213 lines
26 KiB
XML
213 lines
26 KiB
XML
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<mods:title id="CA427F9B63D667E2512C4B656FAE53B5">Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera</mods:title>
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<mods:namePart id="9BA11A5FB6422D0759623FC5A78FE6CD">Winterton, Shaun L.</mods:namePart>
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<mods:namePart id="227F243498786AFD77B50BDC7B1FF6D4">Martins, Caleb Califre</mods:namePart>
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<mods:namePart id="A48605166DECB99E4C93AE23197BE20A">Makarkin, Vladimir</mods:namePart>
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<mods:namePart id="982E30D58BF311C775F2FD50B573C347">Wang, Yongjie</mods:namePart>
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<mods:date id="EF35B3A01F9895C88E784780AD9A8D3B">2019</mods:date>
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<treatment id="03C47176FF898D3E7AD207D8FAFF976E" ID-GBIF-Taxon="156146955" LSID="urn:lsid:plazi:treatment:03C47176FF898D3E7AD207D8FAFF976E" httpUri="http://treatment.plazi.org/id/03C47176FF898D3E7AD207D8FAFF976E" lastPageNumber="11" pageId="10" pageNumber="11">
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<subSubSection id="C37793EBFF898D3E7AD207D8FB97928F" box="[151,1079,152,178]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BD2C060FF898D3E7AD207D8FB97928F" blockId="10.[151,1079,152,178]" box="[151,1079,152,178]" pageId="10" pageNumber="11">
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<heading id="D09A770CFF898D3E7AD207D8FB97928F" bold="true" box="[151,1079,152,178]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="B9191C72FF898D3E7AD207D8FB97928F" bold="true" box="[151,1079,152,178]" pageId="10" pageNumber="11">
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<taxonomicName id="4C6DBBE3FF898D3E7AD207D8FDE1928F" ID-CoL="6NF" authority="Riek, 1953" authorityName="Riek" authorityYear="1953" box="[151,577,152,178]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">
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ARCHEOSMYLIDAE
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<bibRefCitation id="EFFCBD91FF898D3E7BF807D8FDE1928F" author="Riek" box="[445,577,152,178]" firstAuthor="Riek" pageId="10" pageNumber="11" pagination="55 - 87" refId="ref53960" refString="Riek, E. F. (1953) Fossil mecopteroid insects from the Upper Permian of New South Wales. Records of the Australian Museum, 23, 55 - 87. https: // doi. org / 10.3853 / j. 0067 - 1975.23.1953.621" type="journal article" year="1953">Riek, 1953</bibRefCitation>
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</taxonomicName>
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(Permo-Triassic ‘stem lance lacewings’)
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37793EBFF898D3E7AD2079EFDDB92CA" box="[151,635,222,247]" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="8BD2C060FF898D3E7AD2079EFDDB92CA" blockId="10.[151,635,222,247]" box="[151,635,222,247]" pageId="10" pageNumber="11">
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<emphasis id="B9191C72FF898D3E7AD2079EFE8492CA" bold="true" box="[151,292,222,247]" pageId="10" pageNumber="11">
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<typeStatus id="54D67EC2FF898D3E7AD2079EFF7292CA" box="[151,210,222,247]" pageId="10" pageNumber="11">Type</typeStatus>
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genus.
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</emphasis>
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<taxonomicName id="4C6DBBE3FF898D3E7B6E079EFDD692CA" authority="Riek, 1953: 85" authorityName="Riek" authorityPageNumber="85" authorityYear="1953" box="[299,630,222,247]" class="Insecta" family="Archeosmylidae" genus="Archeosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FF898D3E7B6E079EFE6792CA" box="[299,455,222,247]" italics="true" pageId="10" pageNumber="11">Archeosmylus</emphasis>
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<bibRefCitation id="EFFCBD91FF898D3E7B8B079EFDED92CA" author="Riek" box="[462,589,222,247]" firstAuthor="Riek" pageId="10" pageNumber="11" pagination="55 - 87" refId="ref53960" refString="Riek, E. F. (1953) Fossil mecopteroid insects from the Upper Permian of New South Wales. Records of the Australian Museum, 23, 55 - 87. https: // doi. org / 10.3853 / j. 0067 - 1975.23.1953.621" type="journal article" year="1953">Riek, 1953</bibRefCitation>
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: 85
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37793EBFF898D3E7AD20666FEA4902A" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="8BD2C060FF898D3E7AD20666FEA4902A" blockId="10.[151,1437,294,1363]" pageId="10" pageNumber="11">
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<emphasis id="B9191C72FF898D3E7AD20666FEAA9302" bold="true" box="[151,266,294,319]" pageId="10" pageNumber="11">Diagnosis</emphasis>
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. Forewing ovoid, costal area relatively narrow, wider basally, subcostal veinlets mostly simple, more widely spaced basally; subcostal crossveins not detected (due to poor preservation); Sc and RA fused distally and thence slightly curved posteriorly, entering wing margin well before apex; RP with numerous branches (more than nine); crossveins in radial area scarce, dense end-twigging along posterior margin of wing; M fork into MA and MP distal to the origin of RP1; MA and MP mostly dichotomously branched; CuA, CuP and 1A few-branched, poorly pectinate with irregular forking, branches with distal angle relatively acute; HW known only from small fragment.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37793EBFF898D3E7A820562FAFF976E" pageId="10" pageNumber="11" type="discussion">
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<paragraph id="8BD2C060FF898D3E7A820562FC429712" blockId="10.[151,1437,294,1363]" pageId="10" pageNumber="11">
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<emphasis id="B9191C72FF898D3E7A820562FEE89006" bold="true" box="[199,328,546,571]" pageId="10" pageNumber="11">Comments</emphasis>
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.
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<taxonomicName id="4C6DBBE3FF898D3E7B1F0562FDB19006" authorityName="Riek" authorityYear="1953" box="[346,529,546,571]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Archeosmylidae</taxonomicName>
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has been considered a synonym of
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<taxonomicName id="4C6DBBE3FF898D3E79870562FBC69006" authorityName="Tillyard" authorityYear="1922" box="[962,1126,546,571]" class="Insecta" family="Permithonidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Permithonidae</taxonomicName>
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by previous authors (e.g.,
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<bibRefCitation id="EFFCBD91FF898D3E7AD20506FEE59062" author="Whalley" box="[151,325,582,607]" firstAuthor="Whalley" pageId="10" pageNumber="11" pagination="45 - 63" refId="ref55030" refString="Whalley, P. E. S. (1988) Mesozoic Neuroptera and Raphidioptera (Insecta) in Britain. Bulletin of the British Museum of Natural History, Geology, 44, 45 - 63." type="journal article" year="1988">Whalley, 1988</bibRefCitation>
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;
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<bibRefCitation id="EFFCBD91FF898D3E7B1D0506FD199062" author="Makarkin & Archibald" box="[344,697,582,607]" firstAuthor="Makarkin" pageId="10" pageNumber="11" pagination="171 - 180" refId="ref50748" refString="Makarkin, V. N. & Archibald, S. B. (2003) Family affinity of the genus Palaeopsychops Andersen with description of a new species from the Early Eocene of British Columbia (Neuroptera: Polystoechotidae). Annals of the Entomological Society of America, 96, 171 - 180. https: // doi. org / 10.1603 / 0013 - 8746 (2003) 096 [0171: FAOTGP] 2.0. CO; 2" type="journal article" year="2003">Makarkin & Archibald, 2003</bibRefCitation>
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;
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<bibRefCitation id="EFFCBD91FF898D3E78890506FC599062" author="Engel & Grimaldi" box="[716,1017,582,607]" firstAuthor="Engel" pageId="10" pageNumber="11" pagination="1 - 86" refId="ref48270" refString="Engel, M. S. & Grimaldi, D. A. (2008) Diverse Neuropterida in Cretaceous amber, with particular reference to the paleofauna of Myanmar (Insecta). Nova Supplementa Entomologica, 20, 1 - 86." type="journal article" year="2008">Engel & Grimaldi, 2008</bibRefCitation>
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). We agree with the placement of
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<taxonomicName id="4C6DBBE3FF898D3E7AD2052AFEEF90BE" authorityName="Riek" authorityYear="1953" box="[151,335,618,643]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Archeosmylidae</taxonomicName>
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as closely related to
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<taxonomicName id="4C6DBBE3FF898D3E780A052AFD6990BE" authorityName="Leach" authorityYear="1815" box="[591,713,618,643]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Osmylidae</taxonomicName>
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and
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<taxonomicName id="4C6DBBE3FF898D3E794D052AFC1B90BE" authorityName="Ren & Yin" authorityYear="2003" box="[776,955,618,643]" class="Insecta" family="Saucrosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Saucrosmylidae</taxonomicName>
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and it indeed likely represents a Permo- Triassic-aged stem group for these younger crown group families (
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<bibRefCitation id="EFFCBD91FF898D3E79C005CEFBA6909A" author="Riek" box="[901,1030,654,679]" firstAuthor="Riek" pageId="10" pageNumber="11" pagination="755 - 789" refId="ref54051" refString="Riek, E. F. (1976) New Upper Permian insects from Natal, South Africa. Annals of the Natal Museum, 22, 755 - 789." type="journal article" year="1976">Riek, 1976</bibRefCitation>
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;
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<bibRefCitation id="EFFCBD91FF898D3E7E5405CEFAA7909A" author="Makarkin, V. N. & Yang, Q. & Ren, D." box="[1041,1287,654,679]" pageId="10" pageNumber="11" pagination="209 - 214" refId="ref50998" refString="Makarkin, V. N., Yang, Q. & Ren, D. (2014) A new basal osmylid neuropteran insect from the Middle Jurassic of China linking Osmylidae to the Permian-Triassic Archeosmylidae. Acta Palaeontologica Polonica, 59, 209 - 214." type="journal article" year="2014">
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Makarkin
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<emphasis id="B9191C72FF898D3E7EC005CFFB1E909A" box="[1157,1214,654,679]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
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||
, 2014
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</bibRefCitation>
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||
;
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||
<bibRefCitation id="EFFCBD91FF898D3E7F5605CEFF7490F6" author="Engel" firstAuthor="Engel" pageId="10" pageNumber="11" pagination="531 - 551" refId="ref48314" refString="Engel, M. S., Winterton, S. L. & Breitkreuz, L. C. V. (2018) Phylogeny and Evolution of Neuropterida: Where have wings of lace taken us? Annual Review of Entomology, 63, 531 - 551. https: // doi. org / 10.1146 / annurev-ento- 020117 - 043127" type="journal article" year="2018">
|
||
Engel
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||
<emphasis id="B9191C72FF898D3E7F1905CFFA35909A" box="[1372,1429,654,679]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
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, 2018
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</bibRefCitation>
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). All three families have similar venation and differ mainly in the forewing configuration of the posterior vein fields. The veins CuA, CuP and A1 are weakly pectinate (weakly divergent branches) with few branches in
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<taxonomicName id="4C6DBBE3FF898D3E7AD205BAFEEF912E" authorityName="Riek" authorityYear="1953" box="[151,335,762,787]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Archeosmylidae</taxonomicName>
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while in osmylid genera these veins are more strongly pectinate with the branches strongly divergent and frequently arranged in a palisade-like pattern (see
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<bibRefCitation id="EFFCBD91FF898D3E79C3045EFB26910A" author="Makarkin, V. N. & Yang, Q. & Ren, D." box="[902,1158,798,823]" pageId="10" pageNumber="11" pagination="209 - 214" refId="ref50998" refString="Makarkin, V. N., Yang, Q. & Ren, D. (2014) A new basal osmylid neuropteran insect from the Middle Jurassic of China linking Osmylidae to the Permian-Triassic Archeosmylidae. Acta Palaeontologica Polonica, 59, 209 - 214." type="journal article" year="2014">
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Makarkin
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<emphasis id="B9191C72FF898D3E79B9045FFB98910A" box="[1020,1080,798,823]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
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, 2014
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</bibRefCitation>
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). A condition similar to
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<taxonomicName id="4C6DBBE3FF898D3E7AD20402FEEF9166" authorityName="Riek" authorityYear="1953" box="[151,335,834,859]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Archeosmylidae</taxonomicName>
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is found in
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<taxonomicName id="4C6DBBE3FF898D3E7BB60402FD099166" authorityName="Ren & Yin" authorityYear="2003" box="[499,681,834,859]" class="Insecta" family="Saucrosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Saucrosmylidae</taxonomicName>
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, but these families differ widely in other venation characters, principally associated with the larger wings of the latter and the consequent proliferation of secondary veins and crossveins. All three families also share the condition where FW Sc is fused with RA apically, ending on costal margin well before the wing apex. Some fossil and living taxa now placed in families such as
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<taxonomicName id="4C6DBBE3FF898D3E7EA204EEFACA91FA" authorityName="Rambur" authorityYear="1842" box="[1255,1386,942,967]" class="Insecta" family="Nymphidae" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Nymphidae</taxonomicName>
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and
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<taxonomicName id="4C6DBBE3FF898D3E7AD20492FE8091D6" authorityName="Martynova" authorityYear="1949" box="[151,288,978,1003]" class="Insecta" family="Osmylitidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Osmylitidae</taxonomicName>
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were previously considered as being in Osmyloidea (e.g.,
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<taxonomicName id="4C6DBBE3FF898D3E79840492FBEE91D6" box="[961,1102,978,1003]" class="Insecta" family="Osmylidae" genus="Cratosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B9191C72FF898D3E79840492FBEE91D6" box="[961,1102,978,1003]" italics="true" pageId="10" pageNumber="11">Cratosmylus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFFCBD91FF898D3E7E380492FA3091D6" author="Myskowiak" box="[1149,1424,978,1003]" firstAuthor="Myskowiak" pageId="10" pageNumber="11" pagination="27 - 33" refId="ref51837" refString="Myskowiak, J., Escuilli, F. & Nel, A. (2015) A new Osmylidae (Insecta, Neuroptera) from the Lower Cretaceous Crato Formation in Brazil. Cretaceous Research, 54, 27 - 33. https: // doi. org / 10.1016 / j. cretres. 2014.11.012" type="journal article" year="2015">
|
||
Myskowiak
|
||
<emphasis id="B9191C72FF898D3E7F4F0494FAEB91D6" box="[1290,1355,978,1003]" italics="true" pageId="10" pageNumber="11">et al.,</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
), but all have the fused with Sc+RA vein joining the wing margin beyond the wing apex.
|
||
<taxonomicName id="4C6DBBE3FF898D3E7EE004B6FAFD9632" authorityName="Riek" authorityYear="1953" box="[1189,1373,1014,1039]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Archeosmylidae</taxonomicName>
|
||
have relatively few crossveins in the radial field, which are typically numerous in
|
||
<taxonomicName id="4C6DBBE3FF898D3E7E6C035AFB03960E" authorityName="Leach" authorityYear="1815" box="[1065,1187,1050,1075]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Osmylidae</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6DBBE3FF898D3E7EA7035AFA39960E" authorityName="Ren & Yin" authorityYear="2003" box="[1250,1433,1050,1075]" class="Insecta" family="Saucrosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Saucrosmylidae</taxonomicName>
|
||
. Interestingly, the osmylid genus
|
||
<taxonomicName id="4C6DBBE3FF898D3E7841037EFD22966A" box="[516,642,1086,1111]" class="Insecta" family="Osmylidae" genus="Sinosmylus" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FF898D3E7841037EFD22966A" box="[516,642,1086,1111]" italics="true" pageId="10" pageNumber="11">Sinosmylus</emphasis>
|
||
</taxonomicName>
|
||
is a typical member of
|
||
<taxonomicName id="4C6DBBE3FF898D3E79CE037EFBA4966A" authorityName="Leach" authorityYear="1815" box="[907,1028,1086,1111]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subFamily" subFamily="Osmylinae">Osmylinae</taxonomicName>
|
||
but has secondarily convergent lack of radial crossveins except for two gradate series.
|
||
<bibRefCitation id="EFFCBD91FF898D3E78990322FCC59646" author="Riek" box="[732,869,1122,1147]" firstAuthor="Riek" pageId="10" pageNumber="11" pagination="755 - 789" refId="ref54051" refString="Riek, E. F. (1976) New Upper Permian insects from Natal, South Africa. Annals of the Natal Museum, 22, 755 - 789." type="journal article" year="1976">Riek (1976)</bibRefCitation>
|
||
suggested that
|
||
<taxonomicName id="4C6DBBE3FF898D3E7E5E0322FB729646" authorityName="Riek" authorityYear="1953" box="[1051,1234,1122,1147]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Archeosmylidae</taxonomicName>
|
||
were stem group representatives of
|
||
<taxonomicName id="4C6DBBE3FF898D3E7B2303C6FE4296A2" authorityName="Leach" authorityYear="1815" box="[358,482,1158,1183]" class="Insecta" family="Osmylidae" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Osmylidae</taxonomicName>
|
||
, although the absence of nygmata in
|
||
<taxonomicName id="4C6DBBE3FF898D3E79CE03C6FBE296A2" authorityName="Riek" authorityYear="1953" box="[907,1090,1158,1183]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Archeosmylidae</taxonomicName>
|
||
is anomalous.
|
||
<bibRefCitation id="EFFCBD91FF898D3E7EAB03C6FF4296FE" author="Makarkin, V. N. & Yang, Q. & Ren, D." pageId="10" pageNumber="11" pagination="209 - 214" refId="ref50998" refString="Makarkin, V. N., Yang, Q. & Ren, D. (2014) A new basal osmylid neuropteran insect from the Middle Jurassic of China linking Osmylidae to the Permian-Triassic Archeosmylidae. Acta Palaeontologica Polonica, 59, 209 - 214." type="journal article" year="2014">
|
||
Makarkin
|
||
<emphasis id="B9191C72FF898D3E7F2603C7FA3C96A2" box="[1379,1436,1158,1183]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
(2014)
|
||
</bibRefCitation>
|
||
discussed in detail the taxonomic composition and morphological characterisation of
|
||
<taxonomicName id="4C6DBBE3FF898D3E7E9B03EAFA3996FE" authorityName="Riek" authorityYear="1953" box="[1246,1433,1194,1219]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Archeosmylidae</taxonomicName>
|
||
. Nygmata are widely considered as plesiomorphic for
|
||
<taxonomicName id="4C6DBBE3FF898D3E78BD038FFCD996DB" box="[760,889,1231,1254]" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="order">Neuroptera</taxonomicName>
|
||
, being found in many distantly related lacewing families. The lack of nygmata in
|
||
<taxonomicName id="4C6DBBE3FF898D3E785B03B2FD769736" authorityName="Riek" authorityYear="1953" box="[542,726,1266,1291]" class="Insecta" family="Archeosmylidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="family">Archeosmylidae</taxonomicName>
|
||
may represent a secondary loss in the clade, or simply even incomplete preservation in the compression fossil specimens of this family.
|
||
</paragraph>
|
||
<paragraph id="8BD2C060FF898D3E7A82027AFAFF976E" blockId="10.[151,1437,294,1363]" box="[199,1375,1338,1363]" pageId="10" pageNumber="11">
|
||
<emphasis id="B9191C72FF898D3E7A82027AFE2C976E" bold="true" box="[199,396,1338,1363]" pageId="10" pageNumber="11">Genera included</emphasis>
|
||
.
|
||
<taxonomicName id="4C6DBBE3FF898D3E7BDE027AFDD7976E" authority="Riek" authorityName="Riek" authorityYear="1953" box="[411,631,1338,1363]" class="Insecta" family="Archeosmylidae" genus="Archeosmylus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FF898D3E7BDE027AFD97976E" box="[411,567,1338,1363]" italics="true" pageId="10" pageNumber="11">Archeosmylus</emphasis>
|
||
Riek
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C6DBBE3FF898D3E78C7027AFBD6976E" authority="Ponomarenko & Shcherbakov" authorityName="Ponomarenko & Shcherbakov" authorityYear="2004" box="[642,1142,1338,1363]" class="Insecta" family="Archeosmylidae" genus="Babykamenia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FF898D3E78C7027AFCBA976E" box="[642,794,1338,1363]" italics="true" pageId="10" pageNumber="11">Babykamenia</emphasis>
|
||
Ponomarenko & Shcherbakov
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C6DBBE3FF898D3E7EC4027AFAFF976E" authority="Riek." authorityName="Riek." box="[1153,1375,1338,1363]" class="Insecta" family="Archeosmylidae" genus="Lithosmylidia" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9191C72FF898D3E7EC4027AFABC976E" box="[1153,1308,1338,1363]" italics="true" pageId="10" pageNumber="11">Lithosmylidia</emphasis>
|
||
Riek.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |