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304 lines
40 KiB
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<document id="07B2B6BF6BC23B98295A2F6A180F03CB" ID-DOI="10.11646/zootaxa.3736.4.1" ID-GBIF-Dataset="e51b8007-6e2a-49fc-9edb-3b564a610009" ID-ISSN="1175-5326" ID-Zenodo-Dep="248455" ID-ZooBank="28E79EB3-43A6-4257-B396-1C76582C249D" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460698044059" checkinUser="plazi" docAuthor="Souza, Diego De S. & Monné, Marcela L." docDate="2013" docId="039987C17323FFE069A46250AF6CCD78" docLanguage="en" docName="zt03736p337.pdf" docOrigin="Zootaxa 3736 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ateralphus variegatus Mendes 1938" docType="treatment" docVersion="8" lastPageNumber="310" masterDocId="FFA0FFB97324FFE96933676CAE40CA34" masterDocTitle="Revision of the genus Ateralphus Restello, Iannuzzi & Marinoni, 2001 (Coleoptera: Cerambycidae: Lamiinae)" masterLastPageNumber="337" masterPageNumber="301" pageNumber="308" updateTime="1698354006376" updateUser="plazi">
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<mods:titleInfo id="9A5016724AE8E36A96E57458ACEA62C9">
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<mods:title id="4A19E3FF49A582C81EC9A63E2798A7F3">Revision of the genus Ateralphus Restello, Iannuzzi & Marinoni, 2001 (Coleoptera: Cerambycidae: Lamiinae)</mods:title>
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<mods:name id="638CBE210EEAF4AF45401E600296B7EE" type="personal">
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<mods:namePart id="01D0253CCC0DB92C350D6C5D1ECF3BD5">Souza, Diego De S.</mods:namePart>
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<mods:namePart id="96462FE4C624DAF7B6EE895B449794D2">Monné, Marcela L.</mods:namePart>
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<mods:title id="E5E22D1688FA02304FFAECFF0253742F">Zootaxa</mods:title>
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<mods:part id="E86307032B3A09295772928B540BDEF2">
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<mods:date id="59C28A897FA467896DE1BC349534CC31">2013</mods:date>
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<mods:detail id="623D8E2CBC49A1A05CB9B05EDC7D13C0" type="volume">
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<mods:number id="1017C865EFB8B0C8E38BB9470D46EEBE">3736</mods:number>
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<mods:number id="BC9F816AA70ED6E59835383B4018D331">4</mods:number>
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<mods:identifier id="CE6D6A4FD1E367E95915187FD843403B" type="DOI">10.11646/zootaxa.3736.4.1</mods:identifier>
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<treatment id="039987C17323FFE069A46250AF6CCD78" ID-DOI="http://doi.org/10.5281/zenodo.6158701" ID-GBIF-Taxon="119582346" ID-Zenodo-Dep="6158701" LSID="urn:lsid:plazi:treatment:039987C17323FFE069A46250AF6CCD78" httpUri="http://treatment.plazi.org/id/039987C17323FFE069A46250AF6CCD78" lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308">
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<subSubSection id="C32A655C7323FFEE69A46250AF11CFCE" pageId="7" pageNumber="308" type="nomenclature">
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<paragraph id="8B8F36D77323FFEE69A46250AC26CF62" blockId="7.[151,614,1340,1399]" box="[151,614,1340,1366]" pageId="7" pageNumber="308">
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<heading id="D0C781BB7323FFEE69A46250AC26CF62" bold="true" box="[151,614,1340,1366]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
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<taxonomicName id="4C304D547323FFEE69A46250AC26CF62" ID-CoL="J97G" authority="Mendes, 1938" authorityName="Mendes" authorityYear="1938" box="[151,614,1340,1366]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="variegatus">
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<emphasis id="B944EAC57323FFEE69A46250AC26CF62" bold="true" box="[151,614,1340,1366]" pageId="7" pageNumber="308">
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<emphasis id="B944EAC57323FFEE69A46250AFE1CF62" bold="true" box="[151,417,1340,1366]" italics="true" pageId="7" pageNumber="308">Ateralphus variegatus</emphasis>
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(Mendes, 1938)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B8F36D77323FFEE69A46232AC42CF43" blockId="7.[151,614,1340,1399]" box="[151,514,1374,1399]" pageId="7" pageNumber="308">
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(
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<figureCitation id="130B2A527323FFEE69AC6232AF59CF43" box="[159,281,1374,1399]" captionStart="FIGURES 8 – 13" captionStartId="8.[151,264,1625,1648]" captionTargetBox="[205,1381,194,1595]" captionTargetId="figure@8.[205,1381,193,1604]" captionTargetPageId="8" captionText="FIGURES 8 – 13. Ateralphus variegatus (Mendes, 1938). Male: 8 – 9, median lobe, dorsal and lateral view; 10, tegmen. Female: 11, sternite VIII; 12, female terminalia; 13, coxites. Scales = 1 mm. Figures 8 – 9, 11 – 12, respectively, same scale." httpUri="https://zenodo.org/record/248458/files/figure.png" pageId="7" pageNumber="308">Figs. 8–13</figureCitation>
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; 22–25; 31–32; 128)
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</paragraph>
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<paragraph id="8B8F36D77323FFEE69A462CAAF11CFCE" blockId="7.[151,1436,1446,1622]" pageId="7" pageNumber="308">
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<taxonomicName id="4C304D547323FFEE69A462CAAC61CF88" authority="Mendes, 1938: 120" authorityName="Mendes" authorityPageNumber="120" authorityYear="1938" box="[151,545,1446,1469]" class="Insecta" family="Cerambycidae" genus="Alphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="variegatus">
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<emphasis id="B944EAC57323FFEE69A462CAAF10CF89" box="[151,336,1446,1469]" italics="true" pageId="7" pageNumber="308">Alphus variegatus</emphasis>
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Mendes, 1938: 120
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</taxonomicName>
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; Blackwelder, 1946: 609 (cat.); Zikán & Wygodzinsky, 1948: 48 (holot.); Lane, 1965: 315; Gilmour, 1965: 607 (cat.); Martins & Monné, 1993:138; Monné, 1994: 40 (cat., dist.); Monné & Giesbert, 1994: 233 (check, dist.).
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</paragraph>
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</subSubSection>
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<subSubSection id="C32A655C7323FFE069A4616EAA95CEF0" lastPageId="9" lastPageNumber="310" pageId="7" pageNumber="308" type="description">
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<paragraph id="8B8F36D77323FFEE69A4616EABC3CC62" blockId="7.[151,1436,1446,1622]" pageId="7" pageNumber="308">
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<taxonomicName id="4C304D547323FFEE69A4616EAF39CC2C" box="[151,377,1538,1560]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="variegatus">
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<emphasis id="B944EAC57323FFEE69A4616EAF39CC2C" box="[151,377,1538,1560]" italics="true" pageId="7" pageNumber="308">Ateralphus variegatus</emphasis>
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</taxonomicName>
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; Restello
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<emphasis id="B944EAC57323FFEE68D1616FAC50CC2C" box="[482,528,1538,1560]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
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., 2001: 301; Monné, 2005: 168 (cat., dist.); Monné & Hovore, 2005: 204 (check, dist.); Wappes
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<emphasis id="B944EAC57323FFEE6813614EAF10CC03" box="[288,336,1569,1591]" italics="true" pageId="7" pageNumber="308">et al</emphasis>
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., 2006: 32 (dist.); Wappes
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<emphasis id="B944EAC57323FFEE6B43614EACE6CC03" box="[624,678,1569,1591]" italics="true" pageId="7" pageNumber="308">et al.</emphasis>
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, 2007: 35 (dist.); Monné & Bezark, 2009: 219 (check, dist.); Monné & Bezark, 2010: 222 (check, dist.); Monné & Bezark, 2011: 228 (check, dist.); Bezark & Monné, 2013: 235 (check, dist.).
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</paragraph>
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<paragraph id="8B8F36D77323FFEE69A461EEADCFCD33" blockId="7.[151,1436,1666,1799]" pageId="7" pageNumber="308">
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<emphasis id="B944EAC57323FFEE69A461EEAE9BCCAF" bold="true" box="[151,219,1666,1691]" pageId="7" pageNumber="308">Male.</emphasis>
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Integument dark brown. Body covered with dense and short yellowish-grey setae. Frons (
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<figureCitation id="130B2A527323FFEE6C0A61EEABD4CCAF" box="[1337,1428,1666,1691]" captionStart="FIGURES 20 – 28. 20 – 21" captionStartId="12.[151,264,1801,1824]" captionTargetBox="[171,1415,303,1750]" captionTargetId="figure@12.[171,1428,303,1781]" captionTargetPageId="12" captionText="FIGURES 20 – 28. 20 – 21, Ateralphus lacteus Galileo & Martins, 2006: 20, male, elytral apex; 21, female, abdomen, ventral view. 22 – 25, Ateralphus variegatus (Mendes, 1938): 22, frons; 23, basal-crests and punctation on prothorax and elytra; 24, female, abdomen, ventral view; 25, protibia. 26 – 28, Ateralphus tumidus new species: 26 – 27, mesosternal process, ventral and lateral view; 28, tarsômero V. Scales = 1 mm." httpUri="https://zenodo.org/record/248460/files/figure.png" pageId="7" pageNumber="308">Fig. 22</figureCitation>
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) rectangular; slightly convex, almost flat in lateral view. Upper ocular lobes height almost half of the lower ocular lobe; distance between them three times width of lobe. Lower ocular lobe (
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<figureCitation id="130B2A527323FFEE6AC261A6AA04CCD7" box="[1009,1092,1738,1763]" captionStart="FIGURES 20 – 28. 20 – 21" captionStartId="12.[151,264,1801,1824]" captionTargetBox="[171,1415,303,1750]" captionTargetId="figure@12.[171,1428,303,1781]" captionTargetPageId="12" captionText="FIGURES 20 – 28. 20 – 21, Ateralphus lacteus Galileo & Martins, 2006: 20, male, elytral apex; 21, female, abdomen, ventral view. 22 – 25, Ateralphus variegatus (Mendes, 1938): 22, frons; 23, basal-crests and punctation on prothorax and elytra; 24, female, abdomen, ventral view; 25, protibia. 26 – 28, Ateralphus tumidus new species: 26 – 27, mesosternal process, ventral and lateral view; 28, tarsômero V. Scales = 1 mm." httpUri="https://zenodo.org/record/248460/files/figure.png" pageId="7" pageNumber="308">Fig. 22</figureCitation>
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) rectangular; height about 1.4 times smaller than malar area. Genae (
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<figureCitation id="130B2A527323FFEE6B766182ACD7CD33" box="[581,663,1774,1799]" captionStart="FIGURES 20 – 28. 20 – 21" captionStartId="12.[151,264,1801,1824]" captionTargetBox="[171,1415,303,1750]" captionTargetId="figure@12.[171,1428,303,1781]" captionTargetPageId="12" captionText="FIGURES 20 – 28. 20 – 21, Ateralphus lacteus Galileo & Martins, 2006: 20, male, elytral apex; 21, female, abdomen, ventral view. 22 – 25, Ateralphus variegatus (Mendes, 1938): 22, frons; 23, basal-crests and punctation on prothorax and elytra; 24, female, abdomen, ventral view; 25, protibia. 26 – 28, Ateralphus tumidus new species: 26 – 27, mesosternal process, ventral and lateral view; 28, tarsômero V. Scales = 1 mm." httpUri="https://zenodo.org/record/248460/files/figure.png" pageId="7" pageNumber="308">Fig. 22</figureCitation>
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) parallel to divergent.
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</paragraph>
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<caption id="DF4F665F732CFFE169A46135AB4ACCBA" httpUri="https://zenodo.org/record/248458/files/figure.png" pageId="8" pageNumber="309" targetBox="[205,1381,194,1595]" targetPageId="8">
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<paragraph id="8B8F36D7732CFFE169A46135AB4ACCBA" blockId="8.[151,1436,1625,1678]" pageId="8" pageNumber="309">
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<emphasis id="B944EAC5732CFFE169A46135AF07CC5B" bold="true" box="[151,327,1625,1648]" pageId="8" pageNumber="309">FIGURES 8–13.</emphasis>
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<taxonomicName id="4C304D54732CFFE1687F6136AC96CC44" authority="Mendes, 1938" authorityName="Mendes" authorityYear="1938" box="[332,726,1626,1648]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="variegatus">
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<emphasis id="B944EAC5732CFFE1687F6136AC6DCC44" box="[332,557,1626,1648]" italics="true" pageId="8" pageNumber="309">Ateralphus variegatus</emphasis>
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(Mendes, 1938)
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</taxonomicName>
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. Male: 8–9, median lobe, dorsal and lateral view; 10, tegmen. Female: 11, sternite VIII; 12, female terminalia; 13, coxites. Scales = 1mm. Figures 8–9, 11–12, respectively, same scale.
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</paragraph>
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</caption>
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<paragraph id="8B8F36D7732CFFE169F461DAAAE3CD22" blockId="8.[151,1436,1717,2030]" pageId="8" pageNumber="309">
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Antennae (
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<figureCitation id="130B2A52732CFFE1687961D9AFE3CCFA" box="[330,419,1717,1742]" captionStart="FIGURES 29 – 34. 29 – 30" captionStartId="13.[151,264,1690,1713]" captionTargetBox="[167,1417,226,1634]" captionTargetId="figure@13.[154,1439,213,1647]" captionTargetPageId="13" captionText="FIGURES 29 – 34. 29 – 30, Ateralphus lacteus Galileo & Martins, 2006: 29, male; 30, female. 31 – 32. Ateralphus variegatus (Mendes, 1938): 31, male; 32, female. 33 – 34, Ateralphus tumidus new species: 33, holotype, female; 34, paratype, male. Scales = 2 mm." httpUri="https://zenodo.org/record/248461/files/figure.png" pageId="8" pageNumber="309">Fig. 31</figureCitation>
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) reaching elytral apex approximately at antennomeres VIII–IX. Scape with slightly depressed ring on apical fourth. Pedicel with one spot of dark brown setae on apical region. Antennomeres III–XI with one ring of dark brown setae on apical region, gradually longer on distal antennomeres.
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</paragraph>
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<paragraph id="8B8F36D7732CFFE069F4604DAFC5CAE0" blockId="8.[151,1436,1717,2030]" lastBlockId="9.[151,1436,151,1869]" lastPageId="9" lastPageNumber="310" pageId="8" pageNumber="309">
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Prothorax (
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<figureCitation id="130B2A52732CFFE16877604DAFD5CD0E" box="[324,405,1825,1850]" captionStart="FIGURES 20 – 28. 20 – 21" captionStartId="12.[151,264,1801,1824]" captionTargetBox="[171,1415,303,1750]" captionTargetId="figure@12.[171,1428,303,1781]" captionTargetPageId="12" captionText="FIGURES 20 – 28. 20 – 21, Ateralphus lacteus Galileo & Martins, 2006: 20, male, elytral apex; 21, female, abdomen, ventral view. 22 – 25, Ateralphus variegatus (Mendes, 1938): 22, frons; 23, basal-crests and punctation on prothorax and elytra; 24, female, abdomen, ventral view; 25, protibia. 26 – 28, Ateralphus tumidus new species: 26 – 27, mesosternal process, ventral and lateral view; 28, tarsômero V. Scales = 1 mm." httpUri="https://zenodo.org/record/248460/files/figure.png" pageId="8" pageNumber="309">Fig. 23</figureCitation>
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) maximum width almost as long as twice the length; coarsely punctate, with setae encircling the inner edge of the punctation pores; anterior margin subequal to posterior margin. Pronotum with a slight longitudinal elevation usually glabrous, only near anterior margin; depressed between median tubercles; with three tubercles on disc: two median elevated, obtuse and glabrous apex; with the third tubercle on midline, situated posterior to median tubercles, smaller and with glabrous apex. Prosternal process width almost one third the diameter of one procoxa. Mesosternal process flat at posterior half and anterior margin decline at an angle of approximately 45° relative to mesosternum. Mesepisternum, mesepimeron and metepisternum covered with yellowish-gray setae.
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</paragraph>
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<paragraph id="8B8F36D7732DFFE069F467B3AFFECB28" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">Scutellum wider than long, semicircular; one slight median longitudinal depression; finely punctate, covered with yellowish-gray setae.</paragraph>
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<paragraph id="8B8F36D7732DFFE069F4664BAC98CBC0" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
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Elytra length about 3.5 times longer than prothorax; with irregularly distributed coarse punctation and with setae encircling the inner edge of punctures as the prothorax; covered mainly with yellowish-gray setae and irregular patches of whitish setae on dorsum; with elevated carinae distinctly pronounced. Basal-crest (
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<figureCitation id="130B2A52732DFFE06C0C6603ABD3CBBC" box="[1343,1427,367,392]" captionStart="FIGURES 20 – 28. 20 – 21" captionStartId="12.[151,264,1801,1824]" captionTargetBox="[171,1415,303,1750]" captionTargetId="figure@12.[171,1428,303,1781]" captionTargetPageId="12" captionText="FIGURES 20 – 28. 20 – 21, Ateralphus lacteus Galileo & Martins, 2006: 20, male, elytral apex; 21, female, abdomen, ventral view. 22 – 25, Ateralphus variegatus (Mendes, 1938): 22, frons; 23, basal-crests and punctation on prothorax and elytra; 24, female, abdomen, ventral view; 25, protibia. 26 – 28, Ateralphus tumidus new species: 26 – 27, mesosternal process, ventral and lateral view; 28, tarsômero V. Scales = 1 mm." httpUri="https://zenodo.org/record/248460/files/figure.png" pageId="9" pageNumber="310">Fig. 23</figureCitation>
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) elevated, adorned by pronounced tubercles, inordinate and rounded apex; from crests, one “V”-shaped stain of dark brown setae; region between basal-crests with setae colored as the elytron. Epipleura setae coloration similar to dorsum. Apical margin of elytra obliquely truncate.
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</paragraph>
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<paragraph id="8B8F36D7732DFFE069F4656CAA9AC809" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
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Legs entirely covered with yellowish-gray setae. Tibiae (
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<figureCitation id="130B2A52732DFFE06A586693AD82C82C" box="[875,962,511,536]" captionStart="FIGURES 20 – 28. 20 – 21" captionStartId="12.[151,264,1801,1824]" captionTargetBox="[171,1415,303,1750]" captionTargetId="figure@12.[171,1428,303,1781]" captionTargetPageId="12" captionText="FIGURES 20 – 28. 20 – 21, Ateralphus lacteus Galileo & Martins, 2006: 20, male, elytral apex; 21, female, abdomen, ventral view. 22 – 25, Ateralphus variegatus (Mendes, 1938): 22, frons; 23, basal-crests and punctation on prothorax and elytra; 24, female, abdomen, ventral view; 25, protibia. 26 – 28, Ateralphus tumidus new species: 26 – 27, mesosternal process, ventral and lateral view; 28, tarsômero V. Scales = 1 mm." httpUri="https://zenodo.org/record/248460/files/figure.png" pageId="9" pageNumber="310">Fig. 25</figureCitation>
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) with one spot of dark brown setae on subapical region. Tarsomere V bicolorous, base with yellowish-white setae and apex dark brown.
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</paragraph>
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<paragraph id="8B8F36D7732DFFE069F4652BAAB6C8B1" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
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Abdomen (
|
||
<figureCitation id="130B2A52732DFFE06876652BAFD7C854" box="[325,407,583,608]" captionStart="FIGURES 20 – 28. 20 – 21" captionStartId="12.[151,264,1801,1824]" captionTargetBox="[171,1415,303,1750]" captionTargetId="figure@12.[171,1428,303,1781]" captionTargetPageId="12" captionText="FIGURES 20 – 28. 20 – 21, Ateralphus lacteus Galileo & Martins, 2006: 20, male, elytral apex; 21, female, abdomen, ventral view. 22 – 25, Ateralphus variegatus (Mendes, 1938): 22, frons; 23, basal-crests and punctation on prothorax and elytra; 24, female, abdomen, ventral view; 25, protibia. 26 – 28, Ateralphus tumidus new species: 26 – 27, mesosternal process, ventral and lateral view; 28, tarsômero V. Scales = 1 mm." httpUri="https://zenodo.org/record/248460/files/figure.png" pageId="9" pageNumber="310">Fig. 24</figureCitation>
|
||
) covered with yellowish-gray setae, with glabrous maculae on lateral edges of sternites II– IV. Sternite V wider than long; length about equal or less than length of sternite III and IV together.
|
||
</paragraph>
|
||
<paragraph id="8B8F36D7732DFFE069F465E3AAADC921" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
|
||
Terminalia. Median lobe (
|
||
<figureCitation id="130B2A52732DFFE068C365E3AC1BC89C" box="[496,603,655,680]" captionStart="FIGURES 8 – 13" captionStartId="8.[151,264,1625,1648]" captionTargetBox="[205,1381,194,1595]" captionTargetId="figure@8.[205,1381,193,1604]" captionTargetPageId="8" captionText="FIGURES 8 – 13. Ateralphus variegatus (Mendes, 1938). Male: 8 – 9, median lobe, dorsal and lateral view; 10, tegmen. Female: 11, sternite VIII; 12, female terminalia; 13, coxites. Scales = 1 mm. Figures 8 – 9, 11 – 12, respectively, same scale." httpUri="https://zenodo.org/record/248458/files/figure.png" pageId="9" pageNumber="310">Figs. 8–9</figureCitation>
|
||
) curved in lateral view; dorsal and ventral lobes subequal in length, with rounded apex; basal apophysis length subequal to apical portion. Tegmen (
|
||
<figureCitation id="130B2A52732DFFE06D3165D8AA17C8F8" box="[1026,1111,692,717]" captionStart="FIGURES 8 – 13" captionStartId="8.[151,264,1625,1648]" captionTargetBox="[205,1381,194,1595]" captionTargetId="figure@8.[205,1381,193,1604]" captionTargetPageId="8" captionText="FIGURES 8 – 13. Ateralphus variegatus (Mendes, 1938). Male: 8 – 9, median lobe, dorsal and lateral view; 10, tegmen. Female: 11, sternite VIII; 12, female terminalia; 13, coxites. Scales = 1 mm. Figures 8 – 9, 11 – 12, respectively, same scale." httpUri="https://zenodo.org/record/248458/files/figure.png" pageId="9" pageNumber="310">Fig. 10</figureCitation>
|
||
) length slightly longer than median lobe; parameres about 3.6 times less than tegmen length, near each other; with long and sparse setae in inner surface; ring piece with one apical projection elongate and curved and apical margin sinuous.
|
||
</paragraph>
|
||
<paragraph id="8B8F36D7732DFFE069F46473AAE7C903" blockId="9.[151,1436,151,1869]" box="[199,1191,799,824]" pageId="9" pageNumber="310">
|
||
<emphasis id="B944EAC5732DFFE069F46473AF65C90C" bold="true" box="[199,293,799,824]" pageId="9" pageNumber="310">Female.</emphasis>
|
||
Antennae (
|
||
<figureCitation id="130B2A52732DFFE068956473AFB8C90C" box="[422,504,799,824]" captionStart="FIGURES 29 – 34. 29 – 30" captionStartId="13.[151,264,1690,1713]" captionTargetBox="[167,1417,226,1634]" captionTargetId="figure@13.[154,1439,213,1647]" captionTargetPageId="13" captionText="FIGURES 29 – 34. 29 – 30, Ateralphus lacteus Galileo & Martins, 2006: 29, male; 30, female. 31 – 32. Ateralphus variegatus (Mendes, 1938): 31, male; 32, female. 33 – 34, Ateralphus tumidus new species: 33, holotype, female; 34, paratype, male. Scales = 2 mm." httpUri="https://zenodo.org/record/248461/files/figure.png" pageId="9" pageNumber="310">Fig. 32</figureCitation>
|
||
) reaching elytral apex approximately at antennomeres X–XI.
|
||
</paragraph>
|
||
<paragraph id="8B8F36D7732DFFE069F46428AAA5CE24" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
|
||
Terminalia (
|
||
<figureCitation id="130B2A52732DFFE068656428AF99C969" box="[342,473,836,861]" captionStart="FIGURES 8 – 13" captionStartId="8.[151,264,1625,1648]" captionTargetBox="[205,1381,194,1595]" captionTargetId="figure@8.[205,1381,193,1604]" captionTargetPageId="8" captionText="FIGURES 8 – 13. Ateralphus variegatus (Mendes, 1938). Male: 8 – 9, median lobe, dorsal and lateral view; 10, tegmen. Female: 11, sternite VIII; 12, female terminalia; 13, coxites. Scales = 1 mm. Figures 8 – 9, 11 – 12, respectively, same scale." httpUri="https://zenodo.org/record/248458/files/figure.png" pageId="9" pageNumber="310">Figs.11–13</figureCitation>
|
||
). Sternite VIII (
|
||
<figureCitation id="130B2A52732DFFE06BAA6428ACAFC968" box="[665,751,836,861]" captionStart="FIGURES 8 – 13" captionStartId="8.[151,264,1625,1648]" captionTargetBox="[205,1381,194,1595]" captionTargetId="figure@8.[205,1381,193,1604]" captionTargetPageId="8" captionText="FIGURES 8 – 13. Ateralphus variegatus (Mendes, 1938). Male: 8 – 9, median lobe, dorsal and lateral view; 10, tegmen. Female: 11, sternite VIII; 12, female terminalia; 13, coxites. Scales = 1 mm. Figures 8 – 9, 11 – 12, respectively, same scale." httpUri="https://zenodo.org/record/248458/files/figure.png" pageId="9" pageNumber="310">Fig. 11</figureCitation>
|
||
) rounded, with apical region slightly narrower than basal region; sternal apodeme length about three times the sternite VIII. Tergal apodemes concave and narrowing towards basal region. Coxites (
|
||
<figureCitation id="130B2A52732DFFE068C564E0AC08C990" box="[502,584,908,933]" captionStart="FIGURES 8 – 13" captionStartId="8.[151,264,1625,1648]" captionTargetBox="[205,1381,194,1595]" captionTargetId="figure@8.[205,1381,193,1604]" captionTargetPageId="8" captionText="FIGURES 8 – 13. Ateralphus variegatus (Mendes, 1938). Male: 8 – 9, median lobe, dorsal and lateral view; 10, tegmen. Female: 11, sternite VIII; 12, female terminalia; 13, coxites. Scales = 1 mm. Figures 8 – 9, 11 – 12, respectively, same scale." httpUri="https://zenodo.org/record/248458/files/figure.png" pageId="9" pageNumber="310">Fig. 13</figureCitation>
|
||
) parallel and robust, near each other on two basal thirds and away towards apex; basal region width about 2–3 times wider than apex.
|
||
<taxonomicName id="4C304D54732DFFE06A0A64DDADB8C9FC" box="[825,1016,944,968]" class="Gastropoda" family="Bursidae" genus="Bursa" kingdom="Animalia" order="Littorinimorpha" pageId="9" pageNumber="310" phylum="Mollusca" rank="species" species="copulatrix">
|
||
<emphasis id="B944EAC5732DFFE06A0A64DDADB8C9FC" box="[825,1016,944,968]" italics="true" pageId="9" pageNumber="310">Bursa copulatrix</emphasis>
|
||
</taxonomicName>
|
||
elongated and oval aspect, about six times the spermatheca length. Spermathecal duct short. Spermatheca base elongate, gradually wider towards apical region and apex rounded. Spermathecal gland short, length about twice or more than spermatheca.
|
||
</paragraph>
|
||
<paragraph id="8B8F36D7732DFFE069F46370AA82CE6C" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
|
||
<emphasis id="B944EAC5732DFFE069F46370AF11CE01" bold="true" box="[199,337,1052,1077]" pageId="9" pageNumber="310">Variability.</emphasis>
|
||
Some specimens do not have the spot of dark brown setae on apical region of pedicel. The elytral “V”-shaped stain of dark brown setae may be absent or completely filled like a triangular stain.
|
||
</paragraph>
|
||
<paragraph id="8B8F36D7732DFFE069F46308ABD0CE94" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
|
||
<emphasis id="B944EAC5732DFFE069F46308AC26CE49" bold="true" box="[199,614,1124,1149]" pageId="9" pageNumber="310">Measurements (mm), male/female</emphasis>
|
||
. Total length, 18.8–22.7/17.0–23.8; prothorax length, 3.3–4.0/3.2–4.1; prothorax maximum width, 5.4–7.1/5.3–7.6; elytral length, 13.5–17.0/12.8–17.8; humeral width, 7.2–9.0/6.8–9.7.
|
||
</paragraph>
|
||
<paragraph id="8B8F36D7732DFFE069F463C0AA95CEF0" blockId="9.[151,1436,151,1869]" box="[199,1237,1196,1221]" pageId="9" pageNumber="310">
|
||
<emphasis id="B944EAC5732DFFE069F463C0AF39CEF1" bold="true" box="[199,377,1196,1221]" pageId="9" pageNumber="310">Type-material.</emphasis>
|
||
<typeStatus id="548B8875732DFFE068B363C0AFABCEF1" box="[384,491,1196,1221]" pageId="9" pageNumber="310" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<taxonomicName id="4C304D54732DFFE068C563C0AC81CEF0" box="[502,705,1196,1220]" class="Insecta" family="Cerambycidae" genus="Alphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="variegatus">
|
||
<emphasis id="B944EAC5732DFFE068C563C0AC81CEF0" box="[502,705,1196,1220]" italics="true" pageId="9" pageNumber="310">Alphus variegatus</emphasis>
|
||
</taxonomicName>
|
||
, female, Campinas, São Paulo,
|
||
<collectingCountry id="F3277647732DFFE06D1563C0AA2ACEF1" box="[1062,1130,1196,1221]" name="Brazil" pageId="9" pageNumber="310">Brazil</collectingCountry>
|
||
(MZSP).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A655C732DFFE069F463A3AC45CC3C" pageId="9" pageNumber="310" type="discussion">
|
||
<paragraph id="8B8F36D7732DFFE069F463A3AD96CFA8" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
|
||
<emphasis id="B944EAC5732DFFE069F463A3AF0CCEDC" bold="true" box="[199,332,1231,1256]" pageId="9" pageNumber="310">Discussion.</emphasis>
|
||
<taxonomicName id="4C304D54732DFFE0686763BCAC0ECED3" box="[340,590,1232,1256]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="variegatus">
|
||
<emphasis id="B944EAC5732DFFE0686763BCAC0ECED3" box="[340,590,1232,1256]" italics="true" pageId="9" pageNumber="310">Ateralphus variegatus</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C304D54732DFFE06BDB63BCAD22CEDC" box="[744,866,1232,1256]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="tumidus" status="sp. nov.">
|
||
<emphasis id="B944EAC5732DFFE06BDB63BCAD22CEDC" box="[744,866,1232,1256]" italics="true" pageId="9" pageNumber="310">A. tumidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757BE732DFFE06A5863A3ADB6CEDC" box="[875,1014,1231,1256]" pageId="9" pageNumber="310" rank="species">
|
||
<emphasis id="B944EAC5732DFFE06A5863A3ADB6CEDC" bold="true" box="[875,1014,1231,1256]" pageId="9" pageNumber="310">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
and to
|
||
<emphasis id="B944EAC5732DFFE06D7C63BCAA87CEDC" box="[1103,1223,1232,1256]" italics="true" pageId="9" pageNumber="310">
|
||
<taxonomicName id="4C304D54732DFFE06D7C63BCAA83CEDC" box="[1103,1219,1232,1256]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="lacteus">A. lacteus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
It is differentiated from
|
||
<taxonomicName id="4C304D54732DFFE069E76398AF02CF38" box="[212,322,1268,1292]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="lacteus">
|
||
<emphasis id="B944EAC5732DFFE069E76398AF02CF38" box="[212,322,1268,1292]" italics="true" pageId="9" pageNumber="310">A. lacteus</emphasis>
|
||
</taxonomicName>
|
||
by characteristics cited in the above discussion.
|
||
<taxonomicName id="4C304D54732DFFE06A516398AA1ACF38" box="[866,1114,1268,1292]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="variegatus">
|
||
<emphasis id="B944EAC5732DFFE06A516398AA1ACF38" box="[866,1114,1268,1292]" italics="true" pageId="9" pageNumber="310">Ateralphus variegatus</emphasis>
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="4C304D54732DFFE06C176398ABDCCF38" box="[1316,1436,1268,1292]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="tumidus" status="sp. nov.">
|
||
<emphasis id="B944EAC5732DFFE06C176398ABDCCF38" box="[1316,1436,1268,1292]" italics="true" pageId="9" pageNumber="310">A. tumidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757BE732DFFE069A4627BAF64CF04" box="[151,292,1303,1328]" pageId="9" pageNumber="310" rank="species">
|
||
<emphasis id="B944EAC5732DFFE069A4627BAF64CF04" bold="true" box="[151,292,1303,1328]" pageId="9" pageNumber="310">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
, and is easily recognized by mesosternal process being flat at posterior half and tarsomere V with apical third covered with dark brown setae, while
|
||
<taxonomicName id="4C304D54732DFFE06BF66250AD7DCF60" box="[709,829,1340,1364]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="tumidus" status="sp. nov.">
|
||
<emphasis id="B944EAC5732DFFE06BF66250AD7DCF60" box="[709,829,1340,1364]" italics="true" pageId="9" pageNumber="310">A. tumidus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A27757BE732DFFE06A706250AD8DCF61" box="[835,973,1340,1365]" pageId="9" pageNumber="310" rank="species">
|
||
<emphasis id="B944EAC5732DFFE06A706250AD8DCF61" bold="true" box="[835,973,1340,1365]" pageId="9" pageNumber="310">new species</emphasis>
|
||
</taxonomicNameLabel>
|
||
has the mesosternal process tumescent at posterior half and tarsomere V with dark brown setae on base and apical third. Moreover,
|
||
<taxonomicName id="4C304D54732DFFE06D89620CAB12CF43" box="[1210,1362,1376,1399]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="variegatus">
|
||
<emphasis id="B944EAC5732DFFE06D89620CAB12CF43" box="[1210,1362,1376,1399]" italics="true" pageId="9" pageNumber="310">A. variegatus</emphasis>
|
||
</taxonomicName>
|
||
is the largest species of the genus, allowing its easy identification in
|
||
<emphasis id="B944EAC5732DFFE06A6062E8AD96CFA8" box="[851,982,1412,1436]" italics="true" pageId="9" pageNumber="310">
|
||
<taxonomicName id="4C304D54732DFFE06A6062E8AD92CFA8" box="[851,978,1412,1436]" class="Insecta" family="Cerambycidae" genus="Ateralphus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="310" phylum="Arthropoda" rank="genus">Ateralphus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8F36D7732DFFE069F462CBAC45CC3C" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
|
||
<emphasis id="B944EAC5732DFFE069F462CBAC49CFF4" bold="true" box="[199,521,1447,1472]" pageId="9" pageNumber="310">Geographical distribution.</emphasis>
|
||
<collectingCountry id="F3277647732DFFE06B2762CBAC18CFF4" box="[532,600,1447,1472]" name="Brazil" pageId="9" pageNumber="310">Brazil</collectingCountry>
|
||
(Minas Gerais, Espírito Santo, Rio de Janeiro, São Paulo and Paraná),
|
||
<collectingCountry id="F3277647732DFFE069A462A0AEAACFD1" box="[151,234,1484,1509]" name="Bolivia" pageId="9" pageNumber="310">Bolivia</collectingCountry>
|
||
(Santa Cruz (Wappes
|
||
<emphasis id="B944EAC5732DFFE068CB62A1AC6CCFD0" box="[504,556,1484,1508]" italics="true" pageId="9" pageNumber="310">et al</emphasis>
|
||
., 2006)). New state records are added from
|
||
<collectingCountry id="F3277647732DFFE06D0D62A0AAA8CFD1" box="[1086,1256,1484,1509]" name="French Guiana" pageId="9" pageNumber="310">French Guiana</collectingCountry>
|
||
(Cayenne) and
|
||
<collectingCountry id="F3277647732DFFE069A46283AF47CC3C" box="[151,263,1519,1544]" name="Argentina" pageId="9" pageNumber="310">Argentina</collectingCountry>
|
||
(Misiones) (
|
||
<figureCitation id="130B2A52732DFFE068A56283AFB8CC3C" box="[406,504,1519,1544]" captionStart="FIGURE 128" captionStartId="33.[151,250,1905,1927]" captionTargetBox="[189,1398,1130,1883]" captionTargetId="figure@33.[188,1400,1130,1883]" captionTargetPageId="33" captionText="FIGURE 128. Distribution map for Ateralphus lacteus Galileo & Martins, 2006, Ateralphus variegatus (Mendes, 1938) and Ateralphus tumidus new species." httpUri="https://zenodo.org/record/248473/files/figure.png" pageId="9" pageNumber="310">Fig. 128</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32A655C732DFFE069F46178AF6CCD78" pageId="9" pageNumber="310" type="materials_examined">
|
||
<paragraph id="8B8F36D7732DFFE069F46178AF6CCD78" blockId="9.[151,1436,151,1869]" pageId="9" pageNumber="310">
|
||
<emphasis id="B944EAC5732DFFE069F46178AFF4CC19" bold="true" box="[199,436,1556,1581]" pageId="9" pageNumber="310">Material examined.</emphasis>
|
||
<collectingCountry id="F3277647732DFFE0688C6178ACEACC18" box="[447,682,1556,1580]" name="French Guiana" pageId="9" pageNumber="310">FRENCH GUIANA</collectingCountry>
|
||
, Cayenne,
|
||
<specimenCount id="9D36FD5E732DFFE06A1E6178ADD5CC19" box="[813,917,1556,1581]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
, Fry leg. (BMNH).
|
||
<collectingCountry id="F3277647732DFFE06DB06178AAABCC18" box="[1155,1259,1556,1580]" name="Brazil" pageId="9" pageNumber="310">BRAZIL</collectingCountry>
|
||
, Minas Gerais: Teófilo Otoni,
|
||
<specimenCount id="9D36FD5E732DFFE068726154AFE6CC64" box="[321,422,1591,1616]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE068816154AC5BCC64" box="[434,539,1592,1616]" pageId="9" pageNumber="310" value="1970-12">XII.1970</date>
|
||
, F. M. Oliveira leg. (MNRJ);
|
||
<specimenCount id="9D36FD5E732DFFE06A416154AD81CC64" box="[882,961,1591,1616]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE06AFD6154AA6DCC64" box="[974,1069,1592,1616]" pageId="9" pageNumber="310" value="1974-11">XI.1974</date>
|
||
, S. P. Nascimento leg. (MNRJ); Espírito Santo: Linhares,
|
||
<specimenCount id="9D36FD5E732DFFE068866130AC58CC41" box="[437,536,1628,1653]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE06B106130AC37CC41" box="[547,631,1628,1653]" pageId="9" pageNumber="310" value="1969-10">X.1969</date>
|
||
(MNRJ); Rio Bonito,
|
||
<quantity id="4CC89B32732DFFE06A406130ADFCCC40" box="[883,956,1628,1653]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="310" unit="m" value="600.0">600 m</quantity>
|
||
,
|
||
<specimenCount id="9D36FD5E732DFFE06AF46130AA6BCC41" box="[967,1067,1628,1653]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE06D056130AAD3CC41" box="[1078,1171,1628,1653]" pageId="9" pageNumber="310" value="1963-11">XI.1963</date>
|
||
(MNRJ); Santa Teresa,
|
||
<specimenCount id="9D36FD5E732DFFE069A461ECAEBBCCAC" box="[151,251,1663,1688]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE068366113AFD0CCAC" box="[261,400,1663,1688]" pageId="9" pageNumber="310" value="1966-12-12">12.XII.1966</date>
|
||
, C. T. & C. Elias leg. (MZSP);
|
||
<specimenCount id="9D36FD5E732DFFE06BC361ECAD7DCCAC" box="[752,829,1663,1688]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE06A7A6113AD94CCAC" box="[841,980,1663,1688]" pageId="9" pageNumber="310" value="1966-12-27">27.XII.1966</date>
|
||
, C. Elias & C. T. Elias leg. (DZUP); São Paulo:
|
||
<specimenCount id="9D36FD5E732DFFE069DA61C8AF11CC89" box="[233,337,1700,1725]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
, 1985, J. Lane leg. (MZSP); Amparo,
|
||
<specimenCount id="9D36FD5E732DFFE06A2861C8AD2ACC89" box="[795,874,1700,1725]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
(MZSP); Campinas,
|
||
<typeStatus id="548B8875732DFFE06D5561C8AA8BCC89" box="[1126,1227,1700,1725]" pageId="9" pageNumber="310" type="holotype">holotype</typeStatus>
|
||
, female,
|
||
<date id="FF8E1017732DFFE06C0B61C8ABD7CC88" box="[1336,1431,1700,1724]" pageId="9" pageNumber="310" value="1919-11">XI.1919</date>
|
||
, Merbach leg. (MZSP); Ubatuba (Praia do Pulso),
|
||
<specimenCount id="9D36FD5E732DFFE06BE661A4AD7CCCD4" box="[725,828,1735,1760]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE06A7961A4ADF2CCD4" box="[842,946,1736,1760]" pageId="9" pageNumber="310" value="2003-12">XII.2003</date>
|
||
, M. L. Monné, J. R. Mermundes & M. A. Monné leg. (MNRJ); Rio de Janeiro: Rio de Janeiro (Corcovado),
|
||
<specimenCount id="9D36FD5E732DFFE06AAD6180ADB0CD31" box="[926,1008,1772,1797]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE06ACD6180AA37CD31" box="[1022,1143,1772,1797]" pageId="9" pageNumber="310" value="1954-10-28">28.X.1954</date>
|
||
, D. Zajciw leg. (MNRJ); Paraná: Matelândia,
|
||
<specimenCount id="9D36FD5E732DFFE0684E607CAFA1CD1C" box="[381,481,1807,1832]" pageId="9" pageNumber="310" type="female">1 female</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE068D86063AC73CD1C" box="[491,563,1807,1832]" pageId="9" pageNumber="310" value="1962-01">I.1962</date>
|
||
(MNRJ).
|
||
<collectingCountry id="F3277647732DFFE06B96607CAD0BCD1C" box="[677,843,1808,1832]" name="Argentina" pageId="9" pageNumber="310">ARGENTINA</collectingCountry>
|
||
, Misiones: Santa María,
|
||
<specimenCount id="9D36FD5E732DFFE06D52607CAAEECD1C" box="[1121,1198,1807,1832]" pageId="9" pageNumber="310" type="male">1 male</specimenCount>
|
||
,
|
||
<date id="FF8E1017732DFFE06D8A607CAB4FCD1C" box="[1209,1295,1808,1832]" pageId="9" pageNumber="310" value="1943-10">X.1943</date>
|
||
, M. J. Viana leg. (MNRJ).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |