="28" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44211D8CF9A948" box="[176,210,1319,1342]" italics="true" pageId="24" pageNumber="25">GP</emphasis>
c
</th>
<td id="76845DB7F94A62B3FEB8211C8DD6A949" box="[332,509,1318,1343]" gridcol="1" gridrow="28" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE211C8EA5A949" box="[538,654,1318,1343]" gridcol="2" gridrow="28" pageId="24" pageNumber="25">44.2 ± 10.9</td>
<td id="76845DB7F94A62B3FD5E211C8F0BA949" box="[682,800,1318,1343]" gridcol="3" gridrow="28" pageId="24" pageNumber="25">(25.062.1)</td>
<td id="76845DB7F94A62B3FCC8211C887FA949" box="[828,1108,1318,1343]" colspan="2" colspanRight="1" gridcol="4" gridrow="28" pageId="24" pageNumber="25">52.9 ± 12.5 (31.266.7)</td>
<td id="76845DB7F94A62B3FB8F211C8922A949" box="[1147,1289,1318,1343]" gridcol="6" gridrow="28" pageId="24" pageNumber="25">45.9 ± 6.1</td>
<td id="76845DB7F94A62B3FAEF211C89BAA949" box="[1307,1425,1318,1343]" gridcol="7" gridrow="28" pageId="24" pageNumber="25">(37.156.3)</td>
</tr>
<tr id="355534CBF94A62B3FF57217589BAA911" box="[163,1425,1359,1383]" gridrow="29" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5721758D32A911" box="[163,281,1359,1383]" gridcol="0" gridrow="29" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44216A8CCEA911" box="[176,229,1360,1383]" italics="true" pageId="24" pageNumber="25">GCO</emphasis>
d (%)
</th>
<td id="76845DB7F94A62B3FEB821758DD6A911" box="[332,509,1359,1383]" gridcol="1" gridrow="29" pageId="24" pageNumber="25">(Ratio)</td>
<td id="76845DB7F94A62B3FDEE21758EA5A911" box="[538,654,1359,1383]" gridcol="2" gridrow="29" pageId="24" pageNumber="25">11.1</td>
<td id="76845DB7F94A62B3FD5E21758F0BA911" box="[682,800,1359,1383]" gridcol="3" gridrow="29" pageId="24" pageNumber="25">(O1: C8)</td>
<td id="76845DB7F94A62B3FCC821758F99A911" box="[828,946,1359,1383]" gridcol="4" gridrow="29" pageId="24" pageNumber="25">77.8</td>
<td id="76845DB7F94A62B3FC302175887FA911" box="[964,1108,1359,1383]" gridcol="5" gridrow="29" pageId="24" pageNumber="25">(O7: C2)</td>
<td id="76845DB7F94A62B3FB8F21758922A911" box="[1147,1289,1359,1383]" gridcol="6" gridrow="29" pageId="24" pageNumber="25">33.3</td>
<td id="76845DB7F94A62B3FAEF217589BAA911" box="[1307,1425,1359,1383]" gridcol="7" gridrow="29" pageId="24" pageNumber="25">(O4: C8)</td>
</tr>
<tr id="355534CBF94A62B3FF57214D89BAA9E6" box="[163,1425,1399,1424]" gridrow="30" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF57214D8D32A9E6" box="[163,281,1399,1424]" gridcol="0" gridrow="30" pageId="24" pageNumber="25">Body wall</th>
<td id="76845DB7F94A62B3FEB8214D8DD6A9E6" box="[332,509,1399,1424]" gridcol="1" gridrow="30" pageId="24" pageNumber="25">
Anchor plates (
<emphasis id="B911EA9BF94A9D54FE1C21428DDEA9F9" box="[488,501,1400,1423]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE214D8F0BA9E6" box="[538,800,1399,1424]" colspan="2" colspanRight="1" gridcol="2" gridrow="30" pageId="24" pageNumber="25">10</td>
<td id="76845DB7F94A62B3FC30214D887FA9E6" box="[964,1108,1399,1424]" gridcol="5" gridrow="30" pageId="24" pageNumber="25">8</td>
<td id="76845DB7F94A62B3FB8F214D8922A9E6" box="[1147,1289,1399,1424]" gridcol="6" gridrow="30" pageId="24" pageNumber="25">6</td>
</tr>
<tr id="355534CBF94A62B3FF57219A89BAA9CE" box="[163,1425,1440,1464]" gridrow="31" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57219A8D32A9CE" box="[163,281,1440,1464]" gridcol="0" gridrow="31" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB8219A8DD6A9CE" box="[332,509,1440,1464]" gridcol="1" gridrow="31" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE219A8EA5A9CE" box="[538,654,1440,1464]" gridcol="2" gridrow="31" pageId="24" pageNumber="25">126.1 ± 9.4</td>
<td id="76845DB7F94A62B3FD5E219A8F0BA9CE" box="[682,800,1440,1464]" gridcol="3" gridrow="31" pageId="24" pageNumber="25">(109135)</td>
<td id="76845DB7F94A62B3FCC8219A887FA9CE" box="[828,1108,1440,1464]" colspan="2" colspanRight="1" gridcol="4" gridrow="31" pageId="24" pageNumber="25">109.6 ± 8.6 (98128)</td>
<td id="76845DB7F94A62B3FB8F219A8922A9CE" box="[1147,1289,1440,1464]" gridcol="6" gridrow="31" pageId="24" pageNumber="25">99.1 ± 7.3</td>
<td id="76845DB7F94A62B3FAEF219A89BAA9CE" box="[1307,1425,1440,1464]" gridcol="7" gridrow="31" pageId="24" pageNumber="25">(91108)</td>
</tr>
<tr id="355534CBF94A62B3FF5721F289BAA996" box="[163,1425,1480,1504]" gridrow="32" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5721F28D32A996" box="[163,281,1480,1504]" gridcol="0" gridrow="32" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB821F28DD6A996" box="[332,509,1480,1504]" gridcol="1" gridrow="32" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE21F28EA5A996" box="[538,654,1480,1504]" gridcol="2" gridrow="32" pageId="24" pageNumber="25">94.1 ± 9.4</td>
<td id="76845DB7F94A62B3FD5E21F28F0BA996" box="[682,800,1480,1504]" gridcol="3" gridrow="32" pageId="24" pageNumber="25">(76106)</td>
<td id="76845DB7F94A62B3FCC821F28F99A996" box="[828,946,1480,1504]" gridcol="4" gridrow="32" pageId="24" pageNumber="25">67.5 ± 7.6</td>
<td id="76845DB7F94A62B3FC3021F2887FA996" box="[964,1108,1480,1504]" gridcol="5" gridrow="32" pageId="24" pageNumber="25">(5580)</td>
<td id="76845DB7F94A62B3FB8F21F28922A996" box="[1147,1289,1480,1504]" gridcol="6" gridrow="32" pageId="24" pageNumber="25">66.8 ±11.5</td>
<td id="76845DB7F94A62B3FAEF21F289BAA996" box="[1307,1425,1480,1504]" gridcol="7" gridrow="32" pageId="24" pageNumber="25">(5379)</td>
</tr>
<tr id="355534CBF94A62B3FF5721D589BAAA7E" box="[163,1425,1519,1544]" gridrow="33" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5721D58D32AA7E" box="[163,281,1519,1544]" gridcol="0" gridrow="33" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4421CB8CF7AA7E" box="[176,220,1521,1544]" italics="true" pageId="24" pageNumber="25">APS</emphasis>
e
</th>
<td id="76845DB7F94A62B3FEB821D58DD6AA7E" box="[332,509,1519,1544]" gridcol="1" gridrow="33" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE21D58EA5AA7E" box="[538,654,1519,1544]" gridcol="2" gridrow="33" pageId="24" pageNumber="25">14.9 ±15.1</td>
<td id="76845DB7F94A62B3FD5E21D58F0BAA7E" box="[682,800,1519,1544]" gridcol="3" gridrow="33" pageId="24" pageNumber="25">(2.743.0)</td>
<td id="76845DB7F94A62B3FCC821D58F99AA7E" box="[828,946,1519,1544]" gridcol="4" gridrow="33" pageId="24" pageNumber="25">8.9 ± 6.6</td>
<td id="76845DB7F94A62B3FC3021D5887FAA7E" box="[964,1108,1519,1544]" gridcol="5" gridrow="33" pageId="24" pageNumber="25">(4.522.4)</td>
<td id="76845DB7F94A62B3FB8F21D58922AA7E" box="[1147,1289,1519,1544]" gridcol="6" gridrow="33" pageId="24" pageNumber="25">4.1 ± 4.4</td>
<td id="76845DB7F94A62B3FAEF21D589BAAA7E" box="[1307,1425,1519,1544]" gridcol="7" gridrow="33" pageId="24" pageNumber="25">(0.612.5)</td>
</tr>
<tr id="355534CBF94A62B3FF57222289BAAA47" box="[163,1425,1560,1585]" gridrow="34" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5722228D32AA47" box="[163,281,1560,1585]" gridcol="0" gridrow="34" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4422238CFEAA46" box="[176,213,1561,1584]" italics="true" pageId="24" pageNumber="25">Ntp</emphasis>
f
</th>
<td id="76845DB7F94A62B3FEB822228DD6AA47" box="[332,509,1560,1585]" gridcol="1" gridrow="34" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE22228EA5AA47" box="[538,654,1560,1585]" gridcol="2" gridrow="34" pageId="24" pageNumber="25">3.5 ± 1.1</td>
<td id="76845DB7F94A62B3FD5E22228F0BAA47" box="[682,800,1560,1585]" gridcol="3" gridrow="34" pageId="24" pageNumber="25">(26)</td>
<td id="76845DB7F94A62B3FCC822228F99AA47" box="[828,946,1560,1585]" gridcol="4" gridrow="34" pageId="24" pageNumber="25">2.9 ± 1.2</td>
<td id="76845DB7F94A62B3FC302222887FAA47" box="[964,1108,1560,1585]" gridcol="5" gridrow="34" pageId="24" pageNumber="25">(15)</td>
<td id="76845DB7F94A62B3FB8F22228922AA47" box="[1147,1289,1560,1585]" gridcol="6" gridrow="34" pageId="24" pageNumber="25">3.2 ± 0.8</td>
<td id="76845DB7F94A62B3FAEF222289BAAA47" box="[1307,1425,1560,1585]" gridcol="7" gridrow="34" pageId="24" pageNumber="25">(24)</td>
</tr>
<tr id="355534CBF94A62B3FF57227A89BAAA2F" box="[163,1425,1600,1625]" gridrow="35" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF57227A8D32AA2F" box="[163,281,1600,1625]" gridcol="0" gridrow="35" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF44227B8CF6AA2E" box="[176,221,1601,1624]" italics="true" pageId="24" pageNumber="25">APT</emphasis>
g (%)
</th>
<td id="76845DB7F94A62B3FEB8227A8DD6AA2F" box="[332,509,1600,1625]" gridcol="1" gridrow="35" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE227A8EA5AA2F" box="[538,654,1600,1625]" gridcol="2" gridrow="35" pageId="24" pageNumber="25">10.5±2.4</td>
<td id="76845DB7F94A62B3FD5E227A8F0BAA2F" box="[682,800,1600,1625]" gridcol="3" gridrow="35" pageId="24" pageNumber="25">(6.914.6)</td>
<td id="76845DB7F94A62B3FCC8227A8F99AA2F" box="[828,946,1600,1625]" gridcol="4" gridrow="35" pageId="24" pageNumber="25">10.4 ± 5.7</td>
<td id="76845DB7F94A62B3FC30227A887FAA2F" box="[964,1108,1600,1625]" gridcol="5" gridrow="35" pageId="24" pageNumber="25">(2.820.8)</td>
<td id="76845DB7F94A62B3FB8F227A8922AA2F" box="[1147,1289,1600,1625]" gridcol="6" gridrow="35" pageId="24" pageNumber="25">17.4 ± 4.1</td>
<td id="76845DB7F94A62B3FAEF227A89BAAA2F" box="[1307,1425,1600,1625]" gridcol="7" gridrow="35" pageId="24" pageNumber="25">(14.325.0)</td>
</tr>
<tr id="355534CBF94A62B3FF57225389BAAAF7" box="[163,1425,1641,1665]" gridrow="36" pageId="24" pageNumber="25" rowspan-4="1" rowspan-7="1">
<th id="76845DB7F94A62B3FF5722538D32AAF7" box="[163,281,1641,1665]" gridcol="0" gridrow="36" pageId="24" pageNumber="25">Body wall</th>
<td id="76845DB7F94A62B3FEB822538DD6AAF7" box="[332,509,1641,1665]" gridcol="1" gridrow="36" pageId="24" pageNumber="25">
Anchor (
<emphasis id="B911EA9BF94A9D54FE3322508DFFAAF7" box="[455,468,1642,1665]" italics="true" pageId="24" pageNumber="25">n</emphasis>
)
</td>
<td id="76845DB7F94A62B3FDEE22538F0BAAF7" box="[538,800,1641,1665]" colspan="2" colspanRight="1" gridcol="2" gridrow="36" pageId="24" pageNumber="25">9</td>
<td id="76845DB7F94A62B3FC302253887FAAF7" box="[964,1108,1641,1665]" gridcol="5" gridrow="36" pageId="24" pageNumber="25">8</td>
<td id="76845DB7F94A62B3FB8F22538922AAF7" box="[1147,1289,1641,1665]" gridcol="6" gridrow="36" pageId="24" pageNumber="25">6</td>
</tr>
<tr id="355534CBF94A62B3FF5722AB89BAAADF" box="[163,1425,1681,1705]" gridrow="37" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5722AB8D32AADF" box="[163,281,1681,1705]" gridcol="0" gridrow="37" pageId="24" pageNumber="25">Length</th>
<td id="76845DB7F94A62B3FEB822AB8DD6AADF" box="[332,509,1681,1705]" gridcol="1" gridrow="37" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE22AB8EA5AADF" box="[538,654,1681,1705]" gridcol="2" gridrow="37" pageId="24" pageNumber="25">165.6 ± 9.3</td>
<td id="76845DB7F94A62B3FD5E22AB8F0BAADF" box="[682,800,1681,1705]" gridcol="3" gridrow="37" pageId="24" pageNumber="25">(151178)</td>
<td id="76845DB7F94A62B3FCC822AB887FAADF" box="[828,1108,1681,1705]" colspan="2" colspanRight="1" gridcol="4" gridrow="37" pageId="24" pageNumber="25">168.5 ± 10.3 (153178)</td>
<td id="76845DB7F94A62B3FB8F22AB8922AADF" box="[1147,1289,1681,1705]" gridcol="6" gridrow="37" pageId="24" pageNumber="25">160.5 ± 14.6</td>
<td id="76845DB7F94A62B3FAEF22AB89BAAADF" box="[1307,1425,1681,1705]" gridcol="7" gridrow="37" pageId="24" pageNumber="25">(144181)</td>
</tr>
<tr id="355534CBF94A62B3FF57228389BAAAA4" box="[163,1425,1721,1746]" gridrow="38" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5722838D32AAA4" box="[163,281,1721,1746]" gridcol="0" gridrow="38" pageId="24" pageNumber="25">Width</th>
<td id="76845DB7F94A62B3FEB822838DD6AAA4" box="[332,509,1721,1746]" gridcol="1" gridrow="38" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE22838EA5AAA4" box="[538,654,1721,1746]" gridcol="2" gridrow="38" pageId="24" pageNumber="25">86.4 ± 6.7</td>
<td id="76845DB7F94A62B3FD5E22838F0BAAA4" box="[682,800,1721,1746]" gridcol="3" gridrow="38" pageId="24" pageNumber="25">(7495)</td>
<td id="76845DB7F94A62B3FCC822838F99AAA4" box="[828,946,1721,1746]" gridcol="4" gridrow="38" pageId="24" pageNumber="25">90.9 ± 6.9</td>
<td id="76845DB7F94A62B3FC302283887FAAA4" box="[964,1108,1721,1746]" gridcol="5" gridrow="38" pageId="24" pageNumber="25">(81103)</td>
<td id="76845DB7F94A62B3FB8F22838922AAA4" box="[1147,1289,1721,1746]" gridcol="6" gridrow="38" pageId="24" pageNumber="25">67.2 ± 11.2</td>
<td id="76845DB7F94A62B3FAEF228389BAAAA4" box="[1307,1425,1721,1746]" gridcol="7" gridrow="38" pageId="24" pageNumber="25">(5587)</td>
</tr>
<tr id="355534CBF94A62B3FF5722D889BAAA8C" box="[163,1425,1762,1786]" gridrow="39" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5722D88D32AA8C" box="[163,281,1762,1786]" gridcol="0" gridrow="39" pageId="24" pageNumber="25">Teeth</th>
<td id="76845DB7F94A62B3FEB822D88DD6AA8C" box="[332,509,1762,1786]" gridcol="1" gridrow="39" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE22D88EA5AA8C" box="[538,654,1762,1786]" gridcol="2" gridrow="39" pageId="24" pageNumber="25">2.6±1.9</td>
<td id="76845DB7F94A62B3FD5E22D88F0BAA8C" box="[682,800,1762,1786]" gridcol="3" gridrow="39" pageId="24" pageNumber="25">(24)</td>
<td id="76845DB7F94A62B3FCC822D88F99AA8C" box="[828,946,1762,1786]" gridcol="4" gridrow="39" pageId="24" pageNumber="25">3.4±0.7</td>
<td id="76845DB7F94A62B3FC3022D8887FAA8C" box="[964,1108,1762,1786]" gridcol="5" gridrow="39" pageId="24" pageNumber="25">(24)</td>
<td id="76845DB7F94A62B3FB8F22D88922AA8C" box="[1147,1289,1762,1786]" gridcol="6" gridrow="39" pageId="24" pageNumber="25">6.2 ± 1.6</td>
<td id="76845DB7F94A62B3FAEF22D889BAAA8C" box="[1307,1425,1762,1786]" gridcol="7" gridrow="39" pageId="24" pageNumber="25">(48)</td>
</tr>
<tr id="355534CBF94A62B3FF57233089BAAB54" box="[163,1425,1802,1826]" gridrow="40" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5723308D32AB54" box="[163,281,1802,1826]" gridcol="0" gridrow="40" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4423308CCAAB57" box="[176,225,1802,1825]" italics="true" pageId="24" pageNumber="25">ASW</emphasis>
h(%)
</th>
<td id="76845DB7F94A62B3FEB823308DD6AB54" box="[332,509,1802,1826]" gridcol="1" gridrow="40" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE23308EA5AB54" box="[538,654,1802,1826]" gridcol="2" gridrow="40" pageId="24" pageNumber="25">22.4 ±1.6</td>
<td id="76845DB7F94A62B3FD5E23308F0BAB54" box="[682,800,1802,1826]" gridcol="3" gridrow="40" pageId="24" pageNumber="25">(19.725.0)</td>
<td id="76845DB7F94A62B3FCC823308F99AB54" box="[828,946,1802,1826]" gridcol="4" gridrow="40" pageId="24" pageNumber="25">20.3 ± 0.8</td>
<td id="76845DB7F94A62B3FC302330887FAB54" box="[964,1108,1802,1826]" gridcol="5" gridrow="40" pageId="24" pageNumber="25">(19.121.3)</td>
<td id="76845DB7F94A62B3FB8F23308922AB54" box="[1147,1289,1802,1826]" gridcol="6" gridrow="40" pageId="24" pageNumber="25">25.8 ± 3.5</td>
<td id="76845DB7F94A62B3FAEF233089BAAB54" box="[1307,1425,1802,1826]" gridcol="7" gridrow="40" pageId="24" pageNumber="25">(20.730.9)</td>
</tr>
<tr id="355534CBF94A62B3FF57230889BAAB3C" box="[163,1425,1842,1866]" gridrow="41" pageId="24" pageNumber="25">
<th id="76845DB7F94A62B3FF5723088D32AB3C" box="[163,281,1842,1866]" gridcol="0" gridrow="41" pageId="24" pageNumber="25">
<emphasis id="B911EA9BF94A9D54FF4423098CCFAB3C" box="[176,228,1843,1866]" italics="true" pageId="24" pageNumber="25">AEW</emphasis>
i (%)
</th>
<td id="76845DB7F94A62B3FEB823088DD6AB3C" box="[332,509,1842,1866]" gridcol="1" gridrow="41" pageId="24" pageNumber="25">(Range)</td>
<td id="76845DB7F94A62B3FDEE23088EA5AB3C" box="[538,654,1842,1866]" gridcol="2" gridrow="41" pageId="24" pageNumber="25">42.6 ± 3.1</td>
<td id="76845DB7F94A62B3FD5E23088F0BAB3C" box="[682,800,1842,1866]" gridcol="3" gridrow="41" pageId="24" pageNumber="25">(38.247.7)</td>
<td id="76845DB7F94A62B3FCC823088F99AB3C" box="[828,946,1842,1866]" gridcol="4" gridrow="41" pageId="24" pageNumber="25">44.9 ± 2.8</td>
<td id="76845DB7F94A62B3FC302308887FAB3C" box="[964,1108,1842,1866]" gridcol="5" gridrow="41" pageId="24" pageNumber="25">(40.848.3)</td>
<td id="76845DB7F94A62B3FB8F23088922AB3C" box="[1147,1289,1842,1866]" gridcol="6" gridrow="41" pageId="24" pageNumber="25">53.2 ± 7.4</td>
<td id="76845DB7F94A62B3FAEF230889BAAB3C" box="[1307,1425,1842,1866]" gridcol="7" gridrow="41" pageId="24" pageNumber="25">(43.763.8)</td>
</tr>
</table>
</paragraph>
<paragraph id="8BDA3689F94A9D54FF63236289A1AB06" blockId="24.[151,1418,1880,1904]" box="[151,1418,1880,1904]" pageId="24" pageNumber="25">
<tableNote id="76833707F94A9D54FF63236289A1AB06" box="[151,1418,1880,1904]" pageId="24" pageNumber="25" targetBox="[163,1425,192,1866]" targetPageId="24">
ai, (see, Materials and Methods) Indicators for ossicles shape, and one parameter (
<emphasis id="B911EA9BF94A9D54FBE623628812AB19" box="[1042,1081,1880,1903]" italics="true" pageId="24" pageNumber="25">Ntp</emphasis>
, number s teethed perforation).
</tableNote>
</paragraph>
<caption id="DF1A6601F94B9D55FF6322998EA7AB51" ID-DOI="http://doi.org/10.5281/zenodo.14703560" ID-Zenodo-Dep="14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="25" pageNumber="26" startId="25.[151,250,1699,1723]" targetBox="[151,1436,181,1667]" targetPageId="25" targetType="figure">
<paragraph id="8BDA3689F94B9D55FF6322998EA7AB51" blockId="25.[151,1437,1699,1831]" pageId="25" pageNumber="26">
<emphasis id="B911EA9BF94B9D55FF6322998D0BAACD" bold="true" box="[151,288,1699,1723]" pageId="25" pageNumber="26">FIGURE 13.</emphasis>
SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as
<taxonomicName id="4C654D0AF94B9D55FF4722F28D5DAAA9" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[179,374,1735,1759]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="25" pageNumber="26" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF94B9D55FF4722F28D5DAAA9" box="[179,374,1735,1759]" italics="true" pageId="25" pageNumber="26">Patinapta deformis</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF94B9D55FE8822FD8DE7AAA9" bold="true" box="[380,460,1735,1759]" pageId="25" pageNumber="26">
<taxonomicNameLabel id="A22257E0F94B9D55FE8822FD8DE7AAA9" box="[380,460,1735,1759]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle.
</paragraph>
</caption>
<caption id="DF1A6601F9489D56FF63263E895FAEDA" ID-DOI="http://doi.org/10.5281/zenodo.14703562" ID-Zenodo-Dep="14703562" httpUri="https://zenodo.org/record/14703562/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,250,516,540]" targetBox="[164,1423,188,485]" targetPageId="26" targetType="figure">
<paragraph id="8BDA3689F9489D56FF63263E895FAEDA" blockId="26.[151,1437,516,685]" pageId="26" pageNumber="27">
<emphasis id="B911EA9BF9489D56FF63263E8D35AE6B" bold="true" box="[151,286,516,541]" pageId="26" pageNumber="27">FIGURE 14.</emphasis>
Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima (holotype specimen, WMNH-2015-INV-106), described as
<taxonomicName id="4C654D0AF9489D56FD7A26138F78AE37" authorityName="Yamana &amp; Hirashima &amp; Sato &amp; Yamamori" authorityYear="2025" box="[654,851,553,577]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="26" pageNumber="27" phylum="Echinodermata" rank="species" species="deformis" status="sp. nov.">
<emphasis id="B911EA9BF9489D56FD7A26138F78AE37" box="[654,851,553,577]" italics="true" pageId="26" pageNumber="27">Patinapta deformis</emphasis>
</taxonomicName>
<emphasis id="B911EA9BF9489D56FCAE26138F80AE37" bold="true" box="[858,939,553,577]" pageId="26" pageNumber="27">
<taxonomicNameLabel id="A22257E0F9489D56FCAE26138F80AE37" box="[858,939,553,577]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, in the present study. The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system:
<emphasis id="B911EA9BF9489D56FCF626778F6FAE12" box="[770,836,589,612]" italics="true" pageId="26" pageNumber="27">RIRV</emphasis>
and
<emphasis id="B911EA9BF9489D56FC8226778FE7AE12" box="[886,972,589,613]" italics="true" pageId="26" pageNumber="27">IR1IR5</emphasis>
where
<emphasis id="B911EA9BF9489D56FBE32677880DAE12" box="[1047,1062,589,612]" italics="true" pageId="26" pageNumber="27">R</emphasis>
and
<emphasis id="B911EA9BF9489D56FBAD2677885AAE12" box="[1113,1137,589,612]" italics="true" pageId="26" pageNumber="27">IR</emphasis>
stand for positions for radius and interradius, respectively. Hence
<emphasis id="B911EA9BF9489D56FDFD264B8E0AAEFE" box="[521,545,625,648]" italics="true" pageId="26" pageNumber="27">RI</emphasis>
, mid-ventral;
<emphasis id="B911EA9BF9489D56FD45264B8EFAAEFE" box="[689,721,625,648]" italics="true" pageId="26" pageNumber="27">RII</emphasis>
, left ventral;
<emphasis id="B911EA9BF9489D56FCAF264B8FAFAEFE" box="[859,900,625,648]" italics="true" pageId="26" pageNumber="27">RIII</emphasis>
, right ventral;
<emphasis id="B911EA9BF9489D56FBEF264B8869AEFE" box="[1051,1090,625,648]" italics="true" pageId="26" pageNumber="27">RIV</emphasis>
, left dorsal;
<emphasis id="B911EA9BF9489D56FB37264B88CAAEFE" box="[1219,1249,625,648]" italics="true" pageId="26" pageNumber="27">RV</emphasis>
, right dorsal; and:
<emphasis id="B911EA9BF9489D56FF6326AF8C90AEDB" box="[151,187,661,685]" italics="true" pageId="26" pageNumber="27">IR1</emphasis>
, left ventral;
<emphasis id="B911EA9BF9489D56FEB026AF8D42AEDB" box="[324,361,661,685]" italics="true" pageId="26" pageNumber="27">IR2</emphasis>
, right ventral;
<emphasis id="B911EA9BF9489D56FE0B26AF8E72AEDB" box="[511,601,661,685]" italics="true" pageId="26" pageNumber="27">IR3-IR3</emphasis>
, left dorso-lateral;
<emphasis id="B911EA9BF9489D56FCE826AF8F5DAEDB" box="[796,886,661,685]" italics="true" pageId="26" pageNumber="27">IR4-IR4</emphasis>
, right dorso-lateral;
<emphasis id="B911EA9BF9489D56FBB226AF8841AEDA" box="[1094,1130,661,684]" italics="true" pageId="26" pageNumber="27">IR5</emphasis>
, mid-dorsal. Anterior top.
</paragraph>
</caption>
<paragraph id="8BDA3689F9489D56FF3326C58F4BAFBB" blockId="26.[151,1437,767,1190]" pageId="26" pageNumber="27">
Anchor ossicles ranged within 144178 μm in length (
<figureCitation id="135E2A0CF9489D56FCED26C58F88AF6F" box="[793,931,767,793]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13 B, D</figureCitation>
;
<tableCitation id="C6E70332F9489D56FC5826C58FD4AF6F" box="[940,1023,767,793]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="26" pageNumber="27">Table 6</tableCitation>
). All the anchor arms (bills) equipped with minute teeth on their outer tips, numbers of teeth variable within 27 (mostly
<quantity id="4C9D9B6CF9489D56FBBB271988BCAF4B" box="[1103,1175,803,829]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="7.62" metricValueMin="2.54" pageId="26" pageNumber="27" unit="in" value="2.0" valueMax="3.0" valueMin="1.0">13 in</quantity>
one arm of the anchor ossicle). Anchor ossicles of this specimen had broad stem and relatively wide anchor arms, with
<emphasis id="B911EA9BF9489D56FB33277D88D6AF17" box="[1223,1277,839,865]" italics="true" pageId="26" pageNumber="27">ASW</emphasis>
=19.121.3%;
<emphasis id="B911EA9BF9489D56FF6327518CFAAFF3" box="[151,209,875,901]" italics="true" pageId="26" pageNumber="27">AEW</emphasis>
=40.851.1%, and T-shaped basal end of the anchor ossicle of this species strongly widened and semicircular in shape (
<figureCitation id="135E2A0CF9489D56FEF527B58DA3AFDF" box="[257,392,911,937]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13B, D</figureCitation>
). Such features were have not been reported in both previously described congeneric Japanese species (both species possess crescent-like T-shaped basal end).
</paragraph>
<paragraph id="8BDA3689F9489D56FF3327ED88A9A82B" blockId="26.[151,1437,767,1190]" pageId="26" pageNumber="27">
Body longitudinal muscle, granule ossicles of this specimen ranged within 2336 μm in length (
<figureCitation id="135E2A0CF9489D56FAFB27ED89BCAF87" box="[1295,1431,983,1009]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13C, E</figureCitation>
;
<tableCitation id="C6E70332F9489D56FF6327C18CC0A863" box="[151,235,1019,1045]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="26" pageNumber="27">Table 6</tableCitation>
). Calculated indicator for granule proportions was variated within
<emphasis id="B911EA9BF9489D56FC2527C18FDDA863" box="[977,1014,1019,1045]" italics="true" pageId="26" pageNumber="27">GP</emphasis>
=31.282.6%;
<emphasis id="B911EA9BF9489D56FB6F27C188FDA863" box="[1179,1238,1019,1045]" italics="true" pageId="26" pageNumber="27">GCO</emphasis>
=44.477.8% and were not noticeably unbalanced between O-shaped or C-shaped granules, however some granule ossicles had one or two minute perforations, resulting in turtle-face-like characteristic shape (
<figureCitation id="135E2A0CF9489D56FC1B2079885EA82B" box="[1007,1141,1091,1117]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13C, E</figureCitation>
).
</paragraph>
<paragraph id="8BDA3689F9489D56FF33205D8815A8D0" blockId="26.[151,1437,767,1190]" pageId="26" pageNumber="27">
Tentacle ossicles rods, ranged within 5285 μm in length (
<figureCitation id="135E2A0CF9489D56FCA5205D8F9CA8F7" box="[849,951,1127,1153]" captionStart="FIGURE 13" captionStartId="25.[151,250,1699,1723]" captionTargetBox="[151,1436,181,1667]" captionTargetId="figure-20@25.[151,1436,181,1675]" captionTargetPageId="25" captionText="FIGURE 13. SEM images of the ossicles extracted from a specimen collected from Yoron-jima Island, Kagoshima, described as Patinapta deformis sp. nov., in the present study (holotype specimen, WMNH-2015-INV-106): A, ossicles from tentacle; B, ossicles from mid-dorsal body wall; C, ossicles from mid-dorsal longitudinal muscle; D, ossicles from mid-ventral body wall; E, ossicles from mid-ventral longitudinal muscle." figureDoi="http://doi.org/10.5281/zenodo.14703560" httpUri="https://zenodo.org/record/14703560/files/figure.png" pageId="26" pageNumber="27">Fig. 13A</figureCitation>
;
<tableCitation id="C6E70332F9489D56FC37205D883CA8F7" box="[963,1047,1127,1153]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="26" pageNumber="27">Table 6</tableCitation>
). Two calculated indicators for rod proportions and complexities were variated within
<emphasis id="B911EA9BF9489D56FD2620B68EDFA8D3" box="[722,756,1164,1189]" italics="true" pageId="26" pageNumber="27">RP</emphasis>
=10.720.3%;
<emphasis id="B911EA9BF9489D56FC5520B18FEEA8D3" box="[929,965,1163,1189]" italics="true" pageId="26" pageNumber="27">RC</emphasis>
=5.112.2.
</paragraph>
<paragraph id="8BDA3689F9489D56FF6320CD8F20A967" blockId="26.[151,779,1271,1297]" box="[151,779,1271,1297]" pageId="26" pageNumber="27">
<heading id="D09281E5F9489D56FF6320CD8F20A967" bold="true" box="[151,779,1271,1297]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B911EA9BF9489D56FF6320CD8F20A967" bold="true" box="[151,779,1271,1297]" pageId="26" pageNumber="27">Observation on the morphology of calcareous ring</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9489D56FF6321058824A9B3" blockId="26.[151,1436,1343,1477]" pageId="26" pageNumber="27">
Calcareous plates 12, firmly bound (
<figureCitation id="135E2A0CF9489D56FDC921058EB9A92F" box="[573,658,1343,1369]" captionStart="FIGURE 14" captionStartId="26.[151,250,516,540]" captionTargetBox="[164,1423,188,485]" captionTargetId="figure-29@26.[151,1435,181,493]" captionTargetPageId="26" captionText="FIGURE 14. Outer-surface view of calcareous ring plates of a specimen collected from Yoron-jima Island, Kagoshima (holotype specimen, WMNH-2015-INV-106), described as Patinapta deformis sp. nov., in the present study. The positions of the plates are denoted by Ludwigs [18891892 (1889), p156] system: RIRV and IR1IR5 where R and IR stand for positions for radius and interradius, respectively. Hence RI, mid-ventral; RII, left ventral; RIII, right ventral; RIV, left dorsal; RV, right dorsal; and: IR1, left ventral; IR2, right ventral; IR3-IR3, left dorso-lateral; IR4-IR4, right dorso-lateral; IR5, mid-dorsal. Anterior top." figureDoi="http://doi.org/10.5281/zenodo.14703562" httpUri="https://zenodo.org/record/14703562/files/figure.png" pageId="26" pageNumber="27">Fig. 14</figureCitation>
), with slight anterior projection and slight posterior depression. All radial plates perforated in each anterior half. Two ventro-lateral plates (
<emphasis id="B911EA9BF9489D56FC36215E880EA90B" box="[962,1061,1380,1405]" italics="true" pageId="26" pageNumber="27">RII; RIII</emphasis>
) protruding strongly, and exhibit pentagonal shape, resulting in both suddenly turning sharply on these ventro-lateral plates. There is no space between the additional interradial plate (
<emphasis id="B911EA9BF9489D56FE0C21918E47A9B3" box="[504,620,1451,1477]" italics="true" pageId="26" pageNumber="27">IR4; IR3</emphasis>
) and adjacent radial plate (
<emphasis id="B911EA9BF9489D56FC6921968FD4A9B3" box="[925,1023,1452,1477]" italics="true" pageId="26" pageNumber="27">RIII; RII</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9489D57FF63222D8878AFBF" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="description">
<paragraph id="8BDA3689F9489D56FF63222D8EC6AA47" blockId="26.[151,749,1559,1585]" box="[151,749,1559,1585]" pageId="26" pageNumber="27">
<heading id="D09281E5F9489D56FF63222D8EC6AA47" bold="true" box="[151,749,1559,1585]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B911EA9BF9489D56FF63222D8EC6AA47" bold="true" box="[151,749,1559,1585]" pageId="26" pageNumber="27">Variation of external and internal morphologies</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9489D56FF6322658E37AAB7" blockId="26.[151,1436,1631,1729]" pageId="26" pageNumber="27">
<specimenCount id="9D63FD00F9489D56FF6322658D7DAA0F" box="[151,342,1631,1657]" count="3" pageId="26" pageNumber="27" type="generic">Three specimens</specimenCount>
were all large-size specimens of about
<quantity id="4C9D9B6CF9489D56FCE822658F8EAA0F" box="[796,933,1631,1657]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="26" pageNumber="27" unit="mm" value="105.0" valueMax="120.0" valueMin="90.0">90120 mm</quantity>
(
<tableCitation id="C6E70332F9489D56FC4122658821AA0F" box="[949,1034,1631,1657]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="26" pageNumber="27">Table 1</tableCitation>
), all
<specimenCount id="9D63FD00F9489D56FBBD226588D4AA0F" box="[1097,1279,1631,1657]" count="3" pageId="26" pageNumber="27" type="generic">three specimens</specimenCount>
well-matured possessing fully equipped gonad tubules in their body cavities.Their external and internal morphologies approximately agreed with each others (
<tableCitation id="C6E70332F9489D56FE43229D8E20AAB7" box="[439,523,1703,1729]" captionStart="TABLE 1" captionStartId="6.[152,244,152,176]" captionTargetPageId="6" captionText="TABLE 1. Measurements (mm) of body size and internal organs and their positions from specimens examined." pageId="26" pageNumber="27">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8BDA3689F9489D56FF6323298E55AB5B" blockId="26.[151,638,1811,1837]" box="[151,638,1811,1837]" pageId="26" pageNumber="27">
<heading id="D09281E5F9489D56FF6323298E55AB5B" bold="true" box="[151,638,1811,1837]" fontSize="11" level="1" pageId="26" pageNumber="27" reason="1">
<emphasis id="B911EA9BF9489D56FF6323298E55AB5B" bold="true" box="[151,638,1811,1837]" pageId="26" pageNumber="27">
Ossicle variation among the
<typeStatus id="54DE882BF9489D56FE0E23298E1BAB5B" box="[506,560,1811,1837]" pageId="26" pageNumber="27">type</typeStatus>
series
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3689F9489D57FF6323618E3AAD6B" blockId="26.[151,1436,1883,2018]" lastBlockId="27.[151,1437,151,969]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Body wall ossicles anchors and anchor plates. Anchor plate ossicle length and calculated indicators of
<specimenCount id="9D63FD00F9489D56FA9023618D27ABEF" count="3" pageId="26" pageNumber="27" type="generic">three specimens</specimenCount>
ranged within: 83138 μm in length,
<emphasis id="B911EA9BF9489D56FD3523458EDAABEF" box="[705,753,1919,1945]" italics="true" pageId="26" pageNumber="27">APS</emphasis>
=0.643.0,
<emphasis id="B911EA9BF9489D56FC8723BA8FB0ABEF" box="[883,923,1920,1945]" italics="true" pageId="26" pageNumber="27">Ntp</emphasis>
=16,
<emphasis id="B911EA9BF9489D56FC112345883CABEF" box="[997,1047,1919,1945]" italics="true" pageId="26" pageNumber="27">APT</emphasis>
=2.325.0%. Among these values, length, frequency of teethed perforations
<emphasis id="B911EA9BF9489D56FD9D23998EB0ABCB" box="[617,667,1955,1981]" italics="true" pageId="26" pageNumber="27">APT</emphasis>
and their parameter
<emphasis id="B911EA9BF9489D56FC70239E8F87ABCB" box="[900,940,1956,1981]" italics="true" pageId="26" pageNumber="27">Ntp</emphasis>
, and non-size dependance indicator of plate skewness
<emphasis id="B911EA9BF9489D56FEF823FD8D17AB97" box="[268,316,1991,2017]" italics="true" pageId="26" pageNumber="27">APS</emphasis>
of anchor plate ossicles (both ventrally and dorsally) was not significantly different (
<emphasis id="B911EA9BF9489D56FAD923F28962AB97" box="[1325,1353,1992,2017]" italics="true" pageId="26" pageNumber="27">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test). From these results, the anchor plate ossicles in the body wall of the present
<specimenCount id="9D63FD00F9499D57FB1C24AD89B6ACC7" box="[1256,1437,151,177]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
were not different in ossicle length or the skewness of the layout for perforations. Therefore, the length (about 80140 μm), numbers of teethed perforations (16), and strongly distorted (up to 43) anchor plate ossicles can be appropriate diagnostic characters.
</paragraph>
<paragraph id="8BDA3689F9499D57FF33251D8820AD83" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
Anchor ossicle length and calculated indicators ranged within: 144181 μm in length,
<emphasis id="B911EA9BF9499D57FB3C251D88D5AD37" box="[1224,1278,295,321]" italics="true" pageId="27" pageNumber="28">ASW</emphasis>
=19.130.9%,
<emphasis id="B911EA9BF9499D57FF6325718CFAAD13" box="[151,209,331,357]" italics="true" pageId="27" pageNumber="28">AEW</emphasis>
=38.263.8% (
<tableCitation id="C6E70332F9499D57FE8D25718DF9AD13" box="[377,466,331,357]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="27" pageNumber="28">Table 6</tableCitation>
). Among these values, lengths of anchor ossicles (both ventrally and dorsally) were not significantly different (
<emphasis id="B911EA9BF9499D57FE3025558DCBADFE" box="[452,480,367,392]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test), while the indicator for anchor stem breadth
<emphasis id="B911EA9BF9499D57FAE42555896DADFF" box="[1296,1350,367,393]" italics="true" pageId="27" pageNumber="28">ASW</emphasis>
and for basal / distal ends width
<emphasis id="B911EA9BF9499D57FE5C25A98DC9ADDB" box="[424,482,403,429]" italics="true" pageId="27" pageNumber="28">AEW</emphasis>
were significantly different (
<emphasis id="B911EA9BF9499D57FCD325A98F68ADDA" box="[807,835,403,428]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&lt;0.005, Kruskal-Wallis test). From these results, the anchor ossicles in the body wall of the present
<specimenCount id="9D63FD00F9499D57FD51258D8F71ADA7" box="[677,858,439,465]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
were not different in ossicle length, but different in the shape. Therefore, proportion of the anchor ossicles can be the accurate key.
</paragraph>
<paragraph id="8BDA3689F9499D57FF3325C58D67AE87" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
Longitudinal muscle involved both O-shaped and C-shaped granule ossicles, length and calculated indicators ranged within: 2336 μm length,
<emphasis id="B911EA9BF9499D57FDDD26198E65AE4B" box="[553,590,547,573]" italics="true" pageId="27" pageNumber="28">GP</emphasis>
=31.282.6%,
<emphasis id="B911EA9BF9499D57FD0E26198F1EAE4B" box="[762,821,547,573]" italics="true" pageId="27" pageNumber="28">GCO</emphasis>
=11.177.8% (
<tableCitation id="C6E70332F9499D57FC2B26198810AE4B" box="[991,1083,547,573]" captionStart="TABLE 6" captionStartId="24.[152,244,151,175]" captionTargetPageId="24" captionText="TABLE 6. Measurements (mean±SD μm) and counts of ossicles of three specimens of Patinapta deformis sp. nov." pageId="27" pageNumber="28">Table 6</tableCitation>
). Among these values, length of granule ossicles was not different significantly (
<emphasis id="B911EA9BF9499D57FD12267D8F29AE16" box="[742,770,583,608]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test), also calculated indicator for proportions
<emphasis id="B911EA9BF9499D57FEEB26518D6FAEF3" box="[287,324,619,645]" italics="true" pageId="27" pageNumber="28">GP</emphasis>
was not different significantly in
<emphasis id="B911EA9BF9499D57FD4F26518ECBAEF3" box="[699,736,619,645]" italics="true" pageId="27" pageNumber="28">GP</emphasis>
(
<emphasis id="B911EA9BF9499D57FD0426518F27AEF2" box="[752,780,619,644]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test, both in ventrally and dorsally), but different significantly in
<emphasis id="B911EA9BF9499D57FE2A26B58E32AEDF" box="[478,537,655,681]" italics="true" pageId="27" pageNumber="28">GCO</emphasis>
(
<emphasis id="B911EA9BF9499D57FDDE26B58E6DAEDE" box="[554,582,655,680]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&lt;0.05, χ
<superScript id="7C109BC1F9499D57FD5726B48E80AEEA" attach="right" box="[675,683,654,668]" fontSize="6" pageId="27" pageNumber="28">2</superScript>
-multiple test). Therefore, the length and proportion of the granule ossicles can be the appropriate keys, while the ratio C- or O-shaped granule ossicles of this species cannot be the appropriate key.
</paragraph>
<paragraph id="8BDA3689F9499D57FF3326C18878AFBF" blockId="27.[151,1437,151,969]" pageId="27" pageNumber="28">
Tentacle ossicles rods, length and calculated indicators of
<specimenCount id="9D63FD00F9499D57FCA026C18823AF63" box="[852,1032,763,789]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
ranged within: 3585 μm in length,
<emphasis id="B911EA9BF9499D57FF6327258C92AF4E" box="[151,185,799,824]" italics="true" pageId="27" pageNumber="28">RP</emphasis>
=10.126.3%, and
<emphasis id="B911EA9BF9499D57FE6227258D91AF4F" box="[406,442,799,825]" italics="true" pageId="27" pageNumber="28">RC</emphasis>
=5.017.3. Among these values, length of rod ossicles was not significantly different (
<emphasis id="B911EA9BF9499D57FF5527798C99AF2A" box="[161,178,835,860]" italics="true" pageId="27" pageNumber="28">P</emphasis>
&gt;0.05, Kruskal-Wallis test), also non-size dependence indicator for proportion (
<emphasis id="B911EA9BF9499D57FBBD27798840AF2A" box="[1097,1131,835,860]" italics="true" pageId="27" pageNumber="28">RP</emphasis>
) and non-size dependence complexities (
<emphasis id="B911EA9BF9499D57FEC0275D8D73AFF7" box="[308,344,871,897]" italics="true" pageId="27" pageNumber="28">RC</emphasis>
) were not significantly different (
<emphasis id="B911EA9BF9499D57FD3C275D8ECFAFF6" box="[712,740,871,896]" italics="true" pageId="27" pageNumber="28">Ps</emphasis>
&gt;0.05, Kruskal-Wallis test). From these results, the rod ossicles in the tentacles of the present
<specimenCount id="9D63FD00F9499D57FE0427B18E8DAFD3" box="[496,678,907,933]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
have similar lengths, proportions and complexities. Therefore, the morphologies of the rod ossicles of this species are appropriate diagnostic characters.
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9499D57FF6320218F8CA9EB" pageId="27" pageNumber="28" type="discussion">
<paragraph id="8BDA3689F9499D57FF6320218D22A843" blockId="27.[151,265,1051,1077]" box="[151,265,1051,1077]" pageId="27" pageNumber="28">
<emphasis id="B911EA9BF9499D57FF6320218D22A843" bold="true" box="[151,265,1051,1077]" pageId="27" pageNumber="28">Remarks</emphasis>
</paragraph>
<paragraph id="8BDA3689F9499D57FF6320598F8CA9EB" blockId="27.[151,1437,1123,1437]" pageId="27" pageNumber="28">
Prior specimens of this species have been misidentified as
<taxonomicName id="4C654D0AF9499D57FCBC20598FD8A80A" authorityName="Chao, Rowe &amp; Chang" authorityYear="1988" box="[840,1011,1123,1148]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="taiwaniensis">
<emphasis id="B911EA9BF9499D57FCBC20598FD8A80A" box="[840,1011,1123,1148]" italics="true" pageId="27" pageNumber="28">P. taiwaniensis</emphasis>
</taxonomicName>
, based on the distorted anchor plate ossicles. However, observations herein showed that all
<specimenCount id="9D63FD00F9499D57FD0720BD8F8EA8D7" box="[755,933,1159,1185]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
possessed ciliated funnels in their right dorsal interradius (
<emphasis id="B911EA9BF9499D57FED420918D62A8B3" box="[288,329,1195,1221]" italics="true" pageId="27" pageNumber="28">IR4</emphasis>
), while
<taxonomicName id="4C654D0AF9499D57FE5320918E64A8B2" authorityName="Chao, Rowe &amp; Chang" authorityYear="1988" box="[423,591,1195,1220]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="taiwaniensis">
<emphasis id="B911EA9BF9499D57FE5320918E64A8B2" box="[423,591,1195,1220]" italics="true" pageId="27" pageNumber="28">P. taiwaniensis</emphasis>
</taxonomicName>
possesses ciliated funnels in left dorsal interradius (
<emphasis id="B911EA9BF9499D57FB54209188E3A8B3" box="[1184,1224,1195,1221]" italics="true" pageId="27" pageNumber="28">IR3</emphasis>
). Such differences are a critical morphological character (Table 7-1). Presently, genus
<taxonomicName id="4C654D0AF9499D57FC5E20F588EBA89F" authority="Heding, 1928" authorityName="Heding" authorityYear="1928" box="[938,1216,1231,1257]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="genus">
<emphasis id="B911EA9BF9499D57FC5E20F58833A89E" box="[938,1048,1231,1256]" italics="true" pageId="27" pageNumber="28">Patinapta</emphasis>
<bibRefCitation id="EFF44B78F9499D57FBD720F588EBA89F" author="Heding, S. G." box="[1059,1216,1231,1257]" pageId="27" pageNumber="28" pagination="105 - 323" refId="ref31634" refString="Heding, S. G. (1928) Synaptidae. Papers from Dr. Th. Mortensen's Pacific Expedition 1914 - 1916. No. 66. Videnskabelige Meddeleser fra Dansk naturhistorisk Forening i KObenhavn, 85, 105 - 323." type="journal article" year="1928">Heding, 1928</bibRefCitation>
</taxonomicName>
accommodates six species from Indo-West Pacific waters (WoRMS 2024, Table 7-1; 7-2). Four other species had been described in this genus before:
<taxonomicName id="4C654D0AF9499D57FEC3212D8E59A947" authority="Heding, 1929" authorityName="Heding" authorityYear="1929" box="[311,626,1303,1329]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="crosslandii">
<emphasis id="B911EA9BF9499D57FEC3212D8DE5A947" box="[311,462,1303,1329]" italics="true" pageId="27" pageNumber="28">P. crosslandii</emphasis>
<bibRefCitation id="EFF44B78F9499D57FE22212D8E59A947" author="Heding, S. G." box="[470,626,1303,1329]" pageId="27" pageNumber="28" pagination="139 - 154" refId="ref31677" refString="Heding, S. G. (1929 [dated 1930]) Contributions to the Kowledge of the Synaputidae. I. Videnskabelige Meddeleser fra Dansk naturhistorisk Forening i KObenhavn, 88, 139 - 154." type="journal article" year="1929">Heding, 1929</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C654D0AF9499D57FD8B212D8FE3A947" authority="Cherbonnier, 1954" authorityName="Cherbonnier" authorityYear="1954" box="[639,968,1302,1329]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="dumasi">
<emphasis id="B911EA9BF9499D57FD8B212D8EC7A947" box="[639,748,1303,1329]" italics="true" pageId="27" pageNumber="28">P. dumasi</emphasis>
<bibRefCitation id="EFF44B78F9499D57FD07212D8FE3A947" author="Cherbonnier, G." box="[755,968,1302,1329]" pageId="27" pageNumber="28" pagination="252 - 260" refId="ref31597" refString="Cherbonnier, G. (1954) Note preliminarie sur les holothuries de la Mer Rouge. Bulletin Museum National Histoire Naturelle, Paris, Serie 2, 26 (2), 252 - 260." type="journal article" year="1954">Cherbonnier, 1954</bibRefCitation>
</taxonomicName>
;
<taxonomicName id="4C654D0AF9499D57FC20212D88DAA947" authority="(Bedford, 1899)" baseAuthorityName="Bedford" baseAuthorityYear="1899" box="[980,1265,1303,1329]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="laevis">
<emphasis id="B911EA9BF9499D57FC20212D881AA947" box="[980,1073,1303,1329]" italics="true" pageId="27" pageNumber="28">P. laevis</emphasis>
(
<bibRefCitation id="EFF44B78F9499D57FBB5212D88C2A947" author="Bedford, F. P." box="[1089,1257,1303,1329]" pageId="27" pageNumber="28" pagination="141 - 150" refId="ref31264" refString="Bedford, F. P. (1899) Holothurians. In: Willey, A. (Ed.), Zoological Results based on material from New Britain, New Guinea, Loyalty Islands and elsewhere, collected during the years 1895, 1896 and 1897. Part II. University Press, Cambridge, pp. 141 - 150." type="book chapter" year="1899">Bedford, 1899</bibRefCitation>
)
</taxonomicName>
; and
<taxonomicName id="4C654D0AF9499D57FADA212D8DF1A923" authority="Cherbonnier, 1953" authorityName="Cherbonnier" authorityYear="1953" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="vaughani">
<emphasis id="B911EA9BF9499D57FADA212D8CD4A923" italics="true" pageId="27" pageNumber="28">Patinapta vaughani</emphasis>
<bibRefCitation id="EFF44B78F9499D57FEF221018DF1A923" author="Cherbonnier, G." box="[262,474,1338,1365]" pageId="27" pageNumber="28" pagination="501 - 504" refId="ref31556" refString="Cherbonnier, G. (1953) Note sur une nouvelle espece de Synapte de l'Ie Maurice: Patinapta vaughani n. sp. Bulletin Museum National Histoire Naturelle, Paris, Serie 2, 25, 501 - 504." type="journal article" year="1953">Cherbonnier, 1953</bibRefCitation>
</taxonomicName>
. Among these species, only
<taxonomicName id="4C654D0AF9499D57FCE221018FB2A923" authorityName="Cherbonnier" authorityYear="1953" box="[790,921,1339,1365]" class="Holothuroidea" family="Synaptidae" genus="Patinapta" kingdom="Animalia" order="Apodida" pageId="27" pageNumber="28" phylum="Echinodermata" rank="species" species="vaughani">
<emphasis id="B911EA9BF9499D57FCE221018FB2A923" box="[790,921,1339,1365]" italics="true" pageId="27" pageNumber="28">P. vaughani</emphasis>
</taxonomicName>
was reported as possessing ciliated funnels in the right dorsal interradius (
<emphasis id="B911EA9BF9499D57FE2021658DD6A90F" box="[468,509,1375,1401]" italics="true" pageId="27" pageNumber="28">IR4</emphasis>
). However, each species is distinguishable by the differences of the shapes of the longitudinal muscle granules and body wall anchor plates (Table 7-2).
</paragraph>
</subSubSection>
<subSubSection id="C37F6502F9499D57FF6321D48936AB73" pageId="27" pageNumber="28" type="distribution">
<paragraph id="8BDA3689F9499D57FF6321D48D1BAA7E" blockId="27.[151,304,1518,1544]" box="[151,304,1518,1544]" pageId="27" pageNumber="28">
<emphasis id="B911EA9BF9499D57FF6321D48D1BAA7E" bold="true" box="[151,304,1518,1544]" pageId="27" pageNumber="28">Distribution</emphasis>
</paragraph>
<paragraph id="8BDA3689F9499D57FF63220D8936AB73" blockId="27.[151,1437,1591,1797]" pageId="27" pageNumber="28">
Southern
<collectingCountry id="F3727619F9499D57FEF3220D8D6CAA27" box="[263,327,1591,1617]" name="Japan" pageId="27" pageNumber="28">Japan</collectingCountry>
(Kyushu Island and the Ryukyu Islands). The present
<specimenCount id="9D63FD00F9499D57FC32220D8856AA27" box="[966,1149,1591,1617]" count="3" pageId="27" pageNumber="28" type="generic">three specimens</specimenCount>
collected from intertidal sandy beach with boulders originated from dead corals, in the
<typeStatus id="54DE882BF9499D57FC9F22618FB0AA03" box="[875,923,1627,1653]" pageId="27" pageNumber="28">type</typeStatus>
locality, Yoron-jima Island, Ryukyu Islands,
<collectingRegion id="49A1F86BF9499D57FF6322458D32AAEF" box="[151,281,1663,1689]" country="Japan" name="Kagoshima" pageId="27" pageNumber="28">Kagoshima</collectingRegion>
,
<collectingCountry id="F3727619F9499D57FEDE22458D46AAEF" box="[298,365,1663,1689]" name="Japan" pageId="27" pageNumber="28">Japan</collectingCountry>
. Among the specimens housed in WMNH-INV, including the present specimens, 4 bottles including
<specimenCount id="9D63FD00F9499D57FEFC22998DB9AACB" box="[264,402,1699,1725]" count="9" pageId="27" pageNumber="28" type="generic">9 specimens</specimenCount>
were identified as this species (
<tableCitation id="C6E70332F9499D57FD0222998F67AACB" box="[758,844,1699,1725]" captionStart="TABLE 8" captionStartId="31.[152,244,152,176]" captionTargetPageId="31" captionText="TABLE 8. List of WMNH-INV specimens of Patinapta deformis sp. nov. as of 2024." pageId="27" pageNumber="28">Table 8</tableCitation>
). Other than the
<typeStatus id="54DE882BF9499D57FBFD22998812AACB" box="[1033,1081,1699,1725]" pageId="27" pageNumber="28">type</typeStatus>
locality,
<specimenCount id="9D63FD00F9499D57FB542299890BAACB" box="[1184,1312,1699,1725]" count="6" pageId="27" pageNumber="28" type="immature">6 immature</specimenCount>
specimens collected from Yokaku bay and Hirose-gawa river mouth Amakusa City,
<collectingRegion id="49A1F86BF9499D57FC3B22FD887BAA97" box="[975,1104,1735,1761]" country="Japan" name="Kumamoto" pageId="27" pageNumber="28">Kumamoto</collectingRegion>
,
<collectingCountry id="F3727619F9499D57FBA822FD88B7AA97" box="[1116,1180,1735,1761]" name="Japan" pageId="27" pageNumber="28">Japan</collectingCountry>
were identified as this species. In both localities in Amakusa City, the specimens were collected from estuarine intertidal flats.
</paragraph>
</subSubSection>
</treatment>
</document>