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<document id="EA940FCBBEF337620DC150DE374883AC" ID-DOI="10.11646/zootaxa.4747.3.11" ID-GBIF-Dataset="70f148ee-1a74-40d7-a11d-e30eb4f3f5e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="3696198" ID-ZooBank="194F0901-51B8-41BD-BAB6-195F607274B2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583304130844" checkinUser="plazi" docAuthor="Barros, Lurdiana Dayse De, Barão, Kim Ribeiro &amp; Grazia, Jocelia" docDate="2020" docId="03B84F22080AFFF260B5F956FB486F75" docLanguage="en" docName="zootaxa.4747.3.11.pdf" docOrigin="Zootaxa 4747 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Elanela colombiana Barros &amp; Barão &amp; Grazia 2020, sp. nov." docType="treatment" docUuid="048B2685-8861-4954-A502-F5D04C979356" docUuidSource="ZooBank" docVersion="8" lastPageNumber="588" masterDocId="FF81375A080BFFF16022FFF1FFE86A22" masterDocTitle="New species and new distributional records of Elanela Rolston, 1980 (Heteroptera: Pentatomidae)" masterLastPageNumber="592" masterPageNumber="585" pageNumber="586" updateTime="1698724510451" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="100B7FB14FD72B32AA299747FB68D4A0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="062FF31B005947FADF8E63F57387DE0B">New species and new distributional records of Elanela Rolston, 1980 (Heteroptera: Pentatomidae)</mods:title>
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<mods:role id="398D81C45B8E44982B3F53B2E2FD3958">
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<mods:namePart id="D579AC3E495BC76D297498AB249B5FF2">Barros, Lurdiana Dayse De</mods:namePart>
<mods:affiliation id="6137FFFA63D8E8845501B55ACF7E4222">Departamento de Zoologia, Instituto de Biociências, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, 91501 - 970, Porto Alegre, RS, Brazil. &amp; Programa de Pós-Graduação em Biologia Animal, Universidade Federal do Rio Grande do Sul.</mods:affiliation>
</mods:name>
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<mods:role id="A4E25D61E3CFC48D56D6711F17C89709">
<mods:roleTerm id="187F274212C257F4A5BE279AC2CE299A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B0B2DB005466A81CB71246351046773F">Barão, Kim Ribeiro</mods:namePart>
<mods:affiliation id="363078752A82ED0343A0F3B2E177717A">Unidade Educacional Penedo, Campus Arapiraca, Universidade Federal de Alagoas (UFAL), AL, Brazil. &amp; PPG Diversidade Biológica e Conservação nos Trópicos, Universidade Federal de Alagoas.</mods:affiliation>
</mods:name>
<mods:name id="799664174B007F64B62539A630A05E63" type="personal">
<mods:role id="5EDBBC82EC745BF01DE8E0437AD31B4E">
<mods:roleTerm id="8685EF47DC4C7B59D29B8A7B7A5EFAEF">Author</mods:roleTerm>
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<mods:namePart id="4770F6A5E4BAA69380F9342BDD57AB8F">Grazia, Jocelia</mods:namePart>
<mods:affiliation id="5F677C140D752B0FFF11EA67BD1ED21F">Departamento de Zoologia, Instituto de Biociências, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves 9500, 91501 - 970, Porto Alegre, RS, Brazil. &amp; Programa de Pós-Graduação em Biologia Animal, Universidade Federal do Rio Grande do Sul.</mods:affiliation>
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<mods:title id="7359C757FC17BA91F9909852274835FF">Zootaxa</mods:title>
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<mods:part id="E790528029D2CD4D2E6DBF06E894CAE9">
<mods:date id="FE4B0158E2082ACA2925DAABC515B4EE">2020</mods:date>
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<mods:number id="7B6A885181AE93FC9F0DB713146A8A2B">2020-03-04</mods:number>
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<mods:detail id="DBB2E57028FAA68F7134D8E28C3A013E" type="volume">
<mods:number id="50151711730C769FCDFBB8FFD951FA4B">4747</mods:number>
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<mods:classification id="163FC959F9FFE0DBAD18720792BAB54D">journal article</mods:classification>
<mods:identifier id="2CCE779B1760D8850361ACB0B7EDD2A7" type="DOI">10.11646/zootaxa.4747.3.11</mods:identifier>
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<treatment id="03B84F22080AFFF260B5F956FB486F75" ID-DOI="http://doi.org/10.5281/zenodo.3703948" ID-GBIF-Taxon="162383799" ID-Zenodo-Dep="3703948" LSID="urn:lsid:zoobank.org:act:048B2685-8861-4954-A502-F5D04C979356" httpUri="http://treatment.plazi.org/id/03B84F22080AFFF260B5F956FB486F75" lastPageId="3" lastPageNumber="588" pageId="1" pageNumber="586">
<subSubSection id="C30BADBF080AFFF060B5F956FE186CE0" box="[151,496,1703,1730]" pageId="1" pageNumber="586" type="nomenclature">
<paragraph id="8BAEFE34080AFFF060B5F956FE186CE0" blockId="1.[151,496,1703,1766]" box="[151,496,1703,1730]" pageId="1" pageNumber="586">
<heading id="D0E64958080AFFF060B5F956FE186CE0" bold="true" box="[151,496,1703,1730]" fontSize="11" level="1" pageId="1" pageNumber="586" reason="1">
<emphasis id="B9652226080AFFF060B5F956FE186CE0" bold="true" box="[151,496,1703,1730]" pageId="1" pageNumber="586">
<taxonomicName id="4C1185B7080AFFF060B5F956FE636CE0" authority="Barros &amp; Barão &amp; Grazia, 2020" authorityName="Barros &amp; Barão &amp; Grazia" authorityYear="2020" box="[151,395,1703,1730]" class="Insecta" family="Pentatomidae" genus="Elanela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="586" phylum="Arthropoda" rank="species" species="colombiana" status="sp. nov.">
<emphasis id="B9652226080AFFF060B5F956FE636CE0" bold="true" box="[151,395,1703,1730]" italics="true" pageId="1" pageNumber="586">Elanela colombiana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2569F5D080AFFF061B0F959FE186CE0" box="[402,496,1704,1730]" pageId="1" pageNumber="586" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30BADBF080AFFF060B5F93AFEB16CC4" box="[151,345,1739,1766]" pageId="1" pageNumber="586" type="description">
<paragraph id="8BAEFE34080AFFF060B5F93AFEB16CC4" blockId="1.[151,496,1703,1766]" box="[151,345,1739,1766]" pageId="1" pageNumber="586">
(
<figureCitation id="132AE2B1080AFFF060BDF93AFEC06CC4" box="[159,296,1739,1766]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1787,1812]" captionTargetBox="[248,1339,185,1762]" captionTargetId="figure@2.[248,1339,181,1762]" captionTargetPageId="2" captionText="FIGURES 1–7. Elanela colombiana sp. nov. 1–2, habitus; 1, dorsal view; 2, ventral view; 3, head and pronotum in dorsal view; 4, external scent efferent system, arrow showing ostiolar area; 5–6, female genital plates in posteroventral view; 7, female receptaculum seminis and ausenwand. Abbreviations: aaf, anterior annular flange; cs, capsula seminalis; ddr, distal ductus receptaculi; pdr, proximal ductus receptaculi; ev, evaporatorium; g9, gonapophyses IX; gc8, gonocoxites VIII; gc9, gonocoxites IX; la8, laterotergites VIII; la9, laterotergites IX; ms, mesopleura; mt, metapleura; o, ostiole; paf, posterior annular flange; pi, pars intermedialis; tvi, thickening of vaginal intima; va, vesicular area; X, abdominal segment X. Scale bars:1–3= 0.5 mm; 5–6= 0.5 mm; 7= 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3696202" httpUri="https://zenodo.org/record/3696202/files/figure.png" pageId="1" pageNumber="586">Figures 1–7</figureCitation>
,
<figureCitation id="132AE2B1080AFFF06116F93AFEB96CC4" box="[308,337,1739,1766]" captionStart="FIGURE 16" captionStartId="6.[151,250,1052,1077]" captionTargetBox="[151,1436,181,1026]" captionTargetId="figure@6.[151,1436,181,1026]" captionTargetPageId="6" captionText="FIGURE 16. Known occurrence records of species of Elanela Rolston, 1980. Map adapted from Grazia et al. (2016) to include E. colombiana sp. nov. and E. ecuatoriana sp. nov. (highlighted in red) and the new record of E. jordi." figureDoi="http://doi.org/10.5281/zenodo.3696209" httpUri="https://zenodo.org/record/3696209/files/figure.png" pageId="1" pageNumber="586">16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30BADBF080AFFF060B5F8E5FE036DBB" pageId="1" pageNumber="586" type="materials_examined">
<paragraph id="8BAEFE34080AFFF060B5F8E5FE036DBB" blockId="1.[151,1437,1811,1982]" pageId="1" pageNumber="586">
<emphasis id="B9652226080AFFF060B5F8E5FEA26D0C" bold="true" box="[151,330,1811,1838]" pageId="1" pageNumber="586">Type material.</emphasis>
<materialsCitation id="3B79F469080AFFF06177F8E5FB276D70" ID-GBIF-Occurrence="2573309212" collectingDate="2013-08-23" collectionCode="UNAB" collectorName="Colecta &amp; L. Perez" country="Colombia" county="Cali" elevation="984" latitude="3.3423889" location="Microestacion" longLatPrecision="1" longitude="-76.533714" municipality="Universidad del Valle" pageId="1" pageNumber="586" specimenCount="1" specimenCount-female="1" stateProvince="Valle del Cauca" typeStatus="holotype">
<typeStatus id="54AA4096080AFFF06177F8E5FE046D0C" box="[341,492,1812,1838]" pageId="1" pageNumber="586" type="holotype">HOLOTYPE</typeStatus>
,
<specimenCount id="9D1735BD080AFFF061DEF8E5FDF76D0C" box="[508,543,1812,1838]" pageId="1" pageNumber="586" type="female">1♀</specimenCount>
,
<collectingCountry id="F306BEA4080AFFF06210F8E5FD226D0C" box="[562,714,1812,1838]" name="Colombia" pageId="1" pageNumber="586">COLOMBIA</collectingCountry>
,
<collectingRegion id="49D530D6080AFFF062FBF8E2FC7D6D0C" box="[729,917,1811,1838]" country="Colombia" name="Valle del Cauca" pageId="1" pageNumber="586">Valle del Cauca</collectingRegion>
,
<collectingCounty id="62CF86B8080AFFF06387F8E2FC3E6D0F" box="[933,982,1811,1837]" pageId="1" pageNumber="586">Cali</collectingCounty>
,
<collectingMunicipality id="6BCA644E080AFFF063C5F8E2FB0D6D0F" box="[999,1253,1811,1837]" pageId="1" pageNumber="586">Universidad del Valle</collectingMunicipality>
,
<location id="8ECEA8EF080AFFF064D7F8E2FA706D0F" LSID="urn:lsid:plazi:treatment:03B84F22080AFFF260B5F956FB486F75:8ECEA8EF080AFFF064D7F8E2FA706D0F" box="[1269,1432,1811,1837]" country="Colombia" county="Cali" latitude="3.3423889" longLatPrecision="1" longitude="-76.533714" municipality="Universidad del Valle" name="Microestacion" pageId="1" pageNumber="586" stateProvince="Valle del Cauca">Microestación</location>
,
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<geoCoordinate id="EE2598F3080AFFF0611EF8C6FE076D73" box="[316,495,1847,1873]" degrees="76" direction="west" minutes="32" orientation="longitude" pageId="1" pageNumber="586" precision="1" seconds="01.37" value="-76.533714">76°32’01.37”W</geoCoordinate>
,
<quantity id="4CE953D1080AFFF061D8F8C9FDD26D70" box="[506,570,1848,1874]" metricMagnitude="2" metricUnit="m" metricValue="9.84" pageId="1" pageNumber="586" unit="m" value="984.0">
<elevation id="003C1907080AFFF061D8F8C9FDD26D70" box="[506,570,1848,1874]" metricMagnitude="2" metricUnit="m" metricValue="9.84" pageId="1" pageNumber="586" unit="m" value="984.0">984m</elevation>
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alt.,
<collectorName id="26E49BE2080AFFF06250F8C6FD206D73" box="[626,712,1847,1873]" pageId="1" pageNumber="586">Colecta</collectorName>
manual,
<date id="FFAFD8F4080AFFF0630DF8C9FC216D70" box="[815,969,1848,1874]" pageId="1" pageNumber="586" value="2013-08-23">
<collectingDate id="EFEB211C080AFFF0630DF8C9FC216D70" box="[815,969,1848,1874]" pageId="1" pageNumber="586" value="2013-08-23">23-VIII-2013</collectingDate>
</date>
,
<collectorName id="26E49BE2080AFFF063F6F8C9FBD96D70" box="[980,1073,1848,1874]" pageId="1" pageNumber="586">L. Perez</collectorName>
leg. (
<collectionCode id="ED0066F1080AFFF06453F8C9FB2F6D70" box="[1137,1223,1848,1874]" country="Colombia" httpUri="http://grbio.org/cool/hx5r-ym26" name="Universidad Nacional, Facultad de Agronomia" pageId="1" pageNumber="586">UNAB</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B79F469080AFFF064FBF8C9FE306DBB" ID-GBIF-Occurrence="2573309214" collectingDate="2009-12-26" collectingMethod="captura jama" collectionCode="UNAB" collectorName="L. Mardharst" country="Colombia" county="Valle" elevation="990" location="Felidia" municipality="Cali" pageId="1" pageNumber="586" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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,
<specimenCount id="9D1735BD080AFFF06550F8C9FA7D6D70" box="[1394,1429,1848,1874]" pageId="1" pageNumber="586" type="female">1♀</specimenCount>
,
<collectingCountry id="F306BEA4080AFFF060B5F8ADFEC76D54" box="[151,303,1884,1910]" name="Colombia" pageId="1" pageNumber="586">COLOMBIA</collectingCountry>
,
<collectingCounty id="62CF86B8080AFFF06118F8AAFE9E6D57" box="[314,374,1883,1909]" pageId="1" pageNumber="586">Valle</collectingCounty>
,
<collectingMunicipality id="6BCA644E080AFFF061A3F8AAFE5A6D57" box="[385,434,1883,1909]" pageId="1" pageNumber="586">Cali</collectingMunicipality>
,
<location id="8ECEA8EF080AFFF0619CF8AAFDE76D57" LSID="urn:lsid:plazi:treatment:03B84F22080AFFF260B5F956FB486F75:8ECEA8EF080AFFF0619CF8AAFDE76D57" box="[446,527,1883,1909]" country="Colombia" county="Valle" municipality="Cali" name="Felidia" pageId="1" pageNumber="586">Felidia</location>
,
<location id="8ECEA8EF080AFFF0623BF8AAFD416D57" LSID="urn:lsid:plazi:treatment:03B84F22080AFFF260B5F956FB486F75:8ECEA8EF080AFFF0623BF8AAFD416D57" box="[537,681,1883,1909]" country="Colombia" county="Valle" municipality="Cali" name="El Diamante" pageId="1" pageNumber="586">El Diamante</location>
,
<quantity id="4CE953D1080AFFF06296F8ADFD1C6D54" box="[692,756,1884,1910]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="1" pageNumber="586" unit="m" value="990.0">
<elevation id="003C1907080AFFF06296F8ADFD1C6D54" box="[692,756,1884,1910]" metricMagnitude="2" metricUnit="m" metricValue="9.9" pageId="1" pageNumber="586" unit="m" value="990.0">990m</elevation>
</quantity>
alt.,
<collectingMethod id="52508623080AFFF06309F8ADFC556D57" box="[811,957,1883,1910]" pageId="1" pageNumber="586">captura jama</collectingMethod>
,
<location id="8ECEA8EF080AFFF063EBF8ADFBDC6D54" LSID="urn:lsid:plazi:treatment:03B84F22080AFFF260B5F956FB486F75:8ECEA8EF080AFFF063EBF8ADFBDC6D54" box="[969,1076,1884,1910]" country="Colombia" county="Valle" municipality="Cali" name="Carretera" pageId="1" pageNumber="586">Carretera</location>
,
<date id="FFAFD8F4080AFFF0641DF8AAFB386D57" box="[1087,1232,1883,1909]" pageId="1" pageNumber="586" value="2009-12-26">
<collectingDate id="EFEB211C080AFFF0641DF8AAFB386D57" box="[1087,1232,1883,1909]" pageId="1" pageNumber="586" value="2009-12-26">26-XII-2009</collectingDate>
</date>
,
<collectorName id="26E49BE2080AFFF064F8F8ADFA846D57" box="[1242,1388,1883,1910]" pageId="1" pageNumber="586">L. Mardharst</collectorName>
leg. (
<collectionCode id="ED0066F1080AFFF060BDF871FF1A6DB8" box="[159,242,1920,1946]" collectionName="Colombia, Bogota, Universidad Nacional, Facultad de Agronomia" country="Colombia" httpUri="http://grbio.org/cool/hx5r-ym26" name="Universidad Nacional, Facultad de Agronomia" pageId="1" pageNumber="586">UNAB</collectionCode>
) &lt;illustrated female
</materialsCitation>
&gt;.
</paragraph>
</subSubSection>
<subSubSection id="C30BADBF080AFFF060E5F855FABB6D9F" box="[199,1363,1955,1982]" pageId="1" pageNumber="586" type="etymology">
<paragraph id="8BAEFE34080AFFF060E5F855FABB6D9F" blockId="1.[151,1437,1811,1982]" box="[199,1363,1955,1982]" pageId="1" pageNumber="586">
<emphasis id="B9652226080AFFF060E5F855FEA66D9C" bold="true" box="[199,334,1956,1982]" pageId="1" pageNumber="586">Etymology.</emphasis>
Genitive singular; name in reference to the collection locality in Vale del
<collectingRegion id="49D530D6080AFFF064ABF855FB3A6D9C" box="[1161,1234,1956,1982]" country="Colombia" name="Cauca" pageId="1" pageNumber="586">Cauca</collectingRegion>
,
<collectingCountry id="F306BEA4080AFFF064FFF852FAA76D9F" box="[1245,1359,1955,1981]" name="Colombia" pageId="1" pageNumber="586">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="DF6EAEBC0809FFF360B5F90AFE9D6DC9" ID-DOI="http://doi.org/10.5281/zenodo.3696202" ID-Zenodo-Dep="3696202" httpUri="https://zenodo.org/record/3696202/files/figure.png" pageId="2" pageNumber="587" startId="2.[151,264,1787,1812]" targetBox="[248,1339,185,1762]" targetPageId="2">
<paragraph id="8BAEFE340809FFF360B5F90AFE9D6DC9" blockId="2.[151,1437,1787,2028]" pageId="2" pageNumber="587">
<emphasis id="B96522260809FFF360B5F90AFED56D36" bold="true" box="[151,317,1787,1812]" pageId="2" pageNumber="587">FIGURES 1–7.</emphasis>
<taxonomicName id="4C1185B70809FFF36167F90DFDFD6D36" authority="Barros &amp; Barão &amp; Grazia, 2020" authorityName="Barros &amp; Barão &amp; Grazia" authorityYear="2020" box="[325,533,1788,1812]" class="Insecta" family="Pentatomidae" genus="Elanela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="587" phylum="Arthropoda" rank="species" species="colombiana" status="sp. nov.">
<emphasis id="B96522260809FFF36167F90DFDFD6D36" box="[325,533,1788,1812]" italics="true" pageId="2" pageNumber="587">Elanela colombiana</emphasis>
</taxonomicName>
<emphasis id="B96522260809FFF3623FF90DFD986D36" bold="true" box="[541,624,1788,1812]" pageId="2" pageNumber="587">
<taxonomicNameLabel id="A2569F5D0809FFF3623FF90DFD986D36" box="[541,624,1788,1812]" pageId="2" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
1–2, habitus; 1, dorsal view; 2, ventral view; 3, head and pronotum in dorsal view; 4, external scent efferent system, arrow showing ostiolar area; 5–6, female genital plates in posteroventral view; 7, female receptaculum seminis and ausenwand. Abbreviations: aaf, anterior annular flange; cs,
<emphasis id="B96522260809FFF36424F8B5FB546D7E" box="[1030,1212,1860,1884]" italics="true" pageId="2" pageNumber="587">capsula seminalis</emphasis>
; ddr, distal
<emphasis id="B96522260809FFF36514F8B5FF1C6DA2" italics="true" pageId="2" pageNumber="587">ductus receptaculi</emphasis>
; pdr, proximal
<emphasis id="B96522260809FFF361B0F899FDA46DA2" box="[402,588,1896,1920]" italics="true" pageId="2" pageNumber="587">ductus receptaculi</emphasis>
; ev, evaporatorium; g9, gonapophyses IX; gc8, gonocoxites VIII; gc9, gonocoxites IX; la8, laterotergites VIII; la9, laterotergites IX; ms, mesopleura; mt, metapleura; o, ostiole; paf, posterior annular flange; pi,
<emphasis id="B96522260809FFF360B5F840FEA76DEA" box="[151,335,1968,1992]" italics="true" pageId="2" pageNumber="587">pars intermedialis</emphasis>
; tvi, thickening of vaginal intima; va, vesicular area; X, abdominal segment X. Scale bars:1–3= 0.5 mm; 5–6= 0.5 mm; 7= 0.25 mm.
</paragraph>
</caption>
<subSubSection id="C30BADBF0808FFF260E5FF69FEF36B45" pageId="3" pageNumber="588" type="diagnosis">
<paragraph id="8BAEFE340808FFF260E5FF69FEF36B45" blockId="3.[151,1437,152,1367]" pageId="3" pageNumber="588">
<emphasis id="B96522260808FFF260E5FF69FEAA6A91" bold="true" box="[199,322,152,179]" pageId="3" pageNumber="588">Diagnosis.</emphasis>
Anterior and anterolateral margins of pronotum whitish, impunctate; anterior and anterolateral margins of pronotum with black punctuations. Disk of scutellum callused, ivory. Abdominal sternite VII immaculate medially. Each laterotergite VIII subtriangular, lacking spiracle; posterior margin of each laterotergite VIII with an acute projection. Anterior margin of each gonocoxite IX straight, posterior margin notched medially.
<emphasis id="B96522260808FFF26520FEF7FF3D6B61" italics="true" pageId="3" pageNumber="588">Pars intermedialis</emphasis>
short, straight basally and shorter than
<emphasis id="B96522260808FFF262B3FED8FCB26B61" box="[657,858,297,323]" italics="true" pageId="3" pageNumber="588">capsula seminalis</emphasis>
.
<emphasis id="B96522260808FFF2634AFED8FBDF6B61" box="[872,1079,297,323]" italics="true" pageId="3" pageNumber="588">Capsula seminalis</emphasis>
globose, with lateral and apical projections.
</paragraph>
</subSubSection>
<subSubSection id="C30BADBF0808FFF260E5FE81FE066E79" pageId="3" pageNumber="588" type="description">
<paragraph id="8BAEFE340808FFF260E5FE81FE87681D" blockId="3.[151,1437,152,1367]" pageId="3" pageNumber="588">
<emphasis id="B96522260808FFF260E5FE81FEB16BA9" bold="true" box="[199,345,368,395]" pageId="3" pageNumber="588">Description.</emphasis>
Head wider than long; head surface irregular, with few brownish punctuations; antennomeres I–III entirely brownish, IV–V brownish basally. Proportion of antennomeres I &lt;II&gt; III&lt;IV &lt;V. Anterior and anterolateral margins of pronotum whitish, impunctate, bordered by a line of dark punctuations (
<figureCitation id="132AE2B10808FFF264BAFE48FB366BF1" box="[1176,1246,441,467]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1787,1812]" captionTargetBox="[248,1339,185,1762]" captionTargetId="figure@2.[248,1339,181,1762]" captionTargetPageId="2" captionText="FIGURES 1–7. Elanela colombiana sp. nov. 1–2, habitus; 1, dorsal view; 2, ventral view; 3, head and pronotum in dorsal view; 4, external scent efferent system, arrow showing ostiolar area; 5–6, female genital plates in posteroventral view; 7, female receptaculum seminis and ausenwand. Abbreviations: aaf, anterior annular flange; cs, capsula seminalis; ddr, distal ductus receptaculi; pdr, proximal ductus receptaculi; ev, evaporatorium; g9, gonapophyses IX; gc8, gonocoxites VIII; gc9, gonocoxites IX; la8, laterotergites VIII; la9, laterotergites IX; ms, mesopleura; mt, metapleura; o, ostiole; paf, posterior annular flange; pi, pars intermedialis; tvi, thickening of vaginal intima; va, vesicular area; X, abdominal segment X. Scale bars:1–3= 0.5 mm; 5–6= 0.5 mm; 7= 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3696202" httpUri="https://zenodo.org/record/3696202/files/figure.png" pageId="3" pageNumber="588">Fig. 3</figureCitation>
). Evaporatorium brownish (
<figureCitation id="132AE2B10808FFF2612DFE2CFEBD6BD5" box="[271,341,477,503]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1787,1812]" captionTargetBox="[248,1339,185,1762]" captionTargetId="figure@2.[248,1339,181,1762]" captionTargetPageId="2" captionText="FIGURES 1–7. Elanela colombiana sp. nov. 1–2, habitus; 1, dorsal view; 2, ventral view; 3, head and pronotum in dorsal view; 4, external scent efferent system, arrow showing ostiolar area; 5–6, female genital plates in posteroventral view; 7, female receptaculum seminis and ausenwand. Abbreviations: aaf, anterior annular flange; cs, capsula seminalis; ddr, distal ductus receptaculi; pdr, proximal ductus receptaculi; ev, evaporatorium; g9, gonapophyses IX; gc8, gonocoxites VIII; gc9, gonocoxites IX; la8, laterotergites VIII; la9, laterotergites IX; ms, mesopleura; mt, metapleura; o, ostiole; paf, posterior annular flange; pi, pars intermedialis; tvi, thickening of vaginal intima; va, vesicular area; X, abdominal segment X. Scale bars:1–3= 0.5 mm; 5–6= 0.5 mm; 7= 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3696202" httpUri="https://zenodo.org/record/3696202/files/figure.png" pageId="3" pageNumber="588">Fig. 4</figureCitation>
). Disk of scutellum callused, ivory (
<figureCitation id="132AE2B10808FFF262CEFE2CFCC66BD5" box="[748,814,477,503]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1787,1812]" captionTargetBox="[248,1339,185,1762]" captionTargetId="figure@2.[248,1339,181,1762]" captionTargetPageId="2" captionText="FIGURES 1–7. Elanela colombiana sp. nov. 1–2, habitus; 1, dorsal view; 2, ventral view; 3, head and pronotum in dorsal view; 4, external scent efferent system, arrow showing ostiolar area; 5–6, female genital plates in posteroventral view; 7, female receptaculum seminis and ausenwand. Abbreviations: aaf, anterior annular flange; cs, capsula seminalis; ddr, distal ductus receptaculi; pdr, proximal ductus receptaculi; ev, evaporatorium; g9, gonapophyses IX; gc8, gonocoxites VIII; gc9, gonocoxites IX; la8, laterotergites VIII; la9, laterotergites IX; ms, mesopleura; mt, metapleura; o, ostiole; paf, posterior annular flange; pi, pars intermedialis; tvi, thickening of vaginal intima; va, vesicular area; X, abdominal segment X. Scale bars:1–3= 0.5 mm; 5–6= 0.5 mm; 7= 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3696202" httpUri="https://zenodo.org/record/3696202/files/figure.png" pageId="3" pageNumber="588">Fig. 1</figureCitation>
). Posterolateral angles of connexivum black and acute. Abdominal sternites with a sublateral band of brownish punctuations; a rounded, yellowish spot laterally on each abdominal sternite.
</paragraph>
<paragraph id="8BAEFE340808FFF260E5FDB8FA9569A1" blockId="3.[151,1437,152,1367]" pageId="3" pageNumber="588">
<emphasis id="B96522260808FFF260E5FDB8FDF06841" bold="true" box="[199,536,584,611]" pageId="3" pageNumber="588">
Female genitalia (
<figureCitation id="132AE2B10808FFF26182FDB9FDE36841" box="[416,523,584,611]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1787,1812]" captionTargetBox="[248,1339,185,1762]" captionTargetId="figure@2.[248,1339,181,1762]" captionTargetPageId="2" captionText="FIGURES 1–7. Elanela colombiana sp. nov. 1–2, habitus; 1, dorsal view; 2, ventral view; 3, head and pronotum in dorsal view; 4, external scent efferent system, arrow showing ostiolar area; 5–6, female genital plates in posteroventral view; 7, female receptaculum seminis and ausenwand. Abbreviations: aaf, anterior annular flange; cs, capsula seminalis; ddr, distal ductus receptaculi; pdr, proximal ductus receptaculi; ev, evaporatorium; g9, gonapophyses IX; gc8, gonocoxites VIII; gc9, gonocoxites IX; la8, laterotergites VIII; la9, laterotergites IX; ms, mesopleura; mt, metapleura; o, ostiole; paf, posterior annular flange; pi, pars intermedialis; tvi, thickening of vaginal intima; va, vesicular area; X, abdominal segment X. Scale bars:1–3= 0.5 mm; 5–6= 0.5 mm; 7= 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3696202" httpUri="https://zenodo.org/record/3696202/files/figure.png" pageId="3" pageNumber="588">Figs 5–7</figureCitation>
).
</emphasis>
Genital plates setose (
<figureCitation id="132AE2B10808FFF26302FDB8FC616841" box="[800,905,584,611]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1787,1812]" captionTargetBox="[248,1339,185,1762]" captionTargetId="figure@2.[248,1339,181,1762]" captionTargetPageId="2" captionText="FIGURES 1–7. Elanela colombiana sp. nov. 1–2, habitus; 1, dorsal view; 2, ventral view; 3, head and pronotum in dorsal view; 4, external scent efferent system, arrow showing ostiolar area; 5–6, female genital plates in posteroventral view; 7, female receptaculum seminis and ausenwand. Abbreviations: aaf, anterior annular flange; cs, capsula seminalis; ddr, distal ductus receptaculi; pdr, proximal ductus receptaculi; ev, evaporatorium; g9, gonapophyses IX; gc8, gonocoxites VIII; gc9, gonocoxites IX; la8, laterotergites VIII; la9, laterotergites IX; ms, mesopleura; mt, metapleura; o, ostiole; paf, posterior annular flange; pi, pars intermedialis; tvi, thickening of vaginal intima; va, vesicular area; X, abdominal segment X. Scale bars:1–3= 0.5 mm; 5–6= 0.5 mm; 7= 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3696202" httpUri="https://zenodo.org/record/3696202/files/figure.png" pageId="3" pageNumber="588">Figs 5–6</figureCitation>
). Gonocoxites VIII (gc8) concave, wider than long, posterior margins sinuous, sutural margins parallel not juxtaposed. Gonapophyses VIII (g8) visible between gonocoxites VIII. Laterotergites VIII (la8) subtriangular, lacking spiracles; posterior margin of each laterotergite VIII with an acute projection. Gonocoxites IX (gc9) strongly convex; anterior margin straight; posterior margin notched medially. Laterotergites IX (la9) depressed basally; laterotergites IX extending beyond posterior margin of tergite VIII. Segment X rectangular.
<emphasis id="B96522260808FFF26209FD0CFD206935" box="[555,712,765,791]" italics="true" pageId="3" pageNumber="588">Chitinellipsen</emphasis>
(ch) elliptical; thickening of vaginal intima (tvi) globose (
<figureCitation id="132AE2B10808FFF26569FD0CFA666935" box="[1355,1422,765,791]" captionStart="FIGURES 1–7" captionStartId="2.[151,264,1787,1812]" captionTargetBox="[248,1339,185,1762]" captionTargetId="figure@2.[248,1339,181,1762]" captionTargetPageId="2" captionText="FIGURES 1–7. Elanela colombiana sp. nov. 1–2, habitus; 1, dorsal view; 2, ventral view; 3, head and pronotum in dorsal view; 4, external scent efferent system, arrow showing ostiolar area; 5–6, female genital plates in posteroventral view; 7, female receptaculum seminis and ausenwand. Abbreviations: aaf, anterior annular flange; cs, capsula seminalis; ddr, distal ductus receptaculi; pdr, proximal ductus receptaculi; ev, evaporatorium; g9, gonapophyses IX; gc8, gonocoxites VIII; gc9, gonocoxites IX; la8, laterotergites VIII; la9, laterotergites IX; ms, mesopleura; mt, metapleura; o, ostiole; paf, posterior annular flange; pi, pars intermedialis; tvi, thickening of vaginal intima; va, vesicular area; X, abdominal segment X. Scale bars:1–3= 0.5 mm; 5–6= 0.5 mm; 7= 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.3696202" httpUri="https://zenodo.org/record/3696202/files/figure.png" pageId="3" pageNumber="588">Fig. 7</figureCitation>
). Proximal
<emphasis id="B96522260808FFF26126FCD0FE386919" box="[260,464,801,827]" italics="true" pageId="3" pageNumber="588">ductus receptaculi</emphasis>
(pdr) longer than vesicular area (va); distal
<emphasis id="B96522260808FFF26397FCD0FB696919" box="[949,1153,801,827]" italics="true" pageId="3" pageNumber="588">ductus receptaculi</emphasis>
(ddr) as long as vesicular area;
<emphasis id="B96522260808FFF260F6FCB7FE48697D" box="[212,416,837,863]" italics="true" pageId="3" pageNumber="588">pars intermedialis</emphasis>
(pi) narrower basally than apically, shorter than
<emphasis id="B96522260808FFF2639EFCB4FB6C697D" box="[956,1156,837,863]" italics="true" pageId="3" pageNumber="588">capsula seminalis</emphasis>
; posterior annular flange (paf) wider than anterior annular flange (aaf);
<emphasis id="B96522260808FFF262BEFC98FC8C69A1" box="[668,868,873,899]" italics="true" pageId="3" pageNumber="588">capsula seminalis</emphasis>
(cs) globose with lateral and apical projections.
</paragraph>
<paragraph id="8BAEFE340808FFF260E5FC7CFE066E79" blockId="3.[151,1437,152,1367]" pageId="3" pageNumber="588">
<emphasis id="B96522260808FFF260E5FC7CFE966985" bold="true" box="[199,382,909,935]" pageId="3" pageNumber="588">Measurements.</emphasis>
Total length, 7.31 ± 0.26 (7.12–7.50); length of head, 1.13 ± 0.01 (1.12–1.15); head length ahead of eyes, 0.875; interocular distance, 1.03 ± 0.05 (1.00–1.07); length of antennal segments: I, 0.35 ± 0.03 (0.32–0.37); II, 0.85 ± 0.03 (0.82–0.85); III, 0.78 ± 0.05 (0.75–0.82); IV, 1.37; V, 1.62; width of head, 1.87 ± 0.03 (1.85–1.90); width of pronotum, 4.32 ± 0.24 (4.15–4.50); length of pronotum, 1.67 ± 0.03 (1.65–1.70); width of scutellum, 2.45 ± 0.08 (2.39–2.50); length of scutellum, 2.79 ± 0.08 (2.73–2.85); width of abdomen at third segment, 4.31 ± 0.08 (4.25–4.37).
</paragraph>
</subSubSection>
<subSubSection id="C30BADBF0808FFF260E5FB95FDAE6EA2" box="[199,582,1124,1152]" pageId="3" pageNumber="588" type="distribution">
<paragraph id="8BAEFE340808FFF260E5FB95FDAE6EA2" blockId="3.[151,1437,152,1367]" box="[199,582,1124,1152]" pageId="3" pageNumber="588">
<emphasis id="B96522260808FFF260E5FB95FE896E5D" bold="true" box="[199,353,1124,1151]" pageId="3" pageNumber="588">Distribution.</emphasis>
<collectingCountry id="F306BEA40808FFF26145FB94FE3D6E5D" box="[359,469,1125,1151]" name="Colombia" pageId="3" pageNumber="588">Colombia</collectingCountry>
(
<figureCitation id="132AE2B10808FFF261C7FB94FDD16EA2" box="[485,569,1125,1152]" captionStart="FIGURE 16" captionStartId="6.[151,250,1052,1077]" captionTargetBox="[151,1436,181,1026]" captionTargetId="figure@6.[151,1436,181,1026]" captionTargetPageId="6" captionText="FIGURE 16. Known occurrence records of species of Elanela Rolston, 1980. Map adapted from Grazia et al. (2016) to include E. colombiana sp. nov. and E. ecuatoriana sp. nov. (highlighted in red) and the new record of E. jordi." figureDoi="http://doi.org/10.5281/zenodo.3696209" httpUri="https://zenodo.org/record/3696209/files/figure.png" pageId="3" pageNumber="588">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30BADBF0808FFF260E5FB79FB486F75" pageId="3" pageNumber="588" type="discussion">
<paragraph id="8BAEFE340808FFF260E5FB79FB486F75" blockId="3.[151,1437,152,1367]" pageId="3" pageNumber="588">
<emphasis id="B96522260808FFF260E5FB79FEA76E81" bold="true" box="[199,335,1160,1187]" pageId="3" pageNumber="588">Comments.</emphasis>
The dorsal habitus of
<taxonomicName id="4C1185B70808FFF26269FB7BFD076E81" authority="Barros &amp; Barão &amp; Grazia, 2020" authorityName="Barros &amp; Barão &amp; Grazia" authorityYear="2020" box="[587,751,1161,1187]" class="Insecta" family="Pentatomidae" genus="Elanela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="colombiana" status="sp. nov.">
<emphasis id="B96522260808FFF26269FB7BFDB46E81" box="[587,604,1162,1187]" italics="true" pageId="3" pageNumber="588">E</emphasis>
.
<emphasis id="B96522260808FFF26249FB78FD076E81" box="[619,751,1161,1187]" italics="true" pageId="3" pageNumber="588">colombiana</emphasis>
</taxonomicName>
<emphasis id="B96522260808FFF262D4FB78FCB86E81" bold="true" box="[758,848,1161,1187]" pageId="3" pageNumber="588">
<taxonomicNameLabel id="A2569F5D0808FFF262D4FB78FCB86E81" box="[758,848,1161,1187]" pageId="3" pageNumber="588" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C1185B70808FFF263A5FB7BFBA16E81" authorityName="Grazia, Barros &amp; Barao" authorityYear="2016" box="[903,1097,1161,1187]" class="Insecta" family="Pentatomidae" genus="Elanela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="scutomaculata">
<emphasis id="B96522260808FFF263A5FB7BFBA16E81" box="[903,1097,1161,1187]" italics="true" pageId="3" pageNumber="588">E. scutomaculata</emphasis>
</taxonomicName>
are similar by the presence of an ivory spot on the disk of scutellum. Both species can be distinguished from each other by the coloration of the antennae, by the vestiture of the anterior and lateral margins of pronotum, and presence or absence of a black spot on abdominal sternite VII medially. The female genitalia of
<taxonomicName id="4C1185B70808FFF26364FB07FC036F2D" authority="Barros &amp; Barão &amp; Grazia, 2020" authorityName="Barros &amp; Barão &amp; Grazia" authorityYear="2020" box="[838,1003,1269,1295]" class="Insecta" family="Pentatomidae" genus="Elanela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="colombiana" status="sp. nov.">
<emphasis id="B96522260808FFF26364FB07FCBF6F2D" box="[838,855,1270,1295]" italics="true" pageId="3" pageNumber="588">E</emphasis>
.
<emphasis id="B96522260808FFF26345FB04FC036F2D" box="[871,1003,1269,1295]" italics="true" pageId="3" pageNumber="588">colombiana</emphasis>
</taxonomicName>
<emphasis id="B96522260808FFF263D6FB04FBA76F2D" bold="true" box="[1012,1103,1269,1295]" pageId="3" pageNumber="588">
<taxonomicNameLabel id="A2569F5D0808FFF263D6FB04FBA76F2D" box="[1012,1103,1269,1295]" pageId="3" pageNumber="588" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C1185B70808FFF264ABFB07FE666F11" authority="Grazia, Barros &amp; Barao, 2016" authorityName="Grazia, Barros &amp; Barao" authorityYear="2016" class="Insecta" family="Pentatomidae" genus="Elanela" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="588" phylum="Arthropoda" rank="species" species="diamphidiosa">
<emphasis id="B96522260808FFF264ABFB07FB726F2D" box="[1161,1178,1270,1295]" italics="true" pageId="3" pageNumber="588">E</emphasis>
.
<emphasis id="B96522260808FFF2648BFB04FAA96F2D" box="[1193,1345,1269,1295]" italics="true" pageId="3" pageNumber="588">diamphidiosa</emphasis>
<bibRefCitation id="EF8083C50808FFF26569FB04FE666F11" author="Grazia, J. &amp; Barros, L. D. &amp; Barao, K. R." pageId="3" pageNumber="588" pagination="561 - 571" refId="ref5065" refString="Grazia, J., Barros, L. D. &amp; Barao, K. R. (2016) Elanela Rolston revisited (Heteroptera: Pentatomidae): new distributional records and description of new species. Zootaxa, 4092 (4), 561 - 571. https: // doi. org / 10.11646 / zootaxa. 4092.4.6" type="journal article" year="2016">Grazia, Barros &amp; Barão, 2016</bibRefCitation>
</taxonomicName>
are alike, but in the former la8 lack spiracles, posterolateral angles of la8 acute, gc9 swollen, and
<emphasis id="B96522260808FFF260E5FACCFE676F75" box="[199,399,1341,1367]" italics="true" pageId="3" pageNumber="588">capsula seminalis</emphasis>
globose, bearing two projections, and shorter than
<emphasis id="B96522260808FFF263EEFACFFB716F75" box="[972,1177,1341,1367]" italics="true" pageId="3" pageNumber="588">pars intermedialis</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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