575 lines
73 KiB
XML
575 lines
73 KiB
XML
<document id="2D995CC41F488104A9D71DBE8B7A0564" ID-DOI="10.5281/zenodo.176298" ID-GBIF-Dataset="7f1966f0-92ea-4b0f-9e5c-a9492a71c57d" ID-ISSN="1175-5326" ID-Zenodo-Dep="176298" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459830639136" checkinUser="plazi" docAuthor="Pincheira-Donoso, Daniel & Scolaro, Alejandro" docDate="2007" docId="0399D46DFFA73B63FF3680DC7A108656" docLanguage="en" docName="zt01452p068.pdf" docOrigin="Zootaxa 1452" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Liolaemus tregenzai Pincheira-Donoso & Scolaro, 2007, sp. nov." docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="FFA0AC15FFA33B69FFA1804B795D8704" masterDocTitle="Iguanian species-richness in the Andes of boreal Patagonia: Evidence for an additional new Liolaemus lizard from Argentina lacking precloacal glands (Iguania, Liolaeminae)" masterLastPageNumber="68" masterPageNumber="55" pageNumber="59" updateTime="1698222916122" updateUser="plazi">
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<mods:mods id="8066DFA743EF743C17F1A5E28AD4588C" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5C6F7FD01049ABCF34BB0BBA3F4D01CF">
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<mods:title id="14FCC263F263F238B37B525C7858AFD4">Iguanian species-richness in the Andes of boreal Patagonia: Evidence for an additional new Liolaemus lizard from Argentina lacking precloacal glands (Iguania, Liolaeminae)</mods:title>
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</mods:titleInfo>
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<mods:name id="B26F54444D0FD8CC3416A6C2E17DA5D9" type="personal">
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<mods:roleTerm id="062F4F0A0022BD39227ED2DB319C505E">Author</mods:roleTerm>
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<mods:namePart id="B873FAC48D1DC5CF38FAD1C73FD459BE">Pincheira-Donoso, Daniel</mods:namePart>
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<mods:name id="2EB505B06034CFC761E2110B43FA8C3C" type="personal">
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<mods:roleTerm id="71D0B5136AB26CA2449FB68715B75006">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0E1DEB7735E673B00BE7C2EB8F698AA2">Scolaro, Alejandro</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="7A4615579C405263C7AA81608A0AC0CB">text</mods:typeOfResource>
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<mods:titleInfo id="72D3541E0C3378E06AD1F3B585BB8912">
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<mods:title id="A69A48F343394147AD19C794F12BCF15">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="23004E078FF882E5D4061E661DD014EF">
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<mods:date id="5901E50A39DDD3F240CFC32CE3124521">2007</mods:date>
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<mods:detail id="2AC96068CB744BD02ACDC4510E70443F" type="volume">
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<mods:number id="D2958A14E88891F766D5082647E65598">1452</mods:number>
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<mods:end id="612DE4064645FF7B9BCCCE29D8557FF8">68</mods:end>
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<mods:classification id="5BE5142C25EAAE7E3D85CECECF648A13">journal article</mods:classification>
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<mods:identifier id="D691FB1A024E10B4187C39FE725D5D1C" type="DOI">10.5281/zenodo.176298</mods:identifier>
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<mods:identifier id="CD9C35793D9270F97197F051166B4215" type="GBIF-Dataset">7f1966f0-92ea-4b0f-9e5c-a9492a71c57d</mods:identifier>
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<mods:identifier id="D04053B5DADC9C365702F2E79DA8AE9C" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D3A5D815D4BDE557C574A3B509BB9A59" type="Zenodo-Dep">176298</mods:identifier>
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<treatment id="0399D46DFFA73B63FF3680DC7A108656" ID-DOI="http://doi.org/10.5281/zenodo.6237641" ID-GBIF-Taxon="119357898" ID-Zenodo-Dep="6237641" LSID="urn:lsid:plazi:treatment:0399D46DFFA73B63FF3680DC7A108656" httpUri="http://treatment.plazi.org/id/0399D46DFFA73B63FF3680DC7A108656" lastPageId="10" lastPageNumber="65" pageId="4" pageNumber="59">
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<subSubSection id="C32A36F0FFA73B6DFF3680DC785987D0" pageId="4" pageNumber="59" type="nomenclature">
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<paragraph id="8B8F657BFFA73B6DFF3680DC78AD87B5" blockId="4.[151,496,151,212]" box="[151,496,151,177]" pageId="4" pageNumber="59">
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<heading id="D0C7D217FFA73B6DFF3680DC78AD87B5" bold="true" box="[151,496,151,177]" fontSize="11" level="1" pageId="4" pageNumber="59" reason="1">
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<emphasis id="B944B969FFA73B6DFF3680DC78AD87B5" bold="true" box="[151,496,151,177]" pageId="4" pageNumber="59">
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<taxonomicName id="4C301EF8FFA73B6DFF3680DC78D187B5" box="[151,396,151,177]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="tregenzai" status="sp. nov.">
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<emphasis id="B944B969FFA73B6DFF3680DC78D187B5" bold="true" box="[151,396,151,177]" italics="true" pageId="4" pageNumber="59">Liolaemus tregenzai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2770412FFA73B6DFE3280DC78AD87B5" box="[403,496,151,177]" pageId="4" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8B8F657BFFA73B6DFF3680F1785987D0" blockId="4.[151,496,151,212]" box="[151,260,186,212]" pageId="4" pageNumber="59">
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<figureCitation id="130B79FEFFA73B6DFF3680F1785987D0" box="[151,260,186,212]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="4.[151,255,1997,2021]" captionStartId-1="5.[151,255,1979,2003]" captionTargetBox-0="[264,1268,610,1983]" captionTargetBox-1="[179,1418,198,1954]" captionTargetId-0="figure@4.[264,1323,598,1983]" captionTargetId-1="figure@5.[176,1420,194,1956]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 1. Holotype (a; MLP. R. 5274) and female paratype (b; MLP. R. 5276) of Liolaemus tregenzai in dorsal view." captionText-1="FIGURE 2. Male (a) and female (b) of Liolaemus tregenzai in life." httpUri-0="https://zenodo.org/record/176299/files/figure.png" httpUri-1="https://zenodo.org/record/176300/files/figure.png" pageId="4" pageNumber="59">Figs. 1–2</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32A36F0FFA73B6DFF36814E7A9C8693" pageId="4" pageNumber="59" type="materials_examined">
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<paragraph id="8B8F657BFFA73B6DFF36814E7AB6866B" blockId="4.[151,1437,261,567]" pageId="4" pageNumber="59">
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<emphasis id="B944B969FFA73B6DFF36814E780B861B" bold="true" box="[151,342,261,287]" pageId="4" pageNumber="59">
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<typeStatus id="548BDBD9FFA73B6DFF36814E798F861B" box="[151,210,261,287]" pageId="4" pageNumber="59">Type</typeStatus>
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Material.
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</emphasis>
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<typeStatus id="548BDBD9FFA73B6DFEC9814E788E861A" box="[360,467,261,286]" pageId="4" pageNumber="59" type="holotype">
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<emphasis id="B944B969FFA73B6DFEC9814E788E861A" box="[360,467,261,286]" italics="true" pageId="4" pageNumber="59">Holotype</emphasis>
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</typeStatus>
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.
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<collectionCode id="ED21FDBEFFA73B6DFE4A814E7B76861B" box="[491,555,261,287]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="4" pageNumber="59">MLP</collectionCode>
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.R. 5274, adult male. Collected in Copahue Volcano slopes (
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<geoCoordinate id="EE0403BCFFA73B6DFA92814E7CCB861B" box="[1331,1430,261,287]" direction="south" orientation="latitude" pageId="4" pageNumber="59" precision="925" value="-37.833332">37°50’S</geoCoordinate>
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;
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<geoCoordinate id="EE0403BCFFA73B6DFF36816678588643" box="[151,261,301,327]" direction="west" orientation="longitude" pageId="4" pageNumber="59" precision="925" value="-71.1">71°06’W</geoCoordinate>
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),
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<quantity id="4CC8C89EFFA73B6DFEBC816678268643" box="[285,379,301,327]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="4" pageNumber="59" unit="m" value="2150.0">2150 m</quantity>
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, near Termas del Copahue, Departamento de Ñorquín, Neuquén Province,
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<collectingCountry id="F32725EBFFA73B6DFB5B81667C2F8643" box="[1274,1394,301,327]" name="Argentina" pageId="4" pageNumber="59">Argentina</collectingCountry>
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, by J.A. Scolaro, N. Scolaro, G. Scolaro and F. Tavera,
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<date id="FF8E43BBFFA73B6DFD4E811E7ABA866B" box="[751,999,341,367]" pageId="4" pageNumber="59" value="2006-02-06" valueMax="2006-02-10" valueMin="2006-02-06">February 6–10, 2006</date>
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.
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</paragraph>
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<paragraph id="8B8F657BFFA73B6DFF6781367A9C8693" blockId="4.[151,1437,261,567]" box="[198,961,381,407]" pageId="4" pageNumber="59">
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<emphasis id="B944B969FFA73B6DFF67813678758692" box="[198,296,381,406]" italics="true" pageId="4" pageNumber="59">Allotype</emphasis>
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.
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<collectionCode id="ED21FDBEFFA73B6DFE978136782B8693" box="[310,374,381,407]" httpUri="http://grbio.org/cool/66si-588n" name="Museo de La Plata" pageId="4" pageNumber="59">MLP</collectionCode>
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.R. 5275, adult female. The same data as
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<typeStatus id="548BDBD9FFA73B6DFCF381367AE18693" box="[850,956,381,407]" pageId="4" pageNumber="59" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A36F0FFA73B6DFF3681EE798C8533" pageId="4" pageNumber="59" type="description">
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<paragraph id="8B8F657BFFA73B6DFF3681EE798C8533" blockId="4.[151,1437,261,567]" pageId="4" pageNumber="59">
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<emphasis id="B944B969FFA73B6DFF3681EE785386BA" box="[151,270,421,446]" italics="true" pageId="4" pageNumber="59">
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<typeStatus id="548BDBD9FFA73B6DFF3681EE785386BA" box="[151,270,421,446]" pageId="4" pageNumber="59" type="paratype">Paratypes</typeStatus>
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</emphasis>
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. MLP.R. 5276-5277, both adult females. The same data detailed for the
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<typeStatus id="548BDBD9FFA73B6DFB2E81EE7DA886BB" box="[1167,1269,421,447]" pageId="4" pageNumber="59" type="holotype">holotype</typeStatus>
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and allotype. CHDPD 0 1098, adult male, CHDPD 0 1099, adult female. The same data detailed for the
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<typeStatus id="548BDBD9FFA73B6DFB6581867C7786E3" box="[1220,1322,461,487]" pageId="4" pageNumber="59" type="holotype">holotype</typeStatus>
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and allotype. JAS-DC 806, adult male, JAS-DC 807, adult female. The same data detailed for the
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<typeStatus id="548BDBD9FFA73B6DFB6481BE7C71850B" box="[1221,1324,501,527]" pageId="4" pageNumber="59" type="holotype">holotype</typeStatus>
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and allotype.
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</paragraph>
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||
</subSubSection>
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||
<caption id="DF4F35F3FFA73B6DFF3687867CD880E1" box="[151,1413,1997,2021]" httpUri="https://zenodo.org/record/176299/files/figure.png" pageId="4" pageNumber="59" targetBox="[264,1268,610,1983]" targetPageId="4">
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||
<paragraph id="8B8F657BFFA73B6DFF3687867CD880E1" blockId="4.[151,1413,1997,2021]" box="[151,1413,1997,2021]" pageId="4" pageNumber="59">
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||
<emphasis id="B944B969FFA73B6DFF368786784480E1" bold="true" box="[151,281,1997,2021]" pageId="4" pageNumber="59">FIGURE 1.</emphasis>
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Holotype (a; MLP.R. 5274) and female paratype (b; MLP.R. 5276) of
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<taxonomicName id="4C301EF8FFA73B6DFBA887857DBC80E1" box="[1033,1249,1998,2021]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="59" phylum="Chordata" rank="species" species="tregenzai">
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<emphasis id="B944B969FFA73B6DFBA887857DBC80E1" box="[1033,1249,1998,2021]" italics="true" pageId="4" pageNumber="59">Liolaemus tregenzai</emphasis>
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</taxonomicName>
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in dorsal view.
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</paragraph>
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</caption>
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<caption id="DF4F35F3FFA63B6CFF3687F07A3880D6" box="[151,869,1979,2003]" httpUri="https://zenodo.org/record/176300/files/figure.png" pageId="5" pageNumber="60" targetBox="[179,1418,198,1954]" targetPageId="5">
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<paragraph id="8B8F657BFFA63B6CFF3687F07A3880D6" blockId="5.[151,869,1979,2003]" box="[151,869,1979,2003]" pageId="5" pageNumber="60">
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<emphasis id="B944B969FFA63B6CFF3687F0784480D7" bold="true" box="[151,281,1979,2003]" pageId="5" pageNumber="60">FIGURE 2.</emphasis>
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Male (a) and female (b) of
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<taxonomicName id="4C301EF8FFA63B6CFDE187F07A4580D6" box="[576,792,1979,2002]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="60" phylum="Chordata" rank="species" species="tregenzai">
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<emphasis id="B944B969FFA63B6CFDE187F07A4580D6" box="[576,792,1979,2002]" italics="true" pageId="5" pageNumber="60">Liolaemus tregenzai</emphasis>
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</taxonomicName>
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in life.
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</paragraph>
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</caption>
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<subSubSection id="C32A36F0FFA53B6FFF6780DC78CD862C" pageId="6" pageNumber="61" type="etymology">
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<paragraph id="8B8F657BFFA53B6FFF6780DC78CD862C" blockId="6.[151,1437,151,2036]" pageId="6" pageNumber="61">
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<emphasis id="B944B969FFA53B6FFF6780DC780E87B5" bold="true" box="[198,339,151,177]" pageId="6" pageNumber="61">Etymology.</emphasis>
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This new lizard species is named
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<taxonomicName id="4C301EF8FFA53B6FFD5880D37AB787B5" box="[761,1002,152,177]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
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<emphasis id="B944B969FFA53B6FFD5880D37AB787B5" box="[761,1002,152,177]" italics="true" pageId="6" pageNumber="61">Liolaemus tregenzai</emphasis>
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</taxonomicName>
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after Tom Tregenza (Royal Society Research Fellow, University of Exeter), in recognition of his contributions to evolutionary biology. This dedication is also a personal recognition for the time spent guiding DP-D through a doctoral period of research in the
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<collectingCountry id="F32725EBFFA53B6FFF63814578D1862C" box="[194,396,270,296]" name="United Kingdom" pageId="6" pageNumber="61">United Kingdom</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A36F0FFA53B6FFF67817E7B6C8145" pageId="6" pageNumber="61" type="diagnosis">
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<paragraph id="8B8F657BFFA53B6FFF67817E7B6C8145" blockId="6.[151,1437,151,2036]" pageId="6" pageNumber="61">
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<emphasis id="B944B969FFA53B6FFF67817E781B864B" bold="true" box="[198,326,309,335]" pageId="6" pageNumber="61">Diagnosis.</emphasis>
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A large
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<taxonomicName id="4C301EF8FFA53B6FFE12817D7B70864B" box="[435,557,310,335]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="genus">
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<emphasis id="B944B969FFA53B6FFE12817D7B70864B" box="[435,557,310,335]" italics="true" pageId="6" pageNumber="61">Liolaemus</emphasis>
|
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</taxonomicName>
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species belonging to the
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<taxonomicName id="4C301EF8FFA53B6FFCC2817D7A8F864B" box="[867,978,310,335]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="chiliensis">
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<emphasis id="B944B969FFA53B6FFCC2817D7A8F864B" box="[867,978,310,335]" italics="true" pageId="6" pageNumber="61">chiliensis</emphasis>
|
||
</taxonomicName>
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clade diagnosed by
|
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<bibRefCitation id="EFA1188AFFA53B6FFB6A817D7CCA8654" author="Etheridge" box="[1227,1431,310,336]" pageId="6" pageNumber="67" refString="Etheridge, R. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">Etheridge (1995)</bibRefCitation>
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, because it has a low number (4-4) of narrow supralabial scales, the last one elongated and upturned in the posterior margin, without a bladelike process on the posterior distal tibia, without femoral patch of enlarged scales, without tridentate dorsal scales, and with Meckel’s groove fused (see also
|
||
<bibRefCitation id="EFA1188AFFA53B6FFBF481E77DA886C2" author="Laurent" box="[1109,1269,428,454]" pageId="6" pageNumber="67" refString="Laurent, R. F. (1992) On some overlooked species of the genus Liolaemus Wiegmann (Reptilia Tropiduridae) from Peru. Breviora, 494, 1 - 33." type="journal article" year="1992">Laurent 1992</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA1188AFFA53B6FFAA381E7798886EA" author="Schulte" pageId="6" pageNumber="68" refString="Schulte, J. A., Macey, J. R., Espinoza, R. E. & Larson, A. (2000) Phylogenetic relationships in the iguanid lizard genus Liolaemus: multiple origins of viviparous reproduction and evidence for recurring Andean vicariance and dispersal. Biological Journal of the Linnean Society, 69, 75 - 102." type="journal article" year="2000">
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||
Schulte
|
||
<emphasis id="B944B969FFA53B6FFAC081E67CC186C2" box="[1377,1436,429,454]" italics="true" pageId="6" pageNumber="61">et al.</emphasis>
|
||
2000
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA1188AFFA53B6FFF42819F783A86EA" author="Lobo" box="[227,359,468,494]" pageId="6" pageNumber="67" refString="Lobo, F. (2001) A Phylogenetic analysis of lizards of the Liolaemus chiliensis group (Iguania: Tropiduridae). Herpetological Journal, 11, 137 - 150." type="journal article" year="2001">Lobo 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA1188AFFA53B6FFED4819F7A5A86EA" author="Pincheira-Donoso" box="[373,775,468,494]" pageId="6" pageNumber="68" refString="Pincheira-Donoso, D. & Nunez, H. (2005) Las especies chilenas del genero Liolaemus (Iguania, Tropiduridae, Liolaeminae). Taxonomia, sistematica y evolucion. Publicacion Ocasional del Museo Nacional de Historia Natural de Chile, Santiago, 486 pp." type="book" year="2005">Pincheira-Donoso & Núñez 2005</bibRefCitation>
|
||
). All these traits differentiate
|
||
<taxonomicName id="4C301EF8FFA53B6FFBCA819F7DAA86E9" box="[1131,1271,468,493]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFBCA819F7DAA86E9" box="[1131,1271,468,493]" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<emphasis id="B944B969FFA53B6FFAE3819F78D98511" italics="true" pageId="6" pageNumber="61">
|
||
<taxonomicName id="4C301EF8FFA53B6FFAE3819F79A28511" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="periglacialis">L. periglacialis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C301EF8FFA53B6FFEAB81B778D98511" box="[266,388,508,533]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="kolengh">L. kolengh</taxonomicName>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C301EF8FFA53B6FFE3281B77B398511" box="[403,612,508,533]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="lineomaculatus">
|
||
<emphasis id="B944B969FFA53B6FFE3281B77B398511" box="[403,612,508,533]" italics="true" pageId="6" pageNumber="61">L. lineomaculatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C301EF8FFA53B6FFD3D81B77A418511" box="[668,796,508,533]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="silvanae">
|
||
<emphasis id="B944B969FFA53B6FFD3D81B77A418511" box="[668,796,508,533]" italics="true" pageId="6" pageNumber="61">L. silvanae</emphasis>
|
||
</taxonomicName>
|
||
, which exhibit the diagnostic traits detailed for the
|
||
<taxonomicName id="4C301EF8FFA53B6FFAD281B778768538" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="lineomaculatus">
|
||
<emphasis id="B944B969FFA53B6FFAD281B778768538" italics="true" pageId="6" pageNumber="61">lineomaculatus</emphasis>
|
||
</taxonomicName>
|
||
clade (see
|
||
<bibRefCitation id="EFA1188AFFA53B6FFE0E82687BAA8539" author="Donoso-Barros" box="[431,759,547,573]" pageId="6" pageNumber="67" refString="Donoso-Barros, R. & Cei, J. M. (1971) New lizards from the volcanic Patagonian plateau of Argentina. Journal of Herpetology, 5, 89 - 95." type="journal article" year="1971">Donoso-Barros & Cei 1971</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA1188AFFA53B6FFCA582687A2F8539" author="Cei" box="[772,882,547,573]" pageId="6" pageNumber="67" refString="Cei, J. M. (1986) Reptiles del centro, centro-oeste y sur de la Argentina. Herpetofauna de las zonas aridas y semiaridas, Museo Regionale di Scienze Naturali, Torino, 527 pp." type="book" year="1986">Cei 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA1188AFFA53B6FFCDE82687D6A8539" author="Etheridge" box="[895,1079,547,573]" pageId="6" pageNumber="67" refString="Etheridge, R. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">Etheridge 1995</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFA1188AFFA53B6FFBED82687C498539" author="Etheridge" box="[1100,1300,547,573]" pageId="6" pageNumber="67" refString="Etheridge, R. (1995) Redescription of Ctenoblepharys adspersa Tschudi, 1845, and the taxonomy of Liolaeminae (Reptilia: Squamata: Tropiduridae). American Museum Novitates, 3142, 1 - 34." type="journal article" year="1995">Etheridge (1995)</bibRefCitation>
|
||
recognized four or fewer precloacal glands in males of most of the species belonging to the
|
||
<taxonomicName id="4C301EF8FFA53B6FFBE182007DF28560" box="[1088,1199,587,612]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="chiliensis">
|
||
<emphasis id="B944B969FFA53B6FFBE182007DF28560" box="[1088,1199,587,612]" italics="true" pageId="6" pageNumber="61">chiliensis</emphasis>
|
||
</taxonomicName>
|
||
clade. In
|
||
<taxonomicName id="4C301EF8FFA53B6FFA838200785E858F" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFA838200785E858F" italics="true" pageId="6" pageNumber="61">Liolaemus tregenzai</emphasis>
|
||
</taxonomicName>
|
||
these glands are entirely absents, which differentiate this species from almost every taxa into this clade, except from
|
||
<taxonomicName id="4C301EF8FFA53B6FFEDC82D17B4D85B7" box="[381,528,666,691]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="coeruleus">
|
||
<emphasis id="B944B969FFA53B6FFEDC82D17B4D85B7" box="[381,528,666,691]" italics="true" pageId="6" pageNumber="61">L. coeruleus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C301EF8FFA53B6FFD8182D17BFE85B7" box="[544,675,666,691]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="cristiani">
|
||
<emphasis id="B944B969FFA53B6FFD8182D17BFE85B7" box="[544,675,666,691]" italics="true" pageId="6" pageNumber="61">L. cristiani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C301EF8FFA53B6FFD1582D17A0E85B7" box="[692,851,666,691]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="flavipiceus">
|
||
<emphasis id="B944B969FFA53B6FFD1582D17A0E85B7" box="[692,851,666,691]" italics="true" pageId="6" pageNumber="61">L. flavipiceus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C301EF8FFA53B6FFCC582D17D5A85B7" box="[868,1031,666,691]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="neuquensis">
|
||
<emphasis id="B944B969FFA53B6FFCC582D17D5A85B7" box="[868,1031,666,691]" italics="true" pageId="6" pageNumber="61">L. neuquensis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C301EF8FFA53B6FFBEA82D17DB385B7" box="[1099,1262,666,691]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="thermarum">
|
||
<emphasis id="B944B969FFA53B6FFBEA82D17DB385B7" box="[1099,1262,666,691]" italics="true" pageId="6" pageNumber="61">L. thermarum</emphasis>
|
||
</taxonomicName>
|
||
, which do not exhibit precloacal glands.
|
||
<taxonomicName id="4C301EF8FFA53B6FFE73828A7B9F85DE" box="[466,706,705,730]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFE73828A7B9F85DE" box="[466,706,705,730]" italics="true" pageId="6" pageNumber="61">Liolaemus tregenzai</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C301EF8FFA53B6FFCC5828A7ABA85DE" box="[868,999,705,730]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="cristiani">
|
||
<emphasis id="B944B969FFA53B6FFCC5828A7ABA85DE" box="[868,999,705,730]" italics="true" pageId="6" pageNumber="61">L. cristiani</emphasis>
|
||
</taxonomicName>
|
||
in having a Patagonian distribution, and green-bluish ventral coloration with intense black pigment on the throat, chest and abdomen, whereas
|
||
<taxonomicName id="4C301EF8FFA53B6FFA2582A279A4842E" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="cristiani">
|
||
<emphasis id="B944B969FFA53B6FFA2582A279A4842E" italics="true" pageId="6" pageNumber="61">L. cristiani</emphasis>
|
||
</taxonomicName>
|
||
occurs in the Andes of central
|
||
<collectingCountry id="F32725EBFFA53B6FFDC6835B7BF4842E" box="[615,681,784,810]" name="Chile" pageId="6" pageNumber="61">Chile</collectingCountry>
|
||
, and exhibit a brown-yellowish or copper-yellowish ventral coloration, with total absence of intense black pigmentation in both sexes. Also, in
|
||
<taxonomicName id="4C301EF8FFA53B6FFBE183737D978455" box="[1088,1226,824,849]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFBE183737D978455" box="[1088,1226,824,849]" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
sexual dichromatism is evident, being absent in
|
||
<taxonomicName id="4C301EF8FFA53B6FFD88832B7BF2847D" box="[553,687,864,889]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="cristiani">
|
||
<emphasis id="B944B969FFA53B6FFD88832B7BF2847D" box="[553,687,864,889]" italics="true" pageId="6" pageNumber="61">L. cristiani</emphasis>
|
||
</taxonomicName>
|
||
. Significant differences in the colour pattern differentiate
|
||
<taxonomicName id="4C301EF8FFA53B6FFA25832B785E84A4" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFA25832B785E84A4" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C301EF8FFA53B6FFEEF83CC78B384A4" box="[334,494,903,928]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="flavipiceus">
|
||
<emphasis id="B944B969FFA53B6FFEEF83CC78B384A4" box="[334,494,903,928]" italics="true" pageId="6" pageNumber="61">L. flavipiceus</emphasis>
|
||
</taxonomicName>
|
||
. The olive, brown-blackish or chest-nut dorsal coloration and the green-bluish ventral colour with intense black pigmentation characteristics of
|
||
<taxonomicName id="4C301EF8FFA53B6FFC3583E47D7D84CC" box="[916,1056,943,968]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFC3583E47D7D84CC" box="[916,1056,943,968]" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
, contrast with the yellow dorsal background, black on the head, flanks, throat and chest, and with the greyish or reddish ventral colour with scattered orange or reddish spots observed in
|
||
<taxonomicName id="4C301EF8FFA53B6FFD6383B57A3E8313" box="[706,867,1022,1047]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="flavipiceus">
|
||
<emphasis id="B944B969FFA53B6FFD6383B57A3E8313" box="[706,867,1022,1047]" italics="true" pageId="6" pageNumber="61">L. flavipiceus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C301EF8FFA53B6FFCD583B57D398313" box="[884,1124,1022,1047]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFCD583B57D398313" box="[884,1124,1022,1047]" italics="true" pageId="6" pageNumber="61">Liolaemus tregenzai</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<taxonomicName id="4C301EF8FFA53B6FFAE083B579B7833B" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="thermarum">
|
||
<emphasis id="B944B969FFA53B6FFAE083B579B7833B" italics="true" pageId="6" pageNumber="61">L. thermarum</emphasis>
|
||
</taxonomicName>
|
||
, because this last species does not exhibit black pigmentation on the ventral surface, which is whitish with conspicuous pinkish shades. The
|
||
<taxonomicName id="4C301EF8FFA53B6FFDC084067B868362" box="[609,731,1101,1126]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="genus">
|
||
<emphasis id="B944B969FFA53B6FFDC084067B868362" box="[609,731,1101,1126]" italics="true" pageId="6" pageNumber="61">Liolaemus</emphasis>
|
||
</taxonomicName>
|
||
species lacking precloacal glands that exhibit more phenotypic and geographical similitude with
|
||
<taxonomicName id="4C301EF8FFA53B6FFDD2843E7A5F838A" box="[627,770,1141,1166]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFDD2843E7A5F838A" box="[627,770,1141,1166]" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
are
|
||
<taxonomicName id="4C301EF8FFA53B6FFC9A843E7A8D838A" box="[827,976,1141,1166]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="coeruleus">
|
||
<emphasis id="B944B969FFA53B6FFC9A843E7A8D838A" box="[827,976,1141,1166]" italics="true" pageId="6" pageNumber="61">L. coeruleus</emphasis>
|
||
</taxonomicName>
|
||
, from Zapala and Pino Hachado (
|
||
<bibRefCitation id="EFA1188AFFA53B6FFAD3843F798883B2" author="Cei" pageId="6" pageNumber="67" refString="Cei, J. M. (1986) Reptiles del centro, centro-oeste y sur de la Argentina. Herpetofauna de las zonas aridas y semiaridas, Museo Regionale di Scienze Naturali, Torino, 527 pp." type="book" year="1986">Cei 1986</bibRefCitation>
|
||
; Pincheira-Donoso 2003) and
|
||
<taxonomicName id="4C301EF8FFA53B6FFDE284D77BBB83B1" box="[579,742,1180,1205]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="neuquensis">
|
||
<emphasis id="B944B969FFA53B6FFDE284D77BBB83B1" box="[579,742,1180,1205]" italics="true" pageId="6" pageNumber="61">L. neuquensis</emphasis>
|
||
</taxonomicName>
|
||
, from Copahue (
|
||
<bibRefCitation id="EFA1188AFFA53B6FFC1284D77D7883B2" author="Cei" box="[947,1061,1180,1206]" pageId="6" pageNumber="67" refString="Cei, J. M. (1986) Reptiles del centro, centro-oeste y sur de la Argentina. Herpetofauna de las zonas aridas y semiaridas, Museo Regionale di Scienze Naturali, Torino, 527 pp." type="book" year="1986">Cei 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFA1188AFFA53B6FFB9284D77C4683B2" author="Videla" box="[1075,1307,1180,1206]" pageId="6" pageNumber="68" refString="Videla, F. & Cei, J. M. (1996) A new peculiar Liolaemus species of the " chiliensis " phyletic group from the volcanic Cordilleran landscapes of southern Mendoza Province, Argentina (Iguania, Lacertilia, Reptilia). Bollettino del Museo Regionale di Scienze Naturali di Torino, 14, 505 - 516." type="journal article" year="1996">Videla & Cei 1996</bibRefCitation>
|
||
), the same area where the new species occurs (see above).
|
||
<taxonomicName id="4C301EF8FFA53B6FFD7E848F7A8D83D9" box="[735,976,1220,1245]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFD7E848F7A8D83D9" box="[735,976,1220,1245]" italics="true" pageId="6" pageNumber="61">Liolaemus tregenzai</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C301EF8FFA53B6FFBD5848F7C5483D9" box="[1140,1289,1220,1245]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="coeruleus">
|
||
<emphasis id="B944B969FFA53B6FFBD5848F7C5483D9" box="[1140,1289,1220,1245]" italics="true" pageId="6" pageNumber="61">L. coeruleus</emphasis>
|
||
</taxonomicName>
|
||
in having a body size ranging between
|
||
<quantity id="4CC8C89EFFA53B6FFE4284A07BCD8201" box="[483,656,1259,1285]" metricMagnitude="-2" metricUnit="m" metricValue="7.68" metricValueMax="9.02" metricValueMin="6.34" pageId="6" pageNumber="61" unit="mm" value="76.8" valueMax="90.2" valueMin="63.4">63.4–90.2 mm</quantity>
|
||
(see below for details) which contrasts significantly (see above) with the
|
||
<quantity id="4CC8C89EFFA53B6FFEA0855978F28228" box="[257,431,1298,1324]" metricMagnitude="-2" metricUnit="m" metricValue="5.245" metricValueMax="6.02" metricValueMin="4.47" pageId="6" pageNumber="61" unit="mm" value="52.45" valueMax="60.2" valueMin="44.7">44.7–60.2 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C301EF8FFA53B6FFE7985587B368228" box="[472,619,1299,1324]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="coeruleus">
|
||
<emphasis id="B944B969FFA53B6FFE7985587B368228" box="[472,619,1299,1324]" italics="true" pageId="6" pageNumber="61">L. coeruleus</emphasis>
|
||
</taxonomicName>
|
||
. Also, the number of scales around the midbody (and therefore, the proportional size of body scales in relation to body size) is significantly different between these species, having
|
||
<taxonomicName id="4C301EF8FFA53B6FFF658529780C827F" box="[196,337,1378,1403]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFF658529780C827F" box="[196,337,1378,1403]" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
a mean value of 80.63, whereas the mean value in
|
||
<taxonomicName id="4C301EF8FFA53B6FFC1385297D19827F" box="[946,1092,1378,1403]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="coeruleus">
|
||
<emphasis id="B944B969FFA53B6FFC1385297D19827F" box="[946,1092,1378,1403]" italics="true" pageId="6" pageNumber="61">L. coeruleus</emphasis>
|
||
</taxonomicName>
|
||
is 63.4. In addition, while
|
||
<taxonomicName id="4C301EF8FFA53B6FFA258529785E82A6" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFA258529785E82A6" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
exhibits a green-bluish ventral coloration, the abdominal and pectoral background in
|
||
<taxonomicName id="4C301EF8FFA53B6FFB5185C27CDC82A6" box="[1264,1409,1417,1442]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="coeruleus">
|
||
<emphasis id="B944B969FFA53B6FFB5185C27CDC82A6" box="[1264,1409,1417,1442]" italics="true" pageId="6" pageNumber="61">L. coeruleus</emphasis>
|
||
</taxonomicName>
|
||
is blue or sky-blue. Finally,
|
||
<taxonomicName id="4C301EF8FFA53B6FFE6E85FA7B0182CE" box="[463,604,1457,1482]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFE6E85FA7B0182CE" box="[463,604,1457,1482]" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C301EF8FFA53B6FFD5E85FA7AFE82CE" box="[767,931,1457,1482]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="neuquensis">
|
||
<emphasis id="B944B969FFA53B6FFD5E85FA7AFE82CE" box="[767,931,1457,1482]" italics="true" pageId="6" pageNumber="61">L. neuquensis</emphasis>
|
||
</taxonomicName>
|
||
in having a significantly larger body size (
|
||
<quantity id="4CC8C89EFFA53B6FFF3E8593781082F6" box="[159,333,1496,1522]" metricMagnitude="-2" metricUnit="m" metricValue="5.82" metricValueMax="6.11" metricValueMin="5.529999999999999" pageId="6" pageNumber="61" unit="mm" value="58.2" valueMax="61.1" valueMin="55.3">55.3–61.1 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C301EF8FFA53B6FFED485927B4A82F6" box="[373,535,1497,1522]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="neuquensis">
|
||
<emphasis id="B944B969FFA53B6FFED485927B4A82F6" box="[373,535,1497,1522]" italics="true" pageId="6" pageNumber="61">L. neuquensis</emphasis>
|
||
</taxonomicName>
|
||
; see above), and because
|
||
<taxonomicName id="4C301EF8FFA53B6FFCF185927AAE82F6" box="[848,1011,1497,1522]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="neuquensis">
|
||
<emphasis id="B944B969FFA53B6FFCF185927AAE82F6" box="[848,1011,1497,1522]" italics="true" pageId="6" pageNumber="61">L. neuquensis</emphasis>
|
||
</taxonomicName>
|
||
shows a prominent hemigular fold, completely absent in
|
||
<taxonomicName id="4C301EF8FFA53B6FFE32864B7B42811D" box="[403,543,1536,1561]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFE32864B7B42811D" box="[403,543,1536,1561]" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
. The same ventral differences detailed for
|
||
<taxonomicName id="4C301EF8FFA53B6FFBBB864B7DF1811D" box="[1050,1196,1536,1561]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="coeruleus">
|
||
<emphasis id="B944B969FFA53B6FFBBB864B7DF1811D" box="[1050,1196,1536,1561]" italics="true" pageId="6" pageNumber="61">L. coeruleus</emphasis>
|
||
</taxonomicName>
|
||
in relation to
|
||
<taxonomicName id="4C301EF8FFA53B6FFAF2864B79BE8145" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA53B6FFAF2864B79BE8145" italics="true" pageId="6" pageNumber="61">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
, are valid for
|
||
<taxonomicName id="4C301EF8FFA53B6FFE2986637B748145" box="[392,553,1576,1601]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="61" phylum="Chordata" rank="species" species="neuquensis">
|
||
<emphasis id="B944B969FFA53B6FFE2986637B748145" box="[392,553,1576,1601]" italics="true" pageId="6" pageNumber="61">L. neuquensis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A36F0FFA53B61FF6786047D3D80E9" lastPageId="8" lastPageNumber="63" pageId="6" pageNumber="61" type="description">
|
||
<paragraph id="8B8F657BFFA53B6FFF67860479B78052" blockId="6.[151,1437,151,2036]" pageId="6" pageNumber="61">
|
||
<emphasis id="B944B969FFA53B6FFF6786047B7D816D" bold="true" box="[198,544,1615,1641]" pageId="6" pageNumber="61">
|
||
Description of the
|
||
<typeStatus id="548BDBD9FFA53B6FFE0F86047B7D816D" box="[430,544,1615,1641]" pageId="6" pageNumber="61" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="130B79FEFFA53B6FFD9186047B2B816D" box="[560,630,1615,1641]" captionStart="FIGURE 1" captionStartId="4.[151,255,1997,2021]" captionTargetBox="[264,1268,610,1983]" captionTargetId="figure@4.[264,1323,598,1983]" captionTargetPageId="4" captionText="FIGURE 1. Holotype (a; MLP. R. 5274) and female paratype (b; MLP. R. 5276) of Liolaemus tregenzai in dorsal view." httpUri="https://zenodo.org/record/176299/files/figure.png" pageId="6" pageNumber="61">Fig. 1</figureCitation>
|
||
). Adult male, snout-vent length
|
||
<quantity id="4CC8C89EFFA53B6FFC5786047D3F816D" box="[1014,1122,1615,1641]" metricMagnitude="-2" metricUnit="m" metricValue="8.25" pageId="6" pageNumber="61" unit="mm" value="82.5">82.5 mm</quantity>
|
||
, forelimb length
|
||
<quantity id="4CC8C89EFFA53B6FFA8D86047CC5816D" box="[1324,1432,1615,1641]" metricMagnitude="-2" metricUnit="m" metricValue="3.2299999999999995" pageId="6" pageNumber="61" unit="mm" value="32.3">32.3 mm</quantity>
|
||
, hind limb length
|
||
<quantity id="4CC8C89EFFA53B6FFEC8863D788A8194" box="[361,471,1654,1680]" metricMagnitude="-2" metricUnit="m" metricValue="4.79" pageId="6" pageNumber="61" unit="mm" value="47.9">47.9 mm</quantity>
|
||
. Tail broken. Head length 18.0 mm, head width
|
||
<quantity id="4CC8C89EFFA53B6FFB87863D7DC98194" box="[1062,1172,1654,1680]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="6" pageNumber="61" unit="mm" value="16.5">16.5 mm</quantity>
|
||
, head height
|
||
<quantity id="4CC8C89EFFA53B6FFA99863D7CC58194" box="[1336,1432,1654,1680]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="6" pageNumber="61" unit="mm" value="9.2">9.2 mm</quantity>
|
||
. Stout body, with prominent head.
|
||
<collectingCountry id="F32725EBFFA53B6FFD8E86D57B3781BC" box="[559,618,1694,1720]" name="India" pageId="6" pageNumber="61">Hind</collectingCountry>
|
||
limb adpressed reaches the shoulder. Neck wider than head. Lateral cervical folds, postauricaular, supra-auricular, longitudinal neck, oblique neck, and antehumeral, prominent, forming a V behind the external auditory meatus. Deep prehumeral pocket. Lateral nuchal scales small, granular, convex, nonoverlapping, with numerous tiny granules in the interstitial spaces. Hemigular fold entirely absent.
|
||
</paragraph>
|
||
<paragraph id="8B8F657BFFA53B6FFF67872F7CC180F0" blockId="6.[151,1437,151,2036]" pageId="6" pageNumber="61">Upper head scales smooth, pitted with numerous sensitive organs mainly on snout. Rostral shield pentagonal, about four times as wide as high, and as wide as mental, directly surrounded by six scales (two supralabials, two lorilabials and two postrostrals). Two postrostrals, together with anterior supralabials almost separate nasals from rostral. Nasal shields longer than wide, as large as postrostrals, surrounded by six scales.</paragraph>
|
||
<paragraph id="8B8F657BFFA43B6EFF3680D37BAF847E" blockId="7.[151,1437,152,2036]" pageId="7" pageNumber="62">Nostril rounded, slightly over one half length of nasal, posterior in position. Four internasal scales, the medial pair as large as postrostrals. Two irregularly shaped azygous forming a longitudinal row on the snout, the anterior twice as large. The posterior in direct contact with a prefrontal azygous. A frontal azygous, slightly larger than interparietal. Two postfrontal shields, smaller than interparietal. Interparietal shield small, as large as nasals, pentagonal, slightly enlarged, flanked posterolaterally by a pair of irregularly shaped parietal scales relatively larger than frontal azygous, and anteriorly in direct contact with postfrontals. Parietal eye small and conspicuous, posterior in position. Supratemporal scales relatively convexes, smooth, irregularly shaped and exhibiting high variation in size. Temporal scales more regulars, weakly keeled and subimbricate or juxtaposed. Circum orbitalis conspicuous, formed by 13-13 small and often hexagonal scales. Supraoculars 6-4, strongly hexagonals and transversally expanded, up to eight times larger than circum orbitalis scales. Three rows of smaller scales disposed between supraorbitals and superciliars. Superciliar scales longitudinally expanded, imbricate. Canthal margin inconspicuous. Loreal region weakly deep, covered by 5-5 irregularly shaped scales which exhibit high variability in size. Ciliar scales do not form palpebral “combs”. Upper ciliar scales 15-15, lower ciliars 11-10. A single and longitudinally enlarged subocular shield. Supralabial scales 4- 4, between two and three times longer than high, the fourth scale is evidently upturned on the posterior margin. Between subocular and supralabials there is a single row of lorilabial scales whose size is similar or slightly smaller than supralabials. External auditory meatus as large as the eye surface, oval, and higher than wide. Tympanic scales 3-3, convex, slightly projected backwards over external auditory meatus. Auricular scales 2-2, weakly convexes, as large as tympanics.</paragraph>
|
||
<paragraph id="8B8F657BFFA43B6EFF6783CC7D1D833B" blockId="7.[151,1437,152,2036]" pageId="7" pageNumber="62">Mental shield about twice as wide as high, surrounded by the anterior infralabials and the anterior postmentals, but not in contact with anterior sublabials. Two divergent and decreasing rows of 5-5 postmental scales. Infralabial shields 5-5, between two and three times wider than high, whose surface is similar or slightly larger than supralabials. A single row of sublabial scales between the anterior infralabials and the anterior postmentals. Gular scales smooth, flat, imbricate, with rounded margin.</paragraph>
|
||
<paragraph id="8B8F657BFFA43B6EFF678406788B8228" blockId="7.[151,1437,152,2036]" pageId="7" pageNumber="62">Dorsal body scales small, whose surface represents 45–60% of the interparietal shield, subtriangular, subimbricate, with a conspicuous but blunt keel, and without mucrons. There are scarce tiny and granular scales in the interstitial spaces. Dorsal scales grade laterally into slightly smaller scales, being fundamentally similar, but with abundant tiny granules in the interstices. Scales surrounding forelimb insertion small, convex, granular and juxtaposed. Ventral body scales 1.5–2 larger than dorsal scales, rounded, smooth, flat, and imbricate. Scales around midbody 81.</paragraph>
|
||
<paragraph id="8B8F657BFFA43B6EFF6785717C27807A" blockId="7.[151,1437,152,2036]" pageId="7" pageNumber="62">Forelimb scales as large as the dorsal body scales, subtriangular, imbricate and slightly keeled, becoming smooth on the dorsal surface of the hand. Lower humeral region covered by small, convex, granular, and juxtaposed, becoming larger, rounded, subimbricate and weakly keeled on the forearm. Several scales on this zone exhibit a tridentate margin. Infracarpal scales smaller, strongly imbricate, with well developed keels, and often with a tridentate margin. Subdigital lamellae laterally expanded, with three prominent keels, often with three conspicuous mucrons in the margin, although several of them can exhibit up to five mucrons. Posterior surface of thighs with tiny, granular, convex and juxtaposed scales. Upper hind limb scales slightly larger than dorsal scales, subtriangular, subimbricate, and weakly keeled on the femoral area, grading into more rounded, more keeled and subimbricate-juxtaposed on the tibial area. Supratarsal scales almost smooth, triangular and imbricate. On the lower femoral surface (infrafemorals), scales are smooth, flat, rounded and imbricate, grading into slightly larger scales on the tibial zone (infratibials). Infratarsal scales small, strongly imbricate, with well developed keels, and with a tridentate margin. Subdigital lamellae laterally expanded, with 3–4 strong keels, and with up to six prominent mucrons on the margin. Subdigital lamellae on the fourth toe of the left foot 26 (non-measurable on the right foot). Upper caudals larger than dorsal body scales, keeled and imbricate. On the ventral surface, scales are triangular, slightly keeled and imbricate. Without precloacal glands.</paragraph>
|
||
<paragraph id="8B8F657BFFA43B6EFF6787C0782E80F0" blockId="7.[151,1437,152,2036]" pageId="7" pageNumber="62">
|
||
<emphasis id="B944B969FFA43B6EFF6787C0780880A1" bold="true" box="[198,341,1931,1957]" pageId="7" pageNumber="62">Coloration.</emphasis>
|
||
Based in the whole
|
||
<typeStatus id="548BDBD9FFA43B6EFDE587C07B2B80A1" box="[580,630,1931,1957]" pageId="7" pageNumber="62">type</typeStatus>
|
||
series, including males and females. A sexually dichromatic species (
|
||
<figureCitation id="130B79FEFFA43B6EFF3E87F879AD80C9" box="[159,240,1971,1997]" captionStart="FIGURE 1" captionStartId="4.[151,255,1997,2021]" captionTargetBox="[264,1268,610,1983]" captionTargetId="figure@4.[264,1323,598,1983]" captionTargetPageId="4" captionText="FIGURE 1. Holotype (a; MLP. R. 5274) and female paratype (b; MLP. R. 5276) of Liolaemus tregenzai in dorsal view." httpUri="https://zenodo.org/record/176299/files/figure.png" pageId="7" pageNumber="62">Figs. 1</figureCitation>
|
||
and
|
||
<figureCitation id="130B79FEFFA43B6EFE8A87F8786180C9" box="[299,316,1971,1997]" captionStart="FIGURE 2" captionStartId="5.[151,255,1979,2003]" captionTargetBox="[179,1418,198,1954]" captionTargetId="figure@5.[176,1420,194,1956]" captionTargetPageId="5" captionText="FIGURE 2. Male (a) and female (b) of Liolaemus tregenzai in life." httpUri="https://zenodo.org/record/176300/files/figure.png" pageId="7" pageNumber="62">2</figureCitation>
|
||
). Although the coloration pattern is fundamentally similar in both sexes, pigments in males are more conspicuous.
|
||
</paragraph>
|
||
<paragraph id="8B8F657BFFAB3B61FF6780D37885851E" blockId="8.[151,1437,152,2029]" pageId="8" pageNumber="63">
|
||
<emphasis id="B944B969FFAB3B61FF6780D3785387B5" box="[198,270,152,177]" italics="true" pageId="8" pageNumber="63">Males</emphasis>
|
||
. Background coloration of dorsum and upper head surface dark green olive, with disperse light green scales. Some specimens can exhibit a brown-blackish general colour on the upper head scales. On the back sides the ground colour becomes darker, forming a kind of wide lateral band, which can be brown-blackish or almost completely black in some individuals. The lateral fusion of some small blackish spots can form transversal and fine stripes which contact the dark band of the flanks. Upper surface of limbs ranging from green olive to dark brown, with abundant, contrasting and small light green spots irregularly dispersed. Lower surface of limbs black. Dorsal surface of tail with dark green semilunar spots surrounded by inconspicuous light green shades, black on the lower surface. Background colour of ventral surface light and shiny greenblueish. This coloration reaches the lower flanks. On the throat, chest, and medial surface of abdomen completely and intensely black.
|
||
</paragraph>
|
||
<paragraph id="8B8F657BFFAB3B61FF6782637BD7845E" blockId="8.[151,1437,152,2029]" pageId="8" pageNumber="63">
|
||
<emphasis id="B944B969FFAB3B61FF67826378748545" box="[198,297,552,577]" italics="true" pageId="8" pageNumber="63">Females</emphasis>
|
||
. Dorsal head surface dark green, with a few small light green spots. Dorsal background colour dark chest-nut, with abundant and irregularly dispersed light green scales and irregularly shaped dark brown or blackish spots. On the anterior back sides there is a relatively wide dark brown or blackish lateral band. Background colour of ventral surface pale green-bluish, with abundant and irregularly dispersed light brownish or brown-copper, sometimes forming incipient transversal stripes. The medial colorations of throat, chest and abdomen are dark grey or greyish, never black as in males. Upper surface of limbs and tail colour dark chest-nut with abundant light green and blackish small spots. In juveniles the coloration is more similar to females, with dark greyish ventral surface.
|
||
</paragraph>
|
||
<paragraph id="8B8F657BFFAB3B61FF67832C7DD18109" blockId="8.[151,1437,152,2029]" pageId="8" pageNumber="63">
|
||
<emphasis id="B944B969FFAB3B61FF67832C781C8485" bold="true" box="[198,321,871,897]" pageId="8" pageNumber="63">Variation.</emphasis>
|
||
Based on the entire
|
||
<typeStatus id="548BDBD9FFAB3B61FD9F83237B2C8486" box="[574,625,872,898]" pageId="8" pageNumber="63">type</typeStatus>
|
||
series. Variation in allometric proportions revealed the existence of sexual dimorphism in most of the studied variables. Snout-vent length differs significantly between both sexes (
|
||
<emphasis id="B944B969FFAB3B61FF0083F379F484D5" box="[161,169,952,977]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
-test,
|
||
<emphasis id="B944B969FFAB3B61FF4883F379AC84D5" box="[233,241,952,977]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
9 = 2.978,
|
||
<emphasis id="B944B969FFAB3B61FEC983F3782784D5" box="[360,378,952,977]" italics="true" pageId="8" pageNumber="63">P</emphasis>
|
||
= 0.016), being
|
||
<quantity id="4CC8C89EFFAB3B61FD9B83F37BB984D6" box="[570,740,952,978]" metricMagnitude="-2" metricUnit="m" metricValue="8.295" metricValueMax="9.02" metricValueMin="7.57" pageId="8" pageNumber="63" unit="mm" value="82.95" valueMax="90.2" valueMin="75.7">75.7–90.2 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FCF383F37AE084D6" box="[850,957,952,978]" metricMagnitude="-2" metricUnit="m" metricValue="8.26" pageId="8" pageNumber="63" unit="mm" value="82.6">82.6 mm</quantity>
|
||
; SD = 5.94) in males, and
|
||
<quantity id="4CC8C89EFFAB3B61FB5383F37CC184D6" box="[1266,1436,952,978]" metricMagnitude="-2" metricUnit="m" metricValue="7.209999999999999" metricValueMax="8.08" metricValueMin="6.34" pageId="8" pageNumber="63" unit="mm" value="72.1" valueMax="80.8" valueMin="63.4">63.4–80.8 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FEA083A9782184F8" box="[257,380,994,1020]" metricMagnitude="-2" metricUnit="m" metricValue="7.141" pageId="8" pageNumber="63" unit="mm" value="71.41">71.41 mm</quantity>
|
||
, SD = 6.02) in females. Head length shows also significant sexual differences (
|
||
<emphasis id="B944B969FFAB3B61FA8883A87C6C84F8" box="[1321,1329,995,1020]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
-test,
|
||
<emphasis id="B944B969FFAB3B61FAD283A87C2684F8" box="[1395,1403,995,1020]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
9 = 2.884,
|
||
<emphasis id="B944B969FFAB3B61FF4B844579A18323" box="[234,252,1038,1063]" italics="true" pageId="8" pageNumber="63">P</emphasis>
|
||
= 0.018), ranging between
|
||
<quantity id="4CC8C89EFFAB3B61FDEC84467BA68323" box="[589,763,1037,1063]" metricMagnitude="-2" metricUnit="m" metricValue="1.89" metricValueMax="2.05" metricValueMin="1.73" pageId="8" pageNumber="63" unit="mm" value="18.9" valueMax="20.5" valueMin="17.3">17.3–20.5 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FCD384467AAD8323" box="[882,1008,1037,1063]" metricMagnitude="-2" metricUnit="m" metricValue="1.8829999999999998" pageId="8" pageNumber="63" unit="mm" value="18.83">18.83 mm</quantity>
|
||
, SD = 1.45) in males, and between
|
||
<quantity id="4CC8C89EFFAB3B61FF36847E781C834B" box="[151,321,1077,1103]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" metricValueMax="1.89" metricValueMin="1.45" pageId="8" pageNumber="63" unit="mm" value="16.7" valueMax="18.9" valueMin="14.5">14.5–18.9 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FE10847E7B76834B" box="[433,555,1077,1103]" metricMagnitude="-2" metricUnit="m" metricValue="1.629" pageId="8" pageNumber="63" unit="mm" value="16.29">16.29 mm</quantity>
|
||
, SD = 1.38) in females. Head weight differs significantly between sexes (
|
||
<emphasis id="B944B969FFAB3B61FA2B847D7CCF834B" box="[1418,1426,1078,1103]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
- test,
|
||
<emphasis id="B944B969FFAB3B61FF71841579858373" box="[208,216,1118,1143]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
9 = 2.828,
|
||
<emphasis id="B944B969FFAB3B61FEF6841578348373" box="[343,361,1118,1143]" italics="true" pageId="8" pageNumber="63">P</emphasis>
|
||
= 0.02), being
|
||
<quantity id="4CC8C89EFFAB3B61FD8784157B8E837C" box="[550,723,1118,1144]" metricMagnitude="-2" metricUnit="m" metricValue="1.48" metricValueMax="1.65" metricValueMin="1.31" pageId="8" pageNumber="63" unit="mm" value="14.8" valueMax="16.5" valueMin="13.1">13.1–16.5 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FCE884157AE9837C" box="[841,948,1118,1144]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="8" pageNumber="63" unit="mm" value="15.2">15.2 mm</quantity>
|
||
, SD = 1.6) in males, and
|
||
<quantity id="4CC8C89EFFAB3B61FB4E84157CC0837C" box="[1263,1437,1118,1144]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" metricValueMax="1.44" metricValueMin="1.24" pageId="8" pageNumber="63" unit="mm" value="13.4" valueMax="14.4" valueMin="12.4">12.4–14.4 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FEA384C2783283A7" box="[258,367,1161,1187]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="8" pageNumber="63" unit="mm" value="13.2">13.2 mm</quantity>
|
||
, SD = 0.79) in females. Sexes differ in head height (
|
||
<emphasis id="B944B969FFAB3B61FC4584C27AB183A6" box="[996,1004,1161,1186]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
-test,
|
||
<emphasis id="B944B969FFAB3B61FB8E84C27D6A83A6" box="[1071,1079,1161,1186]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
9 = 2.414,
|
||
<emphasis id="B944B969FFAB3B61FB1284C27D9883A6" box="[1203,1221,1161,1186]" italics="true" pageId="8" pageNumber="63">P</emphasis>
|
||
= 0.039), ranging between
|
||
<quantity id="4CC8C89EFFAB3B61FEA084F878C383C9" box="[257,414,1203,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.065" metricValueMax="1.22" metricValueMin="0.9099999999999999" pageId="8" pageNumber="63" unit="mm" value="10.649999999999999" valueMax="12.2" valueMin="9.1">9.1–12.2 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FDB384F87B2383C9" box="[530,638,1203,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="8" pageNumber="63" unit="mm" value="10.2">10.2 mm</quantity>
|
||
, SD = 1.44) in males, and between
|
||
<quantity id="4CC8C89EFFAB3B61FB8884F87DEA83C9" box="[1065,1207,1203,1229]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" metricValueMax="9.3" metricValueMin="7.1" pageId="8" pageNumber="63" unit="mm" value="8.2" valueMax="9.3" valueMin="7.1">7.1–9.3 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FA8A84F87CC583C9" box="[1323,1432,1203,1229]" metricMagnitude="-3" metricUnit="m" metricValue="8.51" pageId="8" pageNumber="63" unit="mm" value="8.51">8.51 mm</quantity>
|
||
, SD = 0.91) in females. Conversely, tail and limb lengths do not show sexual dimorphism. Tail length in males is
|
||
<quantity id="4CC8C89EFFAB3B61FF15854878428219" box="[180,287,1283,1309]" metricMagnitude="-3" metricUnit="m" metricValue="8.46" pageId="8" pageNumber="63" unit="mm" value="8.46">8.46 mm</quantity>
|
||
(only one specimen with the entire tail), and
|
||
<quantity id="4CC8C89EFFAB3B61FCE985487D5E8219" box="[840,1027,1283,1309]" metricMagnitude="-2" metricUnit="m" metricValue="9.945" metricValueMax="10.54" metricValueMin="9.35" pageId="8" pageNumber="63" unit="mm" value="99.45" valueMax="105.4" valueMin="93.5">93.5–105.4 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FBDB85487DB58219" box="[1146,1256,1283,1309]" metricMagnitude="-2" metricUnit="m" metricValue="9.51" pageId="8" pageNumber="63" unit="mm" value="95.1">95.1 mm</quantity>
|
||
, SD = 1.69) in females. Forelimb length in males is
|
||
<quantity id="4CC8C89EFFAB3B61FDEE85607BA78241" box="[591,762,1323,1349]" metricMagnitude="-2" metricUnit="m" metricValue="3.005" metricValueMax="3.2299999999999995" metricValueMin="2.78" pageId="8" pageNumber="63" unit="mm" value="30.049999999999997" valueMax="32.3" valueMin="27.8">27.8–32.3 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FCCC85607AB58241" box="[877,1000,1323,1349]" metricMagnitude="-2" metricUnit="m" metricValue="2.9379999999999997" pageId="8" pageNumber="63" unit="mm" value="29.38">29.38 mm</quantity>
|
||
, SD = 2.05) in males, and
|
||
<quantity id="4CC8C89EFFAB3B61FA86856079988269" metricMagnitude="-2" metricUnit="m" metricValue="2.73" metricValueMax="3.08" metricValueMin="2.38" pageId="8" pageNumber="63" unit="mm" value="27.3" valueMax="30.8" valueMin="23.8">23.8–30.8 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FE97851878EC8269" box="[310,433,1363,1389]" metricMagnitude="-2" metricUnit="m" metricValue="2.662" pageId="8" pageNumber="63" unit="mm" value="26.62">26.62 mm</quantity>
|
||
, SD = 2.8) in females (
|
||
<emphasis id="B944B969FFAB3B61FD65851F7B8B8269" box="[708,726,1364,1389]" italics="true" pageId="8" pageNumber="63">P</emphasis>
|
||
= 0.123).
|
||
<collectingCountry id="F32725EBFFAB3B61FCF085187AD18269" box="[849,908,1363,1389]" name="India" pageId="8" pageNumber="63">Hind</collectingCountry>
|
||
limb length is
|
||
<quantity id="4CC8C89EFFAB3B61FB9F85187DB48269" box="[1086,1257,1363,1389]" metricMagnitude="-2" metricUnit="m" metricValue="4.484999999999999" metricValueMax="4.79" metricValueMin="4.18" pageId="8" pageNumber="63" unit="mm" value="44.849999999999994" valueMax="47.9" valueMin="41.8">41.8–47.9 mm</quantity>
|
||
(mean =
|
||
<quantity id="4CC8C89EFFAB3B61FAFB851879958291" metricMagnitude="-2" metricUnit="m" metricValue="4.458" pageId="8" pageNumber="63" unit="mm" value="44.58">44.58 mm</quantity>
|
||
; SD = 2.73) in males, and 35.9–46.0 mm (mean =
|
||
<quantity id="4CC8C89EFFAB3B61FC8785307AFF8291" box="[806,930,1403,1429]" metricMagnitude="-2" metricUnit="m" metricValue="4.056" pageId="8" pageNumber="63" unit="mm" value="40.56">40.56 mm</quantity>
|
||
, SD = 3.35) in females (
|
||
<emphasis id="B944B969FFAB3B61FB6685377D848291" box="[1223,1241,1404,1429]" italics="true" pageId="8" pageNumber="63">P</emphasis>
|
||
= 0.073). When contrasting the derived variables forelimb length/snout-vent length (
|
||
<emphasis id="B944B969FFAB3B61FC6685EF7A9282B9" box="[967,975,1444,1469]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
-test,
|
||
<emphasis id="B944B969FFAB3B61FBB285EF7D7882B9" box="[1043,1061,1444,1469]" italics="true" pageId="8" pageNumber="63">P</emphasis>
|
||
= 0.351; 0.36 ±
|
||
<quantity id="4CC8C89EFFAB3B61FB5285E87C1A82B9" box="[1267,1351,1443,1469]" metricMagnitude="-4" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="63" unit="in" value="0.03">0.03 in</quantity>
|
||
males, versus 0.37 ±
|
||
<quantity id="4CC8C89EFFAB3B61FE9A858078D082E1" box="[315,397,1483,1509]" metricMagnitude="-4" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="63" unit="in" value="0.02">0.02 in</quantity>
|
||
females) and hind limb length/snout-vent length (
|
||
<emphasis id="B944B969FFAB3B61FC7885877ABC82E1" box="[985,993,1484,1509]" italics="true" pageId="8" pageNumber="63">t</emphasis>
|
||
-test,
|
||
<emphasis id="B944B969FFAB3B61FB8285877D6882E1" box="[1059,1077,1484,1509]" italics="true" pageId="8" pageNumber="63">P</emphasis>
|
||
= 0.256; 0.54 ±
|
||
<quantity id="4CC8C89EFFAB3B61FB5685807C1482E1" box="[1271,1353,1483,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="63" unit="in" value="0.04">0.04 in</quantity>
|
||
males, versus 0.57 ±
|
||
<quantity id="4CC8C89EFFAB3B61FE9A85B878D08109" box="[315,397,1523,1549]" metricMagnitude="-4" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="63" unit="in" value="0.03">0.03 in</quantity>
|
||
females) they do not show significant differences between sexes.
|
||
</paragraph>
|
||
<paragraph id="8B8F657BFFAB3B61FF6786507D3D80E9" blockId="8.[151,1437,152,2029]" pageId="8" pageNumber="63">
|
||
Variation in scale characteristics shown indistinctly for both males and females. Number of supraocular scales 5–6 (mean = 5.33, SD = 0.52). Scales forming circum orbitalis 13–14 (mean = 13.33, SD = 0.51). Upper ciliar scales 13–15 (mean = 13.83, SD = 0.75). Lower ciliar scales 10–14 (mean = 11.67, SD = 1.63). Number of loreal scales 5–9 (mean = 6.71, SD = 1.38). Number of supralabial scales 4–
|
||
<quantity id="4CC8C89EFFAB3B61FB3786D87D9F81A9" box="[1174,1218,1683,1709]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="8" pageNumber="63" unit="in" value="4.0">4 in</quantity>
|
||
the whole sample. Number of tympanic scales 2–4 (mean = 3.0, SD = 0.63). Number of auricular scales 2–3 (mean = 2.33, SD = 0.51). Number of postmental scales 4–6 (mean = 4.83, SD = 0.75). Number of infralabial scales 5–7 (mean = 6.0, SD = 0.63). Subdigital lamellae on the fourth toe of the left foot 24–26 (mean = 24.75, SD = 0.96; we counted lamellae on the left food because in the
|
||
<typeStatus id="548BDBD9FFAB3B61FD7587787A668049" box="[724,827,1843,1869]" pageId="8" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
the right one is damaged). Variation in dorsal body scales is observed in form and disposition, ranging from rhomboidal to subtriangular (
|
||
<figureCitation id="130B79FEFFAB3B61FB2687107D918071" box="[1159,1228,1883,1909]" captionStart="FIGURE 3" captionStartId="9.[151,255,1664,1688]" captionTargetBox="[152,1434,195,1638]" captionTargetId="figure@9.[151,1436,194,1640]" captionTargetPageId="9" captionText="FIGURE 3. Variation in morphological traits in the holotype male (a, c, e) and allotype female (b, d, f) of Liolaemus tregenzai. Dorsal body scales in the holotype (a) and in the allotype, ventral body scales in the holotype (c) and in the allotype, and cloacal region in the holotype (e) and in the allotype (f)." httpUri="https://zenodo.org/record/176301/files/figure.png" pageId="8" pageNumber="63">Fig. 3</figureCitation>
|
||
a, b), being juxtaposed-subimbricate to subimbricate-imbricate. Ventral body scales do not exhibit remarkable variation in relation to the
|
||
<typeStatus id="548BDBD9FFAB3B61FEBD87E078DE80C1" box="[284,387,1963,1989]" pageId="8" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="130B79FEFFAB3B61FE3487E0788080C1" box="[405,477,1963,1989]" captionStart="FIGURE 3" captionStartId="9.[151,255,1664,1688]" captionTargetBox="[152,1434,195,1638]" captionTargetId="figure@9.[151,1436,194,1640]" captionTargetPageId="9" captionText="FIGURE 3. Variation in morphological traits in the holotype male (a, c, e) and allotype female (b, d, f) of Liolaemus tregenzai. Dorsal body scales in the holotype (a) and in the allotype, ventral body scales in the holotype (c) and in the allotype, and cloacal region in the holotype (e) and in the allotype (f)." httpUri="https://zenodo.org/record/176301/files/figure.png" pageId="8" pageNumber="63">Fig. 3</figureCitation>
|
||
c, d). Number of scales around the midbody 71–85 (mean = 80.63, SD = 4.27). Precloacal glands are absent in males and females of the whole sample (
|
||
<figureCitation id="130B79FEFFAB3B61FC4587987D7480E9" box="[996,1065,2003,2029]" captionStart="FIGURE 3" captionStartId="9.[151,255,1664,1688]" captionTargetBox="[152,1434,195,1638]" captionTargetId="figure@9.[151,1436,194,1640]" captionTargetPageId="9" captionText="FIGURE 3. Variation in morphological traits in the holotype male (a, c, e) and allotype female (b, d, f) of Liolaemus tregenzai. Dorsal body scales in the holotype (a) and in the allotype, ventral body scales in the holotype (c) and in the allotype, and cloacal region in the holotype (e) and in the allotype (f)." httpUri="https://zenodo.org/record/176301/files/figure.png" pageId="8" pageNumber="63">Fig. 3</figureCitation>
|
||
e, f).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF4F35F3FFAA3B60FF3686CB7A0881D9" httpUri="https://zenodo.org/record/176301/files/figure.png" pageId="9" pageNumber="64" targetBox="[152,1434,195,1638]" targetPageId="9">
|
||
<paragraph id="8B8F657BFFAA3B60FF3686CB7A0881D9" blockId="9.[151,1436,1664,1757]" pageId="9" pageNumber="64">
|
||
<emphasis id="B944B969FFAA3B60FF3686CB7844819C" bold="true" box="[151,281,1664,1688]" pageId="9" pageNumber="64">FIGURE 3.</emphasis>
|
||
Variation in morphological traits in the holotype male (a, c, e) and allotype female (b, d, f) of
|
||
<taxonomicName id="4C301EF8FFAA3B60FAA086CA798081BE" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFAA3B60FAA086CA798081BE" italics="true" pageId="9" pageNumber="64">Liolaemus tregenzai</emphasis>
|
||
</taxonomicName>
|
||
. Dorsal body scales in the holotype (a) and in the allotype, ventral body scales in the holotype (c) and in the allotype, and cloacal region in the holotype (e) and in the allotype (f).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C32A36F0FFAA3B63FF67875B7A108656" lastPageId="10" lastPageNumber="65" pageId="9" pageNumber="64" type="distribution">
|
||
<paragraph id="8B8F657BFFAA3B60FF67875B782C807F" blockId="9.[151,1437,1808,2035]" pageId="9" pageNumber="64">
|
||
<emphasis id="B944B969FFAA3B60FF67875B783A802E" bold="true" box="[198,359,1808,1834]" pageId="9" pageNumber="64">Distribution.</emphasis>
|
||
<taxonomicName id="4C301EF8FFAA3B60FECF875A7B01802E" box="[366,604,1809,1834]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFAA3B60FECF875A7B01802E" box="[366,604,1809,1834]" italics="true" pageId="9" pageNumber="64">Liolaemus tregenzai</emphasis>
|
||
</taxonomicName>
|
||
is known from Copahue Volcano slopes (
|
||
<geoCoordinate id="EE0403BCFFAA3B60FBE6875A7DF7802F" box="[1095,1194,1809,1835]" direction="south" orientation="latitude" pageId="9" pageNumber="64" precision="925" value="-37.833332">37°50’S</geoCoordinate>
|
||
;
|
||
<geoCoordinate id="EE0403BCFFAA3B60FB16875A7C7A802F" box="[1207,1319,1809,1835]" direction="west" orientation="longitude" pageId="9" pageNumber="64" precision="925" value="-71.1">71°06’W</geoCoordinate>
|
||
),
|
||
<quantity id="4CC8C89EFFAA3B60FA9A875A7CC5802F" box="[1339,1432,1809,1835]" metricMagnitude="3" metricUnit="m" metricValue="2.15" pageId="9" pageNumber="64" unit="m" value="2150.0">2150 m</quantity>
|
||
, near Termas del Copahue, Departamento de Ñorquín, Neuquén Province, in north-western Patagonia of
|
||
<collectingCountry id="F32725EBFFAA3B60FF36872A7851807F" box="[151,268,1889,1915]" name="Argentina" pageId="9" pageNumber="64">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="130B79FEFFAA3B60FEBD872A7839807F" box="[284,356,1889,1915]" captionStart="FIGURE 4" captionStartId="10.[151,255,1354,1378]" captionTargetBox="[189,1398,401,1318]" captionTargetId="figure@10.[177,1411,394,1330]" captionTargetPageId="10" captionText="FIGURE 4. Map of the North-Western Patagonia of Argentina and Chile, showing the distribution of Liolaemus tregenzai and L. neuquensis (black circle; Copahue Volcano) and L. coeruleus (both black triangles; Pino Hachado, Primeros Pinos)" httpUri="https://zenodo.org/record/176302/files/figure.png" pageId="9" pageNumber="64">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8F657BFFAA3B63FF6787C37BE587DE" blockId="9.[151,1437,1808,2035]" lastBlockId="10.[151,1436,152,338]" lastPageId="10" lastPageNumber="65" pageId="9" pageNumber="64">
|
||
<emphasis id="B944B969FFAA3B60FF6787C378CE80A6" bold="true" box="[198,403,1928,1954]" pageId="9" pageNumber="64">Natural History.</emphasis>
|
||
<taxonomicName id="4C301EF8FFAA3B60FE3B87C27BDA80A6" box="[410,647,1929,1954]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="64" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFAA3B60FE3B87C27BDA80A6" box="[410,647,1929,1954]" italics="true" pageId="9" pageNumber="64">Liolaemus tregenzai</emphasis>
|
||
</taxonomicName>
|
||
is a Patagonian and diurnal species. Occurs in the Andean ecotonal zones between open antarctandic forests formed by
|
||
<taxonomicName id="4C301EF8FFAA3B60FCA187FA7A2980CF" box="[768,884,1969,1995]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="64" phylum="Tracheophyta" rank="genus">Araucaria</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C301EF8FFAA3B60FC2787FA7D2580CE" box="[902,1144,1969,1994]" class="Pinopsida" family="Araucariaceae" genus="Araucaria" kingdom="Plantae" order="Pinales" pageId="9" pageNumber="64" phylum="Tracheophyta" rank="species" species="araucana">
|
||
<emphasis id="B944B969FFAA3B60FC2787FA7D2580CE" box="[902,1144,1969,1994]" italics="true" pageId="9" pageNumber="64">Araucaria araucana</emphasis>
|
||
</taxonomicName>
|
||
) and areas recovered by
|
||
<emphasis id="B944B969FFAA3B60FF368792785180F6" box="[151,268,2009,2034]" italics="true" pageId="9" pageNumber="64">Chasquea</emphasis>
|
||
sp. It is a common species in areas characterized by clay sediments on the slopes of the Copahue volcano. Remarkably,
|
||
<taxonomicName id="4C301EF8FFA93B63FE3F80D37B7787B5" box="[414,554,152,177]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="65" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA93B63FE3F80D37B7787B5" box="[414,554,152,177]" italics="true" pageId="10" pageNumber="65">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
has often been found in the neighbourhoods of snow deposits, and close to thawing snow, which frequently form streams.
|
||
</paragraph>
|
||
<paragraph id="8B8F657BFFA93B63FF6780A37A108656" blockId="10.[151,1436,152,338]" pageId="10" pageNumber="65">
|
||
The diet of
|
||
<taxonomicName id="4C301EF8FFA93B63FEF180A378868605" box="[336,475,232,257]" class="Reptilia" family="Liolaemidae" genus="Liolaemus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="65" phylum="Chordata" rank="species" species="tregenzai">
|
||
<emphasis id="B944B969FFA93B63FEF180A378868605" box="[336,475,232,257]" italics="true" pageId="10" pageNumber="65">L. tregenzai</emphasis>
|
||
</taxonomicName>
|
||
is almost completely herbivorous. One of the studied specimens kept in captivity during more than nine months, was exclusively fed with lettuce. Nothing is known about other aspects of its natural history, remaining a mystery its reproductive mode.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |