594 lines
73 KiB
XML
594 lines
73 KiB
XML
<document id="24945561C69A1BB589EC5FFCD856FA8D" ID-CLB-Dataset="46699" ID-DOI="10.5281/zenodo.277812" ID-GBIF-Dataset="96327166-d6b8-47d3-893f-517c372c5bcd" ID-ISSN="1175-5326" ID-Zenodo-Dep="277812" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460366908329" checkinUser="plazi" docAuthor="Perger, Robert, Vargas, Rita & Wall, Adam" docDate="2011" docId="03CC878FFFA9FF90FF50FB13997F0573" docLanguage="en" docName="zt02911p068.pdf" docOrigin="Zootaxa 2911" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Johngarthia cocoensis Perger, Vargas & Wall, 2011, n. sp." docType="treatment" docVersion="10" lastPageNumber="63" masterDocId="FFF5FFF7FFA8FF96FFC7FF899B2A064B" masterDocTitle="Johngarthia cocoensis, a new species of Gecarcinidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura) from Cocos Island, Costa Rica" masterLastPageNumber="68" masterPageNumber="57" pageNumber="58" updateTime="1698295298436" updateUser="ExternalLinkService">
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<mods:mods id="340FB6E4B7CB9846045D4CE3DE1006AC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2DBD38DA2AD8E4EB25E24D2DBA41B64B">
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<mods:title id="57030E64CF016F084C78AC76CEA266A6">Johngarthia cocoensis, a new species of Gecarcinidae MacLeay, 1838 (Crustacea, Decapoda, Brachyura) from Cocos Island, Costa Rica</mods:title>
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</mods:titleInfo>
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<mods:name id="833A9C61FA1F0C1536C8FC1698AF8AFC" type="personal">
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<mods:role id="2C7A69B83090046DB0298CD9189E6ECD">
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<mods:roleTerm id="4E726526C9B0A2BCD046F1DE767973C7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="19A7E0F45FD3FDC1B040601C58E8F92F">Perger, Robert</mods:namePart>
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</mods:name>
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<mods:name id="46B49D11EDF00439FCA0FA32886F50BA" type="personal">
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<mods:role id="6A69F872D8542D18B8869C7A4929E324">
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<mods:roleTerm id="2CDBEAC9A120907CDBD589914C58ECFA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AEC2B7DCB46197E7334C6BC43A3F0A69">Vargas, Rita</mods:namePart>
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</mods:name>
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<mods:name id="B9C9660F8B09E6BB0ED97C139C241F08" type="personal">
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<mods:role id="9194E2035F149601D78843C55D7004DF">
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<mods:roleTerm id="5CB26147CEA060FA9BDE59B23251D437">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="25B9067616CDE9F8FE89B0BBF14783EA">Wall, Adam</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="663E01590299D185B8AF89245487183C">text</mods:typeOfResource>
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<mods:relatedItem id="E67065D74522F3E10E40DAC3A727947A" type="host">
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<mods:titleInfo id="7D7DBCC8B9F1C34228231A2B996569E7">
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<mods:title id="E818E200F1425F7F45198F56D6982CB3">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="7BB21B19816BBDF117A7106FFE85EE01">
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<mods:date id="F64972E2D1CEEECF7470D7BF533EB83E">2011</mods:date>
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<mods:detail id="0BEA4D6F77034F45A9D057DDC3B0514A" type="volume">
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<mods:number id="9C32D92CC32141FB769065A1FA8E9437">2911</mods:number>
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</mods:detail>
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<mods:end id="E824CFA664E26B75AD76F80D27EFEE3A">68</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="5CB6FCB4F327F0C1162C4F14B015A65C">journal article</mods:classification>
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<mods:identifier id="F965C580D18C1A19C87E8282F347C331" type="CLB-Dataset">46699</mods:identifier>
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<mods:identifier id="EBF4039B723C1A75C12D504415485885" type="DOI">10.5281/zenodo.277812</mods:identifier>
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<mods:identifier id="112F0D7689A3AB6BFA035CDB620CCBEF" type="GBIF-Dataset">96327166-d6b8-47d3-893f-517c372c5bcd</mods:identifier>
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<mods:identifier id="C54C98E8FE9CB1D8B713A544446C687A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="01C524CB0B3A84EEC53B30E34B4232F9" type="Zenodo-Dep">277812</mods:identifier>
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</mods:mods>
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<treatment id="03CC878FFFA9FF90FF50FB13997F0573" ID-DOI="http://doi.org/10.5281/zenodo.5696982" ID-GBIF-Taxon="119410993" ID-Zenodo-Dep="5696982" LSID="urn:lsid:plazi:treatment:03CC878FFFA9FF90FF50FB13997F0573" httpUri="http://treatment.plazi.org/id/03CC878FFFA9FF90FF50FB13997F0573" lastPageId="6" lastPageNumber="63" pageId="1" pageNumber="58">
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<subSubSection id="C37F6512FFA9FF97FF50FB139AD5029E" pageId="1" pageNumber="58" type="nomenclature">
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||
<paragraph id="8BDA3699FFA9FF97FF50FB139ADB02FF" blockId="1.[151,511,1178,1237]" box="[151,497,1178,1204]" pageId="1" pageNumber="58">
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||
<heading id="D09281F5FFA9FF97FF50FB139ADB02FF" bold="true" box="[151,497,1178,1204]" fontSize="11" level="1" pageId="1" pageNumber="58" reason="1">
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||
<emphasis id="B911EA8BFFA9FF97FF50FB139ADB02FF" bold="true" box="[151,497,1178,1204]" pageId="1" pageNumber="58">
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||
<taxonomicName id="4C654D1AFFA9FF97FF50FB139A8202FF" ID-CoL="3QQ3D" box="[151,424,1178,1204]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
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<emphasis id="B911EA8BFFA9FF97FF50FB139A8202FF" bold="true" box="[151,424,1178,1204]" italics="true" pageId="1" pageNumber="58">Johngarthia cocoensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257F0FFA9FF97FE68FB139ADB02FF" box="[431,497,1178,1204]" pageId="1" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDA3699FFA9FF97FF50FB349AD5029E" blockId="1.[151,511,1178,1237]" box="[151,511,1213,1237]" pageId="1" pageNumber="58">
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(
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<figureCitation id="135E2A1CFFA9FF97FF58FB349BC0029E" box="[159,234,1213,1237]" captionStart="FIGURE 1" captionStartId="2.[151,250,1863,1886]" captionTargetBox="[169,1417,593,1836]" captionTargetId="figure@2.[169,1417,593,1842]" captionTargetPageId="2" captionText="FIGURE 1. Live color, A, Johngarthia cocoensis n. sp., male (Cocos I., Costa Rica, photo by Michel Montoya); B, idem, female, juvenile (Cocos I., Costa Rica), photo by Michel Montoya); C, Gecarcoidea lalandii, male (Taiwan, photo by Hung- Chang Liu, Providence University, Taiwan); D, idem, female (Taiwan, photo by Hung-Chang Liu, Providence University, Taiwan); E, Johngarthia planatus (Clipperton Island, March 2008, photo by Julian P. Sachs, University of Washington); F, idem (Puerto Escondido, Oaxaca, Mexico)." httpUri="https://zenodo.org/record/277813/files/figure.png" pageId="1" pageNumber="58">Figs. 1</figureCitation>
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A, B; 2; 3; 4; 5B; 6D–F)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F6512FFA9FF97FF50FA8C981C015E" pageId="1" pageNumber="58" type="materials_examined">
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<paragraph id="8BDA3699FFA9FF97FF50FA8C992B03C2" blockId="1.[151,1437,1285,2030]" pageId="1" pageNumber="58">
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||
<emphasis id="B911EA8BFFA9FF97FF50FA8C9A6C0356" bold="true" box="[151,326,1285,1310]" pageId="1" pageNumber="58">
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<typeStatus id="54DE883BFFA9FF97FF50FA8C9BFA0356" box="[151,208,1285,1309]" pageId="1" pageNumber="58">Type</typeStatus>
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material.
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</emphasis>
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<collectingCountry id="F3727609FFA9FF97FE88FA8C9AE40356" box="[335,462,1285,1309]" name="Costa Rica" pageId="1" pageNumber="58">Costa Rica</collectingCountry>
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, Cocos Island: male,
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<typeStatus id="54DE883BFFA9FF97FD02FA8C98000356" box="[709,810,1285,1309]" pageId="1" pageNumber="58" type="holotype">holotype</typeStatus>
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, MZ-UCR-622-01,
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<date id="FFDB1059FFA9FF97FBCBFA8C9FA60356" box="[1036,1164,1285,1309]" pageId="1" pageNumber="58" value="1973-08-13">13.08.1973</date>
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, C. Villalobos and D.G. Robinson leg.;
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<specimenCount id="9D63FD10FFA9FF97FE86FAA09AA5030A" box="[321,399,1321,1345]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
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,
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<typeStatus id="54DE883BFFA9FF97FE5DFAA39AD70309" box="[410,509,1322,1346]" pageId="1" pageNumber="58" type="paratype">paratype</typeStatus>
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, same data;
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<collectingCountry id="F3727609FFA9FF97FD41FAA09829030A" box="[646,771,1321,1345]" name="Costa Rica" pageId="1" pageNumber="58">Costa Rica</collectingCountry>
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, Manuelita Island:
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<specimenCount id="9D63FD10FFA9FF97FC1EFAA09F18030A" box="[985,1074,1321,1345]" pageId="1" pageNumber="58" type="male">3 males</specimenCount>
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,
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<specimenCount id="9D63FD10FFA9FF97FBFAFAA09F88030A" box="[1085,1186,1321,1345]" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
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,
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<typeStatus id="54DE883BFFA9FF97FB6AFAA39E310309" box="[1197,1307,1322,1346]" pageId="1" pageNumber="58" type="paratype">paratypes</typeStatus>
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, MZ-UCR- 623-01,
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<date id="FFDB1059FFA9FF97FF33FAC49A4D032E" box="[244,359,1357,1381]" pageId="1" pageNumber="58" value="1973-08-14">14.8.1973</date>
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, N. Scott leg.;
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<collectingCountry id="F3727609FFA9FF97FDC8FAC499A6032E" box="[527,652,1357,1381]" name="Costa Rica" pageId="1" pageNumber="58">Costa Rica</collectingCountry>
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, Cocos Island, Bahía Wafer:
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<specimenCount id="9D63FD10FFA9FF97FC13FAC49F75032E" box="[980,1119,1357,1381]" pageId="1" pageNumber="58" type="juvenile">10 juveniles</specimenCount>
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,
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<typeStatus id="54DE883BFFA9FF97FBADFAC79FF2032D" box="[1130,1240,1358,1382]" pageId="1" pageNumber="58" type="paratype">paratypes</typeStatus>
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, MZ-UCR-2401- 0 6,
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<date id="FFDB1059FFA9FF97FF06FAF89A1403C2" box="[193,318,1393,1417]" pageId="1" pageNumber="58" value="2001-11-28">28.11.2001</date>
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M. Montoya leg.
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</paragraph>
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<paragraph id="8BDA3699FFA9FF97FF00FA1C981C015E" blockId="1.[151,1437,1285,2030]" pageId="1" pageNumber="58">
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<emphasis id="B911EA8BFFA9FF97FF00FA1C997D03E5" bold="true" box="[199,599,1429,1454]" pageId="1" pageNumber="58">Comparative material examined.</emphasis>
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<taxonomicName id="4C654D1AFFA9FF97FDA7FA1C987E03E6" box="[608,852,1429,1453]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="planatus">
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<emphasis id="B911EA8BFFA9FF97FDA7FA1C987E03E6" box="[608,852,1429,1453]" italics="true" pageId="1" pageNumber="58">Johngarthia planatus</emphasis>
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||
</taxonomicName>
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:
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<collectingCountry id="F3727609FFA9FF97FCA2FA1C989703E6" box="[869,957,1429,1453]" name="Mexico" pageId="1" pageNumber="58">Mexico</collectingCountry>
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, Socorro I.:
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<specimenCount id="9D63FD10FFA9FF97FB88FA1C9F8A03E6" box="[1103,1184,1429,1453]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
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, MNHN- B13154, A. Anthony leg.;
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<specimenCount id="9D63FD10FFA9FF97FEFDFA309ABE039A" box="[314,404,1465,1489]" pageId="1" pageNumber="58" type="male">2 males</specimenCount>
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,
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<collectionCode id="ED74AE5CFFA9FF97FE66FA309ADF039A" LSID="urn:lsid:biocol.org:col:34802" box="[417,501,1465,1489]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="58">LACM</collectionCode>
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,
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<date id="FFDB1059FFA9FF97FDC5FA3099B3039A" box="[514,665,1465,1489]" pageId="1" pageNumber="58" value="1971-02-16">16 Feb. 1971</date>
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, J. Garth det.;
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<specimenCount id="9D63FD10FFA9FF97FC85FA3098A5039A" box="[834,911,1465,1489]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
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<collectionCode id="ED74AE5CFFA9FF97FC50FA3098C2039A" LSID="urn:lsid:biocol.org:col:34871" box="[919,1000,1465,1489]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="58">USNM</collectionCode>
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20691, A. Anthony leg.;
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<collectingCountry id="F3727609FFA9FF97FACAFA309E4F039A" box="[1293,1381,1465,1489]" name="Mexico" pageId="1" pageNumber="58">Mexico</collectingCountry>
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, San Benedicto I.:
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<specimenCount id="9D63FD10FFA9FF97FEF7FA549AA303BE" box="[304,393,1501,1525]" pageId="1" pageNumber="58" type="male">2 males</specimenCount>
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, LACM-170, J. Garth det.;
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<specimenCount id="9D63FD10FFA9FF97FD05FA54983B03BE" box="[706,785,1501,1525]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
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,
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<collectionCode id="ED74AE5CFFA9FF97FCDAFA54984403BE" LSID="urn:lsid:biocol.org:col:34871" box="[797,878,1501,1525]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="58">USNM</collectionCode>
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20690, A. Anthony leg.;
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<collectingCountry id="F3727609FFA9FF97FB48FA549FCD03BE" box="[1167,1255,1501,1525]" name="Mexico" pageId="1" pageNumber="58">Mexico</collectingCountry>
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, Lower California:
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<specimenCount id="9D63FD10FFA9FF97FF0DF9889A3D0052" box="[202,279,1537,1561]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
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||
(most likely
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<typeStatus id="54DE883BFFA9FF97FE74F988993F0052" box="[435,533,1537,1561]" pageId="1" pageNumber="58" type="holotype">holotype</typeStatus>
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||
of
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<taxonomicName id="4C654D1AFFA9FF97FDF9F98B98E80052" authority="Bouvier, 1895" authorityName="Bouvier" authorityYear="1895" box="[574,962,1537,1561]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="digueti">
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||
<emphasis id="B911EA8BFFA9FF97FDF9F98B983F0052" box="[574,789,1537,1561]" italics="true" pageId="1" pageNumber="58">Gecarcinus digueti</emphasis>
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||
Bouvier, 1895
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||
</taxonomicName>
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||
), MNHN-B10951, Diguet leg.;
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<specimenCount id="9D63FD10FFA9FF97FAF7F9889EAA0052" box="[1328,1408,1537,1561]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D63FD10FFA9FF97FA49F9889BCC0076" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="ED74AE5CFFA9FF97FF36F9AC9A680076" LSID="urn:lsid:biocol.org:col:34871" box="[241,322,1573,1597]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="58">USNM</collectionCode>
|
||
12465;
|
||
<collectingCountry id="F3727609FFA9FF97FE5AF9AC9ADF0076" box="[413,501,1573,1597]" name="Mexico" pageId="1" pageNumber="58">Mexico</collectingCountry>
|
||
, Tres Marias I., María Cleofas:
|
||
<specimenCount id="9D63FD10FFA9FF97FC9EF9AC988D0076" box="[857,935,1573,1597]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
|
||
,
|
||
<collectionCode id="ED74AE5CFFA9FF97FC75F9AC9F290076" LSID="urn:lsid:biocol.org:col:34871" box="[946,1027,1573,1597]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="58">USNM</collectionCode>
|
||
20650, Nelson & Goldman leg.,
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||
<date id="FFDB1059FFA9FF97FAB1F9AC9BDD002A" pageId="1" pageNumber="58" value="1897-05-30">30- 05-1897</date>
|
||
;
|
||
<collectingCountry id="F3727609FFA9FF97FEC1F9C09A54002A" box="[262,382,1609,1633]" name="Clipperton Island" pageId="1" pageNumber="58">Clipperton</collectingCountry>
|
||
I.:
|
||
<specimenCount id="9D63FD10FFA9FF97FE6EF9C0992E002A" box="[425,516,1609,1633]" pageId="1" pageNumber="58" type="male">4 males</specimenCount>
|
||
,
|
||
<collectionCode id="ED74AE5CFFA9FF97FDD5F9C0994C002A" LSID="urn:lsid:biocol.org:col:34802" box="[530,614,1609,1633]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="58">LACM</collectionCode>
|
||
,
|
||
<date id="FFDB1059FFA9FF97FDB3F9C099DE002A" box="[628,756,1609,1633]" pageId="1" pageNumber="58" value="1958-12-09">12.09.1958</date>
|
||
, Limbaugh leg., J. Garth det.;
|
||
<specimenCount id="9D63FD10FFA9FF97FB9DF9C09F80002A" box="[1114,1194,1609,1633]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
|
||
, MNHN-B13156, D. Guinot det.;
|
||
<specimenCount id="9D63FD10FFA9FF97FEEFF9E49A5200CE" box="[296,376,1645,1669]" pageId="1" pageNumber="58" type="male">1 male</specimenCount>
|
||
,
|
||
<specimenCount id="9D63FD10FFA9FF97FE41F9E49AC600CE" box="[390,492,1645,1669]" pageId="1" pageNumber="58" type="female">1 female</specimenCount>
|
||
,
|
||
<collectionCode id="ED74AE5CFFA9FF97FE3EF9E4996000CE" LSID="urn:lsid:biocol.org:col:34871" box="[505,586,1645,1669]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="58">USNM</collectionCode>
|
||
19646, J. Arnheim leg;
|
||
<taxonomicName id="4C654D1AFFA9FF97FCA2F9E49F5D00CE" box="[869,1143,1645,1669]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="malpilensis">
|
||
<emphasis id="B911EA8BFFA9FF97FCA2F9E49F5D00CE" box="[869,1143,1645,1669]" italics="true" pageId="1" pageNumber="58">Johngarthia malpilensis</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<collectingCountry id="F3727609FFA9FF97FB40F9E49FD200CE" box="[1159,1272,1645,1669]" name="Colombia" pageId="1" pageNumber="58">Colombia</collectingCountry>
|
||
, Malpelo I.:
|
||
<specimenCount id="9D63FD10FFA9FF97FA49F9E49BF100E2" pageId="1" pageNumber="58" type="male">3 males</specimenCount>
|
||
,
|
||
<specimenCount id="9D63FD10FFA9FF97FF21F9189A7E00E2" box="[230,340,1681,1705]" pageId="1" pageNumber="58" type="female">4 females</specimenCount>
|
||
,
|
||
<collectionCode id="ED74AE5CFFA9FF97FE99F9189A9800E2" LSID="urn:lsid:biocol.org:col:34802" box="[350,434,1681,1705]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="1" pageNumber="58">LACM</collectionCode>
|
||
,
|
||
<date id="FFDB1059FFA9FF97FE7BF918991700E2" box="[444,573,1681,1705]" pageId="1" pageNumber="58" value="1933-01-16">16.01.1933</date>
|
||
, Hancock Pacific Expeditions, J. Garth det. Additional high resolution photographs of live
|
||
<taxonomicName id="4C654D1AFFA9FF97FEFEF93F9A9E0086" box="[313,436,1717,1741]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFA9FF97FEFEF93F9A9E0086" box="[313,436,1717,1741]" italics="true" pageId="1" pageNumber="58">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3727609FFA9FF97FE3DF93C99580086" box="[506,626,1717,1741]" name="Clipperton Island" pageId="1" pageNumber="58">Clipperton</collectingCountry>
|
||
(provided by Michel Montoya, Fundación Amigos
|
||
<collectingCountry id="F3727609FFA9FF97FB7EF93C9FCB0086" box="[1209,1249,1717,1741]" name="Iceland" pageId="1" pageNumber="58">Isla</collectingCountry>
|
||
del Coco,
|
||
<collectingCountry id="F3727609FFA9FF97FA9BF93C9BE700BA" name="Costa Rica" pageId="1" pageNumber="58">Costa Rica</collectingCountry>
|
||
, and Julian P. Sachs, University of Washington) and
|
||
<taxonomicName id="4C654D1AFFA9FF97FCDDF953989900BA" box="[794,947,1753,1777]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="malpilensis">
|
||
<emphasis id="B911EA8BFFA9FF97FCDDF953989900BA" box="[794,947,1753,1777]" italics="true" pageId="1" pageNumber="58">J. malpilensis</emphasis>
|
||
</taxonomicName>
|
||
(provided by Mateo Lopéz-Victoria, Justus Liebig Universität, Giessen,
|
||
<collectingCountry id="F3727609FFA9FF97FE1FF9749968015E" box="[472,578,1789,1813]" name="Germany" pageId="1" pageNumber="58">Germany</collectingCountry>
|
||
) were also examined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6512FFA9FF94FF00F8AB9A8B0453" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58" type="discussion">
|
||
<paragraph id="8BDA3699FFA9FF94FF00F8AB9A8B0453" blockId="1.[151,1437,1285,2030]" lastBlockId="2.[151,1436,151,536]" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58">
|
||
<emphasis id="B911EA8BFFA9FF97FF00F8AB9A1A0171" bold="true" box="[199,304,1826,1850]" pageId="1" pageNumber="58">Remark.</emphasis>
|
||
We could not locate the
|
||
<typeStatus id="54DE883BFFA9FF97FD85F8AB99580171" box="[578,626,1826,1850]" pageId="1" pageNumber="58">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C654D1AFFA9FF97FD3EF8AB98590172" box="[761,883,1825,1849]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFA9FF97FD3EF8AB98590172" box="[761,883,1825,1849]" italics="true" pageId="1" pageNumber="58">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
in the USNM, the Zoological Museum in Copenhagen or the Natural History Museum, London, where some
|
||
<typeStatus id="54DE883BFFA9FF97FC94F8CF98A90115" box="[851,899,1862,1886]" pageId="1" pageNumber="58">type</typeStatus>
|
||
material of species described by Stimpson has been deposited (see
|
||
<bibRefCitation id="EFF44B68FFA9FF97FEBDF8E3992B01CA" author="Evans" box="[378,513,1897,1921]" pageId="1" pageNumber="58" refString="Evans, A. C. (1967) Syntypes of Decapoda described by William Stimpson and James Dana in the collections of the British Museum (Natural History). Journal of Natural History, 1 (3), 399 - 411" type="journal article" year="1967">Evans 1967</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF44B68FFA9FF97FDC9F8E099FE01CA" author="Vasile" box="[526,724,1897,1921]" pageId="1" pageNumber="58" refString="Vasile, R. S., Manning, R. B. & Lemaitre, R. (2005) William Stimpson's Journal from the North Pacific Exploring Expedition, 1853 - 1856. Crustacean Research Special, 5, 1 - 220." type="journal article" year="2005">
|
||
Vasile
|
||
<emphasis id="B911EA8BFFA9FF97FD9EF8E399B801CA" box="[601,658,1897,1921]" italics="true" pageId="1" pageNumber="58">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFF44B68FFA9FF97FD27F8E098C201CA" author="Manning" box="[736,1000,1897,1921]" pageId="1" pageNumber="58" refString="Manning, R. B. & Reed, K. J. (2006) Decapod crustaceans deposited in the Zoological Museum of Copenhagen by William Stimpson in 1859. Raffles Bulletin of Zoology, 54 (2), 283 - 293." type="journal article" year="2006">Manning & Reed 2006</bibRefCitation>
|
||
). The material of
|
||
<taxonomicName id="4C654D1AFFA9FF97FB77F8E39E0101CA" box="[1200,1323,1897,1921]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFA9FF97FB77F8E39E0101CA" box="[1200,1323,1897,1921]" italics="true" pageId="1" pageNumber="58">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
was most likely destroyed by the Great Chicago Fire of 1871 (see
|
||
<bibRefCitation id="EFF44B68FFA9FF97FCE3F80498D801EE" author="Vasile" box="[804,1010,1933,1957]" pageId="1" pageNumber="58" refString="Vasile, R. S., Manning, R. B. & Lemaitre, R. (2005) William Stimpson's Journal from the North Pacific Exploring Expedition, 1853 - 1856. Crustacean Research Special, 5, 1 - 220." type="journal article" year="2005">
|
||
Vasile
|
||
<emphasis id="B911EA8BFFA9FF97FCB5F807988601EE" box="[882,940,1933,1957]" italics="true" pageId="1" pageNumber="58">et al.</emphasis>
|
||
2005
|
||
</bibRefCitation>
|
||
). The possible
|
||
<typeStatus id="54DE883BFFA9FF97FB66F8049E2901EE" box="[1185,1283,1933,1957]" pageId="1" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C654D1AFFA9FF97FAEAF8079A120182" authority="Bouvier, 1895" authorityName="Bouvier" authorityYear="1895" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="digueti">
|
||
<emphasis id="B911EA8BFFA9FF97FAEAF8079EB601EE" box="[1325,1436,1933,1957]" italics="true" pageId="1" pageNumber="58">G. digueti</emphasis>
|
||
Bouvier, 1895
|
||
</taxonomicName>
|
||
(MNHN-B10951) was labeled as
|
||
<taxonomicName id="4C654D1AFFA9FF97FD02F83B98850182" box="[709,943,1969,1993]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFA9FF97FD02F83B98850182" box="[709,943,1969,1993]" italics="true" pageId="1" pageNumber="58">Gecarcinus planatus</emphasis>
|
||
</taxonomicName>
|
||
, but it is the only specimen in the MNHN concordant with the
|
||
<typeStatus id="54DE883BFFA9FF97FEB8F85F9A8501A5" box="[383,431,2006,2030]" pageId="1" pageNumber="58">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C654D1AFFA9FF97FD8FF85F999C01A6" box="[584,694,2005,2029]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="58" phylum="Arthropoda" rank="species" species="digueti">
|
||
<emphasis id="B911EA8BFFA9FF97FD8FF85F999C01A6" box="[584,694,2005,2029]" italics="true" pageId="1" pageNumber="58">G. digueti</emphasis>
|
||
</taxonomicName>
|
||
described by Bouvier (1895) and subsequently reported by
|
||
<bibRefCitation id="EFF44B68FFA9FF94FA99F85C9A3006E4" author="Rathbun" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58" refString="Rathbun, M. J. (1899) Notes on the Crustacea of the Tres Marias Islands. North American Fauna, United States Department of Agriculture, 14, 73 - 75." type="journal article" year="1899">Rathbun (1899)</bibRefCitation>
|
||
. Bouvier apparently only had one specimen from “Basse-Californie” (Lower California), which he noted was measuring
|
||
<quantity id="4C9D9B7CFFAAFF94FE4FFF359AFD069F" box="[392,471,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="6.7" pageId="2" pageNumber="59" unit="mm" value="67.0">67 mm</quantity>
|
||
(sex not specified) (
|
||
<bibRefCitation id="EFF44B68FFAAFF94FD7EFF359877069F" author="Bouvier" box="[697,861,188,212]" pageId="2" pageNumber="59" refString="Bouvier, E. L. (1898) Sur quelques Crustaces anomoures et brachyures recuellis par M. Diguet en Basse-Californie. Bulletin du Museum d'Histoire Naturelle (Paris), 4, 371 - 384." type="journal article" year="1898">Bouvier, 1898</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFF44B68FFAAFF94FCA8FF359F35069F" author="Rathbun" box="[879,1055,188,212]" pageId="2" pageNumber="59" refString="Rathbun, M. J. (1899) Notes on the Crustacea of the Tres Marias Islands. North American Fauna, United States Department of Agriculture, 14, 73 - 75." type="journal article" year="1899">Rathbun (1899)</bibRefCitation>
|
||
recognized
|
||
<taxonomicName id="4C654D1AFFAAFF94FB60FF349E39069F" box="[1191,1299,188,212]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="digueti">
|
||
<emphasis id="B911EA8BFFAAFF94FB60FF349E39069F" box="[1191,1299,188,212]" italics="true" pageId="2" pageNumber="59">G. digueti</emphasis>
|
||
</taxonomicName>
|
||
as a distinct species, recording it from María Cleofa Island off Lower California, and apparently examined the
|
||
<typeStatus id="54DE883BFFAAFF94FB21FF699E3C06B3" box="[1254,1302,224,248]" pageId="2" pageNumber="59">type</typeStatus>
|
||
in the Paris Museum which she stated was a male measuring 69.0 by
|
||
<quantity id="4C9D9B7CFFAAFF94FCD9FE8D98AF0757" box="[798,901,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="4.63" pageId="2" pageNumber="59" unit="mm" value="46.3">46.3 mm</quantity>
|
||
. Maria Cleofa Island is situated relatively close to Cabo San Lucas (Cape St. Lucas, the most southern tip of Lower California peninsula), the
|
||
<typeStatus id="54DE883BFFAAFF94FB14FEA19E29070B" box="[1235,1283,296,320]" pageId="2" pageNumber="59">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="4C654D1AFFAAFF94FA4FFEA09A91072F" authority="Stimpson 1860" authorityName="Stimpson" authorityYear="1860" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAAFF94FA4FFEA09BD2072F" italics="true" pageId="2" pageNumber="59">J. planatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF44B68FFAAFF94FEC2FEC59A9E072F" author="Stimpson" box="[261,436,332,356]" pageId="2" pageNumber="59" refString="Stimpson, W. (1860) Notes on North American Crustacea, in the museum of the Smithsonian Institution, No II. Annals of Lyceum of Natural History of New York, 7, 176 - 246, pls. T. 2, 5." type="journal article" year="1860">Stimpson 1860</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. On the basis of the available data, this MNHN specimen is almost certainly the
|
||
<typeStatus id="54DE883BFFAAFF94FAFDFEC59EB6072F" box="[1338,1436,332,356]" pageId="2" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C654D1AFFAAFF94FF70FEF99AFB07CC" authority="Bouvier, 1895" authorityName="Bouvier" authorityYear="1895" box="[183,465,367,392]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="digueti">
|
||
<emphasis id="B911EA8BFFAAFF94FF70FEF99A0F07C3" box="[183,293,368,392]" italics="true" pageId="2" pageNumber="59">G. digueti</emphasis>
|
||
Bouvier, 1895
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFF44B68FFAAFF94FE19FEE699BA07CC" author="Rathbun" box="[478,656,367,391]" pageId="2" pageNumber="59" refString="Rathbun, M. J. (1918) The grapsoid crabs of America. Bulletin of the United States National Museum, 97, 1 - 461." type="journal article" year="1918">Rathbun (1918)</bibRefCitation>
|
||
later synonymized both
|
||
<taxonomicName id="4C654D1AFFAAFF94FC6CFEF99FDE07CC" authority="Faxon, 1893" authorityName="Faxon" authorityYear="1893" box="[939,1268,367,392]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="malpilensis">
|
||
<emphasis id="B911EA8BFFAAFF94FC6CFEF99F6107C3" box="[939,1099,368,392]" italics="true" pageId="2" pageNumber="59">G. malpilensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF44B68FFAAFF94FB9BFEE69FC607CC" author="Faxon" box="[1116,1260,367,391]" pageId="2" pageNumber="59" refString="Faxon, W. (1893) Preliminary descriptions of new species of Crustacea. Bulletin of the Museum of Comparative Zoology at Harvard College, 24,149 - 220." type="book" year="1893">Faxon, 1893</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C654D1AFFAAFF94FAE9FEF99A6007E7" authority="Bouvier, 1895" authorityName="Bouvier" authorityYear="1895" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="digueti">
|
||
<emphasis id="B911EA8BFFAAFF94FAE9FEF99EB607C3" box="[1326,1436,368,392]" italics="true" pageId="2" pageNumber="59">G. digueti</emphasis>
|
||
(Bouvier, 1895)
|
||
</taxonomicName>
|
||
with
|
||
<taxonomicName id="4C654D1AFFAAFF94FE4BFE1C99F007E7" authority="Stimpson, 1860" authorityName="Stimpson" authorityYear="1860" box="[396,730,404,428]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAAFF94FE4BFE1C992607E7" box="[396,524,404,428]" italics="true" pageId="2" pageNumber="59">G. planatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFF44B68FFAAFF94FDDBFE1D99F807E7" author="Stimpson" box="[540,722,404,428]" pageId="2" pageNumber="59" refString="Stimpson, W. (1860) Notes on North American Crustacea, in the museum of the Smithsonian Institution, No II. Annals of Lyceum of Natural History of New York, 7, 176 - 246, pls. T. 2, 5." type="journal article" year="1860">Stimpson, 1860</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EFF44B68FFAAFF94FD20FE1D98A007E7" author="Turkay" box="[743,906,404,428]" pageId="2" pageNumber="59" refString="Turkay, M. (1970) Die Gecarcinidae Amerikas. Mit einem Anhang uber Ucides Rathbun (Crustacea; Decapoda). Senckenbergiana biologica, 51, 333? 354." type="journal article" year="1970">Türkay (1970)</bibRefCitation>
|
||
recognized
|
||
<taxonomicName id="4C654D1AFFAAFF94FBD2FE1C9F9907E7" box="[1045,1203,404,428]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="malpilensis">
|
||
<emphasis id="B911EA8BFFAAFF94FBD2FE1C9F9907E7" box="[1045,1203,404,428]" italics="true" pageId="2" pageNumber="59">G. malpilensis</emphasis>
|
||
</taxonomicName>
|
||
as a distinct species but retained
|
||
<taxonomicName id="4C654D1AFFAAFF94FEE5FE319AA5079B" box="[290,399,440,464]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="digueti">
|
||
<emphasis id="B911EA8BFFAAFF94FEE5FE319AA5079B" box="[290,399,440,464]" italics="true" pageId="2" pageNumber="59">G. digueti</emphasis>
|
||
</taxonomicName>
|
||
under the synonymy of
|
||
<taxonomicName id="4C654D1AFFAAFF94FD58FE319837079B" box="[671,797,440,464]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAAFF94FD58FE319837079B" box="[671,797,440,464]" italics="true" pageId="2" pageNumber="59">G. planatus</emphasis>
|
||
</taxonomicName>
|
||
. The presumptive
|
||
<typeStatus id="54DE883BFFAAFF94FC37FE3E9F780784" box="[1008,1106,439,463]" pageId="2" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C654D1AFFAAFF94FBB0FE319FC9079B" box="[1143,1251,440,464]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="digueti">
|
||
<emphasis id="B911EA8BFFAAFF94FBB0FE319FC9079B" box="[1143,1251,440,464]" italics="true" pageId="2" pageNumber="59">G. digueti</emphasis>
|
||
</taxonomicName>
|
||
has all the diagnostic characters which we here identify with
|
||
<taxonomicName id="4C654D1AFFAAFF94FD63FE54980A07BF" box="[676,800,476,500]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAAFF94FD63FE54980A07BF" box="[676,800,476,500]" italics="true" pageId="2" pageNumber="59">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
and we are confident they are conspecific and different from
|
||
<taxonomicName id="4C654D1AFFAAFF94FF13FD889A760453" box="[212,348,513,536]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFAAFF94FF13FD889A760453" box="[212,348,513,536]" italics="true" pageId="2" pageNumber="59">J. cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFAAFF94FEA4FD899A8B0453" box="[355,417,512,536]" pageId="2" pageNumber="59" rank="species">
|
||
<emphasis id="B911EA8BFFAAFF94FEA4FD899A8B0453" bold="true" box="[355,417,512,536]" pageId="2" pageNumber="59">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1A6611FFAAFF94FF50F8CE993F0193" httpUri="https://zenodo.org/record/277813/files/figure.png" pageId="2" pageNumber="59" targetBox="[169,1417,593,1836]" targetPageId="2">
|
||
<paragraph id="8BDA3699FFAAFF94FF50F8CE993F0193" blockId="2.[151,1436,1863,2008]" pageId="2" pageNumber="59">
|
||
<emphasis id="B911EA8BFFAAFF94FF50F8CE9A3F0116" bold="true" box="[151,277,1863,1886]" pageId="2" pageNumber="59">FIGURE 1.</emphasis>
|
||
Live color, A,
|
||
<taxonomicName id="4C654D1AFFAAFF94FE7FF8C199B50115" box="[440,671,1864,1886]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFAAFF94FE7FF8C199B50115" box="[440,671,1864,1886]" italics="true" pageId="2" pageNumber="59">Johngarthia cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFAAFF94FD6FF8C199C90116" box="[680,739,1864,1885]" pageId="2" pageNumber="59" rank="species">
|
||
<emphasis id="B911EA8BFFAAFF94FD6FF8C199C90116" bold="true" box="[680,739,1864,1885]" pageId="2" pageNumber="59">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male (Cocos I., Costa Rica, photo by Michel Montoya); B, idem, female, juvenile (Cocos I., Costa Rica), photo by Michel Montoya); C,
|
||
<taxonomicName id="4C654D1AFFAAFF94FCBEF8EE9F780136" box="[889,1106,1895,1917]" class="Malacostraca" family="Gecarcinidae" genus="Gecarcoidea" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="lalandii">
|
||
<emphasis id="B911EA8BFFAAFF94FCBEF8EE9F780136" box="[889,1106,1895,1917]" italics="true" pageId="2" pageNumber="59">Gecarcoidea lalandii</emphasis>
|
||
</taxonomicName>
|
||
, male (Taiwan, photo by Hung- Chang Liu, Providence University, Taiwan); D, idem, female (Taiwan, photo by Hung-Chang Liu, Providence University, Taiwan); E,
|
||
<taxonomicName id="4C654D1AFFAAFF94FF32F82D9AFA01F1" box="[245,464,1956,1978]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAAFF94FF32F82D9AFA01F1" box="[245,464,1956,1978]" italics="true" pageId="2" pageNumber="59">Johngarthia planatus</emphasis>
|
||
</taxonomicName>
|
||
(Clipperton Island, March 2008, photo by Julian P. Sachs, University of Washington); F, idem (Puerto Escondido, Oaxaca, Mexico).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1A6611FFABFF95FF50FBDF991D02E2" httpUri="https://zenodo.org/record/277814/files/figure.png" pageId="3" pageNumber="60" targetBox="[169,1415,220,1075]" targetPageId="3">
|
||
<paragraph id="8BDA3699FFABFF95FF50FBDF991D02E2" blockId="3.[151,1437,1110,1194]" pageId="3" pageNumber="60">
|
||
<emphasis id="B911EA8BFFABFF95FF50FBDF9A380227" bold="true" box="[151,274,1110,1133]" pageId="3" pageNumber="60">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C654D1AFFABFF95FEDFFBDE9AD60226" box="[280,508,1111,1133]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFABFF95FEDFFBDE9AD60226" box="[280,508,1111,1133]" italics="true" pageId="3" pageNumber="60">Johngarthia cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFABFF95FDC4FBDE99160227" box="[515,572,1111,1132]" pageId="3" pageNumber="60" rank="species">
|
||
<emphasis id="B911EA8BFFABFF95FDC4FBDE99160227" bold="true" box="[515,572,1111,1132]" pageId="3" pageNumber="60">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, A, holotype male, W 89.85 mm, MZ-UCR-622-01 (Cocos I., Costa Rica), preserved in alcohol 70%, dorsal view; B, idem, ventral view; C, paratype female, W 57.35 mm, MZ-UCR-623-01, preserved in alcohol 70%, dorsal view; D, idem, ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37F6512FFABFF95FF00FB5F99A30335" pageId="3" pageNumber="60" type="diagnosis">
|
||
<paragraph id="8BDA3699FFABFF95FF00FB5F99A30335" blockId="3.[151,1437,1238,2024]" pageId="3" pageNumber="60">
|
||
<emphasis id="B911EA8BFFABFF95FF00FB5F9A6B02A4" bold="true" box="[199,321,1238,1263]" pageId="3" pageNumber="60">Diagnosis.</emphasis>
|
||
P about half as wide as orbital width; juvenile, subadults (W
|
||
<quantity id="4C9D9B7CFFABFF95FC37FB5F9F4302A5" box="[1008,1129,1238,1262]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="3" pageNumber="60" unit="mm" value="40.0" valueMax="60.0" valueMin="20.0">20–60 mm</quantity>
|
||
) with well developed exorbital carapace tooth, 6–16 anterolateral carapace teeth; ILs mostly separated from each other when resting against buccal cavity (
|
||
<figureCitation id="135E2A1CFFABFF95FEFAFA979AA8037D" box="[317,386,1310,1334]" captionStart="FIGURE 3" captionStartId="4.[151,250,1240,1263]" captionTargetBox="[169,1407,260,1201]" captionTargetId="figure@4.[169,1417,185,1227]" captionTargetPageId="4" captionText="FIGURE 3. Frontal view, A, Johngarthia cocoensis n. sp., holotype male, W 89.85 mm, MZ-UCR- 622 - 01 (Cocos I., Costa Rica), preserved in alcohol 70 %, dorsal view; B, idem, paratype female, W 57.35 mm, MZ-UCR- 623 - 01, preserved in alcohol 70 %." httpUri="https://zenodo.org/record/277815/files/figure.png" pageId="3" pageNumber="60">Fig. 3</figureCitation>
|
||
); palp of MXP3 merus partly exposed, epistomial tooth completely exposed (
|
||
<figureCitation id="135E2A1CFFABFF95FB30FA979E17037D" box="[1271,1341,1310,1334]" captionStart="FIGURE 3" captionStartId="4.[151,250,1240,1263]" captionTargetBox="[169,1407,260,1201]" captionTargetId="figure@4.[169,1417,185,1227]" captionTargetPageId="4" captionText="FIGURE 3. Frontal view, A, Johngarthia cocoensis n. sp., holotype male, W 89.85 mm, MZ-UCR- 622 - 01 (Cocos I., Costa Rica), preserved in alcohol 70 %, dorsal view; B, idem, paratype female, W 57.35 mm, MZ-UCR- 623 - 01, preserved in alcohol 70 %." httpUri="https://zenodo.org/record/277815/files/figure.png" pageId="3" pageNumber="60">Fig. 3</figureCitation>
|
||
; 6D–F); MXP3 exopodite not reaching ischium-merus joint, terminal setae protruding beyond ischium-merus joint; male first gonopod relatively straight (
|
||
<figureCitation id="135E2A1CFFABFF95FDCFFAEF99610335" box="[520,587,1382,1406]" captionStart="FIGURE 4" captionStartId="5.[151,250,1046,1069]" captionTargetBox="[193,1381,234,991]" captionTargetId="figure@5.[169,1417,193,1025]" captionTargetPageId="5" captionText="FIGURE 4. Johngarthia cocoensis n. sp., holotype male, MZ-UCR- 622 - 01, A, left male first gonopod, lateral view; Scale bar = 5 mm. B, same, mesial view; C, abdomen (segments 3 – 7), male, Scale bar = 10 mm; D, idem, female, Scale bar = 10 mm; E, right male cheliped (MZ-UCR- 623 - 01, W 112.3 mm); Scale bar = 10 mm." httpUri="https://zenodo.org/record/277816/files/figure.png" pageId="3" pageNumber="60">Fig. 4</figureCitation>
|
||
A, B)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6512FFABFF90FF00FA029E7E069F" lastPageId="6" lastPageNumber="63" pageId="3" pageNumber="60" type="description">
|
||
<paragraph id="8BDA3699FFABFF95FF00FA0299D303BB" blockId="3.[151,1437,1238,2024]" pageId="3" pageNumber="60">
|
||
<emphasis id="B911EA8BFFABFF95FF00FA029A5703E8" bold="true" box="[199,381,1419,1443]" pageId="3" pageNumber="60">Measurements.</emphasis>
|
||
Males
|
||
<quantity id="4C9D9B7CFFABFF95FE12FA0399A103E9" box="[469,651,1418,1442]" metricMagnitude="-2" metricUnit="m" metricValue="8.475" metricValueMax="11.23" metricValueMin="5.72" pageId="3" pageNumber="60" unit="mm" value="84.75" valueMax="112.3" valueMin="57.2">57.2–112.3 mm</quantity>
|
||
; females
|
||
<quantity id="4C9D9B7CFFABFF95FD3DFA03989103E9" box="[762,955,1418,1442]" metricMagnitude="-2" metricUnit="m" metricValue="3.9274999999999998" metricValueMax="5.735" metricValueMin="2.12" pageId="3" pageNumber="60" unit="mm" value="39.275" valueMax="57.35" valueMin="21.2">21.2 –57.35 mm</quantity>
|
||
; W/L = 1.24±0.06; W/H = 2.33±0.06; H/ MXP3 length = 1.52±0.03; MXP3 merus length/width = 1.23±0.05; FO width =
|
||
<geoCoordinate id="EE51505EFFABFF95FBD8FA389F530382" box="[1055,1145,1457,1481]" direction="west" orientation="longitude" pageId="3" pageNumber="60" precision="55" value="-1.353">1.353W</geoCoordinate>
|
||
0,754 (R² = 0,99); F width =
|
||
<geoCoordinate id="EE51505EFFABFF95FF50FA519BDA03BB" box="[151,240,1496,1520]" direction="west" orientation="longitude" pageId="3" pageNumber="60" precision="55" value="-0.445">0.445W</geoCoordinate>
|
||
0,751 (R² = 0.99); PL =
|
||
<geoCoordinate id="EE51505EFFABFF95FE24FA51996003BB" box="[483,586,1496,1520]" direction="west" orientation="longitude" pageId="3" pageNumber="60" precision="5" value="-0.0894">0.0894W</geoCoordinate>
|
||
1,4957 (R² = 0.98)
|
||
</paragraph>
|
||
<paragraph id="8BDA3699FFABFF95FF00FA7599C701A3" blockId="3.[151,1437,1238,2024]" pageId="3" pageNumber="60">
|
||
<emphasis id="B911EA8BFFABFF95FF00FA759A73005E" bold="true" box="[199,345,1532,1557]" pageId="3" pageNumber="60">Description.</emphasis>
|
||
Relatively large-size, robust species; carapace transversely ovate, strongly longitudinally, transversely convex, widest in anterior half; gastric region particularly well defined; median, cervical, urogastric grooves very pronounced; median groove reaches posterior height of greatest width of carapace; extremities of cervical grooves reaching pits at orbital angles; P about half as wide as orbital width; pit median, posterior of each cervical groove, smaller than pits at orbital angles, posterior very close to cervical groove, converging with urogastric groove; irregular lines of smaller pits lateral to cervical grooves; epigastric carinae absent, cardiac lobe strongly backward prolonged between bases of posterior legs; postfrontal crest strongly curved downwards, terminating as prominent carina, carina slightly curved medially. Suborbital, pterygostomial regions sparsely granular laterally; surfaces smooth. Subhepatic region with rounded postero-lateral margins, with rows of 3–9 small granules. In juvenile, subadult specimens (
|
||
<quantity id="4C9D9B7CFFABFF95FE7EF8C9991F0113" box="[441,565,1856,1880]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="3" pageNumber="60" unit="mm" value="40.0" valueMax="60.0" valueMin="20.0">20–60 mm</quantity>
|
||
) carapace in anterior third with small granules, rest of surface smooth; lateral borders swollen, upper border marked by carapace anterolateral border when seen in frontal view, prominently developed exorbital carapace tooth, carapace anterolateral border lined by 6–16 sharp teeth. In adult specimens (>
|
||
<quantity id="4C9D9B7CFFABFF95FF68F8259A2A018F" box="[175,256,1964,1988]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="60" unit="mm" value="60.0">60 mm</quantity>
|
||
) dorsal surface of carapace smooth; protogastric area, branchial regions conspicuously swollen; exorbital tooth weakly developed, anterolateral border smooth.
|
||
</paragraph>
|
||
<caption id="DF1A6611FFACFF92FF50FB519BE60360" httpUri="https://zenodo.org/record/277815/files/figure.png" pageId="4" pageNumber="61" targetBox="[169,1407,260,1201]" targetPageId="4">
|
||
<paragraph id="8BDA3699FFACFF92FF50FB519BE60360" blockId="4.[151,1436,1240,1323]" pageId="4" pageNumber="61">
|
||
<emphasis id="B911EA8BFFACFF92FF50FB519A3E02A5" bold="true" box="[151,276,1240,1263]" pageId="4" pageNumber="61">FIGURE 3.</emphasis>
|
||
Frontal view, A,
|
||
<taxonomicName id="4C654D1AFFACFF92FE0BFB51999802A4" box="[460,690,1240,1263]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFACFF92FE0BFB51999802A4" box="[460,690,1240,1263]" italics="true" pageId="4" pageNumber="61">Johngarthia cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFACFF92FD7CFB5099DF02A5" box="[699,757,1241,1262]" pageId="4" pageNumber="61" rank="species">
|
||
<emphasis id="B911EA8BFFACFF92FD7CFB5099DF02A5" bold="true" box="[699,757,1241,1262]" pageId="4" pageNumber="61">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male, W 89.85 mm, MZ-UCR-622-01 (Cocos I., Costa Rica), preserved in alcohol 70%, dorsal view; B, idem, paratype female, W 57.35 mm, MZ-UCR-623-01, preserved in alcohol 70%.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA3699FFACFF92FF02FADA9A8C03B0" blockId="4.[151,1436,1363,2035]" pageId="4" pageNumber="61">Orbit deep, broadly oval, height about 3/4 width, eye not completely filling orbit when folded; dorsal border slightly raised, outer angle distinctly defined, outer ventral, dorsal orbital borders converge with carapace anterolateral border at height of widest width of orbit when seen in frontal view; wide gap in ventral orbital border, border weakly carinate, projected, with row of 5 or 6 teeth, inner angle reaches front, separates antennae from orbit, furnished with 2 or 3 teeth.</paragraph>
|
||
<paragraph id="8BDA3699FFACFF92FF02F98E9BC30098" blockId="4.[151,1436,1363,2035]" pageId="4" pageNumber="61">Epistome sunken, densely covered with short setae, separated from basal segments of antennules by transverse ridge, ridge aligned with suborbital-border; median with band of setae; epistomial tooth exposed, triangular; antennules folded obliquely; antennae very short, nearly longitudinal, almost concealed by front, inter-antennular septum of moderate width; buccal cavern wide, widest in middle, laterally arched, densely covered with long, thick hairs, setae forming dense brush fitting with lateral border of MXP3, not visible when MXP3 rest against buccal cavern.</paragraph>
|
||
<paragraph id="8BDA3699FFACFF92FF02F95699AF01B8" blockId="4.[151,1436,1363,2035]" pageId="4" pageNumber="61">
|
||
MXP3 leaving between them wide rhomboidal gap when closed, mandibles exposed, internal border of ischium laterally lined with setae, ischium with well pronounced vertical sulcus; ILs mostly separated from each other when resting against buccal cavity, considerably narrower, shorter than ischium, longer than wide, not reaching epistome, leaving gap between lateral borders of buccal cavern, exposing epistomial tooth, borders of MXP3 merus relatively straight; apex apically divided by wide, V-shaped fissure, deepest point of emargination lateral, palp articulating at this point, palp partly exposed in resting position;
|
||
<collectingCountry id="F3727609FFACFF92FBC7F81D9F3001E0" box="[1024,1050,1940,1963]" name="Iceland" pageId="4" pageNumber="61">IL</collectingCountry>
|
||
longer than OL, rounded; exopod concealed by ischium, apex not reaching ischium-merus joint (size about 80% of ischium), terminal setae protrude from ischium-merus joint, flagellum absent.
|
||
</paragraph>
|
||
<caption id="DF1A6611FFADFF93FF50FB9F98A30221" httpUri="https://zenodo.org/record/277816/files/figure.png" pageId="5" pageNumber="62" targetBox="[193,1381,234,991]" targetPageId="5">
|
||
<paragraph id="8BDA3699FFADFF93FF50FB9F98A30221" blockId="5.[151,1436,1046,1130]" pageId="5" pageNumber="62">
|
||
<emphasis id="B911EA8BFFADFF93FF50FB9F9A390267" bold="true" box="[151,275,1046,1069]" pageId="5" pageNumber="62">FIGURE 4.</emphasis>
|
||
<taxonomicName id="4C654D1AFFADFF93FEDEFB9E9AD40266" box="[281,510,1047,1069]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFADFF93FEDEFB9E9AD40266" box="[281,510,1047,1069]" italics="true" pageId="5" pageNumber="62">Johngarthia cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFADFF93FDC3FB9E99170267" box="[516,573,1047,1068]" pageId="5" pageNumber="62" rank="species">
|
||
<emphasis id="B911EA8BFFADFF93FDC3FB9E99170267" bold="true" box="[516,573,1047,1068]" pageId="5" pageNumber="62">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male, MZ-UCR-622-01, A, left male first gonopod, lateral view; Scale bar = 5 mm. B, same, mesial view; C, abdomen (segments 3–7), male, Scale bar = 10 mm; D, idem, female, Scale bar = 10 mm; E, right male cheliped (MZ-UCR-623-01, W 112.3 mm); Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA3699FFADFF93FF02FB1F9F8B0351" blockId="5.[151,1436,1174,2026]" pageId="5" pageNumber="62">
|
||
Meri and chela of chelipeds allometric, in specimens W ≥
|
||
<quantity id="4C9D9B7CFFADFF93FCABFB1F989702E5" box="[876,957,1174,1198]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="62" unit="mm" value="60.0">60 mm</quantity>
|
||
slightly surpassing carapace anterolateral border, in large males abnormally elongated, widely surpassing carapace anterolateral border; in specimens ≥
|
||
<quantity id="4C9D9B7CFFADFF93FA47FB339BE902BD" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="62" unit="mm" value="60.0">60 mm</quantity>
|
||
surface of merus smooth, anterior proximal border furnished with 6 or 7 distinct teeth increasing in size distally, 3 sub-distal, 2 distal carpal teeth; surface, border of merus smooth in larger specimens.
|
||
</paragraph>
|
||
<paragraph id="8BDA3699FFADFF93FF02FAAF98BA0385" blockId="5.[151,1436,1174,2026]" pageId="5" pageNumber="62">
|
||
Specimens <W
|
||
<quantity id="4C9D9B7CFFADFF93FE45FAAF9ACB0375" box="[386,481,1318,1342]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="62" unit="mm" value="100.0">100 mm</quantity>
|
||
homochelous, males> W
|
||
<quantity id="4C9D9B7CFFADFF93FCD3FAAF98590375" box="[788,883,1318,1342]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="62" unit="mm" value="100.0">100 mm</quantity>
|
||
slightly heterochelous (left palm bigger, biggest difference between palm length 20%), palms elongated, length may exceed W, flattened in height, width, interiorproximal with longitudinal furrow; dactyl, pollex elongated, drop shaped in cross section, gaping moderate in largest chela when closed, not crossing, with triangular teeth, terminal with 7 rows of nearly uniform, oblique conical teeth, arrangement of teeth more diffuse proximally, manus smooth.
|
||
</paragraph>
|
||
<paragraph id="8BDA3699FFADFF93FF02FA53992E0159" blockId="5.[151,1436,1174,2026]" pageId="5" pageNumber="62">Ambulatory legs stout, sparsely furnished with setae; merus triangular in cross section, ventral edges with strongly developed ridges, dentate; carpus pentagonal in cross-section, distal edges with spines; propodus rhomboidal in cross-section, edges strongly dentate; dactylus longitudinally ridged, with 6 rows of spines. Sternum wider than long, S1 triangular, exposed, S2 narrow, S1-S3 fused, indistinct, S2-S3 suture, S3 separated from S4; S4 widest; sulci between S3-S6 distinct, sternites fused at upper border of abdominal cavity, separated medially, fused again near midline; abdominal cavity with longitudinal suture in S6; upper border of abdominal cavity with patches of setae; all abdominal somites and telson distinct, freely articulating; first somite filling space between last pair of ambulatory legs, sixth somite longest, considerably longer than telson, telson sub-triangular, narrow; lateral margins nearly straight, tip rounded.</paragraph>
|
||
<paragraph id="8BDA3699FFADFF93FF02F89798E801A1" blockId="5.[151,1436,1174,2026]" pageId="5" pageNumber="62">
|
||
Male first gonopod (
|
||
<figureCitation id="135E2A1CFFADFF93FE69F8979AD9017D" box="[430,499,1822,1846]" captionStart="FIGURE 4" captionStartId="5.[151,250,1046,1069]" captionTargetBox="[193,1381,234,991]" captionTargetId="figure@5.[169,1417,193,1025]" captionTargetPageId="5" captionText="FIGURE 4. Johngarthia cocoensis n. sp., holotype male, MZ-UCR- 622 - 01, A, left male first gonopod, lateral view; Scale bar = 5 mm. B, same, mesial view; C, abdomen (segments 3 – 7), male, Scale bar = 10 mm; D, idem, female, Scale bar = 10 mm; E, right male cheliped (MZ-UCR- 623 - 01, W 112.3 mm); Scale bar = 10 mm." httpUri="https://zenodo.org/record/277816/files/figure.png" pageId="5" pageNumber="62">Fig. 4</figureCitation>
|
||
A, B) slightly sinuous from lateral view, relatively straight from mesial view, slen- der, stiff, heavily chitinized, surfaces concave, mesial suture indistinct, margin distinctly developed; cephalic surface with band of sparse setae, apical border lined with stiff setae, caudal process shorter than mesial process, close-fitting to mesial process from lateral view, separated by emargination in mesial view; mesial process of first gonopod prominently developed, end piece on a level with terminal setae; diagonal, triangular, convex, amber-colored, aperture sub terminal, opening in an external-longitudinal channel.
|
||
</paragraph>
|
||
<paragraph id="8BDA3699FFAEFF90FF00FF1E9E7E069F" blockId="6.[151,1436,151,824]" pageId="6" pageNumber="63">
|
||
<emphasis id="B911EA8BFFAEFF90FF00FF1E9A3A06FB" bold="true" box="[199,272,151,176]" pageId="6" pageNumber="63">Color.</emphasis>
|
||
In adults, carapace reddish brown, carapace pits creamy white. In juvenile specimens, carapace dark brown-purple, carapace pits white, chelipeds and ambulatory legs reddish, tips of fingers white (
|
||
<figureCitation id="135E2A1CFFAEFF90FB0AFF359E26069F" box="[1229,1292,188,212]" captionStart="FIGURE 1" captionStartId="2.[151,250,1863,1886]" captionTargetBox="[169,1417,593,1836]" captionTargetId="figure@2.[169,1417,593,1842]" captionTargetPageId="2" captionText="FIGURE 1. Live color, A, Johngarthia cocoensis n. sp., male (Cocos I., Costa Rica, photo by Michel Montoya); B, idem, female, juvenile (Cocos I., Costa Rica), photo by Michel Montoya); C, Gecarcoidea lalandii, male (Taiwan, photo by Hung- Chang Liu, Providence University, Taiwan); D, idem, female (Taiwan, photo by Hung-Chang Liu, Providence University, Taiwan); E, Johngarthia planatus (Clipperton Island, March 2008, photo by Julian P. Sachs, University of Washington); F, idem (Puerto Escondido, Oaxaca, Mexico)." httpUri="https://zenodo.org/record/277813/files/figure.png" pageId="6" pageNumber="63">Fig. 1</figureCitation>
|
||
A, B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6512FFAEFF90FF00FF69987206BC" box="[199,856,223,248]" pageId="6" pageNumber="63" type="materials_examined">
|
||
<paragraph id="8BDA3699FFAEFF90FF00FF69987206BC" blockId="6.[151,1436,151,824]" box="[199,856,223,248]" pageId="6" pageNumber="63">
|
||
<emphasis id="B911EA8BFFAEFF90FF00FF699A4F06B3" bold="true" box="[199,357,223,248]" pageId="6" pageNumber="63">
|
||
<typeStatus id="54DE883BFFAEFF90FF00FF699A2806B3" box="[199,258,224,248]" pageId="6" pageNumber="63">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F3727609FFAEFF90FEABFF569AC306BC" box="[364,489,223,247]" name="Costa Rica" pageId="6" pageNumber="63">Costa Rica</collectingCountry>
|
||
, Cocos Island, Manuelita Island.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F6512FFAEFF90FF00FE8D997F0573" pageId="6" pageNumber="63" type="etymology">
|
||
<paragraph id="8BDA3699FFAEFF90FF00FE8D98C10757" blockId="6.[151,1436,151,824]" box="[199,1003,260,285]" pageId="6" pageNumber="63">
|
||
<emphasis id="B911EA8BFFAEFF90FF00FE8D9A640757" bold="true" box="[199,334,260,284]" pageId="6" pageNumber="63">Etymology.</emphasis>
|
||
The species is named after the
|
||
<typeStatus id="54DE883BFFAEFF90FD6AFE8C99F70756" box="[685,733,261,285]" pageId="6" pageNumber="63">type</typeStatus>
|
||
locality (Cocos Island).
|
||
</paragraph>
|
||
<paragraph id="8BDA3699FFAEFF90FF00FEAE9A2907E7" blockId="6.[151,1436,151,824]" pageId="6" pageNumber="63">
|
||
<emphasis id="B911EA8BFFAEFF90FF00FEAE9A47070B" bold="true" box="[199,365,295,320]" pageId="6" pageNumber="63">Comparisons.</emphasis>
|
||
In
|
||
<taxonomicName id="4C654D1AFFAEFF90FE56FEA199A4070B" box="[401,654,296,320]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFAEFF90FE56FEA199A4070B" box="[401,654,296,320]" italics="true" pageId="6" pageNumber="63">Johngarthia cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFAEFF90FD53FEA199F9070B" box="[660,723,296,320]" pageId="6" pageNumber="63" rank="species">
|
||
<emphasis id="B911EA8BFFAEFF90FD53FEA199F9070B" bold="true" box="[660,723,296,320]" pageId="6" pageNumber="63">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
the shape of the third maxilliped merus more closely resembles that of
|
||
<taxonomicName id="4C654D1AFFAEFF90FF2FFEC49AAE072F" box="[232,388,332,356]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="malpilensis">
|
||
<emphasis id="B911EA8BFFAEFF90FF2FFEC49AAE072F" box="[232,388,332,356]" italics="true" pageId="6" pageNumber="63">J. malpilensis</emphasis>
|
||
</taxonomicName>
|
||
than
|
||
<taxonomicName id="4C654D1AFFAEFF90FE03FEC4996A072F" box="[452,576,332,356]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAEFF90FE03FEC4996A072F" box="[452,576,332,356]" italics="true" pageId="6" pageNumber="63">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C654D1AFFAEFF90FDA8FEC499D2072F" box="[623,760,333,356]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFAEFF90FDA8FEC499D2072F" box="[623,760,333,356]" italics="true" pageId="6" pageNumber="63">J. cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFAEFF90FCC7FEC59815072F" box="[768,831,332,356]" pageId="6" pageNumber="63" rank="species">
|
||
<emphasis id="B911EA8BFFAEFF90FCC7FEC59815072F" bold="true" box="[768,831,332,356]" pageId="6" pageNumber="63">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C654D1AFFAEFF90FCBEFEC49F3E072F" box="[889,1044,332,356]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="malpilensis">
|
||
<emphasis id="B911EA8BFFAEFF90FCBEFEC49F3E072F" box="[889,1044,332,356]" italics="true" pageId="6" pageNumber="63">J. malpilensis</emphasis>
|
||
</taxonomicName>
|
||
, the V-shaped emargination of the third maxilliped merus is positioned laterally and the outer apical lobe is shorter than the inner apical lobe (
|
||
<figureCitation id="135E2A1CFFAEFF90FA86FEE69EA807CC" box="[1345,1410,367,391]" captionStart="FIGURE 5" captionStartId="7.[151,250,1194,1217]" captionTargetBox="[299,1379,213,1150]" captionTargetId="figure@7.[169,1417,167,1173]" captionTargetPageId="7" captionText="FIGURE 5. Frontal view (P, pit at orbital angle; M, merus of third maxilliped; OL, outer apical lobe of M; IL, inner apical lobe of M), A, Gecarcoidea lalandii, female, LACM, JWK- 326 (Marshall Islands, Enewetak Atoll, Igurin I.); B, Johngarthia cocoensis n. sp., paratype, female, MZ-UCR- 623 - 01 (Manuelita I., Costa Rica); C, Johngarthia malpilensis, female, LACM (Malpelo I., Colombia); D, Johngarthia planatus, female, LACM (Clipperton I., Mexico)." httpUri="https://zenodo.org/record/277817/files/figure.png" pageId="6" pageNumber="63">Fig. 5</figureCitation>
|
||
B, C; 6D–I).
|
||
</paragraph>
|
||
<paragraph id="8BDA3699FFAEFF90FF02FE31997F0573" blockId="6.[151,1436,151,824]" pageId="6" pageNumber="63">
|
||
<taxonomicName id="4C654D1AFFAEFF90FF02FE319AE8079B" box="[197,450,440,464]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFAEFF90FF02FE319AE8079B" box="[197,450,440,464]" italics="true" pageId="6" pageNumber="63">Johngarthia cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFAEFF90FE0DFE319923079B" box="[458,521,440,464]" pageId="6" pageNumber="63" rank="species">
|
||
<emphasis id="B911EA8BFFAEFF90FE0DFE319923079B" bold="true" box="[458,521,440,464]" pageId="6" pageNumber="63">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distinguished from
|
||
<taxonomicName id="4C654D1AFFAEFF90FCC0FE3098A8079B" box="[775,898,440,464]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAEFF90FCC0FE3098A8079B" box="[775,898,440,464]" italics="true" pageId="6" pageNumber="63">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
by having the third maxilliped merus considerably longer than wide (
|
||
<figureCitation id="135E2A1CFFAEFF90FE58FE559ACF07BF" box="[415,485,476,500]" captionStart="FIGURE 3" captionStartId="4.[151,250,1240,1263]" captionTargetBox="[169,1407,260,1201]" captionTargetId="figure@4.[169,1417,185,1227]" captionTargetPageId="4" captionText="FIGURE 3. Frontal view, A, Johngarthia cocoensis n. sp., holotype male, W 89.85 mm, MZ-UCR- 622 - 01 (Cocos I., Costa Rica), preserved in alcohol 70 %, dorsal view; B, idem, paratype female, W 57.35 mm, MZ-UCR- 623 - 01, preserved in alcohol 70 %." httpUri="https://zenodo.org/record/277815/files/figure.png" pageId="6" pageNumber="63">Fig. 3</figureCitation>
|
||
; 5B; 6D–F), in
|
||
<taxonomicName id="4C654D1AFFAEFF90FD5CFE54983C07BF" box="[667,790,476,500]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAEFF90FD5CFE54983C07BF" box="[667,790,476,500]" italics="true" pageId="6" pageNumber="63">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
the third maxilliped merus is as long as wide or slightly longer (
|
||
<figureCitation id="135E2A1CFFAEFF90FF37FE769A1F045C" box="[240,309,511,535]" captionStart="FIGURE 5" captionStartId="7.[151,250,1194,1217]" captionTargetBox="[299,1379,213,1150]" captionTargetId="figure@7.[169,1417,167,1173]" captionTargetPageId="7" captionText="FIGURE 5. Frontal view (P, pit at orbital angle; M, merus of third maxilliped; OL, outer apical lobe of M; IL, inner apical lobe of M), A, Gecarcoidea lalandii, female, LACM, JWK- 326 (Marshall Islands, Enewetak Atoll, Igurin I.); B, Johngarthia cocoensis n. sp., paratype, female, MZ-UCR- 623 - 01 (Manuelita I., Costa Rica); C, Johngarthia malpilensis, female, LACM (Malpelo I., Colombia); D, Johngarthia planatus, female, LACM (Clipperton I., Mexico)." httpUri="https://zenodo.org/record/277817/files/figure.png" pageId="6" pageNumber="63">Fig. 5</figureCitation>
|
||
D; 6A–C). In
|
||
<taxonomicName id="4C654D1AFFAEFF90FE1EFD88997D0453" box="[473,599,512,536]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAEFF90FE1EFD88997D0453" box="[473,599,512,536]" italics="true" pageId="6" pageNumber="63">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
the carapace is more flattened than in
|
||
<taxonomicName id="4C654D1AFFAEFF90FBDBFD889F8D0453" box="[1052,1191,513,536]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFAEFF90FBDBFD889F8D0453" box="[1052,1191,513,536]" italics="true" pageId="6" pageNumber="63">J. cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EA8BFFAEFF90FB77FD899FD80453" bold="true" box="[1200,1266,512,536]" pageId="6" pageNumber="63">
|
||
<taxonomicNameLabel id="A22257F0FFAEFF90FB77FD899FD80453" box="[1200,1266,512,536]" pageId="6" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<tableCitation id="C6E70322FFAEFF90FAC3FE769E66045C" box="[1284,1356,511,535]" captionStart="TABLE 1" captionStartId="6.[151,235,872,893]" captionTargetBox="[151,1436,983,1218]" captionTargetPageId="6" captionText="TABLE 1. Comparison of characters and measurements for J. cocoensis n. sp., J. malpilensis and J. planatus; W, carapace width; H, carapace height; O, Orbit; P, pit at orbital angle; M, MXP 3 merus; OL, M outer apical lobe; IL, M inner apical lobe; a, López-Victoria & Werding (2008); b, Rathbun (1918)." httpUri="http://table.plazi.org/id/DF1A6611FFAEFF90FF50FCE1982505F0" pageId="6" pageNumber="63" tableUuid="DF1A6611FFAEFF90FF50FCE1982505F0">Tab. 1</tableCitation>
|
||
). In
|
||
<taxonomicName id="4C654D1AFFAEFF90FA4FFD889A2E0477" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFAEFF90FA4FFD889A2E0477" italics="true" pageId="6" pageNumber="63">J. cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B911EA8BFFAEFF90FECDFDAD9A620477" bold="true" box="[266,328,548,572]" pageId="6" pageNumber="63">
|
||
<taxonomicNameLabel id="A22257F0FFAEFF90FECDFDAD9A620477" box="[266,328,548,572]" pageId="6" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the third maxilliped meri are so short that the inner apical lobes are mostly not overlapping in resting position and the third maxilliped palp is partly exposed (
|
||
<figureCitation id="135E2A1CFFAEFF90FCFBFDCE98AB0414" box="[828,897,583,607]" captionStart="FIGURE 3" captionStartId="4.[151,250,1240,1263]" captionTargetBox="[169,1407,260,1201]" captionTargetId="figure@4.[169,1417,185,1227]" captionTargetPageId="4" captionText="FIGURE 3. Frontal view, A, Johngarthia cocoensis n. sp., holotype male, W 89.85 mm, MZ-UCR- 622 - 01 (Cocos I., Costa Rica), preserved in alcohol 70 %, dorsal view; B, idem, paratype female, W 57.35 mm, MZ-UCR- 623 - 01, preserved in alcohol 70 %." httpUri="https://zenodo.org/record/277815/files/figure.png" pageId="6" pageNumber="63">Fig. 3</figureCitation>
|
||
; 5B; 6D–F), while in
|
||
<taxonomicName id="4C654D1AFFAEFF90FBBFFDC09E3B042B" box="[1144,1297,584,608]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="malpilensis">
|
||
<emphasis id="B911EA8BFFAEFF90FBBFFDC09E3B042B" box="[1144,1297,584,608]" italics="true" pageId="6" pageNumber="63">J. malpilensis</emphasis>
|
||
</taxonomicName>
|
||
, these lobes are enlarged, mostly overlapping and conceal the third maxilliped palp (
|
||
<figureCitation id="135E2A1CFFAEFF90FC71FDE598D204CF" box="[950,1016,620,644]" captionStart="FIGURE 5" captionStartId="7.[151,250,1194,1217]" captionTargetBox="[299,1379,213,1150]" captionTargetId="figure@7.[169,1417,167,1173]" captionTargetPageId="7" captionText="FIGURE 5. Frontal view (P, pit at orbital angle; M, merus of third maxilliped; OL, outer apical lobe of M; IL, inner apical lobe of M), A, Gecarcoidea lalandii, female, LACM, JWK- 326 (Marshall Islands, Enewetak Atoll, Igurin I.); B, Johngarthia cocoensis n. sp., paratype, female, MZ-UCR- 623 - 01 (Manuelita I., Costa Rica); C, Johngarthia malpilensis, female, LACM (Malpelo I., Colombia); D, Johngarthia planatus, female, LACM (Clipperton I., Mexico)." httpUri="https://zenodo.org/record/277817/files/figure.png" pageId="6" pageNumber="63">Fig. 5</figureCitation>
|
||
C; 6G–I). In
|
||
<taxonomicName id="4C654D1AFFAEFF90FB40FDE49E0A04CF" box="[1159,1312,620,644]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="malpilensis">
|
||
<emphasis id="B911EA8BFFAEFF90FB40FDE49E0A04CF" box="[1159,1312,620,644]" italics="true" pageId="6" pageNumber="63">J. malpilensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C654D1AFFAEFF90FA92FDE49BFE04E3" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAEFF90FA92FDE49BFE04E3" italics="true" pageId="6" pageNumber="63">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
the base of the male first gonopod is relatively more compact and the apex relatively more slender from mesial and lateral views, the opposite condition is discerned when viewed from the caudal and cephalic views. In
|
||
<taxonomicName id="4C654D1AFFAEFF90FA4FFD3C9A2E04BB" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFAEFF90FA4FFD3C9A2E04BB" italics="true" pageId="6" pageNumber="63">J. cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFAEFF90FECCFD519A6004BB" box="[267,330,728,752]" pageId="6" pageNumber="63" rank="species">
|
||
<emphasis id="B911EA8BFFAEFF90FECCFD519A6004BB" bold="true" box="[267,330,728,752]" pageId="6" pageNumber="63">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
the male first gonopod is relatively much straighter (
|
||
<figureCitation id="135E2A1CFFAEFF90FC66FD5E98CF04A4" box="[929,997,727,751]" captionStart="FIGURE 4" captionStartId="5.[151,250,1046,1069]" captionTargetBox="[193,1381,234,991]" captionTargetId="figure@5.[169,1417,193,1025]" captionTargetPageId="5" captionText="FIGURE 4. Johngarthia cocoensis n. sp., holotype male, MZ-UCR- 622 - 01, A, left male first gonopod, lateral view; Scale bar = 5 mm. B, same, mesial view; C, abdomen (segments 3 – 7), male, Scale bar = 10 mm; D, idem, female, Scale bar = 10 mm; E, right male cheliped (MZ-UCR- 623 - 01, W 112.3 mm); Scale bar = 10 mm." httpUri="https://zenodo.org/record/277816/files/figure.png" pageId="6" pageNumber="63">Fig. 4</figureCitation>
|
||
A, B). In
|
||
<taxonomicName id="4C654D1AFFAEFF90FB88FD509FC004BB" box="[1103,1258,728,752]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="malpilensis">
|
||
<emphasis id="B911EA8BFFAEFF90FB88FD509FC004BB" box="[1103,1258,728,752]" italics="true" pageId="6" pageNumber="63">J. malpilensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C654D1AFFAEFF90FAE6FD509EB604BB" box="[1313,1436,728,752]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="planatus">
|
||
<emphasis id="B911EA8BFFAEFF90FAE6FD509EB604BB" box="[1313,1436,728,752]" italics="true" pageId="6" pageNumber="63">J. planatus</emphasis>
|
||
</taxonomicName>
|
||
the surfaces of the are considerably more concave and the apex of the end piece is longer in relation to the male first gonopod than in
|
||
<taxonomicName id="4C654D1AFFAEFF90FE4FFCA899250573" box="[392,527,801,824]" class="Malacostraca" family="Gecarcinidae" genus="Johngarthia" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="cocoensis" status="sp. nov.">
|
||
<emphasis id="B911EA8BFFAEFF90FE4FFCA899250573" box="[392,527,801,824]" italics="true" pageId="6" pageNumber="63">J. cocoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257F0FFAEFF90FDD1FCA9997F0573" box="[534,597,800,824]" pageId="6" pageNumber="63" rank="species">
|
||
<emphasis id="B911EA8BFFAEFF90FDD1FCA9997F0573" bold="true" box="[534,597,800,824]" pageId="6" pageNumber="63">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |