286 lines
34 KiB
XML
286 lines
34 KiB
XML
<document id="DD8B78EF412C1541CC7D2CBFFF4E151F" ID-DOI="10.1080/14772019.2021.1968966" ID-ISSN="1478-0941" ID-Zenodo-Dep="10883707" ID-ZooBank="A185883A-68FC-4A7F-A103-889EE22ED134" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1711485782547" checkinUser="felipe" docAuthor="Liu, Juan" docDate="2021" docId="03CC8786FF897A13FEDB88422B55F9F7" docLanguage="en" docName="JSystPalaeontol.19.9.677-689.pdf" docOrigin="Journal of Systematic Palaeontology 19 (9)" docSource="http://dx.doi.org/10.1080/14772019.2021.1968966" docStyle="DocumentStyle:6400638016C1F4AE0D0B6502E032D571.1:JSystPalaeontol.2016-.journal_article.closed" docStyleId="6400638016C1F4AE0D0B6502E032D571" docStyleName="JSystPalaeontol.2016-.journal_article.closed" docStyleVersion="1" docTitle="Wilsonium brevipinne" docType="treatment" docVersion="3" lastPageNumber="681" masterDocId="FFF5FFFEFF8A7A16FFD48F732A1CFFD4" masterDocTitle="Redescription of ‘ Amyzon’ brevipinne and remarks on North American Eocene catostomids (Cypriniformes: Catostomidae)" masterLastPageNumber="689" masterPageNumber="677" pageNumber="679" updateTime="1711486428063" updateUser="ExternalLinkService">
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<mods:title id="620E5223FD3CCF27394A2668F834B7B9">Redescription of ‘ Amyzon’ brevipinne and remarks on North American Eocene catostomids (Cypriniformes: Catostomidae)</mods:title>
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<mods:namePart id="1DDB89290C10BA1025EF854AC280834E">Liu, Juan</mods:namePart>
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<mods:title id="7FB6B15BBEED1CDC6DC486DCF92C6469">Journal of Systematic Palaeontology</mods:title>
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<mods:part id="09A354FE690F4DCDAA40F71CC80E689C">
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<mods:date id="B2190C1ED91F5ABD83D62E9B9B43BB7F">2021</mods:date>
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<mods:number id="3BB0655CC7A7D331882978164424AD43">2021-09-20</mods:number>
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<mods:number id="2A6C5B644295A6192CC9AC69CBCE92D4">19</mods:number>
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<mods:identifier id="42624278A52CCF06FCC3180443BC031D" type="DOI">10.1080/14772019.2021.1968966</mods:identifier>
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<mods:identifier id="6EC5631F25FC1CE68DE3CCFDA65F0A6A" type="ISSN">1478-0941</mods:identifier>
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<treatment id="03CC8786FF897A13FEDB88422B55F9F7" ID-DOI="http://doi.org/10.5281/zenodo.10883647" ID-Zenodo-Dep="10883647" LSID="urn:lsid:plazi:treatment:03CC8786FF897A13FEDB88422B55F9F7" httpUri="http://treatment.plazi.org/id/03CC8786FF897A13FEDB88422B55F9F7" lastPageId="5" lastPageNumber="681" pageId="3" pageNumber="679">
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<subSubSection id="C37F651BFF897A15FEDB88422881F89C" box="[271,669,1841,1865]" pageId="3" pageNumber="679" type="nomenclature">
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<paragraph id="8BDA3690FF897A15FEDB88422881F89C" blockId="3.[271,669,1841,1899]" box="[271,669,1841,1865]" pageId="3" pageNumber="679">
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<heading id="D09281FCFF897A15FEDB88422881F89C" box="[271,669,1841,1865]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="679" reason="8">
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<taxonomicName id="4C654D13FF897A15FEDB88422881F89C" authority="(Cope, 1893)" baseAuthorityName="Cope" baseAuthorityYear="1893" box="[271,669,1841,1865]" family="Catostomidae" genus="Wilsonium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="679" phylum="Chordata" rank="species" species="brevipinne">
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<emphasis id="B911EA82FF897A15FEDB88422819F89D" bold="true" box="[271,517,1841,1865]" italics="true" pageId="3" pageNumber="679">Wilsonium brevipinne</emphasis>
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||
(
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<bibRefCitation id="EFF44B61FF897A15FDC18842288FF89C" author="Cope, E. D." box="[533,659,1841,1864]" pageId="3" pageNumber="679" pagination="401 - 402" refId="ref8642" refString="Cope, E. D. 1893. Fossil fishes from British Columbia. Proceedings of the Academy of Natural Sciences of Philadelphia, 45, 401 - 402." type="journal article" year="1893">Cope, 1893</bibRefCitation>
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||
)
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37F651BFF897A15FEA288272ECAFC80" box="[374,1238,829,1899]" pageId="3" pageNumber="679" type="description">
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||
<paragraph id="8BDA3690FF897A15FEA28827282AF8BF" blockId="3.[271,669,1841,1899]" box="[374,566,1876,1899]" pageId="3" pageNumber="679">
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||
(
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<figureCitation id="135E2A15FF897A15FEAB88272BD9F8BF" box="[383,453,1876,1899]" captionStart="Figure 2" captionStartId="3.[159,226,726,747]" captionTargetBox="[189,1437,188,705]" captionTargetId="figure-422@3.[189,1437,188,705]" captionTargetPageId="3" captionText="Figure 2. The holotype (CMNFV 6189) of Wilsonium brevipinne from the Eocene Allenby Formation, probably from the Pleasant Valley locality, British Columbia, Canada. Arrow indicates the frontal bone. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10883711" httpUri="https://zenodo.org/record/10883711/files/figure.png" pageId="3" pageNumber="679">Figs 2</figureCitation>
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,
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<figureCitation id="135E2A15FF897A15FE0688272BFDF8BF" box="[466,481,1876,1899]" captionStart="Figure 3" captionStartId="4.[159,226,1277,1298]" captionTargetBox="[189,1437,188,1256]" captionTargetId="figure-171@4.[189,1437,188,1256]" captionTargetPageId="4" captionText="Figure 3. A juvenile or young adult and skulls of Wilsonium brevipinne from the Eocene Allenby Formation, Blakeburn Mine, British Columbia, Canada. A, juvenile or young adult, UALVP 12159; B, skull of UALVP 12658; C, line drawing of composite skull based on UALVP 12159 and UALVP 12658. Arrows indicate the frontal bones. Abbreviations: aa, anguloarticular; c1–4, the space for centra 1 through 4; d, dentary; e, ethmoid; fr, frontal; hm, hyomandibular; iob, infraorbitals, iob 1 (lacrimal) through iob 5 numbered from anterior to posterior; iop, interopercle; ke, kinethmoid; le, lateral ethmoid; m, maxilla; nc, neural complex; ns4, neural spine of the 4th vertebra; op, opercular; p, palatine; pa, parietal; pm, premaxilla; pop, preopercle; pt, pterotic; q, quadrate; r4, the 4th rib or rib of the 4th vertebra (also known as outer arm of os suspensorium in some taxa); ra, retroarticular; so, supraorbital; soc, supraoccipital; sop, sub-opercle; sp, sphenoid. Scale bars: A = 10 mm; B = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10883713" httpUri="https://zenodo.org/record/10883713/files/figure.png" pageId="3" pageNumber="679">3</figureCitation>
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,
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<figureCitation id="135E2A15FF897A15FE3B8827280CF8BF" box="[495,528,1876,1899]" captionStart="Figure 4" captionStartId="5.[159,226,713,734]" captionTargetBox="[190,1422,188,692]" captionTargetId="figure-419@5.[189,1437,188,692]" captionTargetPageId="5" captionText="Figure 4. Comparison of the frontal bones of Wilsonium, Amyzon and Plesiomyxocyprinus. A, frontal of Wilsonium brevipinne from the Eocene Allenby Formation, Blakeburn Mine, British Columbia, Canada, UALVP 12610; B, frontal of Plesiomyxocyprinus arratiae from the Eocene Huadian Formation, Huadian, Jilin, China, IVPP V 15711.29 (images modified from Liu & Chang 2009, fig. 2a, b); C, frontal of Amyzon aggregatum from the Eocene of Horsefly, British Columbia, Canada, UALVP 32931. A, B, dorsal view; C, medial view. Anterior is to the top. Abbreviations: fpf, fronto-parietal fontanelle; obp, orbital process of frontal; on, orbital notch of frontal. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10883715" httpUri="https://zenodo.org/record/10883715/files/figure.png" pageId="3" pageNumber="679">4A</figureCitation>
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,
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<figureCitation id="135E2A15FF897A15FDCB88272830F8BF" box="[543,556,1876,1899]" captionStart="Figure 5" captionStartId="6.[159,226,1375,1396]" captionTargetBox="[189,1436,189,1354]" captionTargetId="figure-144@6.[189,1437,188,1354]" captionTargetPageId="6" captionText="Figure 5. Disarticulated bones and reconstructions of Wilsonium brevipinne from the Eocene Allenby Formation. A, C, anterior to the left; B, D–F, anterior to the right. C and D in medial view; A and C in posteroventral view; and the rest in lateral view. A, pharyngeal bone and teeth, UALVP 31585a; A’, line drawing of pharyngeal bone and teeth based on UALVP 31585a; B, opercle, UALVP 12796; C, interopercle, UALVP 12616; D, vomer; ROM 11161a; E, caudal skeleton, ROM 11161a; E’, line drawing of caudal skeleton, ROM 11161a; F, mandible including dentary, articular, and retroarticular, ROM 11161. Abbreviations: aa, anguloarticular; aaf, anguloarticular fossa for articulation with the quadrate; arp, auricular process of opercle; avp, anteroventral process; cc, compound centrum of caudal skeleton; cp, coronoid process of dentary; den, dentary; emp, preethmoid process; ep, epural; gr, gnathic ramus of dentary; hyp1–5, hypural 1 through 5; opa, opercular arm; opf, opercular fossa; pls, pleurostyle; pt, pharyngeal teeth; r4, rib 4; ra, retroarticular. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.10883717" httpUri="https://zenodo.org/record/10883717/files/figure.png" pageId="3" pageNumber="679">5</figureCitation>
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||
)
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</paragraph>
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<paragraph id="8BDA3690FF897A15FC938C4E2ECAFC80" blockId="3.[839,1466,829,949]" box="[839,1238,829,852]" pageId="3" pageNumber="679">
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||
1893
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<taxonomicName id="4C654D13FF897A15FC518C4E2E86FC80" authority="Cope" authorityName="Cope" authorityYear="1893" box="[901,1178,829,852]" family="Catostomidae" genus="Amyzon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="679" phylum="Chordata" rank="species" species="brevipinne">
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<emphasis id="B911EA82FF897A15FC518C4E2E49FC80" box="[901,1109,829,852]" italics="true" pageId="3" pageNumber="679">Amyzon brevipinne</emphasis>
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Cope
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</taxonomicName>
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: 402.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C37F651BFF897A15FC938C2E29BFFC61" pageId="3" pageNumber="679" type="reference_group">
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||
<paragraph id="8BDA3690FF897A15FC938C2E2F69FCA0" blockId="3.[839,1466,829,949]" box="[839,1397,861,884]" pageId="3" pageNumber="679">
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||
1906
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<taxonomicName id="4C654D13FF897A15FC518C2E2EC7FCA0" authority="Cope, 1893" authorityName="Cope" authorityYear="1893" box="[901,1243,861,884]" family="Catostomidae" genus="Amyzon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="679" phylum="Chordata" rank="species" species="brevipinne">
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<emphasis id="B911EA82FF897A15FC518C2E2E49FCA0" box="[901,1109,861,884]" italics="true" pageId="3" pageNumber="679">Amyzon brevipinne</emphasis>
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<bibRefCitation id="EFF44B61FF897A15FB8A8C2E2EC7FCA0" author="Cope, E. D." box="[1118,1243,861,884]" pageId="3" pageNumber="679" pagination="401 - 402" refId="ref8642" refString="Cope, E. D. 1893. Fossil fishes from British Columbia. Proceedings of the Academy of Natural Sciences of Philadelphia, 45, 401 - 402." type="journal article" year="1893">Cope, 1893</bibRefCitation>
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</taxonomicName>
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; Lambe: 154.
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</paragraph>
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<paragraph id="8BDA3690FF897A15FC938C0E29BFFC61" blockId="3.[839,1466,829,949]" pageId="3" pageNumber="679">
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1977
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<taxonomicName id="4C654D13FF897A15FC588C0E2EEAFC40" authority="Cope, 1893" authorityName="Cope" authorityYear="1893" box="[908,1270,893,916]" family="Catostomidae" genus="Amyzon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="679" phylum="Chordata" rank="species" species="brevipinne">
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<emphasis id="B911EA82FF897A15FC588C0E2E7EFC40" box="[908,1122,893,916]" italics="true" pageId="3" pageNumber="679">Amyzon brevipinne</emphasis>
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<bibRefCitation id="EFF44B61FF897A15FBA68C0E2EEAFC40" author="Cope, E. D." box="[1138,1270,893,916]" pageId="3" pageNumber="679" pagination="401 - 402" refId="ref8642" refString="Cope, E. D. 1893. Fossil fishes from British Columbia. Proceedings of the Academy of Natural Sciences of Philadelphia, 45, 401 - 402." type="journal article" year="1893">Cope, 1893</bibRefCitation>
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</taxonomicName>
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; Wilson: 23–32, fig. 8.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F651BFF897A13FC988CBD2B55F9F7" lastPageId="5" lastPageNumber="681" pageId="3" pageNumber="679" type="materials_examined">
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<paragraph id="8BDA3690FF897A15FC988CBD2F15FBD2" blockId="3.[844,1467,974,1030]" pageId="3" pageNumber="679">
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<emphasis id="B911EA82FF897A15FC988CBD29A7FC31" bold="true" box="[844,955,974,997]" pageId="3" pageNumber="679">
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<typeStatus id="54DE8832FF897A15FC988CBD29AAFC31" box="[844,950,974,997]" pageId="3" pageNumber="679" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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CMNFV 6189, a fish preserved in right lateral view lacking the caudal fin (
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<figureCitation id="135E2A15FF897A15FB6D8C9C2EE5FBD2" box="[1209,1273,1007,1030]" captionStart="Figure 2" captionStartId="3.[159,226,726,747]" captionTargetBox="[189,1437,188,705]" captionTargetId="figure-422@3.[189,1437,188,705]" captionTargetPageId="3" captionText="Figure 2. The holotype (CMNFV 6189) of Wilsonium brevipinne from the Eocene Allenby Formation, probably from the Pleasant Valley locality, British Columbia, Canada. Arrow indicates the frontal bone. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.10883711" httpUri="https://zenodo.org/record/10883711/files/figure.png" pageId="3" pageNumber="679">Fig. 2</figureCitation>
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).
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</paragraph>
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<paragraph id="8BDA3690FF897A15FC988B6C2F47FB0F" blockId="3.[844,1467,1055,1902]" pageId="3" pageNumber="679">
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<emphasis id="B911EA82FF897A15FC988B6C2E33FBE2" bold="true" box="[844,1071,1055,1078]" pageId="3" pageNumber="679">Referred materials.</emphasis>
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Well-preserved skeletal specimens:
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<collectionCode id="ED74AE55FF897A15FC988B3329B9FB83" box="[844,933,1088,1111]" pageId="3" pageNumber="679">UALVP</collectionCode>
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12159, a nearly complete fish;
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<collectionCode id="ED74AE55FF897A15FADA8B332F7BFB83" box="[1294,1383,1088,1111]" pageId="3" pageNumber="679">UALVP</collectionCode>
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12658, a nearly complete fish lacking the tail;
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<collectionCode id="ED74AE55FF897A15FAD98B122F7AFBAC" box="[1293,1382,1121,1144]" pageId="3" pageNumber="679">UALVP</collectionCode>
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12610, a complete right frontal;
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<collectionCode id="ED74AE55FF897A15FBB58BF12EA6FB4D" box="[1121,1210,1154,1177]" pageId="3" pageNumber="679">UALVP</collectionCode>
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31585, a right pharyngeal bone with teeth;
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<collectionCode id="ED74AE55FF897A15FB9F8BD02E94FB6E" box="[1099,1160,1187,1210]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="3" pageNumber="679" type="Museum">ROM</collectionCode>
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11160, a pharyngeal bone with teeth;
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<collectionCode id="ED74AE55FF897A15FC1D8BB72E1AFB0F" box="[969,1030,1220,1243]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="3" pageNumber="679" type="Museum">ROM</collectionCode>
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11163, a nearly complete fish.
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</paragraph>
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<paragraph id="8BDA3690FF897A15FCB38B972F36FA36" blockId="3.[844,1467,1055,1902]" pageId="3" pageNumber="679">
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<collectionCode id="ED74AE55FF897A15FCB38B9729B8FB2F" box="[871,932,1252,1275]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="3" pageNumber="679" type="Museum">CMN</collectionCode>
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collection: CMNFV 2042, 2046, 2050, 2054, 2055, 2057, 2066, 2068, 2069, 2071–2074, 2090, 2092, 2097, 2099, 2104, 34480–34492, 34509, 34820, 34821, 348883, 34885, 34888, 34891, 34893–34899, 41745 and 41746. Specimens listed together under NMC
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<quantity id="4C9D9B75FF897A15FAB78A1B2FA7FAAB" box="[1379,1467,1384,1407]" metricMagnitude="1" metricUnit="m" metricValue="7.95528" pageId="3" pageNumber="679" unit="in" value="3132.0">3132 in</quantity>
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<bibRefCitation id="EFF44B61FF897A15FC998AFA29E7FA74" author="Wilson, M. V. H." box="[845,1019,1417,1440]" pageId="3" pageNumber="679" pagination="1 - 61" refId="ref9795" refString="Wilson, M. V. H. 1977. Middle Eocene freshwater fishes from British Columbia. Life Sciences Contributions, Royal Ontario Museum, 113, 1 - 61." type="journal article" year="1977">Wilson (1977)</bibRefCitation>
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are re-catalogued within CMNFV 34480–34899. CMNFV 6195(a, b) and 25266 from Tranquille area are identified as
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<taxonomicName id="4C654D13FF897A15FB658AB82F3FFA36" authorityName="Liu" authorityYear="2021" box="[1201,1315,1483,1506]" family="Catostomidae" genus="Wilsonium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="679" phylum="Chordata" rank="genus">
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<emphasis id="B911EA82FF897A15FB658AB82F3FFA36" box="[1201,1315,1483,1506]" italics="true" pageId="3" pageNumber="679">Wilsonium</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BDA3690FF897A12FCB38A982FB4F900" blockId="3.[844,1467,1055,1902]" lastBlockId="4.[845,1467,1533,1876]" lastPageId="4" lastPageNumber="680" pageId="3" pageNumber="679">
|
||
<collectionCode id="ED74AE55FF897A15FCB38A9829DCF9D6" box="[871,960,1515,1538]" pageId="3" pageNumber="679">UALVP</collectionCode>
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||
collection:
|
||
<collectionCode id="ED74AE55FF897A15FB548A982EC5F9D6" box="[1152,1241,1515,1538]" pageId="3" pageNumber="679">UALVP</collectionCode>
|
||
12104, 12133, 12462–12464, 12480, 12481, 12483, 12487, 12496, 12509, 12512, 12514–12517, 12519–12521, 12525–12529, 12532–12534, 12537–12539, 12541, 12542, 12545–12547, 12549, 12553–12556, 12558, 12561, 12563–12566, 12568–12570, 12572–12577, 12580, 12583, 12584, 12586–12589, 12591, 12593, 12595, 12597–12599, 12601, 12602, 12605, 12606, 12611–12613, 12616, 12617, 12619, 12621–12626, 12631–12636, 12638, 12639, 12641–12645, 12647, 12651–12654, 12661–12680, 12682–12684, 12686–12689, 12691–12698, 12700–12708, 12710, 12711, 12714–12716, 12719–12729, 12731, 12733–12736, 12743–12746, 12750, 12754, 12757, 12762, 12763, 12767, 12768, 12770–12772, 12775, 12777–12779, 12783–12785, 12787–12790, 12794–12798, 12800–12804, 12806, 12807, 12810, 12812, 12813, 12819, 12820, 12825–12828, 12830, 12831, 12835, 12837–12843, 12845–12847, 12850, 12854–12858, 12860–12864, 12866, 12867, 12869, 12878, 12879, 12884, 12888, 12889, 12893–12904, 12906, 12907, 12910, 12912–12921, 12923, 12957, 12969, 12978, 12981, 12987, 12989, 12991, 12992, 13006, 13008, 13012, 13019, 13021, 13037, 13054, 13071, 13072, 13078, 13085, 13089, 13097, 13114–13126, 13129, 13131–13135, 13137–13144, 13146, 13147, 13149, 13150, 13152–13174, 13177, 13182, 13184, 13187, 13435, 14730, 14731, 14735, 14737, 14738, 14745–14748, 14754, 14755, 14759, 14760, 15912–15915, 17836, 21262–21339 and 27140.
|
||
</paragraph>
|
||
<caption id="DF1A6618FF8E7A12FF4B8B8E29EBFA19" ID-DOI="http://doi.org/10.5281/zenodo.10883713" ID-Zenodo-Dep="10883713" httpUri="https://zenodo.org/record/10883713/files/figure.png" pageId="4" pageNumber="680" startId="4.[159,226,1277,1298]" targetBox="[189,1437,188,1256]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BDA3690FF8E7A12FF4B8B8E29EBFA19" blockId="4.[159,1467,1277,1485]" pageId="4" pageNumber="680">
|
||
<emphasis id="B911EA82FF8E7A12FF4B8B8E2B1CFAC6" bold="true" box="[159,256,1277,1298]" pageId="4" pageNumber="680">Figure 3.</emphasis>
|
||
A juvenile or young adult and skulls of
|
||
<taxonomicName id="4C654D13FF8E7A12FD668B8E2994FAC6" baseAuthorityName="Cope" baseAuthorityYear="1893" box="[690,904,1277,1298]" family="Catostomidae" genus="Wilsonium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="680" phylum="Chordata" rank="species" species="brevipinne">
|
||
<emphasis id="B911EA82FF8E7A12FD668B8E2994FAC6" box="[690,904,1277,1298]" italics="true" pageId="4" pageNumber="680">Wilsonium brevipinne</emphasis>
|
||
</taxonomicName>
|
||
from the Eocene Allenby Formation, Blakeburn Mine, British Columbia, Canada.
|
||
<emphasis id="B911EA82FF8E7A12FE648A6B2BDBFAF9" bold="true" box="[432,455,1304,1325]" pageId="4" pageNumber="680">A,</emphasis>
|
||
juvenile or young adult, UALVP 12159;
|
||
<emphasis id="B911EA82FF8E7A12FCA08A6B2996FAF9" bold="true" box="[884,906,1304,1325]" pageId="4" pageNumber="680">B,</emphasis>
|
||
skull of UALVP 12658;
|
||
<emphasis id="B911EA82FF8E7A12FB468A6B2EB5FAF9" bold="true" box="[1170,1193,1304,1325]" pageId="4" pageNumber="680">C,</emphasis>
|
||
line drawing of composite skull based on UALVP 12159 and UALVP 12658. Arrows indicate the frontal bones.
|
||
<emphasis id="B911EA82FF8E7A12FC2F8A402E97FA9C" bold="true" box="[1019,1163,1331,1352]" pageId="4" pageNumber="680">Abbreviations</emphasis>
|
||
:
|
||
<emphasis id="B911EA82FF8E7A12FB4E8A402EAEFA9C" bold="true" box="[1178,1202,1331,1352]" pageId="4" pageNumber="680">aa</emphasis>
|
||
, anguloarticular;
|
||
<emphasis id="B911EA82FF8E7A12FAB48A402F92FA9C" bold="true" box="[1376,1422,1330,1352]" pageId="4" pageNumber="680">c1–4</emphasis>
|
||
, the space for centra 1 through 4;
|
||
<emphasis id="B911EA82FF8E7A12FE178A3E2BCCFAB6" bold="true" box="[451,464,1357,1378]" pageId="4" pageNumber="680">d</emphasis>
|
||
, dentary;
|
||
<emphasis id="B911EA82FF8E7A12FDE08A3E2823FAB6" bold="true" box="[564,575,1357,1378]" pageId="4" pageNumber="680">e</emphasis>
|
||
, ethmoid;
|
||
<emphasis id="B911EA82FF8E7A12FD7D8A3E28A7FAB6" bold="true" box="[681,699,1357,1378]" pageId="4" pageNumber="680">fr</emphasis>
|
||
, frontal;
|
||
<emphasis id="B911EA82FF8E7A12FCCC8A3E2925FAB6" bold="true" box="[792,825,1357,1378]" pageId="4" pageNumber="680">hm</emphasis>
|
||
, hyomandibular;
|
||
<emphasis id="B911EA82FF8E7A12FC328A3E2E1AFAB6" bold="true" box="[998,1030,1357,1378]" pageId="4" pageNumber="680">iob</emphasis>
|
||
, infraorbitals, iob 1 (lacrimal) through iob 5 numbered from anterior to posterior;
|
||
<emphasis id="B911EA82FF8E7A12FDC88A1B2820FAA9" bold="true" box="[540,572,1384,1405]" pageId="4" pageNumber="680">iop</emphasis>
|
||
, interopercle;
|
||
<emphasis id="B911EA82FF8E7A12FD048A1B28F4FAA9" bold="true" box="[720,744,1384,1405]" pageId="4" pageNumber="680">ke</emphasis>
|
||
, kinethmoid;
|
||
<emphasis id="B911EA82FF8E7A12FCA38A1B2994FAA9" bold="true" box="[887,904,1384,1405]" pageId="4" pageNumber="680">le</emphasis>
|
||
, lateral ethmoid;
|
||
<emphasis id="B911EA82FF8E7A12FB948A1B2E48FAA9" bold="true" box="[1088,1108,1384,1405]" pageId="4" pageNumber="680">m</emphasis>
|
||
, maxilla;
|
||
<emphasis id="B911EA82FF8E7A12FB6B8A1B2ECBFAA9" bold="true" box="[1215,1239,1384,1405]" pageId="4" pageNumber="680">nc</emphasis>
|
||
, neural complex;
|
||
<emphasis id="B911EA82FF8E7A12FA468A1B2FA8FAA9" bold="true" box="[1426,1460,1384,1405]" pageId="4" pageNumber="680">ns4</emphasis>
|
||
, neural spine of the 4th vertebra;
|
||
<emphasis id="B911EA82FF8E7A12FE0B8AF02BE4FA4C" bold="true" box="[479,504,1411,1432]" pageId="4" pageNumber="680">op</emphasis>
|
||
, opercular;
|
||
<emphasis id="B911EA82FF8E7A12FDBA8AF02867FA4C" bold="true" box="[622,635,1411,1432]" pageId="4" pageNumber="680">p</emphasis>
|
||
, palatine;
|
||
<emphasis id="B911EA82FF8E7A12FD378AF028E0FA4C" bold="true" box="[739,764,1411,1432]" pageId="4" pageNumber="680">pa</emphasis>
|
||
, parietal;
|
||
<emphasis id="B911EA82FF8E7A12FC8B8AF0299CFA4C" bold="true" box="[863,896,1411,1432]" pageId="4" pageNumber="680">pm</emphasis>
|
||
, premaxilla;
|
||
<emphasis id="B911EA82FF8E7A12FBD68AF02E34FA4C" bold="true" box="[1026,1064,1411,1432]" pageId="4" pageNumber="680">pop</emphasis>
|
||
, preopercle;
|
||
<emphasis id="B911EA82FF8E7A12FB7D8AF02EA2FA4C" bold="true" box="[1193,1214,1411,1432]" pageId="4" pageNumber="680">pt</emphasis>
|
||
, pterotic;
|
||
<emphasis id="B911EA82FF8E7A12FAF78AF02F2CFA4C" bold="true" box="[1315,1328,1411,1432]" pageId="4" pageNumber="680">q</emphasis>
|
||
, quadrate;
|
||
<emphasis id="B911EA82FF8E7A12FA4A8AF02FA9FA4C" bold="true" box="[1438,1461,1411,1432]" pageId="4" pageNumber="680">r4</emphasis>
|
||
, the 4th rib or rib of the 4th vertebra (also known as outer arm of os suspensorium in some taxa);
|
||
<emphasis id="B911EA82FF8E7A12FBBB8AEE2E99FA66" bold="true" box="[1135,1157,1437,1458]" pageId="4" pageNumber="680">ra</emphasis>
|
||
, retroarticular;
|
||
<emphasis id="B911EA82FF8E7A12FACB8AEE2F28FA66" bold="true" box="[1311,1332,1437,1458]" pageId="4" pageNumber="680">so</emphasis>
|
||
, supraorbital;
|
||
<emphasis id="B911EA82FF8E7A12FF4B8ACB2AA3FA19" bold="true" box="[159,191,1464,1485]" pageId="4" pageNumber="680">soc</emphasis>
|
||
, supraoccipital;
|
||
<emphasis id="B911EA82FF8E7A12FE8B8ACB2B9DFA19" bold="true" box="[351,385,1464,1485]" pageId="4" pageNumber="680">sop</emphasis>
|
||
, sub-opercle;
|
||
<emphasis id="B911EA82FF8E7A12FDD98ACB283FFA19" bold="true" box="[525,547,1464,1485]" pageId="4" pageNumber="680">sp</emphasis>
|
||
, sphenoid. Scale bars: A = 10 mm; B = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA3690FF8E7A12FCB389AE29D6F880" blockId="4.[845,1467,1533,1876]" pageId="4" pageNumber="680">
|
||
<collectionCode id="ED74AE55FF8E7A12FCB389AE29B9F920" box="[871,933,1757,1780]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="4" pageNumber="680" type="Museum">ROM</collectionCode>
|
||
collection:
|
||
<collectionCode id="ED74AE55FF8E7A12FB9989AE2E97F920" box="[1101,1163,1757,1780]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="4" pageNumber="680" type="Museum">ROM</collectionCode>
|
||
11161, 11162, 11164, 11166, 11167, 11169–11171, 11179, 11182, 11370, 11377, 11379, 11382, 11387, 11389, 11461, 11468 and 19420.
|
||
</paragraph>
|
||
<caption id="DF1A6618FF8F7A13FF4B8DBA2819FCB7" ID-DOI="http://doi.org/10.5281/zenodo.10883715" ID-Zenodo-Dep="10883715" httpUri="https://zenodo.org/record/10883715/files/figure.png" pageId="5" pageNumber="681" startId="5.[159,226,713,734]" targetBox="[190,1422,188,692]" targetPageId="5" targetType="figure">
|
||
<paragraph id="8BDA3690FF8F7A13FF4B8DBA2819FCB7" blockId="5.[159,1467,713,867]" pageId="5" pageNumber="681">
|
||
<emphasis id="B911EA82FF8F7A13FF4B8DBA2AE1FD0A" bold="true" box="[159,253,713,734]" pageId="5" pageNumber="681">Figure 4.</emphasis>
|
||
Comparison of the frontal bones of
|
||
<taxonomicName id="4C654D13FF8F7A13FDB88DBA28CDFD0A" authorityName="Liu" authorityYear="2021" box="[620,721,713,734]" family="Catostomidae" genus="Wilsonium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="681" phylum="Chordata" rank="genus">
|
||
<emphasis id="B911EA82FF8F7A13FDB88DBA28CDFD0A" box="[620,721,713,734]" italics="true" pageId="5" pageNumber="681">Wilsonium</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C654D13FF8F7A13FD0B8DBA2936FD0A" box="[735,810,713,734]" family="Catostomidae" genus="Amyzon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="681" phylum="Chordata" rank="genus">
|
||
<emphasis id="B911EA82FF8F7A13FD0B8DBA2936FD0A" box="[735,810,713,734]" italics="true" pageId="5" pageNumber="681">Amyzon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C654D13FF8F7A13FC8A8DBA2E01FD0A" baseAuthorityName="Liu & Chang" baseAuthorityYear="2009" box="[862,1053,713,734]" family="Catostomidae" genus="Plesiomyxocyprinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="681" phylum="Chordata" rank="genus">
|
||
<emphasis id="B911EA82FF8F7A13FC8A8DBA2E01FD0A" box="[862,1053,713,734]" italics="true" pageId="5" pageNumber="681">Plesiomyxocyprinus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B911EA82FF8F7A13FBF88DBA2E5FFD0A" bold="true" box="[1068,1091,713,734]" pageId="5" pageNumber="681">A,</emphasis>
|
||
frontal of
|
||
<taxonomicName id="4C654D13FF8F7A13FB648DBA2F98FD0A" baseAuthorityName="Cope" baseAuthorityYear="1893" box="[1200,1412,713,734]" family="Catostomidae" genus="Wilsonium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="681" phylum="Chordata" rank="species" species="brevipinne">
|
||
<emphasis id="B911EA82FF8F7A13FB648DBA2F98FD0A" box="[1200,1412,713,734]" italics="true" pageId="5" pageNumber="681">Wilsonium brevipinne</emphasis>
|
||
</taxonomicName>
|
||
from the Eocene Allenby Formation, Blakeburn Mine, British Columbia, Canada, UALVP 12610;
|
||
<emphasis id="B911EA82FF8F7A13FBBE8D902E9CFD2C" bold="true" box="[1130,1152,739,760]" pageId="5" pageNumber="681">B,</emphasis>
|
||
frontal of
|
||
<taxonomicName id="4C654D13FF8F7A13FB2F8D902AF1FCC7" family="Catostomidae" genus="Plesiomyxocyprinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="681" phylum="Chordata" rank="species" species="arratiae">
|
||
<emphasis id="B911EA82FF8F7A13FB2F8D902AF1FCC7" italics="true" pageId="5" pageNumber="681">Plesiomyxocyprinus arratiae</emphasis>
|
||
</taxonomicName>
|
||
from the Eocene Huadian Formation, Huadian, Jilin, China, IVPP V 15711.29 (images modified from
|
||
<bibRefCitation id="EFF44B61FF8F7A13FB228D8D2FA8FCC7" author="Liu, J. & Chang, M. - M." box="[1270,1460,766,787]" pageId="5" pageNumber="681" pagination="189 - 202" refId="ref9011" refString="Liu, J. & Chang, M. - M. 2009. A new Eocene catostomid (Teleostei: Cypriniformes) from northeastern China and early divergence of Catostomidae. Science in China, Series D: Earth Sciences, 52, 189 - 202. doi: 10.1007 / s 11430 - 009 - 0022 - 2" type="journal article" year="2009">Liu & Chang 2009</bibRefCitation>
|
||
, fig. 2a, b);
|
||
<emphasis id="B911EA82FF8F7A13FEC58C6A2B34FCFA" bold="true" box="[273,296,793,814]" pageId="5" pageNumber="681">C,</emphasis>
|
||
frontal of
|
||
<taxonomicName id="4C654D13FF8F7A13FE428C6A2842FCFA" box="[406,606,793,814]" family="Catostomidae" genus="Amyzon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="681" phylum="Chordata" rank="species" species="aggregatum">
|
||
<emphasis id="B911EA82FF8F7A13FE428C6A2842FCFA" box="[406,606,793,814]" italics="true" pageId="5" pageNumber="681">Amyzon aggregatum</emphasis>
|
||
</taxonomicName>
|
||
from the Eocene of Horsefly, British Columbia, Canada, UALVP 32931. A, B, dorsal view; C, medial view. Anterior is to the top.
|
||
<emphasis id="B911EA82FF8F7A13FD8E8C40290BFC9C" bold="true" box="[602,791,819,840]" pageId="5" pageNumber="681">Abbreviations: fpf</emphasis>
|
||
, fronto-parietal fontanelle;
|
||
<emphasis id="B911EA82FF8F7A13FBF78C402E55FC9C" bold="true" box="[1059,1097,819,840]" pageId="5" pageNumber="681">obp</emphasis>
|
||
, orbital process of frontal;
|
||
<emphasis id="B911EA82FF8F7A13FA818C402F72FC9C" bold="true" box="[1365,1390,819,840]" pageId="5" pageNumber="681">on</emphasis>
|
||
, orbital notch of frontal. Scale bars = 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA3690FF8F7A13FF4B8CFC28FDFC72" blockId="5.[159,737,911,934]" box="[159,737,911,934]" pageId="5" pageNumber="681">
|
||
<emphasis id="B911EA82FF8F7A13FF4B8CFC2B7AFC72" bold="true" box="[159,358,911,934]" pageId="5" pageNumber="681">Age and horizon.</emphasis>
|
||
Allenby Formation, early Eocene.
|
||
</paragraph>
|
||
<paragraph id="8BDA3690FF8F7A13FF4B8CCC2B5FFBEC" blockId="5.[159,781,959,1080]" pageId="5" pageNumber="681">
|
||
<emphasis id="B911EA82FF8F7A13FF4B8CCC2B5EFC02" bold="true" box="[159,322,959,982]" pageId="5" pageNumber="681">Type locality.</emphasis>
|
||
North Fork of the Similkameen River (Tulameen Rivers), probably in Pleasant Valley (Princeton area),
|
||
<collectingRegion id="49A1F872FF8F7A13FE738B732896FBC3" box="[423,650,1024,1047]" country="Canada" name="British Columbia" pageId="5" pageNumber="681">British Columbia</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3727600FF8F7A13FD698B732911FBC3" box="[701,781,1024,1047]" name="Canada" pageId="5" pageNumber="681">Canada</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFF44B61FF8F7A13FF728B522B2FFBEC" author="Wilson, M. V. H." box="[166,307,1057,1080]" pageId="5" pageNumber="681" pagination="1 - 61" refId="ref9795" refString="Wilson, M. V. H. 1977. Middle Eocene freshwater fishes from British Columbia. Life Sciences Contributions, Royal Ontario Museum, 113, 1 - 61." type="journal article" year="1977">Wilson 1977</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA3690FF8F7A13FF4B8B2228EEFB5D" blockId="5.[159,780,1105,1161]" pageId="5" pageNumber="681">
|
||
<emphasis id="B911EA82FF8F7A13FF4B8B222B92FBBC" bold="true" box="[159,398,1105,1128]" pageId="5" pageNumber="681">Additional localities.</emphasis>
|
||
Whipsaw Creek, Blakeburn Mine areas, and Tulameen, all in
|
||
<collectingRegion id="49A1F872FF8F7A13FE048B012891FB5D" box="[464,653,1138,1161]" country="Canada" name="British Columbia" pageId="5" pageNumber="681">British Columbia</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3727600FF8F7A13FD4E8B0128F1FB5D" box="[666,749,1138,1161]" name="Canada" pageId="5" pageNumber="681">Canada</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDA3690FF8F7A13FF4B8BD12B55F9F7" blockId="5.[159,781,1186,1571]" pageId="5" pageNumber="681">
|
||
<emphasis id="B911EA82FF8F7A13FF4B8BD12B63FB6D" bold="true" box="[159,383,1186,1209]" pageId="5" pageNumber="681">Revised diagnosis.</emphasis>
|
||
Small-sized and shallow-bodied Eocene catostomid, BD/SL 0.20–0.32, HL/SL 0.30–0.40, D 12–18, A 7–10, pharyngeal teeth about 20 on each pharyngeal bone, frontal bone nearly rectangular with elongated orbital notch, supraorbital present, infraorbital series including shallow and slender lacrimal, posteroventral process of dentary short, opercle broad with OPW slightly larger than OPH, neural complex oval in shape, rib 4 not very robust with length shorter than half the regular rib, PFL about 1/2 PL, PSL sub-equal to PL, hypural 3 not fused to compound centrum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |