233 lines
32 KiB
XML
233 lines
32 KiB
XML
<document id="0FD50FE08F22381096DB04A498F474E1" ID-CLB-Dataset="8107" ID-DOI="10.37520/aemnp.2020.29" ID-GBIF-Dataset="de370c5d-f931-48e0-b4ed-3803bc22f2a4" ID-ISSN="1804-6487" ID-Zenodo-Dep="4489408" ID-ZooBank="5CF7D8D7-2AEF-4AFB-9DB2-80A8FB525144" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612221645004" checkinUser="felipe" docAuthor="Kato, Daichi, Nakamura, Takeyuki & Tachi, Takuji" docDate="2020" docId="03C4F90D745CFFAEFCF244ACFDB2F845" docLanguage="en" docName="ActaEntMusNatPrag.60.2.449-461.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 60 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Epiphragma (Epiphragma) amamiense Kato & Nakamura & Tachi 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="452" masterDocId="FFFD8175745DFFADFF94422CFFA1FD70" masterDocTitle="Taxonomic study of the genus Epiphragma of Japan (Diptera: Limoniidae)" masterLastPageNumber="461" masterPageNumber="449" pageNumber="450" updateTime="1698926196176" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="8E03C9C2241BEF5B3B734BF10F8AB7C1">Taxonomic study of the genus Epiphragma of Japan (Diptera: Limoniidae)</mods:title>
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<mods:name id="37E8CCC1ACA601549710614E198E7462" type="personal">
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<mods:namePart id="E3FDB6C75DE350702F2649468D626DF8">Kato, Daichi</mods:namePart>
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<mods:affiliation id="C563D47C0B06013D2F949B9F2C463530">Echigo-Matsunoyama Museum of Natural Sicences, ‘ Kyororo’, 1712 - & Matsunoyama, Tôkamachi, 942 - 1411, Japan</mods:affiliation>
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<mods:namePart id="5DFA8F325176EBDA3FE39616DE728ACC">Nakamura, Takeyuki</mods:namePart>
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<mods:affiliation id="73D0C0257255F15FF3E307FDB1EDC1FC">The Shirakami Institute for Environmental Sciences, Hirosaki University, Bunkyo-cho</mods:affiliation>
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<mods:namePart id="33B1E226D867B939AE57F0401B29A192">Tachi, Takuji</mods:namePart>
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<mods:affiliation id="2FCBFF0FDB9BD017332BC289847F5BE6">Matsunoyama, Tôkamachi, 942 - 1411, Japan</mods:affiliation>
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<mods:title id="F8299E2CD9B4B823953F3CEE7386DF72">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="2A3A36BB3F1C5877070540DB497DEA51">2020</mods:date>
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<mods:number id="D7D454B15FAA41074657D0038D1F988D">2020-08-01</mods:number>
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<treatment id="03C4F90D745CFFAEFCF244ACFDB2F845" ID-DOI="http://doi.org/10.5281/zenodo.4550933" ID-GBIF-Taxon="177426656" ID-Zenodo-Dep="4550933" LSID="urn:lsid:plazi:treatment:03C4F90D745CFFAEFCF244ACFDB2F845" httpUri="http://treatment.plazi.org/id/03C4F90D745CFFAEFCF244ACFDB2F845" lastPageId="3" lastPageNumber="452" pageId="1" pageNumber="450">
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<subSubSection id="C3771B90745CFFACFCF244ACFAD3FBE8" box="[870,1394,1664,1688]" pageId="1" pageNumber="450" type="nomenclature">
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<paragraph id="8BD2481B745CFFACFCF244ACFAD3FBE8" blockId="1.[870,1394,1664,1688]" box="[870,1394,1664,1688]" pageId="1" pageNumber="450">
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<heading id="D09AFF77745CFFACFCF244ACFAD3FBE8" box="[870,1394,1664,1688]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="450" reason="3">
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<emphasis id="B9199409745CFFACFCF244ACFAD3FBE8" bold="true" box="[870,1394,1664,1688]" italics="true" pageId="1" pageNumber="450">
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<taxonomicName id="4C6D3398745CFFACFCF244ACFAB4FBE8" ID-CoL="8GZWM" authority="Kato & Nakamura & Tachi, 2020" authorityName="Kato & Nakamura & Tachi" authorityYear="2020" box="[870,1301,1664,1688]" class="Insecta" family="Limoniidae" genus="Epiphragma" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="450" phylum="Arthropoda" rank="species" species="amamiense" status="sp. nov." subGenus="Epiphragma">Epiphragma (Epiphragma) amamiense</taxonomicName>
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<taxonomicNameLabel id="A22A2972745CFFACFA8844ADFAD3FBE8" box="[1308,1394,1665,1688]" pageId="1" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3771B90745CFFACFBA444B3FB09FBC2" box="[1072,1192,1695,1714]" pageId="1" pageNumber="450" type="description">
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<paragraph id="8BD2481B745CFFACFBA444B3FB09FBC2" blockId="1.[828,1438,1695,1996]" box="[1072,1192,1695,1714]" pageId="1" pageNumber="450">
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(
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<figureCitation id="1356549E745CFFACFBA244B3FB22FBC2" box="[1078,1155,1695,1714]" captionStart="Figs 1–7" captionStartId="2.[151,188,1229,1248]" captionTargetBox="[154,1427,189,1198]" captionTargetId="figure-13@2.[151,1427,189,1204]" captionTargetPageId="2" captionText="Figs 1–7. Epiphragma amamiense sp. nov. 1 – habitus (male); 2 – wing; 3 – male genitalia in dorsal view; 4 – interbase in dorsal view; 5 – female abdominal sternite 8 and hypogynial valve in ventral view; 6 – furca in dorsal view (upper side direct anteriorly); 7 – female abdominal sternite 10 in ventral view (upper side directs posteriorly). Abbreviations: ad, aedeagus; bs, base of interbase; gc, gonocoxite; hv, hypogynial valve; ib, interbase; ig, inner gonostylus; la, linear apodeme; og, outer gonostylus; rd, rod of interbase; st 8, sternite 8; tg 9, tergite 9. Scale bars = 5 mm (1), 2 mm (2), 0.25 mm (3–7)." figureDoi="http://doi.org/10.5281/zenodo.4489412" httpUri="https://zenodo.org/record/4489412/files/figure.png" pageId="1" pageNumber="450">Figs 1–7</figureCitation>
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,
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<figureCitation id="1356549E745CFFACFB1844B3FB03FBC2" box="[1164,1186,1695,1714]" captionStart="Figs 44–47" captionStartId="11.[151,188,2030,2049]" captionTargetBox="[175,1373,1137,1999]" captionTargetId="figure-553@11.[175,1412,1123,2015]" captionTargetPageId="11" captionText="Figs 44–47. Distributions of Epiphragma species in Japan. 44 – E. gracilistylus Alexander, 1933; 45 – E. amamiense sp. nov. (blue) and E. subfascipenne Alexander, 1920 (red); 46 – E. kempi (blue) and E. subinsigne Alexander, 1920 (red); 47 – E. trichomerum Alexander, 1955." figureDoi="http://doi.org/10.5281/zenodo.4489437" httpUri="https://zenodo.org/record/4489437/files/figure.png" pageId="1" pageNumber="450">45</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3771B90745CFFACFCA844E5FBC7FABC" pageId="1" pageNumber="450" type="materials_examined">
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<paragraph id="8BD2481B745CFFACFCA844E5FBC7FABC" blockId="1.[828,1438,1695,1996]" pageId="1" pageNumber="450">
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<emphasis id="B9199409745CFFACFCA844E5FC66FBAC" bold="true" box="[828,967,1737,1756]" pageId="1" pageNumber="450">
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<typeStatus id="54D6F6B9745CFFACFCA844E5FCCBFBAC" box="[828,874,1737,1756]" pageId="1" pageNumber="450">Type</typeStatus>
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material.
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</emphasis>
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<materialsCitation id="3B054246745CFFACFC5B44E5FA9CFA61" ID-GBIF-Occurrence="3046437301" collectingDate="2013-10-03" collectionCode="BLKU" collectorName="D. Kato" country="Japan" location="Mt. Eboshi" municipality="Amami Island" pageId="1" pageNumber="450" specimenCount="1" typeStatus="hοlοτyρε">
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<typeStatus id="54D6F6B9745CFFACFC5B44E5FB8EFBAC" box="[975,1071,1737,1756]" pageId="1" pageNumber="450">HοLοΤYΡΕ</typeStatus>
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: ♁ (
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),
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<emphasis id="B9199409745CFFACFB3E44E5FBE2FB87" bold="true" pageId="1" pageNumber="450">
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<collectingCountry id="F37A088B745CFFACFB3E44E5FB52FBAC" box="[1194,1267,1737,1756]" name="Japan" pageId="1" pageNumber="450">JAPAN</collectingCountry>
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: NΑΝ SEI I SLΑΝDS: AΜΑΜI I SLΑΝDS:KΑGO SΗIΜΑ:
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</emphasis>
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<collectingMunicipality id="6BB6D261745CFFACFBD044C8FB1DFB87" box="[1092,1212,1764,1783]" pageId="1" pageNumber="450">Amami Island</collectingMunicipality>
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, Setouchi-chô,
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<location id="8EB21EC0745CFFACFAAF44C8FA3BFB87" LSID="urn:lsid:plazi:treatment:03C4F90D745CFFAEFCF244ACFDB2F845:8EB21EC0745CFFACFAAF44C8FA3BFB87" box="[1339,1434,1764,1783]" country="Japan" municipality="Amami Island" name="Mt. Eboshi" pageId="1" pageNumber="450">Mt. Eboshi</location>
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,
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<location id="8EB21EC0745CFFACFCA844D2FC3CFA61" LSID="urn:lsid:plazi:treatment:03C4F90D745CFFAEFCF244ACFDB2F845:8EB21EC0745CFFACFCA844D2FC3CFA61" box="[828,925,1790,1809]" country="Japan" municipality="Amami Island" name="Shinokawa" pageId="1" pageNumber="450">Shinokawa</location>
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,
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<date id="FFD36EDB745CFFACFC3044D2FC50FA61" box="[932,1009,1790,1809]" pageId="1" pageNumber="450" value="2013-10-03">
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<collectingDate id="EF979733745CFFACFC3044D2FC50FA61" box="[932,1009,1790,1809]" pageId="1" pageNumber="450" value="2013-10-03">3.x.2013</collectingDate>
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</date>
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,
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<collectorName id="26982DCD745CFFACFC6C44D2FB9BFA61" box="[1016,1082,1790,1809]" pageId="1" pageNumber="450">D. Kato</collectorName>
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(pinned, one hindleg missing)
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</materialsCitation>
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.PΑ*R ΑΤYΡΕ S (6 ♁♁
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<specimenCount id="9D6B8392745CFFACFCED4535FC09FA5C" box="[889,936,1813,1836]" pageId="1" pageNumber="450" type="female">5 ♀♀</specimenCount>
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):
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<emphasis id="B9199409745CFFACFC284535FCFCFA37" bold="true" pageId="1" pageNumber="450">
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<collectingCountry id="F37A088B745CFFACFC284535FBA5FA5C" box="[956,1028,1817,1836]" name="Japan" pageId="1" pageNumber="450">JAPAN</collectingCountry>
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: NΑΝ SEI I SLΑΝDS: AΜΑΜI I SLΑΝDS: KΑGO SΗIΜΑ:
|
||
</emphasis>
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||
2 ♁♁
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<specimenCount id="9D6B8392745CFFACFC0F4518FC1CFA37" box="[923,957,1840,1863]" pageId="1" pageNumber="450" type="female">1 ♀</specimenCount>
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, Amami Island, Setouchi-chô, Mt. Eboshi, Shinokawa,
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<date id="FFD36EDB745CFFACFCA84562FC29FA11" box="[828,904,1870,1889]" pageId="1" pageNumber="450" value="2013-10-03">3.x.2013</date>
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, D. Kato (
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<collectionCode id="ED7CD0DE745CFFACFC744562FBBEFA11" box="[992,1055,1870,1889]" pageId="1" pageNumber="450">BLKU</collectionCode>
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); 1 ♁
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<specimenCount id="9D6B8392745CFFACFBC74562FBD2FA12" box="[1107,1139,1867,1890]" pageId="1" pageNumber="450" type="female">1 ♀</specimenCount>
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||
, Amami Island, Setouchi-chô, Mt. Yui-dake,
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||
<date id="FFD36EDB745CFFACFC074545FC4DFA0C" box="[915,1004,1897,1916]" pageId="1" pageNumber="450" value="2007-07-05">5.vii.2007</date>
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, H. Makihara (
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<collectionCode id="ED7CD0DE745CFFACFBF84545FB0AFA0C" box="[1132,1195,1897,1916]" pageId="1" pageNumber="450">BLKU</collectionCode>
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||
); 3 ♁♁
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||
<specimenCount id="9D6B8392745CFFACFB7B4545FABFFA0C" box="[1263,1310,1893,1916]" pageId="1" pageNumber="450" type="female">2 ♀♀</specimenCount>
|
||
, Tokunoshima Island, Tokunoshima-chô, Kedoku, Mt. Minada-yama,
|
||
<date id="FFD36EDB745CFFACFA8B45A8FADDFAE7" box="[1311,1404,1924,1943]" pageId="1" pageNumber="450" value="2013-09-30">30.ix.2013</date>
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||
, D. Kato (
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<collectionCode id="ED7CD0DE745CFFACFCE145B2FC14FAC1" box="[885,949,1950,1969]" pageId="1" pageNumber="450">BLKU</collectionCode>
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);
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<specimenCount id="9D6B8392745CFFACFC5C45B2FC4AFAC2" box="[968,1003,1947,1970]" pageId="1" pageNumber="450" type="female">1 ♀</specimenCount>
|
||
, Tokunoshima Island, Amagi-chô, Mikyo Path,
|
||
<date id="FFD36EDB745CFFACFCA84595FC39FABC" box="[828,920,1977,1996]" pageId="1" pageNumber="450" value="1996-04-29">29.iv.1996</date>
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||
, R. Matsumoto (
|
||
<collectionCode id="ED7CD0DE745CFFACFBB24595FBFDFABC" box="[1062,1116,1977,1996]" pageId="1" pageNumber="450">SIHU</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
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<subSubSection id="C3771B90745CFFAEFCA845C8FDB7FEA5" lastPageId="3" lastPageNumber="452" pageId="1" pageNumber="450" type="description">
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<paragraph id="8BD2481B745CFFACFCA845C8FC62F56B" blockId="1.[828,1436,2019,2075]" pageId="1" pageNumber="450">
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<emphasis id="B9199409745CFFACFCA845C8FBA8FA8B" bold="true" box="[828,1033,2019,2043]" pageId="1" pageNumber="450">
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Description.
|
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<emphasis id="B9199409745CFFACFC5E45CFFBA3FA8B" bold="true" box="[970,1026,2019,2043]" italics="true" pageId="1" pageNumber="450">Male</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body length
|
||
<quantity id="4C95E5FE745CFFACFB0745C8FAB5FA8B" box="[1171,1300,2020,2043]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="9.7" metricValueMin="7.8" pageId="1" pageNumber="450" unit="mm" value="8.75" valueMax="9.7" valueMin="7.8">7.8–9.7 mm</quantity>
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||
. Wing length 8.0–
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<quantity id="4C95E5FE745CFFACFCF84A28FC1EF56B" box="[876,959,2052,2075]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="1" pageNumber="450" unit="mm" value="9.1">9.1 mm</quantity>
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.
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</paragraph>
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<caption id="DF121893745FFFAFFF0346E1FA3CF840" ID-DOI="http://doi.org/10.5281/zenodo.4489412" ID-Zenodo-Dep="4489412" httpUri="https://zenodo.org/record/4489412/files/figure.png" pageId="2" pageNumber="451" startId="2.[151,188,1229,1248]" targetBox="[154,1427,189,1198]" targetPageId="2">
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<paragraph id="8BD2481B745FFFAFFF0346E1FA3CF840" blockId="2.[151,1437,1229,1328]" pageId="2" pageNumber="451">
|
||
Figs 1–7.
|
||
<taxonomicName id="4C6D3398745FFFAFFF7F46E1FE1BF990" authority="Kato & Nakamura & Tachi, 2020" authorityName="Kato & Nakamura & Tachi" authorityYear="2020" box="[235,442,1229,1248]" class="Insecta" family="Limoniidae" genus="Epiphragma" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="451" phylum="Arthropoda" rank="species" species="amamiense" status="sp. nov.">
|
||
<emphasis id="B9199409745FFFAFFF7F46E1FE1BF990" box="[235,442,1229,1248]" italics="true" pageId="2" pageNumber="451">Epiphragma amamiense</emphasis>
|
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</taxonomicName>
|
||
<taxonomicNameLabel id="A22A2972745FFFAFFE2B46E1FDA0F990" box="[447,513,1229,1248]" pageId="2" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
|
||
1 – habitus (male); 2 – wing; 3 – male genitalia in dorsal view; 4 – interbase in dorsal view; 5 – female abdominal sternite 8 and hypogynial valve in ventral view; 6 – furca in dorsal view (upper side direct anteriorly); 7 – female abdominal sternite 10 in ventral view (upper side directs posteriorly). Abbreviations: ad, aedeagus; bs, base of interbase; gc, gonocoxite; hv, hypogynial valve; ib, interbase; ig, inner gonostylus; la, linear apodeme; og, outer gonostylus; rd, rod of interbase; st 8, sternite 8; tg 9, tergite 9. Scale bars = 5 mm (1), 2 mm (2), 0.25 mm (3–7).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BD2481B745FFFAFFF214753FDB9FB46" blockId="2.[151,760,1407,2070]" pageId="2" pageNumber="451">Head brown to dark brown with yellowish grey pruinescence; vertex widely darkened along central line, especially at middle; antenna with scape and pedicel dark brown, pedicel sometimes slightly yellowish; flagellomeres 1–2 orange-yellow, subsequent flagellomeres evenly darkened; palpus and mouth parts dark brown.</paragraph>
|
||
<paragraph id="8BD2481B745FFFAFFF214413FB57FAC6" blockId="2.[151,760,1407,2070]" lastBlockId="2.[828,1437,1407,2070]" pageId="2" pageNumber="451">
|
||
Thorax with mesonotum brown to dark brown, anterior end of mesonotum with darker area extending posteriorly and often narrowed into capillary line; lateral sides of prescutum darker, these dark areas extending posteromedially; pruinescence on mesonotum grey and weak, indistinct on anterior half and lateral sides in dorsal view; prescutum with four darker areas on posterior half, sometimes fused to one another; postsutural area largely darkened at middle, outer anterior edge of scutal lobe with small dark spot; scutellum dark brown; mediotergite dark brown with indistinctly paler band at middle; pleuron brown to dark brown variegated with darker areas and pruinescence. Legs with coxae and trochanters yellowish brown, fore coxa with dark area at middle, mid and hind coxae with dark area on lower half; femora, tibiae and tarsi yellow to yellowish brown, each femur with brown band near tip, 0.8–1.0 times as long as width of femur in specimens from Amami Is. and 3.2–4.0 times in those from Tokunoshima Is.; distal segments of tarsomeres slightly darker. Wing (
|
||
<figureCitation id="1356549E745FFFAFFCD047D3FC22FB66" box="[836,899,1535,1558]" captionStart="Figs 1–7" captionStartId="2.[151,188,1229,1248]" captionTargetBox="[154,1427,189,1198]" captionTargetId="figure-13@2.[151,1427,189,1204]" captionTargetPageId="2" captionText="Figs 1–7. Epiphragma amamiense sp. nov. 1 – habitus (male); 2 – wing; 3 – male genitalia in dorsal view; 4 – interbase in dorsal view; 5 – female abdominal sternite 8 and hypogynial valve in ventral view; 6 – furca in dorsal view (upper side direct anteriorly); 7 – female abdominal sternite 10 in ventral view (upper side directs posteriorly). Abbreviations: ad, aedeagus; bs, base of interbase; gc, gonocoxite; hv, hypogynial valve; ib, interbase; ig, inner gonostylus; la, linear apodeme; og, outer gonostylus; rd, rod of interbase; st 8, sternite 8; tg 9, tergite 9. Scale bars = 5 mm (1), 2 mm (2), 0.25 mm (3–7)." figureDoi="http://doi.org/10.5281/zenodo.4489412" httpUri="https://zenodo.org/record/4489412/files/figure.png" pageId="2" pageNumber="451">Fig. 2</figureCitation>
|
||
) 3.7–4.5 times as long as wide, greyish subhyaline with dark brown markings, evenly darkened on costal regions, markings on other areas sometimes with darker margin; four adjacent circular markings present anterior to CuA, second circle enclosing origin of Rs, third circle enclosing around cord, fourth circle enclosing outer end of cell d, first and fourth circles smaller, markings inside second and third circles restricted on veins; cell sc often with small subhyaline spot at distal end; cell d with two subhyaline areas on inner and outer sides; spots at tips of veins near wing tip usually fused at bases with adjacent ones, except for one at tip of R 2; variation of development of wing marking small in same locality. Halter dark brown, apex of knob and base of stem yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BD2481B745FFFAEFCCE4593FE59FCA5" blockId="2.[828,1437,1407,2070]" lastBlockId="3.[151,760,190,1333]" lastPageId="3" lastPageNumber="452" pageId="2" pageNumber="451">
|
||
Abdomen brown, lateral margins of tergites and sternites darker, posterior margin of each segment paler with weak grey pruinescence, widened laterally on tergites; abdominal segments 5–8 darkened, sternites 1–4 paler than tergites 1–4. Male genitalia (
|
||
<figureCitation id="1356549E745EFFAEFEE142F2FE15FD85" box="[373,436,222,245]" captionStart="Figs 1–7" captionStartId="2.[151,188,1229,1248]" captionTargetBox="[154,1427,189,1198]" captionTargetId="figure-13@2.[151,1427,189,1204]" captionTargetPageId="2" captionText="Figs 1–7. Epiphragma amamiense sp. nov. 1 – habitus (male); 2 – wing; 3 – male genitalia in dorsal view; 4 – interbase in dorsal view; 5 – female abdominal sternite 8 and hypogynial valve in ventral view; 6 – furca in dorsal view (upper side direct anteriorly); 7 – female abdominal sternite 10 in ventral view (upper side directs posteriorly). Abbreviations: ad, aedeagus; bs, base of interbase; gc, gonocoxite; hv, hypogynial valve; ib, interbase; ig, inner gonostylus; la, linear apodeme; og, outer gonostylus; rd, rod of interbase; st 8, sternite 8; tg 9, tergite 9. Scale bars = 5 mm (1), 2 mm (2), 0.25 mm (3–7)." figureDoi="http://doi.org/10.5281/zenodo.4489412" httpUri="https://zenodo.org/record/4489412/files/figure.png" pageId="3" pageNumber="452">Fig. 3</figureCitation>
|
||
) with tergite 9 bearing pair of rounded median lobes at posterior margin, bases of lobes somewhat separated; sternite 9 flattened at caudal margin, without conical tubercle; gonocoxite paler brown; gonostyli large, about 1/2 length of gonocoxite; interbase (
|
||
<figureCitation id="1356549E745EFFAEFD244372FD4EFC05" box="[688,751,350,373]" captionStart="Figs 1–7" captionStartId="2.[151,188,1229,1248]" captionTargetBox="[154,1427,189,1198]" captionTargetId="figure-13@2.[151,1427,189,1204]" captionTargetPageId="2" captionText="Figs 1–7. Epiphragma amamiense sp. nov. 1 – habitus (male); 2 – wing; 3 – male genitalia in dorsal view; 4 – interbase in dorsal view; 5 – female abdominal sternite 8 and hypogynial valve in ventral view; 6 – furca in dorsal view (upper side direct anteriorly); 7 – female abdominal sternite 10 in ventral view (upper side directs posteriorly). Abbreviations: ad, aedeagus; bs, base of interbase; gc, gonocoxite; hv, hypogynial valve; ib, interbase; ig, inner gonostylus; la, linear apodeme; og, outer gonostylus; rd, rod of interbase; st 8, sternite 8; tg 9, tergite 9. Scale bars = 5 mm (1), 2 mm (2), 0.25 mm (3–7)." figureDoi="http://doi.org/10.5281/zenodo.4489412" httpUri="https://zenodo.org/record/4489412/files/figure.png" pageId="3" pageNumber="452">Fig. 4</figureCitation>
|
||
) with rod short, not reaching distal end of aedeagus, slightly curved dorsally in whole length, tapered toward tip; base of interbase about as long as rod.
|
||
</paragraph>
|
||
<paragraph id="8BD2481B745EFFAEFF2143F1FD7AFF65" blockId="3.[151,760,190,1333]" pageId="3" pageNumber="452">
|
||
<emphasis id="B9199409745EFFAEFF2143F1FEACFC85" bold="true" box="[181,269,477,501]" pageId="3" pageNumber="452">
|
||
<emphasis id="B9199409745EFFAEFF2143F1FEA7FC85" bold="true" box="[181,262,477,501]" italics="true" pageId="3" pageNumber="452">Female</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body length 8.7–11.0 mm. Wing length 8.0–
|
||
<quantity id="4C95E5FE745EFFAEFD4D43F2FF65FF65" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="3" pageNumber="452" unit="mm" value="9.1">9.1 mm</quantity>
|
||
. Generally resembling male except for terminalia.
|
||
</paragraph>
|
||
<paragraph id="8BD2481B745EFFAEFF214032FDB7FEA5" blockId="3.[151,760,190,1333]" pageId="3" pageNumber="452">
|
||
Ovipositor with sternite 8 (
|
||
<figureCitation id="1356549E745EFFAEFE5A4032FDADFF45" box="[462,524,542,565]" captionStart="Figs 1–7" captionStartId="2.[151,188,1229,1248]" captionTargetBox="[154,1427,189,1198]" captionTargetId="figure-13@2.[151,1427,189,1204]" captionTargetPageId="2" captionText="Figs 1–7. Epiphragma amamiense sp. nov. 1 – habitus (male); 2 – wing; 3 – male genitalia in dorsal view; 4 – interbase in dorsal view; 5 – female abdominal sternite 8 and hypogynial valve in ventral view; 6 – furca in dorsal view (upper side direct anteriorly); 7 – female abdominal sternite 10 in ventral view (upper side directs posteriorly). Abbreviations: ad, aedeagus; bs, base of interbase; gc, gonocoxite; hv, hypogynial valve; ib, interbase; ig, inner gonostylus; la, linear apodeme; og, outer gonostylus; rd, rod of interbase; st 8, sternite 8; tg 9, tergite 9. Scale bars = 5 mm (1), 2 mm (2), 0.25 mm (3–7)." figureDoi="http://doi.org/10.5281/zenodo.4489412" httpUri="https://zenodo.org/record/4489412/files/figure.png" pageId="3" pageNumber="452">Fig. 5</figureCitation>
|
||
) bearing linear apodeme on lateral side of posterior half; when viewed ventrally, sternite 8 excluding hypogynial valve usually slightly shorter than wide, with dark markings on anterior half of lateral margin, and with pair of bacilliform dark markings near base of hypogynial valve, about twice as long as wide in specimens from Amami Is. and more than four times in those from Tokunoshima Is.; furca (
|
||
<figureCitation id="1356549E745EFFAEFDBB40D2FDCEFE65" box="[559,623,766,789]" captionStart="Figs 1–7" captionStartId="2.[151,188,1229,1248]" captionTargetBox="[154,1427,189,1198]" captionTargetId="figure-13@2.[151,1427,189,1204]" captionTargetPageId="2" captionText="Figs 1–7. Epiphragma amamiense sp. nov. 1 – habitus (male); 2 – wing; 3 – male genitalia in dorsal view; 4 – interbase in dorsal view; 5 – female abdominal sternite 8 and hypogynial valve in ventral view; 6 – furca in dorsal view (upper side direct anteriorly); 7 – female abdominal sternite 10 in ventral view (upper side directs posteriorly). Abbreviations: ad, aedeagus; bs, base of interbase; gc, gonocoxite; hv, hypogynial valve; ib, interbase; ig, inner gonostylus; la, linear apodeme; og, outer gonostylus; rd, rod of interbase; st 8, sternite 8; tg 9, tergite 9. Scale bars = 5 mm (1), 2 mm (2), 0.25 mm (3–7)." figureDoi="http://doi.org/10.5281/zenodo.4489412" httpUri="https://zenodo.org/record/4489412/files/figure.png" pageId="3" pageNumber="452">Fig. 6</figureCitation>
|
||
) forked near middle, widest at sinuous posterior part, anterior plate slightly narrowed toward and rounded at anterior end, in specimens from Tokunoshima Is. anterior part wider and weakly sclerotized on lateral part; sternite 10 (
|
||
<figureCitation id="1356549E745EFFAEFD114152FD65FEE5" box="[645,708,894,917]" captionStart="Figs 1–7" captionStartId="2.[151,188,1229,1248]" captionTargetBox="[154,1427,189,1198]" captionTargetId="figure-13@2.[151,1427,189,1204]" captionTargetPageId="2" captionText="Figs 1–7. Epiphragma amamiense sp. nov. 1 – habitus (male); 2 – wing; 3 – male genitalia in dorsal view; 4 – interbase in dorsal view; 5 – female abdominal sternite 8 and hypogynial valve in ventral view; 6 – furca in dorsal view (upper side direct anteriorly); 7 – female abdominal sternite 10 in ventral view (upper side directs posteriorly). Abbreviations: ad, aedeagus; bs, base of interbase; gc, gonocoxite; hv, hypogynial valve; ib, interbase; ig, inner gonostylus; la, linear apodeme; og, outer gonostylus; rd, rod of interbase; st 8, sternite 8; tg 9, tergite 9. Scale bars = 5 mm (1), 2 mm (2), 0.25 mm (3–7)." figureDoi="http://doi.org/10.5281/zenodo.4489412" httpUri="https://zenodo.org/record/4489412/files/figure.png" pageId="3" pageNumber="452">Fig. 7</figureCitation>
|
||
) largely roughly squarish, weakly trilobed at posterior margin, middle lobe wider than lateral ones.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3771B90745EFFAEFF0341F2FE0EF965" pageId="3" pageNumber="452" type="etymology">
|
||
<paragraph id="8BD2481B745EFFAEFF0341F2FE0EF965" blockId="3.[151,760,190,1333]" pageId="3" pageNumber="452">
|
||
<emphasis id="B9199409745EFFAEFF0341F2FEB9FE85" bold="true" box="[151,280,990,1013]" pageId="3" pageNumber="452">Etymology.</emphasis>
|
||
This species is named after the
|
||
<typeStatus id="54D6F6B9745EFFAEFDFA41F2FD3DFE85" box="[622,668,990,1013]" pageId="3" pageNumber="452">type</typeStatus>
|
||
locality, Amami Islands; adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3771B90745EFFAEFF034632FF63F925" pageId="3" pageNumber="452" type="distribution">
|
||
<paragraph id="8BD2481B745EFFAEFF034632FF63F925" blockId="3.[151,760,190,1333]" pageId="3" pageNumber="452">
|
||
<emphasis id="B9199409745EFFAEFF034632FE8BF945" bold="true" box="[151,298,1054,1077]" pageId="3" pageNumber="452">Distribution.</emphasis>
|
||
<collectingCountry id="F37A088B745EFFAEFEBB4632FECFF945" box="[303,366,1054,1077]" name="Japan" pageId="3" pageNumber="452">Japan</collectingCountry>
|
||
, Nansei Islands, Amami Islands (
|
||
<figureCitation id="1356549E745EFFAEFD584632FF14F925" captionStart="Figs 44–47" captionStartId="11.[151,188,2030,2049]" captionTargetBox="[175,1373,1137,1999]" captionTargetId="figure-553@11.[175,1412,1123,2015]" captionTargetPageId="11" captionText="Figs 44–47. Distributions of Epiphragma species in Japan. 44 – E. gracilistylus Alexander, 1933; 45 – E. amamiense sp. nov. (blue) and E. subfascipenne Alexander, 1920 (red); 46 – E. kempi (blue) and E. subinsigne Alexander, 1920 (red); 47 – E. trichomerum Alexander, 1955." figureDoi="http://doi.org/10.5281/zenodo.4489437" httpUri="https://zenodo.org/record/4489437/files/figure.png" pageId="3" pageNumber="452">Fig. 45</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3771B90745EFFAEFF034672FDB2F845" pageId="3" pageNumber="452" type="discussion">
|
||
<paragraph id="8BD2481B745EFFAEFF034672FDB2F845" blockId="3.[151,760,190,1333]" pageId="3" pageNumber="452">
|
||
<emphasis id="B9199409745EFFAEFF034672FEA7F905" bold="true" box="[151,262,1118,1141]" pageId="3" pageNumber="452">Remarks.</emphasis>
|
||
This new species is somewhat similar to
|
||
<taxonomicName id="4C6D3398745EFFAEFD5E4672FDB8F9E5" authority="Mao & Yang, 2009" authorityName="Mao & Yang" authorityYear="2009" class="Insecta" family="Limoniidae" genus="Epiphragma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="mediale">
|
||
<emphasis id="B9199409745EFFAEFD5E4672FEF1F9E5" italics="true" pageId="3" pageNumber="452">Epiphragma mediale</emphasis>
|
||
<bibRefCitation id="EFFC35EA745EFFAEFEC04652FDB8F9E5" author="MAO M. & YANG D." box="[340,537,1150,1173]" pageId="3" pageNumber="452" pagination="44 - 56" refId="ref12173" refString="MAO M. & YANG D. 2009: New species of the genus Epiphragma Osten Sacken from Yunnan, southwest China (Diptera, Limoniidae). Zootaxa 2121: 44 - 56." type="journal article" year="2009">Mao & Yang, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F37A088B745EFFAEFDC34652FD37F9E5" box="[599,662,1150,1173]" name="China" pageId="3" pageNumber="452">China</collectingCountry>
|
||
in regard to wing markings and the shape of male genitalia, but is differentiated from them by the following characters: each femur with one short band (two bands in
|
||
<taxonomicName id="4C6D3398745EFFAEFDAB46F2FD0EF985" authorityName="Mao & Yang" authorityYear="2009" box="[575,687,1246,1269]" class="Insecta" family="Limoniidae" genus="Epiphragma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="mediale">
|
||
<emphasis id="B9199409745EFFAEFDAB46F2FD0EF985" box="[575,687,1246,1269]" italics="true" pageId="3" pageNumber="452">E. mediale</emphasis>
|
||
</taxonomicName>
|
||
); male genitalia with rod of interbase almost straight in whole length (bent near tip in
|
||
<taxonomicName id="4C6D3398745EFFAEFE054732FDA3F845" authorityName="Mao & Yang" authorityYear="2009" box="[401,514,1310,1333]" class="Insecta" family="Limoniidae" genus="Epiphragma" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="452" phylum="Arthropoda" rank="species" species="mediale">
|
||
<emphasis id="B9199409745EFFAEFE054732FDA3F845" box="[401,514,1310,1333]" italics="true" pageId="3" pageNumber="452">E. mediale</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |