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<mods:title id="78D849D92A181F80643F360D6C87D94A">Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera</mods:title>
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<heading id="D09A770CFFB58D027AD207D8FE7A928F" bold="true" box="[151,474,152,178]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<taxonomicName id="4C6DBBE3FFB58D027AD207D8FE7A928F" ID-CoL="63GPY" authority="Cockerell, 1908" authorityName="Cockerell" authorityYear="1908" box="[151,474,152,178]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFB58D027AD207D8FE7A928F" bold="true" box="[151,474,152,178]" pageId="54" pageNumber="55">
<emphasis id="B9191C72FFB58D027AD207D8FEB1928F" bold="true" box="[151,273,152,178]" italics="true" pageId="54" pageNumber="55">Osmylidia</emphasis>
<bibRefCitation id="EFFCBD91FFB58D027B5D07D8FE7A928F" author="Cockerell" box="[280,474,152,178]" firstAuthor="Cockerell" pageId="54" pageNumber="55" pagination="341 - 342" refId="ref47949" refString="Cockerell, T. D. A. (1908) Fossil Osmylidae (Neuroptera) in America. Canadian Entomologist, 40, 341 - 342. https: // doi. org / 10.4039 / Ent 40341 - 10" type="journal article" year="1908">Cockerell, 1908</bibRefCitation>
</emphasis>
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<paragraph id="8BD2C060FFB58D027AD2079EFBA992C9" blockId="54.[151,1033,222,245]" box="[151,1033,222,245]" pageId="54" pageNumber="55">
(=
<taxonomicName id="4C6DBBE3FFB58D027AF1079EFE4392C9" authority="Kruger 1913: 19" authorityName="Kruger" authorityPageNumber="19" authorityYear="1913" box="[180,483,222,244]" class="Insecta" family="Osmylidae" genus="Oligosmylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neuroptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFB58D027AF1079EFE9392C9" box="[180,307,222,244]" italics="true" pageId="54" pageNumber="55">Oligosmylus</emphasis>
<bibRefCitation id="EFFCBD91FFB58D027B7C079FFE1D92C9" author="Kruger" box="[313,445,222,244]" firstAuthor="Kruger" pageId="54" pageNumber="55" pagination="3 - 123" refId="ref49715" refString="Kruger, L. (1913 a) Osmylidae. Beitrage zu einer Monographie der Neuropteren-Familie der Osmyliden. II. Charakteristik der Familie, Unterfamilien und Gattungen auf Grund des Geaders. Stettiner Entomologische Zeitung, 74, 3 - 123." type="journal article" year="1913">Krüger 1913</bibRefCitation>
: 19
</taxonomicName>
.
<typeStatus id="54D67EC2FFB58D027BA8079FFDBF92C9" box="[493,543,223,244]" pageId="54" pageNumber="55">Type</typeStatus>
species:
<taxonomicName id="4C6DBBE3FFB58D027839079EFBA292C9" authority="Scudder, 1890: 164" authorityName="Scudder" authorityPageNumber="164" authorityYear="1890" box="[636,1026,222,245]" class="Insecta" family="Osmylidae" genus="Osmylus" kingdom="Animalia" order="Neuroptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="requietus">
<emphasis id="B9191C72FFB58D027839079EFC9692C8" box="[636,822,222,245]" italics="true" pageId="54" pageNumber="55">Osmylus requietus</emphasis>
<bibRefCitation id="EFFCBD91FFB58D027978079EFC7292C9" author="Scudder" box="[829,978,222,244]" firstAuthor="Scudder" pageId="54" pageNumber="55" pagination="1 - 663" refId="ref54175" refString="Scudder, S. H. (1890) Tertiary Insects of North America. Report of the United States Geological and Geographical Survey of the Territories. Report of the United States Geological [and Geographical] Survey of the Territories, 13, 1 - 663." type="journal article" year="1890">Scudder, 1890</bibRefCitation>
: 164
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)
</paragraph>
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<paragraph id="8BD2C060FFB58D027AD2065CFCF3930F" blockId="54.[151,851,284,306]" box="[151,851,284,306]" pageId="54" pageNumber="55">
<emphasis id="B9191C72FFB58D027AD2065CFE82930F" bold="true" box="[151,290,284,306]" pageId="54" pageNumber="55">
<typeStatus id="54D67EC2FFB58D027AD2065CFF6C930F" box="[151,204,284,306]" pageId="54" pageNumber="55">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6DBBE3FFB58D027B6C065CFD0F930F" authority="Scudder, 1890: 164" authorityName="Scudder" authorityPageNumber="164" authorityYear="1890" box="[297,687,284,306]" class="Insecta" family="Osmylidae" genus="Osmylus" kingdom="Animalia" order="Neuroptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="species" species="requietus">
<emphasis id="B9191C72FFB58D027B6C065CFE44930F" box="[297,484,284,306]" italics="true" pageId="54" pageNumber="55">Osmylus requietus</emphasis>
<bibRefCitation id="EFFCBD91FFB58D027BAF065CFDDF930F" author="Scudder" box="[490,639,284,306]" firstAuthor="Scudder" pageId="54" pageNumber="55" pagination="1 - 663" refId="ref54175" refString="Scudder, S. H. (1890) Tertiary Insects of North America. Report of the United States Geological and Geographical Survey of the Territories. Report of the United States Geological [and Geographical] Survey of the Territories, 13, 1 - 663." type="journal article" year="1890">Scudder, 1890</bibRefCitation>
: 164
</taxonomicName>
(by monotypy).
</paragraph>
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<subSubSection id="C37793EBFFB58D027AD2061EFDDA9072" pageId="54" pageNumber="55" type="diagnosis">
<paragraph id="8BD2C060FFB58D027AD2061EFDDA9072" blockId="54.[151,1437,350,771]" pageId="54" pageNumber="55">
<emphasis id="B9191C72FFB58D027AD2061EFEAA934A" bold="true" box="[151,266,350,375]" pageId="54" pageNumber="55">Diagnosis</emphasis>
. Wings ovoid, mostly hyaline; FW costal area relatively narrow, subcostal veinlets simple throughout, sub-parallel in arrangement, relatively widely spaced basally, closely spaced distally; area behind Sc+RA distal to pterostigma relatively large, lacking crossveins to RP; FW RP with 910 branches, RP1 originating relatively close to origin of RP; both wings with two distinct gradate series, arranged subparallel, outer gradate series slightly longer than inner series, relatively few radial crossveins in basal half of wings; FW M forked just basal to origin of RP1; FW MA and MP with few distal branches (nearly dichotomous); FW CuP with more pectinate branches than CuA; FW A1 with relatively few branches.
</paragraph>
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<subSubSection id="C37793EBFFB58D027A82051AFB0D913E" pageId="54" pageNumber="55" type="discussion">
<paragraph id="8BD2C060FFB58D027A82051AFB0D913E" blockId="54.[151,1437,350,771]" pageId="54" pageNumber="55">
<emphasis id="B9191C72FFB58D027A82051AFEE8904E" bold="true" box="[199,328,602,627]" pageId="54" pageNumber="55">Comments</emphasis>
. Known from several, mostly incomplete compression fossils where the forewing is clearly represented, but the body and hind wing are not clear (
<bibRefCitation id="EFFCBD91FFB58D027947053EFC1B90AA" author="Cockerell" box="[770,955,638,663]" firstAuthor="Cockerell" pageId="54" pageNumber="55" pagination="341 - 346" refId="ref47990" refString="Cockerell, T. D. A. (1913) Some fossil insects from Florissant, Colorado. Proceedings of the United States National Museum, 44, 341 - 346. https: // doi. org / 10.5479 / si. 00963801.44 - 1955.341" type="journal article" year="1913">Cockerell, 1913</bibRefCitation>
,
<bibRefCitation id="EFFCBD91FFB58D027982053FFBA290AA" author="Cockerell" box="[967,1026,639,663]" firstAuthor="Cockerell" pageId="54" pageNumber="55" pagination="714 - 724" refId="ref48037" refString="Cockerell, T. D. A. (1914) New and little-known insects from the Miocene of Florissant, Colorado. Journal of Geology, 22, 714 - 724. https: // doi. org / 10.1086 / 622186" type="journal article" year="1914">1914</bibRefCitation>
;
<bibRefCitation id="EFFCBD91FFB58D027E4A053FFB6790AA" author="Carpenter" box="[1039,1223,639,663]" firstAuthor="Carpenter" pageId="54" pageNumber="55" pagination="753 - 760" refId="ref47880" refString="Carpenter, F. M. (1943) Osmylidae of the Florissant shales, Colorado (Insecta-Neuroptera). American Journal of Science, 241, 753 - 760. https: // doi. org / 10.2475 / ajs. 241.12.753" type="journal article" year="1943">Carpenter, 1943</bibRefCitation>
).
<bibRefCitation id="EFFCBD91FFB58D027E9E053FFA3C90AA" author="Carpenter" box="[1243,1436,639,663]" firstAuthor="Carpenter" pageId="54" pageNumber="55" pagination="753 - 760" refId="ref47880" refString="Carpenter, F. M. (1943) Osmylidae of the Florissant shales, Colorado (Insecta-Neuroptera). American Journal of Science, 241, 753 - 760. https: // doi. org / 10.2475 / ajs. 241.12.753" type="journal article" year="1943">Carpenter (1943)</bibRefCitation>
noted a close similarity to
<taxonomicName id="4C6DBBE3FFB58D027B9405E2FDC59086" box="[465,613,674,699]" class="Insecta" family="Osmylidae" genus="Gryposmylus" kingdom="Animalia" order="Neuroptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFB58D027B9405E2FDC59086" box="[465,613,674,699]" italics="true" pageId="54" pageNumber="55">Gryposmylus</emphasis>
</taxonomicName>
, but
<taxonomicName id="4C6DBBE3FFB58D0278E105E2FCB79086" authorityName="Cockerell" authorityYear="1908" box="[676,791,674,699]" class="Insecta" family="Osmylidae" genus="Osmylidia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFB58D0278E105E2FCB79086" box="[676,791,674,699]" italics="true" pageId="54" pageNumber="55">Osmylidia</emphasis>
</taxonomicName>
shows greater similarity to genera such as
<taxonomicName id="4C6DBBE3FFB58D027F5305E4FAC99086" box="[1302,1385,676,699]" class="Insecta" family="Osmylidae" genus="Lysmus" kingdom="Animalia" order="Neuroptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFB58D027F5305E4FAC99086" box="[1302,1385,676,699]" italics="true" pageId="54" pageNumber="55">Lysmus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DBBE3FFB58D027AD20586FE8290E2" authorityName="Kruger" authorityYear="1913" box="[151,290,710,735]" class="Insecta" family="Osmylidae" genus="Protosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="54" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B9191C72FFB58D027AD20586FE8290E2" box="[151,290,710,735]" italics="true" pageId="54" pageNumber="55">Protosmylus</emphasis>
</taxonomicName>
based on characters of the forewing such as absence of forking of subcostal veinlets, number and arrangement of gradate series and relative proportions of branching patterns in CuA and CuP.
</paragraph>
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