treatments-xml/data/03/99/87/039987F8C46B966F6FD3FD9EFEF8FC0C.xml
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<mods:title id="CF07FF3BB4FBE3D043BAB11E91C5F466">Systematics of the keyserlingii group of Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae), with descriptions of three new species from Oaxaca, Mexico</mods:title>
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<taxonomicName id="4C304D6DC46B96696FD3FD9EFCDEFDF4" ID-CoL="36JMY" authority="Francke, 1977" authorityName="Francke" authorityYear="1977" box="[473,871,585,612]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mitlae">
<emphasis id="B944EAFCC46B96696FD3FD9EFD02FDF3" box="[473,699,585,611]" italics="true" pageId="27" pageNumber="28">Diplocentrus mitlae</emphasis>
<bibRefCitation id="EFA14B1FC46B96696CC9FD9EFCDEFDF4" author="Francke, O. F." box="[707,871,585,612]" pageId="27" pageNumber="28" pagination="145 - 200" refId="ref22525" refString="Francke, O. F. 1977. Scorpions of the genus Diplocentrus from Oaxaca, Mexico (Scorpionida, Diplocentridae). Journal of Arachnology 4: 145 - 200." type="journal article" year="1977">Francke, 1977</bibRefCitation>
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<paragraph id="8B8F36EEC46B96696C0CFDB8FC83FD19" blockId="27.[120,1177,623,754]" box="[518,826,623,650]" pageId="27" pageNumber="28">
<heading id="D0C78182C46B96696C0CFDB8FC83FD19" box="[518,826,623,650]" centered="true" fontSize="36" level="3" pageId="27" pageNumber="28" reason="7">
<figureCitation id="130B2A6BC46B96696C0CFDB8FD31FD19" box="[518,648,623,649]" captionStart="FIGURE 1" captionStartId="2.[120,206,1366,1387]" captionTargetBox="[120,1224,224,1340]" captionTargetId="figure-151@2.[120,1224,224,1340]" captionTargetPageId="2" captionText="FIGURE 1. Diplocentrus keyserlingii group, microhabitat and representative species. A. Diplocentrus hoffmanni Francke, 1977, burrow entrance. B. Diplocentrus sissomi, n. sp., female with brood in burrow under stone. C. D. hoffmanni, ♂. D. Diplocentrus mitlae Francke, 1977, ♂. E. Diplocentrus rectimanus Pocock, 1898, ♂. F. D. sissomi, n. sp., ♀." figureDoi="http://doi.org/10.5281/zenodo.5364222" httpUri="https://zenodo.org/record/5364222/files/figure.png" pageId="27" pageNumber="28">Figures 1D</figureCitation>
,
<figureCitation id="130B2A6BC46B96696C9CFDB8FD0CFD19" box="[662,693,623,650]" captionStart="FIGURE 5" captionStartId="9.[120,206,1116,1137]" captionTargetBox="[149,1196,224,1089]" captionTargetId="figure-320@9.[120,1224,224,1090]" captionTargetPageId="9" captionText="FIGURE 5. Diplocentrus keyserlingii group, dextral pedipalp chela, dorsoexternal aspect. A. Diplocentrus keyserlingii Karsch, 1880, holotype ♂ (AMNH) of Diplocentrus reticulatus Francke, 1977. B. Diplocentrus mitlae Francke, 1977, holotype ♂ (AMNH). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5364233" httpUri="https://zenodo.org/record/5364233/files/figure.png" pageId="27" pageNumber="28">5B</figureCitation>
; tables 1, 2
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<emphasis id="B944EAFCC46B96696E72FD41FEFFFD3E" box="[120,326,662,686]" italics="true" pageId="27" pageNumber="28">Diplocentrus mitlae</emphasis>
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, 194198, figs. 4, 16, 23, 5962; Beutelspacher and Trujillo- Olvera, 1999: 9;
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<bibRefCitation id="EFA14B1FC46B96696F5DFD6FFDEDFD40" author="Beutelspacher, C. R." box="[343,596,696,720]" pageId="27" pageNumber="28" refId="ref22081" refString="Beutelspacher, C. R. 2000. Catalogo de los Alacranes de Mexico. Morelia, Michoacan: Universidad Michoacana de San Nicolas de Hidalgo, 175 pp." type="book" year="2000">Beutelspacher, 2000: 33</bibRefCitation>
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;
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<bibRefCitation id="EFA14B1FC46B96696C6AFD6FFCC1FD40" author="Sissom, W. D. &amp; V. Fet" box="[608,888,696,720]" pageId="27" pageNumber="28" pagination="329 - 354" refId="ref24260" refString="Sissom, W. D., and V. Fet. 2000. Family Diplocentridae Karsch, 1880. In V. Fet, W. D. Sissom, G. Lowe, and M. E. Braunwalder, Catalog of the scorpions of the world (1758 - 1998): 329 - 354. New York: New York Entomological Society." type="book chapter" year="2000">Sissom and Fet, 2000: 341</bibRefCitation>
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;
<treatmentCitation id="0A9110FFC46B96696D89FD6FFEBCFD62" author="Armas, L. F. &amp; E. Martin-Frias" page="74" pageId="27" pageNumber="28" year="2003">
<bibRefCitation id="EFA14B1FC46B96696D89FD6FFEBCFD62" author="Armas, L. F. &amp; E. Martin-Frias" pageId="27" pageNumber="28" pagination="71 - 77" refId="ref22019" refString="Armas, L. F., and E. Martin-Frias. 2003. Dos nuevas especies de Diplocentrus Peters, 1861 (Scorpiones: Diplocentridae) de Mexico. Revista Iberica de Aracnologia 7: 71 - 77." type="journal article" year="2003">Armas and Martín-Frías, 2003: 74</bibRefCitation>
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.
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TYPE MATERIAL:
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:
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:
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:
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:
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(
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),
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,
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N [
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,
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, M.
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. Bogert.
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ADDITIONAL MATERIAL:
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<collectingCountry id="F327767EC46B96696FDDFC57FDF6FC0A" box="[471,591,896,922]" name="Mexico" pageId="27" pageNumber="28">MEXICO</collectingCountry>
:
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<collectingRegion id="49F4F80CC46B96696C54FC57FD6EFC0A" box="[606,727,896,922]" country="Mexico" name="Oaxaca" pageId="27" pageNumber="28">OAXACA</collectingRegion>
:
<collectingRegion id="49F4F80CC46B96696CE3FC57FBA5FC0A" box="[745,1052,896,923]" country="Mexico" name="DISTRITO DE TLACOLULA" pageId="27" pageNumber="28">DISTRITO DE TLACOLULA</collectingRegion>
:
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<collectingMunicipality id="6BEBAC94C46B96696A26FC56FE1EFC50" pageId="27" pageNumber="28">Municipio de San Pablo Villa de Mitla</collectingMunicipality>
</emphasis>
:
<location id="8EEF6035C46B96696FB3FC71FD2DFC50" LSID="urn:lsid:plazi:treatment:039987F8C46B966F6FD3FD9EFEF8FC0C:8EEF6035C46B96696FB3FC71FD2DFC50" box="[441,660,934,960]" country="Mexico" latitude="16.932251" longLatPrecision="1" longitude="-96.287" municipality="Municipio de San Pablo Villa de Mitla" name="San Jose del Paso" pageId="27" pageNumber="28" stateProvince="Oaxaca">San José del Paso</location>
,
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N,
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,
<quantity id="4CC89B0BC46B96696A6EFC71FB7AFC50" box="[1124,1219,934,960]" metricMagnitude="3" metricUnit="m" metricValue="1.88" pageId="27" pageNumber="28" unit="m" value="1880.0">
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,
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<collectingDate id="EFCAE9C6C46B96696E72FC1CFF5CFC75" box="[120,229,971,997]" pageId="27" pageNumber="28" value="2007-01-17">17.i.2007</collectingDate>
</date>
,
<collectorName id="26C55338C46B96696EFFFC1CFE22FC75" box="[245,411,971,997]" pageId="27" pageNumber="28">C. Santibáñez</collectorName>
and
<collectorName id="26C55338C46B96696FD4FC1BFDE7FC75" box="[478,606,971,998]" pageId="27" pageNumber="28">N. Gomez</collectorName>
,
<specimenCount id="9D36FD67C46B96696C67FC1CFD27FC75" box="[621,670,970,997]" pageId="27" pageNumber="28" type="female">2 ♀</specimenCount>
,
<specimenCount id="9D36FD67C46B96696CBBFC1CFD4DFC75" box="[689,756,971,997]" pageId="27" pageNumber="28" type="generic">1 juv.</specimenCount>
(
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)
</materialsCitation>
;
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<location id="8EEF6035C46B96696D72FC1CFBE8FC76" LSID="urn:lsid:plazi:treatment:039987F8C46B966F6FD3FD9EFEF8FC0C:8EEF6035C46B96696D72FC1CFBE8FC76" box="[888,1105,971,998]" country="Mexico" latitude="16.928917" longLatPrecision="1" longitude="-96.29778" name="San Jose del Paso" pageId="27" pageNumber="28" stateProvince="DISTRITO DE TLACOLULA">San José del Paso</location>
,
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N,
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,
<quantity id="4CC89B0BC46B96696FBDFC27FDA8FB9B" box="[439,529,1008,1035]" metricMagnitude="3" metricUnit="m" metricValue="1.9369999999999998" pageId="27" pageNumber="28" unit="m" value="1937.0">
<elevation id="001DD1DDC46B96696FBDFC27FDA8FB9B" box="[439,529,1008,1035]" metricMagnitude="3" metricUnit="m" metricValue="1.9369999999999998" pageId="27" pageNumber="28" unit="m" value="1937.0">1937 m</elevation>
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,
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<collectingDate id="EFCAE9C6C46B96696C14FC27FD2EFB9B" box="[542,663,1008,1035]" pageId="27" pageNumber="28" value="2009-09-15">15.ix.2009</collectingDate>
</date>
,
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.
<collectorName id="26C55338C46B96696CCFFC27FCFDFB9A" box="[709,836,1008,1034]" pageId="27" pageNumber="28">Paredes, C</collectorName>
.
<collectorName id="26C55338C46B96696D5BFC27FC6AFB9A" box="[849,979,1008,1034]" pageId="27" pageNumber="28">Santibáñez</collectorName>
, and
<collectorName id="26C55338C46B96696A19FC26FB3EFB9A" box="[1043,1159,1008,1035]" pageId="27" pageNumber="28">A. Valdez</collectorName>
,
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,
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(
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</materialsCitation>
,
<materialsCitation id="3B583CB3C46B96696F3EFBC1FD4AFBA0" box="[308,755,1045,1072]" collectionCode="AMCC" country="Mexico" pageId="27" pageNumber="28" specimenCode="LP 11034" specimenCount="1" specimenCount-male="1" stateProvince="DISTRITO DE TLACOLULA">
<specimenCount id="9D36FD67C46B96696F3EFBC1FE0FFBA0" box="[308,438,1045,1072]" pageId="27" pageNumber="28" type="male">1 subad. ♂</specimenCount>
[leg] (
<collectionCode id="ED21AE2BC46B96696C0CFBC1FDE2FBA0" box="[518,603,1046,1072]" country="USA" httpUri="http://grbio.org/cool/m0b5-nwmg" name="Ambrose Monell Cryo Collection, American Museum of Natural History" pageId="27" pageNumber="28">AMCC</collectionCode>
[
<specimenCode id="DB969E95C46B96696C64FBC1FD5AFBA0" box="[622,739,1045,1072]" collectionCode="LP" country="Russia" name="Laboratory of Palaeontology" pageId="27" pageNumber="28">LP 11034</specimenCode>
])
</materialsCitation>
,
<materialsCitation id="3B583CB3C46B96696CF5FBC1FBD5FBA0" box="[767,1132,1045,1072]" collectionCode="AMCC" country="Mexico" pageId="27" pageNumber="28" specimenCode="LP 11465" specimenCount="1" specimenCount-male="1" stateProvince="DISTRITO DE TLACOLULA">
<specimenCount id="9D36FD67C46B96696CF5FBC1FC97FBA0" box="[767,814,1045,1072]" pageId="27" pageNumber="28" type="male">1 ♂</specimenCount>
[leg] (
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[
<specimenCode id="DB969E95C46B96696DEFFBC1FBE5FBA0" box="[997,1116,1045,1072]" collectionCode="LP" country="Russia" name="Laboratory of Palaeontology" pageId="27" pageNumber="28">LP 11465</specimenCode>
])
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.
</paragraph>
</subSubSection>
<subSubSection id="C32A6565C46B96696EA2FBECFCC0F916" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="8B8F36EEC46B96696EA2FBECFB23FA10" blockId="27.[120,1225,784,1744]" pageId="27" pageNumber="28">
DIAGNOSIS: The following character combination, updated from
<bibRefCitation id="EFA14B1FC46B96696DA7FBECFBDAFBC5" author="Francke, O. F." box="[941,1123,1083,1109]" pageId="27" pageNumber="28" pagination="145 - 200" refId="ref22525" refString="Francke, O. F. 1977. Scorpions of the genus Diplocentrus from Oaxaca, Mexico (Scorpionida, Diplocentridae). Journal of Arachnology 4: 145 - 200." type="journal article" year="1977">Francke (1977)</bibRefCitation>
, is diagnostic for
<taxonomicName id="4C304D6DC46B96696EE5FBB6FEE0FBEA" authorityName="Francke" authorityYear="1977" box="[239,345,1120,1146]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mitlae">
<emphasis id="B944EAFCC46B96696EE5FBB6FEE0FBEA" box="[239,345,1120,1146]" italics="true" pageId="27" pageNumber="28">D. mitlae</emphasis>
</taxonomicName>
. Total length (adult),
<quantity id="4CC89B0BC46B96696C53FBB7FD62FBEB" box="[601,731,1120,1147]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="27" pageNumber="28" unit="mm" value="55.0" valueMax="60.0" valueMin="50.0">5060 mm</quantity>
. Base coloration (adult) reddish to copper red (fig. 1D). Carapace anteromedian notch shallow, V-shaped. Pedipalp femur, dorsal surface sparsely granular (
<specimenCount id="9D36FD67C46B96696F5DFB7DFED6FB55" box="[343,367,1194,1221]" pageId="27" pageNumber="28" type="male"></specimenCount>
) or smooth (
<specimenCount id="9D36FD67C46B96696C1EFB7DFD95FB55" box="[532,556,1194,1221]" pageId="27" pageNumber="28" type="female"></specimenCount>
). Pedipalp patella (
<specimenCount id="9D36FD67C46B96696D12FB7DFC89FB55" box="[792,816,1194,1221]" pageId="27" pageNumber="28" type="male"></specimenCount>
,
<specimenCount id="9D36FD67C46B96696D4BFB7DFCE0FB55" box="[833,857,1194,1221]" pageId="27" pageNumber="28" type="female"></specimenCount>
), dorsomedian, dorsoexternal, externomedian and ventromedian carinae weakly developed to obsolete, smooth. Pedipalp chela manus (
<specimenCount id="9D36FD67C46B96696F2CFB22FE87FA80" box="[294,318,1269,1296]" pageId="27" pageNumber="28" type="male"></specimenCount>
,
<specimenCount id="9D36FD67C46B96696F5BFB22FED0FA80" box="[337,361,1269,1296]" pageId="27" pageNumber="28" type="female"></specimenCount>
) slender, dorsal surface smooth; digital and dorsal secondary carinae weakly developed to obsolete, smooth (fig. 5B); fingers longer than manus. Legs IIV telotarsi, counts of spiniform macrosetae in pro- and retroventral rows, 4/4:4/5:5/5:5/5 (variation in table 2). Pectinal tooth count, 710, mode = 9 (
<specimenCount id="9D36FD67C46B96696C61FAB2FD3AFA10" box="[619,643,1381,1408]" pageId="27" pageNumber="28" type="male"></specimenCount>
) or 78, mode = 8 (
<specimenCount id="9D36FD67C46B96696D7FFAB2FC34FA10" box="[885,909,1381,1408]" pageId="27" pageNumber="28" type="female"></specimenCount>
) (variation in table 1).
</paragraph>
<paragraph id="8B8F36EEC46B96696EA2FA5CFCC0F916" blockId="27.[120,1225,784,1744]" pageId="27" pageNumber="28">
<taxonomicName id="4C304D6DC46B96696EA2FA5CFE35FA35" authorityName="Francke" authorityYear="1977" box="[168,396,1419,1445]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mitlae">
<emphasis id="B944EAFCC46B96696EA2FA5CFE35FA35" box="[168,396,1419,1445]" italics="true" pageId="27" pageNumber="28">Diplocentrus mitlae</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C304D6DC46B96696C18FA5BFD0CFA35" authorityName="Pocock" authorityYear="1898" box="[530,693,1420,1445]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="rectimanus">
<emphasis id="B944EAFCC46B96696C18FA5BFD0CFA35" box="[530,693,1420,1445]" italics="true" pageId="27" pageNumber="28">D. rectimanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D6DC46B96696CF9FA5BFC18FA35" authority="Santibäñez-López &amp; Francke &amp; Prendini, 2013" authorityName="Santibäñez-López &amp; Francke &amp; Prendini" authorityYear="2013" box="[755,929,1419,1445]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sagittipalpus" status="sp. nov.">
<emphasis id="B944EAFCC46B96696CF9FA5BFC18FA35" box="[755,929,1419,1445]" italics="true" pageId="27" pageNumber="28">D sagittipalpus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2775787C46B96696DB8FA5BFC4AFA36" box="[946,1011,1420,1446]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
, in adult size and coloration but can be distinguished as follows. The pedipalp chela dorsal surface is smooth (
<specimenCount id="9D36FD67C46B96696AA3FA67FB78FA5B" box="[1193,1217,1456,1483]" pageId="27" pageNumber="28" type="male"></specimenCount>
,
<specimenCount id="9D36FD67C46B96696E72FA02FF29FA60" box="[120,144,1493,1520]" pageId="27" pageNumber="28" type="female"></specimenCount>
) in
<taxonomicName id="4C304D6DC46B96696ECCFA00FE8AFA60" authorityName="Francke" authorityYear="1977" box="[198,307,1494,1520]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mitlae">
<emphasis id="B944EAFCC46B96696ECCFA00FE8AFA60" box="[198,307,1494,1520]" italics="true" pageId="27" pageNumber="28">D. mitlae</emphasis>
</taxonomicName>
but reticulate (
<specimenCount id="9D36FD67C46B96696FE4FA02FDBFFA60" box="[494,518,1493,1520]" pageId="27" pageNumber="28" type="male"></specimenCount>
) or granular (
<specimenCount id="9D36FD67C46B96696CBFFA02FD74FA60" box="[693,717,1493,1520]" pageId="27" pageNumber="28" type="female"></specimenCount>
) in
<taxonomicName id="4C304D6DC46B96696D09FA00FC1EFA60" authorityName="Pocock" authorityYear="1898" box="[771,935,1495,1520]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="rectimanus">
<emphasis id="B944EAFCC46B96696D09FA00FC1EFA60" box="[771,935,1495,1520]" italics="true" pageId="27" pageNumber="28">D. rectimanus</emphasis>
</taxonomicName>
, and granular-reticulate (
<specimenCount id="9D36FD67C46B96696E88FA2DFF23F985" box="[130,154,1530,1557]" pageId="27" pageNumber="28" type="male"></specimenCount>
) or granular (
<specimenCount id="9D36FD67C46B96696F4FFA2DFEE4F985" box="[325,349,1530,1557]" pageId="27" pageNumber="28" type="female"></specimenCount>
) in
<taxonomicName id="4C304D6DC46B96696F9BFA2BFDFDF985" authority="Santibäñez-López &amp; Francke &amp; Prendini, 2013" authorityName="Santibäñez-López &amp; Francke &amp; Prendini" authorityYear="2013" box="[401,580,1531,1557]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sagittipalpus" status="sp. nov.">
<emphasis id="B944EAFCC46B96696F9BFA2BFDFDF985" box="[401,580,1531,1557]" italics="true" pageId="27" pageNumber="28">D. sagittipalpus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2775787C46B96696C59FA2BFD2AF986" box="[595,659,1532,1558]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
The pedipalp chela digital carina (
<specimenCount id="9D36FD67C46B96696A3BFA2DFBF0F985" box="[1073,1097,1530,1557]" pageId="27" pageNumber="28" type="male"></specimenCount>
) is weakly developed in
<taxonomicName id="4C304D6DC46B96696F1CF9F6FE39F9AA" authorityName="Francke" authorityYear="1977" box="[278,384,1568,1594]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mitlae">
<emphasis id="B944EAFCC46B96696F1CF9F6FE39F9AA" box="[278,384,1568,1594]" italics="true" pageId="27" pageNumber="28">D. mitlae</emphasis>
</taxonomicName>
but strongly developed in
<taxonomicName id="4C304D6DC46B96696CB1F9F6FCE5F9AA" authorityName="Pocock" authorityYear="1898" box="[699,860,1569,1594]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="rectimanus">
<emphasis id="B944EAFCC46B96696CB1F9F6FCE5F9AA" box="[699,860,1569,1594]" italics="true" pageId="27" pageNumber="28">D. rectimanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D6DC46B96696D9DF9F6FBF0F9AA" authority="Santibäñez-López &amp; Francke &amp; Prendini, 2013" authorityName="Santibäñez-López &amp; Francke &amp; Prendini" authorityYear="2013" box="[919,1097,1568,1594]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sagittipalpus" status="sp. nov.">
<emphasis id="B944EAFCC46B96696D9DF9F6FBF0F9AA" box="[919,1097,1568,1594]" italics="true" pageId="27" pageNumber="28">D. sagittipalpus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2775787C46B96696A5DF9F6FB2CF9AB" box="[1111,1173,1569,1595]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
The counts of spiniform macrosetae on the telotarsi of legs III and IV are lower (5/5:5/5) in
<taxonomicName id="4C304D6DC46B96696AA6F990FF79F915" authorityName="Francke" authorityYear="1977" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mitlae">
<emphasis id="B944EAFCC46B96696AA6F990FF79F915" italics="true" pageId="27" pageNumber="28">D. mitlae</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C304D6DC46B96696F2FF9BBFE7EF915" authorityName="Pocock" authorityYear="1898" box="[293,455,1644,1669]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="rectimanus">
<emphasis id="B944EAFCC46B96696F2FF9BBFE7EF915" box="[293,455,1644,1669]" italics="true" pageId="27" pageNumber="28">D. rectimanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D6DC46B96696C09F9BBFD0FF915" authority="Santibäñez-López &amp; Francke &amp; Prendini, 2013" authorityName="Santibäñez-López &amp; Francke &amp; Prendini" authorityYear="2013" box="[515,694,1643,1669]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="sagittipalpus" status="sp. nov.">
<emphasis id="B944EAFCC46B96696C09F9BBFD0FF915" box="[515,694,1643,1669]" italics="true" pageId="27" pageNumber="28">D. sagittipalpus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2775787C46B96696CCFF9BBFCBDF916" box="[709,772,1644,1670]" pageId="27" pageNumber="28" rank="species">n. sp.</taxonomicNameLabel>
(6/6:6/6).
</paragraph>
</subSubSection>
<subSubSection id="C32A6565C46B966F6EA2F946FE1DFD06" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="8B8F36EEC46B96696EA2F946FCF4F940" blockId="27.[120,1225,784,1744]" pageId="27" pageNumber="28">
DESCRIPTION Of THE FEMALE: The original description of
<taxonomicName id="4C304D6DC46B96696D71F946FC53F93A" authorityName="Francke" authorityYear="1977" box="[891,1002,1680,1706]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="mitlae">
<emphasis id="B944EAFCC46B96696D71F946FC53F93A" box="[891,1002,1680,1706]" italics="true" pageId="27" pageNumber="28">D. mitlae</emphasis>
</taxonomicName>
was based on the
<typeStatus id="548B884CC46B96696E72F961FF66F940" box="[120,223,1718,1744]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
male, and the female remained unknown until now.
</paragraph>
<paragraph id="8B8F36EEC46C966E6EA2FF36FB7BFE01" blockId="28.[120,1225,225,1746]" pageId="28" pageNumber="29">
<emphasis id="B944EAFCC46C966E6EA2FF36FEA2FF6B" box="[168,283,225,251]" italics="true" pageId="28" pageNumber="29">Coloration</emphasis>
: Carapace brown to reddish orange; moderately infuscate throughout, uniformly so around median ocelli, variegated elsewhere. Coxosternum pale yellow. Pedipalps orange to reddish brown, carinae darker. Legs pale brown to pale orange, uniformly infuscate.
<taxonomicName id="4C304D6DC46C966E6DD4FEFAFBEEFED7" box="[990,1111,301,327]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="40" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
brown to reddish brown; tergites brown, densely infuscate; sternites pale yellow. Metasoma pale brown to reddish brown; carinae weakly to moderately infuscate.
<taxonomicName id="4C304D6DC46C966E6CCAFEA0FCB0FE01" box="[704,777,375,401]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="28" pageNumber="41" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
orange to reddish, uniformly infuscate.
</paragraph>
<paragraph id="8B8F36EEC46C966E6EA2FE4BFCBFFD91" blockId="28.[120,1225,225,1746]" pageId="28" pageNumber="29">
<emphasis id="B944EAFCC46C966E6EA2FE4BFEAFFE26" box="[168,278,412,438]" italics="true" pageId="28" pageNumber="29">Carapace</emphasis>
: Anterior margin moderately setose; anteromedian notch moderately deep, V-shaped. Frontal lobes and interocular surface weakly granular; other surfaces sparsely granular or shagreened. Three pairs of subequal lateral ocelli.
</paragraph>
<paragraph id="8B8F36EEC46C966E6EA2FDDBFB82FB7C" blockId="28.[120,1225,225,1746]" pageId="28" pageNumber="29">
<emphasis id="B944EAFCC46C966E6EA2FDDBFEACFDB6" box="[168,277,524,550]" italics="true" pageId="28" pageNumber="29">Pedipalps</emphasis>
: Orthobothriotaxic,
<typeStatus id="548B884CC46C966E6C03FDDAFD85FDB7" box="[521,572,525,551]" pageId="28" pageNumber="29">type</typeStatus>
C. Femur height greater than width; dorsal intercarinal surface markedly convex, sparsely granular medially; external intercarinal surface smooth; ventral intercarinal surface flat, smooth; internal intercarinal surface finely and sparsely granular; dorsointernal carina moderately developed, granular; dorsoexternal carina weakly developed, granular proximally and smooth distally; ventroexternal carina obsolete to weakly developed; ventrointernal carina moderately developed, granular proximally, becoming obsolete distally. Patella, all intercarinal surfaces smooth; proximal tubercle moderately developed; dorsointernal carina weakly developed to obsolete; dorsomedian carina weakly developed, smooth; dorsoexternal carina obsolete; externomedian obsolete; ventroexternal carinae obsolete; ventromedian carina obsolete; ventrointernal carina obsolete. Chela manus, rounded, height greater than width, sparsely setose; dorsal and external intercarinal surfaces smooth; dorsal marginal carina weakly developed to obsolete, smooth; digital carina obsolete; dorsal secondary and external secondary carinae obsolete; ventroexternal carina weakly developed to obsolete, smooth distally, becoming obsolete proximally; ventromedian carina weakly developed to obsolete, smooth proximally, becoming obsolete distally, directed toward midpoint of movable finger articulation; ventrointernal carina weakly developed to obsolete, smooth; internodorsal, internomedian and internoventral carinae weakly developed to obsolete, smooth; internal surface with shallow longitudinal depression where chela rests against patella. Chela fixed finger curved; length subequal to femur length and patella length; dorsal surface smooth and moderately setose proximally; external surface flat; internal surface shallowly concave.
</paragraph>
<paragraph id="8B8F36EEC46C966E6EA2FB2FFB35FAA7" blockId="28.[120,1225,225,1746]" pageId="28" pageNumber="29">
<emphasis id="B944EAFCC46C966E6EA2FB2FFF6EFA81" box="[168,215,1272,1297]" italics="true" pageId="28" pageNumber="29">Legs</emphasis>
: Legs IIV femora and tibiae, prolateral surfaces shagreened; telotarsi, counts of spiniform macrosetae in pro- and retroventral rows (dextral/sinistral), 4/4 4/4: 4/5 4/5: 5/5 5/5: 5/5 5/5.
</paragraph>
<paragraph id="8B8F36EEC46C966E6EA2FA95FE5DFACC" blockId="28.[120,1225,225,1746]" box="[168,484,1346,1372]" pageId="28" pageNumber="29">
<emphasis id="B944EAFCC46C966E6EA2FA95FEBDFACB" box="[168,260,1346,1371]" italics="true" pageId="28" pageNumber="29">Pectines</emphasis>
: Tooth count: 78.
</paragraph>
<paragraph id="8B8F36EEC46C966E6EA2FABFFECDFA5C" blockId="28.[120,1225,225,1746]" pageId="28" pageNumber="29">
<taxonomicName id="4C304D6DC46C966E6EA2FABFFE1EFA11" authority=": Tergites I" authorityName="Tergites I" box="[168,423,1383,1409]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="28" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAFCC46C966E6EA2FABFFE9BFA11" box="[168,290,1384,1409]" italics="true" pageId="28" pageNumber="29">Mesosoma</emphasis>
: Tergites I
</taxonomicName>
VI, pretergites smooth to shagreened, posttergites granular; VII granular. Sternites smooth; VII, dorsosubmedian and dorsolateral carinae weakly developed, crenulate to granular.
</paragraph>
<paragraph id="8B8F36EEC46C966F6EA2FA0FFE04FDB6" blockId="28.[120,1225,225,1746]" lastBlockId="29.[120,1225,225,924]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="B944EAFCC46C966E6EA2FA0FFEA6FA61" box="[168,287,1496,1521]" italics="true" pageId="28" pageNumber="29">Metasoma</emphasis>
: Metasomal segments IV, dorsal intercarinal surfaces minutely granular distally on segments I and II; smooth on III and V; lateral intercarinal surfaces smooth; ventral intercarinal surfaces smooth on IV. Segments IIV, dorsolateral carinae weakly developed, granular on I and II; weakly developed to obsolete, smooth on III and IV; lateral supramedian carinae moderately developed, crenulate to slightly granular on I and II, moderately developed, smooth on III and IV; lateral inframedian carinae moderately developed, slightly granular to crenulate on IIII, weakly developed, smooth on IV; ventrolateral carinae moderately to strongly developed, granular on I and II, weakly developed, slightly granular to crenulate on III and IV; ventrosubmedian carinae moderately to strongly developed, granular to slightly crenulate on I and II, weakly developed, slightly crenulate to smooth on III and IV. Segment V length:pedipalp femur length ratio, 1.48; dorsolateral and lateral inframedian carinae weakly developed to obsolete, smooth; ventrolateral carinae moderately developed, granular; ventromedian carina moderately developed, granular, with subspiniform granules posteriorly; ventral transverse carina moderately developed, comprising six subspiniform granules; anal arch semicircular; anal subterminal carina strongly developed, comprising 12 granules; anal terminal carina moderately developed, granular.
</paragraph>
<paragraph id="8B8F36EEC46D966F6EA2FDE6FE1DFD06" blockId="29.[120,1225,225,924]" pageId="29" pageNumber="30">
<taxonomicName id="4C304D6DC46D966F6EA2FDE6FEEBFDDB" authority=": Telson" authorityName="Telson" box="[168,338,561,588]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAFCC46D966F6EA2FDE6FF48FDDB" box="[168,241,561,587]" italics="true" pageId="29" pageNumber="30">Telson</emphasis>
: Telson
</taxonomicName>
, width:length ratio, 0.62. Vesicle, lateral surfaces smooth; ventral surface granular anteriorly. Subaculear tubercle stout, subconical. Aculeus, 1.4 length of subaculear tubercle, strongly curved.
</paragraph>
</subSubSection>
<subSubSection id="C32A6565C46D966F6EA2FD75FC4AFD71" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8B8F36EEC46D966F6EA2FD75FC4AFD71" blockId="29.[120,1225,225,924]" pageId="29" pageNumber="30">
DISTRIBUTION:
<taxonomicName id="4C304D6DC46D966F6F6EFD76FDF0FD2B" authorityName="Francke" authorityYear="1977" box="[356,585,673,699]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mitlae">
<emphasis id="B944EAFCC46D966F6F6EFD76FDF0FD2B" box="[356,585,673,699]" italics="true" pageId="29" pageNumber="30">Diplocentrus mitlae</emphasis>
</taxonomicName>
is known only from the central valleys and adjacent mountains of
<collectingRegion id="49F4F80CC46D966F6F17FD10FECDFD71" box="[285,372,711,737]" country="Mexico" name="Oaxaca" pageId="29" pageNumber="30">Oaxaca</collectingRegion>
in the San Pablo Villa de Mitla municipality (fig. 3B).
</paragraph>
</subSubSection>
<subSubSection id="C32A6565C46D966F6EA2FD3BFEF8FC0C" pageId="29" pageNumber="30" type="biology_ecology">
<paragraph id="8B8F36EEC46D966F6EA2FD3BFEF8FC0C" blockId="29.[120,1225,225,924]" pageId="29" pageNumber="30">
ECOLOGY: This species was collected under stones and from burrows, ca.
<quantity id="4CC89B0BC46D966F6A16FD3BFBDCFC97" box="[1052,1125,748,775]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="cm" value="20.0">20 cm</quantity>
long, at the base of stones, in an area where the dominant vegetation was oak forest.
<taxonomicName id="4C304D6DC46D966F6A38FCC6FF42FCC1" baseAuthorityName="Pocock" baseAuthorityYear="1898" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="nigrimanus">
<emphasis id="B944EAFCC46D966F6A38FCC6FF42FCC1" italics="true" pageId="29" pageNumber="30">Centruroides nigrimanus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C304D6DC46D966F6F31FCEFFC15FCC1" authority="Santibanez-Lopez and Sissom, 2010" authorityName="Santibanez-Lopez and Sissom" authorityYear="2010" box="[315,940,823,849]" class="Arachnida" family="Vaejovidae" genus="Vaejovis" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="oaxaca">
<emphasis id="B944EAFCC46D966F6F31FCEFFE4FFCC1" box="[315,502,824,849]" italics="true" pageId="29" pageNumber="30">Vaejovis oaxaca</emphasis>
Santibáñez-López and Sissom, 2010
</taxonomicName>
, were collected in sympatry. The habitat and habitus of
<taxonomicName id="4C304D6DC46D966F6FF6FC8AFDDFFCE6" authorityName="Francke" authorityYear="1977" box="[508,614,860,886]" class="Arachnida" family="Diplocentridae" genus="Diplocentrus" kingdom="Animalia" order="Scorpiones" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mitlae">
<emphasis id="B944EAFCC46D966F6FF6FC8AFDDFFCE6" box="[508,614,860,886]" italics="true" pageId="29" pageNumber="30">D. mitlae</emphasis>
</taxonomicName>
are consistent with the pelophilous ecomorphotype (
<bibRefCitation id="EFA14B1FC46D966F6E8AFC56FE8DFC0C" author="Prendini, L." box="[128,308,897,924]" pageId="29" pageNumber="30" pagination="113 - 118" refId="ref23784" refString="Prendini, L. 2001. Substratum specialization and speciation in southern African scorpions: the Effect Hypothesis revisited. In V. Fet and P. A Selden (editors.), Scorpions 2001. In memoriam Gary A. Polis: 113 - 118. Burnham Beeches, UK: British Arachnological Society." type="book chapter" year="2001">Prendini, 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>