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<mods:title id="0ADFACED5684C91F8B2A2F5CA4626F1A">Review of the Australian Cyllodini (Coleoptera: Nitidulidae: Nitidulinae), with descriptions of new taxa, and notes on the genus Macleayania (Nitidulini)</mods:title>
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Key to genera of
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occurring in
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1. Mesoventrite with anterior edge strongly oblique, almost vertical, so that most of its surface, including a median carina, is higher (in ventral view, projecting towards the observer) than mesocoxae or metaventrite; metanepisternum less than 3× as long as wide and more or less parallel-sided; mandibular apex unidentate and simple (without serrations or spines); pronotum at least slightly wider than combined elytra (
<figureCitation id="130B2A63F752804450EFFEBBDBC4CEB2" box="[648,733,379,400]" captionStart="FIGURES 111" captionStartId="8.[151,264,1832,1855]" captionTargetBox="[173,1430,227,1810]" captionTargetId="figure@8.[151,1436,223,1810]" captionTargetPageId="8" captionText="FIGURES 111. Habitus: 16, dorsal: 1, Camptodes aff. humeralis (Brullé), bicoloured (5.0 mm); 2, Cyllodes ruficeps (Reitter) (3.1 mm); 3, Camptodes aff. humeralis (Brullé), unicoloured (5.2 mm); 4, Coxollodes loriai (Grouvelle) (1.9 mm); 5, Cyllopallodes limbatus (Reitter) (2.9 mm); 6, Gymnocychramus politus Lea (5.0 mm). 711, lateral: 7, Camptodes aff. humeralis (Brullé), bicoloured; 8, same, unicoloured; 9, Cyllopallodes limbatus (Reitter); 10, Cyllodes ruficeps (Reitter); 11, Coxollodes loriai (Grouvelle). Fig. 10 from Lawrence &amp; Ślipiński (2013). All figures © CSIRO, Australia." httpUri="https://zenodo.org/record/2618375/files/figure.png" pageId="4" pageNumber="305">Figs 1, 3</figureCitation>
); elytra shorter than their combined widths; elytral punctures not forming distinct rows; length at least
<quantity id="4CC89B03F75280445395FE59DB27CE8C" box="[498,574,409,430]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="4" pageNumber="305" unit="mm" value="4.5">4.5 mm</quantity>
; gonocoxites narrowly elongate, parallel-sided and subcontiguous, with distinct, elongate, parallel-sided, apical gonostyli (
<figureCitation id="130B2A63F75280445072FE76DB45CEE9" box="[533,604,438,459]" captionStart="FIGURES 5969" captionStartId="16.[151,264,1804,1827]" captionTargetBox="[151,1436,195,1783]" captionTargetId="figure@16.[151,1436,195,1783]" captionTargetPageId="16" captionText="FIGURES 5969. Genitalia: 59, Camptodes aff. humeralis (Brullé), apex of Ovipositor, ventral; 6061, Gymnocychramus bicolor sp. nov., apex of penis: 60, dorsal (arrow pointing to apex of dorsal lobe); 61, same, lateral; 6263, Gymnocychramus politus Lea, apex of penis: 62, dorsal; 63, lateral; 6467, Macleayania vesca (Olliff): 64, apex of tegmen, dorsal; 65, base of tegmen, dorsal; 66, apex of penis; 67, apex of penile strut; 6869, Macleayania amphotiformis (Reitter): 68, base of tegmen; 69, apex of penile strut. All figures © CSIRO, Australia." httpUri="https://zenodo.org/record/2618385/files/figure.png" pageId="4" pageNumber="305">Fig. 59</figureCitation>
); northern NSW &amp; southern QLD; associated with
<taxonomicName id="4C304D65F7528044562EFE76DD81CEE9" box="[1097,1176,438,459]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" higherTaxonomySource="CoL" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="305" phylum="Magnoliopsida" rank="genus">
<emphasis id="B944EAF4F7528044562EFE76DD81CEE9" box="[1097,1176,438,459]" italics="true" pageId="4" pageNumber="305">Opuntia</emphasis>
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cacti.....................................................................................................
<taxonomicName id="4C304D65F752804456ABFE13DC85CECB" ID-CoL="3GJ7" authority="Erichson" authorityName="Erichson" authorityYear="1843" box="[1228,1436,467,489]" class="Insecta" family="Nitidulidae" genus="Camptodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
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<emphasis id="B944EAF4F752804456ABFE13DC20CECB" bold="true" box="[1228,1337,467,489]" italics="true" pageId="4" pageNumber="305">Camptodes</emphasis>
Erichson
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- Mesoventrite with anterior edge flat or weakly carinate and any change in elevation occurring only at posterior end; metanepisternum more than 3.5× as long as wide and distinctly narrowed posteriorly (
<figureCitation id="130B2A63F752804451D2FDCEDAE4CD01" box="[949,1021,526,547]" captionStart="FIGURES 1223" captionStartId="9.[151,264,1801,1824]" captionTargetBox="[164,1424,205,1764]" captionTargetId="figure@9.[151,1436,193,1780]" captionTargetPageId="9" captionText="FIGURES 1223. Habitus: 12, Gymnocychramus bicolor sp. nov. (4.0 mm), dorsal; 13, Pallodes beccarii Grouvelle (3.0 mm), dorsal; 14, same, ventral; 15, Pallodes nigroapicalis sp. nov. (2.8 mm), dorsal; 16, same, lateral; 17, same, posterodorsal; P. beccarii, dorsal, teneral (2.5 mm); 19, P. beccarii, ventrolateral; 20, Macleayania vesca (Olliff) (3.3 mm), dorsal; 21, M. amphotiformis (Reitter) (4.2 mm), dorsal; 22, same, ventral; 23, G. bicolor, lateral. Fig. 20 from Lawrence &amp; Ślipiński (2013). All figures © CSIRO, Australia." httpUri="https://zenodo.org/record/2618377/files/figure.png" pageId="4" pageNumber="305">Fig. 14</figureCitation>
); pronotum not wider than combined elytra (
<figureCitation id="130B2A63F75280445280FDEBD858CD62" box="[231,321,555,576]" captionStart="FIGURES 111" captionStartId="8.[151,264,1832,1855]" captionTargetBox="[173,1430,227,1810]" captionTargetId="figure@8.[151,1436,223,1810]" captionTargetPageId="8" captionText="FIGURES 111. Habitus: 16, dorsal: 1, Camptodes aff. humeralis (Brullé), bicoloured (5.0 mm); 2, Cyllodes ruficeps (Reitter) (3.1 mm); 3, Camptodes aff. humeralis (Brullé), unicoloured (5.2 mm); 4, Coxollodes loriai (Grouvelle) (1.9 mm); 5, Cyllopallodes limbatus (Reitter) (2.9 mm); 6, Gymnocychramus politus Lea (5.0 mm). 711, lateral: 7, Camptodes aff. humeralis (Brullé), bicoloured; 8, same, unicoloured; 9, Cyllopallodes limbatus (Reitter); 10, Cyllodes ruficeps (Reitter); 11, Coxollodes loriai (Grouvelle). Fig. 10 from Lawrence &amp; Ślipiński (2013). All figures © CSIRO, Australia." httpUri="https://zenodo.org/record/2618375/files/figure.png" pageId="4" pageNumber="305">Figs 2, 4</figureCitation>
); mandibular apex, IF unidentate, THEN armed with serrations or spines; IF length greater than
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, elytra distinctly longer than wide; ovipositor never with elongate, parallel-sided, apical gonostyli; associated with various fungi.....
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2 (1). Labrum wider than clypeus, consisting of a pair of diverging, rounded lobes flanking a deep notch widest at its opening, narrowing posteriorly and almost reaching the labral base; outer apical angle of protibia with distinct tooth; length always greater than
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; associated with the fruiting bodies of
<taxonomicName id="4C304D65F752804450A8FD61DA3CCD94" ID-CoL="3T3" box="[719,805,673,694]" class="Agaricomycetes" higherTaxonomySource="GBIF" kingdom="Fungi" order="Phallales" pageId="4" pageNumber="305" phylum="Basidiomycota" rank="order">Phallales</taxonomicName>
.................................
<taxonomicName id="4C304D65F752804456DCFD60DC82CD94" authority="Lea" authorityName="Lea" authorityYear="1921" box="[1211,1435,672,694]" class="Insecta" family="Nitidulidae" genus="Gymnocychramus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
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<emphasis id="B944EAF4F752804456DCFD60DC76CD94" bold="true" box="[1211,1391,672,694]" italics="true" pageId="4" pageNumber="305">Gymnocychramus</emphasis>
Lea
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Labrum never as above, widest at base and not wider than clypeus, its apex usually with small median cleft with parallel sides; outer apical angle of protibia without distinct tooth; length never greater than
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; associated with the non-woody fruiting bodies of Basidiomycetes (mushrooms)....................................................................
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3 (2). Mesoventrite with anterior edge on a distinctly different plane than metaventrite, its posterior end more or less vertical and emarginate forming a pair of mesothoracic lobes flanking the truncate anterior edge of metaventrite and visible in ventral view (
<figureCitation id="130B2A63F75280445280FC91D841CC44" box="[231,344,849,870]" captionStart="FIGURES 5058" captionStartId="12.[151,264,1364,1387]" captionTargetBox="[154,1422,236,1336]" captionTargetId="figure@12.[151,1436,193,1344]" captionTargetPageId="12" captionText="FIGURES 5058. Various structures: 5051, junction of mesoventrite and metaventrite: 50, Cyllodes ruficeps (Reitter), showing large metathoracic lobes; 51, Cyllopallodes limbicollis (Reitter), showing small mesothoracic lobes (arrow); 52, Coxollodes loriai (Grouvelle), mesoventrite, showing large procoxal rests separated by pair of diverging carinae; 53, Pallodes nigroapicalis sp. nov., mesoventrite showing short median carina and lack of procoxal rests (arrow points to curved line connecting mesocoxal cavities); 54, P. nigroapicalis, anterolateral portion of metaventrite, showing postcoxal line and axillary space; 55, Cyllodes ruficeps (Reitter), base of tegmen, showing highly reduced, broadly rounded strut; 5658, mesotibial apex: 56, Coxollodes loriai; 57, Pallodes nigroapicalis; 58, Pallodes beccarii (Grouvelle). Figs 4349 © Zoological Institute, Russian Academy of Sciences, St. Petersburg. Figs 5058 © CSIRO, Australia." httpUri="https://zenodo.org/record/2618383/files/figure.png" pageId="4" pageNumber="305">Figs 5051</figureCitation>
); posterior edges of mesocoxal cavities not joined by curved line extending across anterior process of metaventrite....................................................................................................
<emphasis id="B944EAF4F752804457F7FCAEDC85CCA6" bold="true" box="[1424,1436,878,900]" pageId="4" pageNumber="305">4</emphasis>
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- Mesoventrite with anterior edge on the same plane as or a slightly different plane than metaventrite, its posterior end emarginate but horizontal or only slightly oblique (
<figureCitation id="130B2A63F7528044501BFC69DBF4CC9C" box="[636,749,937,958]" captionStart="FIGURES 5058" captionStartId="12.[151,264,1364,1387]" captionTargetBox="[154,1422,236,1336]" captionTargetId="figure@12.[151,1436,193,1344]" captionTargetPageId="12" captionText="FIGURES 5058. Various structures: 5051, junction of mesoventrite and metaventrite: 50, Cyllodes ruficeps (Reitter), showing large metathoracic lobes; 51, Cyllopallodes limbicollis (Reitter), showing small mesothoracic lobes (arrow); 52, Coxollodes loriai (Grouvelle), mesoventrite, showing large procoxal rests separated by pair of diverging carinae; 53, Pallodes nigroapicalis sp. nov., mesoventrite showing short median carina and lack of procoxal rests (arrow points to curved line connecting mesocoxal cavities); 54, P. nigroapicalis, anterolateral portion of metaventrite, showing postcoxal line and axillary space; 55, Cyllodes ruficeps (Reitter), base of tegmen, showing highly reduced, broadly rounded strut; 5658, mesotibial apex: 56, Coxollodes loriai; 57, Pallodes nigroapicalis; 58, Pallodes beccarii (Grouvelle). Figs 4349 © Zoological Institute, Russian Academy of Sciences, St. Petersburg. Figs 5058 © CSIRO, Australia." httpUri="https://zenodo.org/record/2618383/files/figure.png" pageId="4" pageNumber="305">Figs 5253</figureCitation>
); posterior edges of mesocoxal cavities joined by curved line extending across anterior process of metaventrite (
<figureCitation id="130B2A63F75280445007FC06DBCECCF9" box="[608,727,966,987]" captionStart="FIGURES 5058" captionStartId="12.[151,264,1364,1387]" captionTargetBox="[154,1422,236,1336]" captionTargetId="figure@12.[151,1436,193,1344]" captionTargetPageId="12" captionText="FIGURES 5058. Various structures: 5051, junction of mesoventrite and metaventrite: 50, Cyllodes ruficeps (Reitter), showing large metathoracic lobes; 51, Cyllopallodes limbicollis (Reitter), showing small mesothoracic lobes (arrow); 52, Coxollodes loriai (Grouvelle), mesoventrite, showing large procoxal rests separated by pair of diverging carinae; 53, Pallodes nigroapicalis sp. nov., mesoventrite showing short median carina and lack of procoxal rests (arrow points to curved line connecting mesocoxal cavities); 54, P. nigroapicalis, anterolateral portion of metaventrite, showing postcoxal line and axillary space; 55, Cyllodes ruficeps (Reitter), base of tegmen, showing highly reduced, broadly rounded strut; 5658, mesotibial apex: 56, Coxollodes loriai; 57, Pallodes nigroapicalis; 58, Pallodes beccarii (Grouvelle). Figs 4349 © Zoological Institute, Russian Academy of Sciences, St. Petersburg. Figs 5058 © CSIRO, Australia." httpUri="https://zenodo.org/record/2618383/files/figure.png" pageId="4" pageNumber="305">Figs. 5253</figureCitation>
)..........................................................
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<paragraph id="8B8F36E6F752804452F0FC24DC82CB16" blockId="4.[151,1437,291,1428]" pageId="4" pageNumber="305">
<keyLead id="30C195D3F752804452F0FC24DC82CB16" pageId="4" pageNumber="305">
4(3). Upper portion of body strongly convex (
<figureCitation id="130B2A63F75280445003FC24DBB5CCDB" box="[612,684,996,1017]" captionStart="FIGURES 111" captionStartId="8.[151,264,1832,1855]" captionTargetBox="[173,1430,227,1810]" captionTargetId="figure@8.[151,1436,223,1810]" captionTargetPageId="8" captionText="FIGURES 111. Habitus: 16, dorsal: 1, Camptodes aff. humeralis (Brullé), bicoloured (5.0 mm); 2, Cyllodes ruficeps (Reitter) (3.1 mm); 3, Camptodes aff. humeralis (Brullé), unicoloured (5.2 mm); 4, Coxollodes loriai (Grouvelle) (1.9 mm); 5, Cyllopallodes limbatus (Reitter) (2.9 mm); 6, Gymnocychramus politus Lea (5.0 mm). 711, lateral: 7, Camptodes aff. humeralis (Brullé), bicoloured; 8, same, unicoloured; 9, Cyllopallodes limbatus (Reitter); 10, Cyllodes ruficeps (Reitter); 11, Coxollodes loriai (Grouvelle). Fig. 10 from Lawrence &amp; Ślipiński (2013). All figures © CSIRO, Australia." httpUri="https://zenodo.org/record/2618375/files/figure.png" pageId="4" pageNumber="305">Fig. 10</figureCitation>
); lateral margins of elytra not visible from above (
<figureCitation id="130B2A63F752804456E9FC24DDD1CCDB" box="[1166,1224,996,1017]" captionStart="FIGURES 111" captionStartId="8.[151,264,1832,1855]" captionTargetBox="[173,1430,227,1810]" captionTargetId="figure@8.[151,1436,223,1810]" captionTargetPageId="8" captionText="FIGURES 111. Habitus: 16, dorsal: 1, Camptodes aff. humeralis (Brullé), bicoloured (5.0 mm); 2, Cyllodes ruficeps (Reitter) (3.1 mm); 3, Camptodes aff. humeralis (Brullé), unicoloured (5.2 mm); 4, Coxollodes loriai (Grouvelle) (1.9 mm); 5, Cyllopallodes limbatus (Reitter) (2.9 mm); 6, Gymnocychramus politus Lea (5.0 mm). 711, lateral: 7, Camptodes aff. humeralis (Brullé), bicoloured; 8, same, unicoloured; 9, Cyllopallodes limbatus (Reitter); 10, Cyllodes ruficeps (Reitter); 11, Coxollodes loriai (Grouvelle). Fig. 10 from Lawrence &amp; Ślipiński (2013). All figures © CSIRO, Australia." httpUri="https://zenodo.org/record/2618375/files/figure.png" pageId="4" pageNumber="305">Fig. 2</figureCitation>
); frontoclypeal suture absent; vertexal line present at sides; mandible unidentate with dorsal and ventral serrations; all basal tarsomeres lobed; abdominal ventrite 1 without postcoxal lines..................................................
<taxonomicName id="4C304D65F75280445681FBDEDC82CB16" ID-CoL="3YDJ" authority="Erichson" authorityName="Erichson" authorityYear="1843" box="[1254,1435,1054,1076]" class="Insecta" family="Nitidulidae" genus="Cyllodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAF4F75280445681FBDEDC82CB16" bold="true" box="[1254,1435,1054,1076]" pageId="4" pageNumber="305">
<emphasis id="B944EAF4F75280445681FBDEDC23CB16" bold="true" box="[1254,1338,1054,1076]" italics="true" pageId="4" pageNumber="305">Cyllodes</emphasis>
Erichson
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8F36E6F752804452F0FBFCDC8FCBAE" blockId="4.[151,1437,291,1428]" pageId="4" pageNumber="305">
<keyLead id="30C195D3F752804452F0FBFCDC8FCBAE" pageId="4" pageNumber="305">
- Upper portion of body only slightly convex (
<figureCitation id="130B2A63F752804450F3FBFCDBC9CB73" box="[660,720,1084,1105]" captionStart="FIGURES 111" captionStartId="8.[151,264,1832,1855]" captionTargetBox="[173,1430,227,1810]" captionTargetId="figure@8.[151,1436,223,1810]" captionTargetPageId="8" captionText="FIGURES 111. Habitus: 16, dorsal: 1, Camptodes aff. humeralis (Brullé), bicoloured (5.0 mm); 2, Cyllodes ruficeps (Reitter) (3.1 mm); 3, Camptodes aff. humeralis (Brullé), unicoloured (5.2 mm); 4, Coxollodes loriai (Grouvelle) (1.9 mm); 5, Cyllopallodes limbatus (Reitter) (2.9 mm); 6, Gymnocychramus politus Lea (5.0 mm). 711, lateral: 7, Camptodes aff. humeralis (Brullé), bicoloured; 8, same, unicoloured; 9, Cyllopallodes limbatus (Reitter); 10, Cyllodes ruficeps (Reitter); 11, Coxollodes loriai (Grouvelle). Fig. 10 from Lawrence &amp; Ślipiński (2013). All figures © CSIRO, Australia." httpUri="https://zenodo.org/record/2618375/files/figure.png" pageId="4" pageNumber="305">Fig. 9</figureCitation>
); lateral margins of elytra visible for their entire lengths from above (
<figureCitation id="130B2A63F75280445712FBFCD9F6CB4C" captionStart="FIGURES 111" captionStartId="8.[151,264,1832,1855]" captionTargetBox="[173,1430,227,1810]" captionTargetId="figure@8.[151,1436,223,1810]" captionTargetPageId="8" captionText="FIGURES 111. Habitus: 16, dorsal: 1, Camptodes aff. humeralis (Brullé), bicoloured (5.0 mm); 2, Cyllodes ruficeps (Reitter) (3.1 mm); 3, Camptodes aff. humeralis (Brullé), unicoloured (5.2 mm); 4, Coxollodes loriai (Grouvelle) (1.9 mm); 5, Cyllopallodes limbatus (Reitter) (2.9 mm); 6, Gymnocychramus politus Lea (5.0 mm). 711, lateral: 7, Camptodes aff. humeralis (Brullé), bicoloured; 8, same, unicoloured; 9, Cyllopallodes limbatus (Reitter); 10, Cyllodes ruficeps (Reitter); 11, Coxollodes loriai (Grouvelle). Fig. 10 from Lawrence &amp; Ślipiński (2013). All figures © CSIRO, Australia." httpUri="https://zenodo.org/record/2618375/files/figure.png" pageId="4" pageNumber="305">Fig. 5</figureCitation>
); frontoclypeal suture present (sometimes weakly indicated externally); vertexal line absent; mandible bidentate with dorsal lobe serrate; no basal tarsomeres lobed; abdominal ventrite 1 with postcoxal lines................
<emphasis id="B944EAF4F752804456CCFBB6DC8FCBAE" bold="true" box="[1195,1430,1142,1164]" pageId="4" pageNumber="305">
<taxonomicName id="4C304D65F752804456CCFBB6DC29CBAE" authority="Lawrence &amp; Kirejtshuk, 2019" authorityName="Lawrence &amp; Kirejtshuk" authorityYear="2019" box="[1195,1328,1142,1164]" class="Insecta" family="Nitidulidae" genus="Cyllopallodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B944EAF4F752804456CCFBB6DC29CBAE" bold="true" box="[1195,1328,1142,1164]" italics="true" pageId="4" pageNumber="305">Cyllopallodes</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A277578FF7528044575BFBB6DC8FCBAE" box="[1340,1430,1142,1164]" pageId="4" pageNumber="305" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C42E43F752804452F0FB54DC82CAB6" pageId="4" pageNumber="305">
<paragraph id="8B8F36E6F752804452F0FB54DC82CA3C" blockId="4.[151,1437,291,1428]" pageId="4" pageNumber="305">
<keyLead id="30C195D3F752804452F0FB54DC82CA3C" pageId="4" pageNumber="305">
5(3). Prosternum in front of coxa very short, less than 0.3× as long as mid length of a coxal cavity; mesal portion of prosternum with median carina extending beyond middle of prosternal process; anterior portion of mesoventrite (
<figureCitation id="130B2A63F752804456F7FB71DDCECBE4" box="[1168,1239,1201,1222]" captionStart="FIGURES 5058" captionStartId="12.[151,264,1364,1387]" captionTargetBox="[154,1422,236,1336]" captionTargetId="figure@12.[151,1436,193,1344]" captionTargetPageId="12" captionText="FIGURES 5058. Various structures: 5051, junction of mesoventrite and metaventrite: 50, Cyllodes ruficeps (Reitter), showing large metathoracic lobes; 51, Cyllopallodes limbicollis (Reitter), showing small mesothoracic lobes (arrow); 52, Coxollodes loriai (Grouvelle), mesoventrite, showing large procoxal rests separated by pair of diverging carinae; 53, Pallodes nigroapicalis sp. nov., mesoventrite showing short median carina and lack of procoxal rests (arrow points to curved line connecting mesocoxal cavities); 54, P. nigroapicalis, anterolateral portion of metaventrite, showing postcoxal line and axillary space; 55, Cyllodes ruficeps (Reitter), base of tegmen, showing highly reduced, broadly rounded strut; 5658, mesotibial apex: 56, Coxollodes loriai; 57, Pallodes nigroapicalis; 58, Pallodes beccarii (Grouvelle). Figs 4349 © Zoological Institute, Russian Academy of Sciences, St. Petersburg. Figs 5058 © CSIRO, Australia." httpUri="https://zenodo.org/record/2618383/files/figure.png" pageId="4" pageNumber="305">Fig. 52</figureCitation>
) with a pair of procoxal rests divided by a pair of posteriorly diverging carinae; metaventrite with short discrimen; axillary spaces absent; outer edge of mesotibia (
<figureCitation id="130B2A63F752804453FCFB2CD8FBCA23" box="[411,482,1260,1281]" captionStart="FIGURES 5058" captionStartId="12.[151,264,1364,1387]" captionTargetBox="[154,1422,236,1336]" captionTargetId="figure@12.[151,1436,193,1344]" captionTargetPageId="12" captionText="FIGURES 5058. Various structures: 5051, junction of mesoventrite and metaventrite: 50, Cyllodes ruficeps (Reitter), showing large metathoracic lobes; 51, Cyllopallodes limbicollis (Reitter), showing small mesothoracic lobes (arrow); 52, Coxollodes loriai (Grouvelle), mesoventrite, showing large procoxal rests separated by pair of diverging carinae; 53, Pallodes nigroapicalis sp. nov., mesoventrite showing short median carina and lack of procoxal rests (arrow points to curved line connecting mesocoxal cavities); 54, P. nigroapicalis, anterolateral portion of metaventrite, showing postcoxal line and axillary space; 55, Cyllodes ruficeps (Reitter), base of tegmen, showing highly reduced, broadly rounded strut; 5658, mesotibial apex: 56, Coxollodes loriai; 57, Pallodes nigroapicalis; 58, Pallodes beccarii (Grouvelle). Figs 4349 © Zoological Institute, Russian Academy of Sciences, St. Petersburg. Figs 5058 © CSIRO, Australia." httpUri="https://zenodo.org/record/2618383/files/figure.png" pageId="4" pageNumber="305">Fig. 56</figureCitation>
) evenly curved and usuallly widest just beyond middle; ovipositor (
<figureCitation id="130B2A63F7528044561FFB2CDDA7CA23" box="[1144,1214,1260,1281]" captionStart="FIGURES 4349" captionStartId="11.[151,264,1133,1156]" captionTargetBox="[208,1377,200,1107]" captionTargetId="figure@11.[206,1381,193,1113]" captionTargetPageId="11" captionText="FIGURES 4349. Ovipositors, ventral: 43, Coxollodes loriai (Grouvelle); 44, Cyllodes ruficeps (Reitter); 45, Cyllopallodes limbatus (Reitter); 46. Gymnocychramus bicolor sp. nov.; 47, Gymnocychramus politus Lea; 48, Pallodes beccarii Grouvelle; 49, Pallodes nigroapicalis sp. nov." httpUri="https://zenodo.org/record/2618381/files/figure.png" pageId="4" pageNumber="305">Fig. 43</figureCitation>
) with broadly truncate apex..............................................................................
<taxonomicName id="4C304D65F752804456DEFAC8DC82CA3C" authority="Kirejtshuk" authorityName="Kirejtshuk" authorityYear="1987" box="[1209,1435,1288,1310]" class="Insecta" family="Nitidulidae" genus="Coxollodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAF4F752804456DEFAC8DC82CA3C" bold="true" box="[1209,1435,1288,1310]" pageId="4" pageNumber="305">
<emphasis id="B944EAF4F752804456DEFAC8DC3FCA3C" bold="true" box="[1209,1318,1288,1310]" italics="true" pageId="4" pageNumber="305">Coxollodes</emphasis>
Kirejtshuk
</emphasis>
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</paragraph>
<paragraph id="8B8F36E6F752804452F0FAE6DC82CAB6" blockId="4.[151,1437,291,1428]" pageId="4" pageNumber="305">
<keyLead id="30C195D3F752804452F0FAE6DC82CAB6" pageId="4" pageNumber="305">
- Prosternum about 0.5× as long as mid length of coxal cavity, without or with weak median carina and sometimes with acute anterior projection; anterior portion of mesoventrite (
<figureCitation id="130B2A63F752804450BCFA84DA39CA7B" box="[731,800,1348,1369]" captionStart="FIGURES 5058" captionStartId="12.[151,264,1364,1387]" captionTargetBox="[154,1422,236,1336]" captionTargetId="figure@12.[151,1436,193,1344]" captionTargetPageId="12" captionText="FIGURES 5058. Various structures: 5051, junction of mesoventrite and metaventrite: 50, Cyllodes ruficeps (Reitter), showing large metathoracic lobes; 51, Cyllopallodes limbicollis (Reitter), showing small mesothoracic lobes (arrow); 52, Coxollodes loriai (Grouvelle), mesoventrite, showing large procoxal rests separated by pair of diverging carinae; 53, Pallodes nigroapicalis sp. nov., mesoventrite showing short median carina and lack of procoxal rests (arrow points to curved line connecting mesocoxal cavities); 54, P. nigroapicalis, anterolateral portion of metaventrite, showing postcoxal line and axillary space; 55, Cyllodes ruficeps (Reitter), base of tegmen, showing highly reduced, broadly rounded strut; 5658, mesotibial apex: 56, Coxollodes loriai; 57, Pallodes nigroapicalis; 58, Pallodes beccarii (Grouvelle). Figs 4349 © Zoological Institute, Russian Academy of Sciences, St. Petersburg. Figs 5058 © CSIRO, Australia." httpUri="https://zenodo.org/record/2618383/files/figure.png" pageId="4" pageNumber="305">Fig. 53</figureCitation>
) with short longitudinal carina and no procoxal rests; metaventrite without discrimen; axillary spaces (
<figureCitation id="130B2A63F75280445053FAA1DB63CA54" box="[564,634,1377,1398]" captionStart="FIGURES 5058" captionStartId="12.[151,264,1364,1387]" captionTargetBox="[154,1422,236,1336]" captionTargetId="figure@12.[151,1436,193,1344]" captionTargetPageId="12" captionText="FIGURES 5058. Various structures: 5051, junction of mesoventrite and metaventrite: 50, Cyllodes ruficeps (Reitter), showing large metathoracic lobes; 51, Cyllopallodes limbicollis (Reitter), showing small mesothoracic lobes (arrow); 52, Coxollodes loriai (Grouvelle), mesoventrite, showing large procoxal rests separated by pair of diverging carinae; 53, Pallodes nigroapicalis sp. nov., mesoventrite showing short median carina and lack of procoxal rests (arrow points to curved line connecting mesocoxal cavities); 54, P. nigroapicalis, anterolateral portion of metaventrite, showing postcoxal line and axillary space; 55, Cyllodes ruficeps (Reitter), base of tegmen, showing highly reduced, broadly rounded strut; 5658, mesotibial apex: 56, Coxollodes loriai; 57, Pallodes nigroapicalis; 58, Pallodes beccarii (Grouvelle). Figs 4349 © Zoological Institute, Russian Academy of Sciences, St. Petersburg. Figs 5058 © CSIRO, Australia." httpUri="https://zenodo.org/record/2618383/files/figure.png" pageId="4" pageNumber="305">Fig. 54</figureCitation>
) usually well developed; outer edge of mesotibia in Australian species (
<figureCitation id="130B2A63F7528044574FFAA1DC8CCA54" box="[1320,1429,1377,1398]" captionStart="FIGURES 5058" captionStartId="12.[151,264,1364,1387]" captionTargetBox="[154,1422,236,1336]" captionTargetId="figure@12.[151,1436,193,1344]" captionTargetPageId="12" captionText="FIGURES 5058. Various structures: 5051, junction of mesoventrite and metaventrite: 50, Cyllodes ruficeps (Reitter), showing large metathoracic lobes; 51, Cyllopallodes limbicollis (Reitter), showing small mesothoracic lobes (arrow); 52, Coxollodes loriai (Grouvelle), mesoventrite, showing large procoxal rests separated by pair of diverging carinae; 53, Pallodes nigroapicalis sp. nov., mesoventrite showing short median carina and lack of procoxal rests (arrow points to curved line connecting mesocoxal cavities); 54, P. nigroapicalis, anterolateral portion of metaventrite, showing postcoxal line and axillary space; 55, Cyllodes ruficeps (Reitter), base of tegmen, showing highly reduced, broadly rounded strut; 5658, mesotibial apex: 56, Coxollodes loriai; 57, Pallodes nigroapicalis; 58, Pallodes beccarii (Grouvelle). Figs 4349 © Zoological Institute, Russian Academy of Sciences, St. Petersburg. Figs 5058 © CSIRO, Australia." httpUri="https://zenodo.org/record/2618383/files/figure.png" pageId="4" pageNumber="305">Figs 5758</figureCitation>
) subapically angulate; ovipositor with acute apex (
<figureCitation id="130B2A63F752804450DBFABEDA33CAB1" box="[700,810,1406,1427]" captionStart="FIGURES 4349" captionStartId="11.[151,264,1133,1156]" captionTargetBox="[208,1377,200,1107]" captionTargetId="figure@11.[206,1381,193,1113]" captionTargetPageId="11" captionText="FIGURES 4349. Ovipositors, ventral: 43, Coxollodes loriai (Grouvelle); 44, Cyllodes ruficeps (Reitter); 45, Cyllopallodes limbatus (Reitter); 46. Gymnocychramus bicolor sp. nov.; 47, Gymnocychramus politus Lea; 48, Pallodes beccarii Grouvelle; 49, Pallodes nigroapicalis sp. nov." httpUri="https://zenodo.org/record/2618381/files/figure.png" pageId="4" pageNumber="305">Figs 4849</figureCitation>
)....................................
<taxonomicName id="4C304D65F75280445681FABEDC82CAB6" ID-CoL="6CT4" authority="Erichson" authorityName="Erichson" authorityYear="1843" box="[1254,1435,1406,1428]" class="Insecta" family="Nitidulidae" genus="Pallodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAF4F75280445681FABEDC82CAB6" bold="true" box="[1254,1435,1406,1428]" pageId="4" pageNumber="305">
<emphasis id="B944EAF4F75280445681FABEDC23CAB6" bold="true" box="[1254,1338,1406,1428]" italics="true" pageId="4" pageNumber="305">Pallodes</emphasis>
Erichson
</emphasis>
</taxonomicName>
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</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>