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<document id="B5A75BFFA8E3F45D7B46433C8B95EF4F" ID-DOI="10.11646/zootaxa.4027.2.7" ID-GBIF-Dataset="25778e16-714a-4100-9277-04f5c8257bf6" ID-ISSN="1175-5326" ID-Zenodo-Dep="240633" ID-ZooBank="74D1205F-CB9F-472A-9FAE-A87E5D10FCD9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461282265536" checkinUser="plazi" docAuthor="Krupitsky, Anatoly V., Pljushtch, Igor G. &amp; Pak, Oleg V." docDate="2015" docId="039987BAFFA1FF94FF1DDD1F105CFC3F" docLanguage="en" docName="zt04027p286.pdf" docOrigin="Zootaxa 4027 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Callophrys succuba Krupitsky, Pljushtch &amp; Pak, sp. n." docType="treatment" docVersion="8" lastPageNumber="285" masterDocId="FFA0FFC2FFA0FF90FF8ADE53131FFF9F" masterDocTitle="A new species of the Callophrys paulae Pfeiffer, 1932 species group from Afghanistan (Lepidoptera, Lycaenidae)" masterLastPageNumber="286" masterPageNumber="281" pageNumber="282" updateTime="1698614341531" updateUser="plazi">
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<mods:title id="8BC8B6C74FB10A265612BD74E78D9446">A new species of the Callophrys paulae Pfeiffer, 1932 species group from Afghanistan (Lepidoptera, Lycaenidae)</mods:title>
</mods:titleInfo>
<mods:name id="879F33CA72B2D5DDD4C5EB400D8BCBA3" type="personal">
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<mods:namePart id="2506DC9110C2E32BC7CADB4DD5944791">Krupitsky, Anatoly V.</mods:namePart>
</mods:name>
<mods:name id="8F3DC210135A4CA39A8AD1B0CD9C3E2C" type="personal">
<mods:role id="5836DF018D862C6041E9059C207DF9A7">
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<mods:namePart id="704F4A672D54546B49CCFD909BAE1327">Pljushtch, Igor G.</mods:namePart>
</mods:name>
<mods:name id="F2FE6C17B1BBD6F217F1D8249B483AAD" type="personal">
<mods:role id="662BB3CAD8E65445C296E12B10D5A47A">
<mods:roleTerm id="C68AB628C4FD2A7C156241D6D2F0C577">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ED0EA2C124831DA35B14BE7F21F75395">Pak, Oleg V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="A7BF07F74763037831E147A9479A1ABB">text</mods:typeOfResource>
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<mods:title id="48B5303D0ECFEE6896CB0AB720854404">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="1FE42D801CC471FF9E9A0DBD6B03B791">
<mods:date id="AC79005A5CAF4BAC105103A57CC27C3E">2015</mods:date>
<mods:detail id="2B7696484C0C2A66BEE56C00D2F09960" type="volume">
<mods:number id="77A3BD43FA0D2B9831D090F0010168C5">4027</mods:number>
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<mods:detail id="D785A019EA711520CA7DA2572036D4E0" type="issue">
<mods:number id="390DDF178F04AC89DFC1A435BF4C06B7">2</mods:number>
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<mods:classification id="D198C1848714A2AAEB2170F546619163">journal article</mods:classification>
<mods:identifier id="1E4641D900F12C9141E79E92A44EE4F3" type="DOI">10.11646/zootaxa.4027.2.7</mods:identifier>
<mods:identifier id="C4025B35205390D5A6102E7522D8A4C6" type="GBIF-Dataset">25778e16-714a-4100-9277-04f5c8257bf6</mods:identifier>
<mods:identifier id="7A0B7E5ECBECB2CDB7A5CE504A25EBED" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A82295F1E24B1B8AD125FCEBF767F64B" type="Zenodo-Dep">240633</mods:identifier>
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<treatment id="039987BAFFA1FF94FF1DDD1F105CFC3F" ID-DOI="http://doi.org/10.5281/zenodo.6103376" ID-GBIF-Taxon="119640867" ID-Zenodo-Dep="6103376" LSID="urn:lsid:plazi:treatment:039987BAFFA1FF94FF1DDD1F105CFC3F" httpUri="http://treatment.plazi.org/id/039987BAFFA1FF94FF1DDD1F105CFC3F" lastPageId="4" lastPageNumber="285" pageId="1" pageNumber="282">
<subSubSection id="C32A6527FFA1FF91FF1DDD1F1102FC18" pageId="1" pageNumber="282" type="nomenclature">
<paragraph id="8B8F36ACFFA1FF91FF1DDD1F1025FCF8" blockId="1.[151,826,844,903]" box="[151,826,844,871]" pageId="1" pageNumber="282">
<heading id="D0C781C0FFA1FF91FF1DDD1F1025FCF8" bold="true" box="[151,826,844,871]" fontSize="11" level="1" pageId="1" pageNumber="282" reason="1">
<emphasis id="B944EABEFFA1FF91FF1DDD1F1025FCF8" bold="true" box="[151,826,844,871]" pageId="1" pageNumber="282">
<taxonomicName id="4C304D2FFFA1FF91FF1DDD1F11F4FCF8" authority="Krupitsky, Pljushtch &amp; Pak" authorityName="Krupitsky, Pljushtch &amp; Pak" box="[151,747,844,871]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="282" phylum="Arthropoda" rank="species" species="succuba" status="sp. nov.">
<emphasis id="B944EABEFFA1FF91FF1DDD1F1299FCF9" bold="true" box="[151,390,844,870]" italics="true" pageId="1" pageNumber="282">Callophrys succuba</emphasis>
Krupitsky, Pljushtch &amp; Pak
</taxonomicName>
,
<taxonomicNameLabel id="A27757C5FFA1FF91FD72DD1E1025FCF8" box="[760,826,845,871]" pageId="1" pageNumber="282" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36ACFFA1FF91FF1DDD231102FC18" blockId="1.[151,826,844,903]" box="[151,541,880,903]" pageId="1" pageNumber="282">(Plate 1, figs. 1–4; plate 2, figs. 1, 4)</paragraph>
</subSubSection>
<subSubSection id="C32A6527FFA1FF91FF1DDDE61750FC70" pageId="1" pageNumber="282" type="materials_examined">
<paragraph id="8B8F36ACFFA1FF91FF1DDDE61750FC70" blockId="1.[151,1436,949,2012]" pageId="1" pageNumber="282">
<emphasis id="B944EABEFFA1FF91FF1DDDE6121EFC53" bold="true" box="[151,257,949,972]" pageId="1" pageNumber="282">Material.</emphasis>
<typeStatus id="548B880EFFA1FF91FE85DDE6126DFC53" box="[271,370,949,972]" pageId="1" pageNumber="282" type="holotype">Holotype</typeStatus>
(
<collectionCode id="ED21AE69FFA1FF91FE02DDE612FDFC53" box="[392,482,949,972]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="1" pageNumber="282">ZMMU</collectionCode>
): ♂,
<collectingCountry id="F327763CFFA1FF91FDAFDDE611B5FC53" box="[549,682,949,972]" name="Afghanistan" pageId="1" pageNumber="282">Afghanistan</collectingCountry>
, Bamyan Province, Band-e Amir,
<geoCoordinate id="EE04506BFFA1FF91FBCADDE617B1FC53" box="[1088,1198,949,972]" direction="north" orientation="latitude" pageId="1" pageNumber="282" precision="925" value="34.8">34°48’ N</geoCoordinate>
,
<geoCoordinate id="EE04506BFFA1FF91FB4ADDE61636FC53" box="[1216,1321,949,972]" direction="east" orientation="longitude" pageId="1" pageNumber="282" precision="925" value="67.183334">67°11’ E</geoCoordinate>
,
<quantity id="4CC89B49FFA1FF91FAB6DDE61688FC53" box="[1340,1431,949,972]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="282" unit="m" value="3100.0">3100 m</quantity>
,
<date id="FF8E106CFFA1FF91FF1DDD8B120CFC70" box="[151,275,984,1007]" pageId="1" pageNumber="282" value="2012-06-01">
01.
<bibRefCitation id="EFA14B5DFFA1FF91FF3FDD8B120CFC70" author="VI" box="[181,275,984,1007]" pageId="1" pageNumber="282" refString="Davidson, J. P. (2012) Early modern supernatural: the dark side of European culture, 1400 - 1700. Santa Barbara, Praeger, 248 pp." type="book chapter" year="2012">VI.2012</bibRefCitation>
</date>
, I.G. Pljushtch leg.
<typeStatus id="548B880EFFA1FF91FE68DD8B1151FC70" box="[482,590,984,1007]" pageId="1" pageNumber="282" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D36FD25FFA1FF91FDD2DD8B119DFC70" box="[600,642,984,1007]" pageId="1" pageNumber="282" type="generic">5 ♂</specimenCount>
, 2 ♀ (AKM), same data, I.G. Pljushtch leg.
</paragraph>
</subSubSection>
<subSubSection id="C32A6527FFA1FF91FF4DDDA9113CF893" pageId="1" pageNumber="282" type="description">
<paragraph id="8B8F36ACFFA1FF91FF4DDDA9121BFAF3" blockId="1.[151,1436,949,2012]" pageId="1" pageNumber="282">
<emphasis id="B944EABEFFA1FF91FF4DDDA9124DFB8E" bold="true" box="[199,338,1018,1041]" pageId="1" pageNumber="282">Description.</emphasis>
<emphasis id="B944EABEFFA1FF91FED0DDA9128EFB8E" box="[346,401,1018,1041]" italics="true" pageId="1" pageNumber="282">Male</emphasis>
(plate 1, figs. 1–2). Head: antenna black, white-ringed at base of segments, club dark, its base white below, apiculus brown. Eye brown with small pale brown hairs, surrounded with green scales. Frons green with pale hairs on the sides, top of head with green scales and brownish hairs. Palpus: 2nd segment greenish-white with green scales and white, black and pale green hairs below; 3rd segment black with white hairs and scales. Thorax: upperside grey with grey hairs, underside with green scales and greenish-white hairs. Leg white with black scales and white hairs. Abdomen: upperside grey, underside grey with white hairs. Forewing oblong, narrow. Hindwing wavy with welldeveloped anal lobe. Upperside of both wings greyish-brown, margin dark-brown, tornus with ochraceous scales and hairs. Forewing with dark contrast oval androconial spot. Fringe white with brownish hairs near veins. Anal lobe of hindwing with small brush of dark hairs. Underside green with ochraceous outer margin. Submarginal line developed only on hindwing, not contrast, greenish-white, wavy. Forewing length 13,0 mm in the
<typeStatus id="548B880EFFA1FF91FBC7DB6117B2FAD6" box="[1101,1197,1330,1353]" pageId="1" pageNumber="282" type="holotype">holotype</typeStatus>
, 12,0–14,0 mm in the
<typeStatus id="548B880EFFA1FF91FF1DDB06121EFAF3" box="[151,257,1365,1388]" pageId="1" pageNumber="282" type="paratype">paratypes</typeStatus>
.
</paragraph>
<paragraph id="8B8F36ACFFA1FF91FF4DDB2B11DDF986" blockId="1.[151,1436,949,2012]" pageId="1" pageNumber="282">
<emphasis id="B944EABEFFA1FF91FF4DDB2B1278FA10" box="[199,359,1400,1423]" italics="true" pageId="1" pageNumber="282">Male genitalia</emphasis>
(plate 2, fig. 1). Annulus very wide—about 1,5x wider than maximum width of uncus, abruptly extended in the middle part of genitalia, has well developed internal lobes nearly reaching valvae. Lobes of uncus with well-developed broad processes rounded at tips. Falx slender, pointed at tip. Valva slightly broadened in membranous part, evenly pointed to rounded tip, nearly reaches tegumen. Saccus broad, rounded at tip. Aedeagus of typical shape for the genus, rather short, about 1,25x genitalia length.
</paragraph>
<paragraph id="8B8F36ACFFA1FF91FF4DD87617FBF9A3" blockId="1.[151,1436,949,2012]" box="[199,1252,1573,1596]" pageId="1" pageNumber="282">
<emphasis id="B944EABEFFA1FF91FF4DD8761208F9A3" box="[199,279,1573,1596]" italics="true" pageId="1" pageNumber="282">Female</emphasis>
(plate 1, figs. 3–4). Forewing length 11,5–12,5 mm. Similar to male, but forewing broader.
</paragraph>
<paragraph id="8B8F36ACFFA1FF91FF4DD81B1201F958" blockId="1.[151,1436,949,2012]" pageId="1" pageNumber="282">
<emphasis id="B944EABEFFA1FF91FF4DD81B1260F9C0" box="[199,383,1608,1631]" italics="true" pageId="1" pageNumber="282">Female genitalia</emphasis>
(plate 2, fig. 4). Papillae anales very thin and long, slightly shorter than apophyses posteriores. Lamella postvaginalis broad, central lobe with deep hollow, lateral lobes pointed to tips, divided from central lobe by rather deep rounded hollows. Ductus bursae short and thick, thinner in medial part, widened anteriorly. Bursa with twospined signi.
</paragraph>
<paragraph id="8B8F36ACFFA1FF91FF4DD881113CF893" blockId="1.[151,1436,949,2012]" pageId="1" pageNumber="282">
<emphasis id="B944EABEFFA1FF91FF4DD88112A2F976" bold="true" box="[199,445,1746,1769]" pageId="1" pageNumber="282">Individual variations.</emphasis>
Shape and contrast of the white submarginal line slightly varies. Significant variations in the male and female genitalia are absent.
</paragraph>
</subSubSection>
<subSubSection id="C32A6527FFA1FF92FF4DD94B13E0F91D" lastPageId="2" lastPageNumber="283" pageId="1" pageNumber="282" type="diagnosis">
<paragraph id="8B8F36ACFFA1FF91FF4DD94B10ACF843" blockId="1.[151,1436,949,2012]" pageId="1" pageNumber="282">
<emphasis id="B944EABEFFA1FF91FF4DD94B1223F8B0" bold="true" box="[199,316,1816,1839]" pageId="1" pageNumber="282">Diagnosis.</emphasis>
Combination of external and genital characters, namely rather narrow uncus and valva in males, trapezoidal distal part of ductus with broad base and broad central lobe in females, and small, but rather prominent anal lobe of hindwing, determine the position of the new species within
<taxonomicName id="4C304D2FFFA1FF91FC1AD90E10E3F8EB" box="[912,1020,1885,1908]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="282" phylum="Arthropoda" rank="species" species="paulae">
<emphasis id="B944EABEFFA1FF91FC1AD90E10E3F8EB" box="[912,1020,1885,1908]" italics="true" pageId="1" pageNumber="282">C. paulae</emphasis>
</taxonomicName>
species group. Differences in genital characters are of specific level as compared with the other
<taxonomicName id="4C304D2FFFA1FF91FCA2D9D31081F808" box="[808,926,1920,1943]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="282" phylum="Arthropoda" rank="genus">
<emphasis id="B944EABEFFA1FF91FCA2D9D31081F808" box="[808,926,1920,1943]" italics="true" pageId="1" pageNumber="282">Callophrys</emphasis>
</taxonomicName>
species—(ten Hagen 2009), for example, and probably perfrom prezygotic isolation. These differences led us to treat the population as the new species.
<taxonomicName id="4C304D2FFFA1FF91FB71D9F1169BF826" box="[1275,1412,1954,1977]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="282" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA1FF91FB71D9F1169BF826" box="[1275,1412,1954,1977]" italics="true" pageId="1" pageNumber="282">C. p. jomuda</emphasis>
</taxonomicName>
is the geographically closest taxon, so the new species was compared with it.
</paragraph>
<paragraph id="8B8F36ACFFA2FF92FF1DDA7513C0FA97" blockId="2.[151,1436,1061,1288]" pageId="2" pageNumber="283">
<emphasis id="B944EABEFFA2FF92FF1DDA7513E0FBA4" bold="true" box="[151,255,1062,1083]" pageId="2" pageNumber="283">PLATE 1.</emphasis>
<taxonomicName id="4C304D2FFFA2FF92FE8CDA76126FFBA4" box="[262,368,1061,1083]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="genus">
<emphasis id="B944EABEFFA2FF92FE8CDA76126FFBA4" box="[262,368,1061,1083]" italics="true" pageId="2" pageNumber="283">Callophrys</emphasis>
</taxonomicName>
spp., imagoes. Scale bar equals 10,0 mm. 1—
<taxonomicName id="4C304D2FFFA2FF92FCB0DA7610E5FBA4" box="[826,1018,1061,1083]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="succuba" status="sp. nov.">
<emphasis id="B944EABEFFA2FF92FCB0DA7610E5FBA4" box="[826,1018,1061,1083]" italics="true" pageId="2" pageNumber="283">Callophrys succuba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757C5FFA2FF92FB89DA751724FBA4" box="[1027,1083,1062,1083]" pageId="2" pageNumber="283" rank="species">
<emphasis id="B944EABEFFA2FF92FB89DA751724FBA4" bold="true" box="[1027,1083,1062,1083]" pageId="2" pageNumber="283">sp. n.</emphasis>
</taxonomicNameLabel>
, ♂,
<typeStatus id="548B880EFFA2FF92FBE2DA7517A1FBA4" box="[1128,1214,1062,1083]" pageId="2" pageNumber="283" type="holotype">holotype</typeStatus>
,
<collectingCountry id="F327763CFFA2FF92FB43DA75165FFBA4" box="[1225,1344,1062,1083]" name="Afghanistan" pageId="2" pageNumber="283">Afghanistan</collectingCountry>
, Bamyan Province, Band-e Amir,
<geoCoordinate id="EE04506BFFA2FF92FE08DA1012C2FBC7" box="[386,477,1091,1112]" direction="north" orientation="latitude" pageId="2" pageNumber="283" precision="925" value="34.8">34°48’ N</geoCoordinate>
,
<geoCoordinate id="EE04506BFFA2FF92FE63DA10115EFBC7" box="[489,577,1091,1112]" direction="east" orientation="longitude" pageId="2" pageNumber="283" precision="925" value="67.183334">67°11’ E</geoCoordinate>
,
<quantity id="4CC89B49FFA2FF92FDC1DA101189FBC7" box="[587,662,1091,1112]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="2" pageNumber="283" unit="m" value="3100.0">3100 m</quantity>
,
<date id="FF8E106CFFA2FF92FD2ADA10100FFBC7" box="[672,784,1091,1112]" pageId="2" pageNumber="283" value="2012-06-01">
01.
<bibRefCitation id="EFA14B5DFFA2FF92FD31DA10100FFBC7" author="VI" box="[699,784,1091,1112]" pageId="2" pageNumber="283" refString="Davidson, J. P. (2012) Early modern supernatural: the dark side of European culture, 1400 - 1700. Santa Barbara, Praeger, 248 pp." type="book chapter" year="2012">VI.2012</bibRefCitation>
</date>
, I.G. Pljushtch leg. (ZMMU); 2—Id., underside; 3—
<taxonomicName id="4C304D2FFFA2FF92FA87DA101667FBC6" box="[1293,1400,1091,1113]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="succuba" status="sp. nov.">
<emphasis id="B944EABEFFA2FF92FA87DA101667FBC6" box="[1293,1400,1091,1113]" italics="true" pageId="2" pageNumber="283">C. succuba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757C5FFA2FF92FAF5DA1013B5FBE9" pageId="2" pageNumber="283" rank="species">
<emphasis id="B944EABEFFA2FF92FAF5DA1013B5FBE9" bold="true" pageId="2" pageNumber="283">sp. n.</emphasis>
</taxonomicNameLabel>
, ♀,
<typeStatus id="548B880EFFA2FF92FF54DA32122CFBE9" box="[222,307,1121,1142]" pageId="2" pageNumber="283" type="paratype">paratype</typeStatus>
, same label as in the
<typeStatus id="548B880EFFA2FF92FD95DA33116CFBE9" box="[543,627,1120,1142]" pageId="2" pageNumber="283" type="holotype">holotype</typeStatus>
(AKM); 4—Id., underside; 5—
<taxonomicName id="4C304D2FFFA2FF92FC31DA33176EFBE9" box="[955,1137,1120,1142]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA2FF92FC31DA33176EFBE9" box="[955,1137,1120,1142]" italics="true" pageId="2" pageNumber="283">C. paulae jomuda</emphasis>
</taxonomicName>
, ♂ (
<typeStatus id="548B880EFFA2FF92FB2CDA3217E5FBE9" box="[1190,1274,1121,1142]" pageId="2" pageNumber="283" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C304D2FFFA2FF92FAAEDA331691FBE9" box="[1316,1422,1120,1142]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="hatuma">
<emphasis id="B944EABEFFA2FF92FAAEDA331691FBE9" box="[1316,1422,1120,1142]" italics="true" pageId="2" pageNumber="283">C. hatuma</emphasis>
</taxonomicName>
), [
<collectingCountry id="F327763CFFA2FF92FF16DA2D1237FB0C" box="[156,296,1150,1171]" name="Turkmenistan" pageId="2" pageNumber="283">Turkmenistan</collectingCountry>
], Kopet-Dagh Mts., Dushak, 30.0 4.1991,
<quantity id="4CC89B49FFA2FF92FD3CDA2D101DFB0C" box="[694,770,1150,1171]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="283" unit="m" value="2000.0">2000 m</quantity>
,. Zhdanko A. leg. (ZISP); 6—Id., underside; 7—
<taxonomicName id="4C304D2FFFA2FF92FB5EDA2E1661FB0C" box="[1236,1406,1149,1171]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA2FF92FB5EDA2E1661FB0C" box="[1236,1406,1149,1171]" italics="true" pageId="2" pageNumber="283">C. paulae jomuda</emphasis>
</taxonomicName>
, ♀ (
<typeStatus id="548B880EFFA2FF92FF17DAC813EEFB2F" box="[157,241,1179,1200]" pageId="2" pageNumber="283" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C304D2FFFA2FF92FE9CDAC81262FB2E" box="[278,381,1179,1201]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="hatuma">
<emphasis id="B944EABEFFA2FF92FE9CDAC81262FB2E" box="[278,381,1179,1201]" italics="true" pageId="2" pageNumber="283">C. hatuma</emphasis>
</taxonomicName>
), same label is in the previous specimen (ZISP); 8—Id., underside; 9—
<emphasis id="B944EABEFFA2FF92FBC0DAC817D4FB2E" box="[1098,1227,1179,1201]" italics="true" pageId="2" pageNumber="283">
С.
<taxonomicName id="4C304D2FFFA2FF92FBE2DACF17D4FB2E" box="[1128,1227,1179,1201]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">p. jomuda</taxonomicName>
</emphasis>
, ♂, W.–
<collectingCountry id="F327763CFFA2FF92FAABDAC81686FB2F" box="[1313,1433,1179,1200]" name="Afghanistan" pageId="2" pageNumber="283">Afghanistan</collectingCountry>
, Herat—Kala Nao [road], Kashka–Pass,
<quantity id="4CC89B49FFA2FF92FD9DDAEA117DFB51" box="[535,610,1209,1230]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="2" pageNumber="283" unit="m" value="2300.0">2300 m</quantity>
, Polsterwuchstformat.[ion], 18.5.[19]56, Amsel leg. (ZSM); 10—Id., underside; 11—
<emphasis id="B944EABEFFA2FF92FF1DDA861206FB74" box="[151,281,1237,1259]" italics="true" pageId="2" pageNumber="283">
С.
<taxonomicName id="4C304D2FFFA2FF92FF3CDA851206FB74" box="[182,281,1237,1259]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">p. jomuda</taxonomicName>
</emphasis>
, ♂, N
<collectingCountry id="F327763CFFA2FF92FEEEDA851292FB74" box="[356,397,1238,1259]" name="Iran" pageId="2" pageNumber="283">Iran</collectingCountry>
, Golestan, Shahkuh Mts., 50 SSE Gorgan, vic. of Tahar,
<quantity id="4CC89B49FFA2FF92FC44DA851747FB74" box="[974,1112,1238,1259]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="2" pageNumber="283" unit="m" value="2200.0" valueMax="2300.0" valueMin="2100.0">2100–2300 m</quantity>
,
<date id="FF8E106CFFA2FF92FBEFDA8517D6FB74" box="[1125,1225,1238,1259]" pageId="2" pageNumber="283" value="2009-05-07">07.V.2009</date>
, A.L. Devyatkin leg. (AKM)
</paragraph>
<paragraph id="8B8F36ACFFA2FF92FF4DDB6013E0F91D" blockId="2.[151,1436,1331,1874]" pageId="2" pageNumber="283">
<taxonomicName id="4C304D2FFFA2FF92FF4DDB601223FAD5" box="[199,316,1331,1354]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="species" species="succuba" status="sp. nov.">
<emphasis id="B944EABEFFA2FF92FF4DDB601223FAD5" box="[199,316,1331,1354]" italics="true" pageId="2" pageNumber="283">C. succuba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757C5FFA2FF92FEC9DB601261FAD5" box="[323,382,1331,1354]" pageId="2" pageNumber="283" rank="species">
<emphasis id="B944EABEFFA2FF92FEC9DB601261FAD5" bold="true" box="[323,382,1331,1354]" pageId="2" pageNumber="283">sp. n.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C304D2FFFA2FF92FD83DB60118DFAD5" box="[521,658,1331,1354]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA2FF92FD83DB60118DFAD5" box="[521,658,1331,1354]" italics="true" pageId="2" pageNumber="283">C. p. jomuda</emphasis>
</taxonomicName>
by the following external characters: narrow oblong forewing, especially in the males (forewing more rounded, broader in
<taxonomicName id="4C304D2FFFA2FF92FD15DB051037FAF2" box="[671,808,1366,1389]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA2FF92FD15DB051037FAF2" box="[671,808,1366,1389]" italics="true" pageId="2" pageNumber="283">C. p. jomuda</emphasis>
</taxonomicName>
), greyish-brown upperside (brown in
<taxonomicName id="4C304D2FFFA2FF92FB31DB05165BFAF2" box="[1211,1348,1366,1389]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA2FF92FB31DB05165BFAF2" box="[1211,1348,1366,1389]" italics="true" pageId="2" pageNumber="283">C. p. jomuda</emphasis>
</taxonomicName>
), nearly black androconial spot (usually of the same colour as ground or lighter in
<taxonomicName id="4C304D2FFFA2FF92FC26DB2A1729FA0F" box="[940,1078,1401,1424]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA2FF92FC26DB2A1729FA0F" box="[940,1078,1401,1424]" italics="true" pageId="2" pageNumber="283">C. p. jomuda</emphasis>
</taxonomicName>
), reduced white submarginal line on a forewing and fully developed but not contrast—on a hindwing, this line is consisted of bracket-shaped strokes and thus has wavy shape (hindwing with very contrast line consisted of broad white strokes in
<taxonomicName id="4C304D2FFFA2FF92FBDCDBED17C0FA4A" box="[1110,1247,1470,1493]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA2FF92FBDCDBED17C0FA4A" box="[1110,1247,1470,1493]" italics="true" pageId="2" pageNumber="283">C. p. jomuda</emphasis>
</taxonomicName>
, and usually only one stroke near anal angle is V-shaped). The male genitalia of the new species are generally thicker and shorter than in
<taxonomicName id="4C304D2FFFA2FF92FA0EDBB2121AF985" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA2FF92FA0EDBB2121AF985" italics="true" pageId="2" pageNumber="283">C. p. jomuda</emphasis>
</taxonomicName>
, with broader rounded vinculum from dorsal and ventral view, and shorter and more slender saccus. The female genitalia of the new species have broad divided lamella postvaginalis with well-developed sharp lateral lobes and a broad central lobe (lobes are poorly developed and not so divided in
<taxonomicName id="4C304D2FFFA2FF92FC31D81A1753F9FF" box="[955,1100,1609,1632]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="283" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">
<emphasis id="B944EABEFFA2FF92FC31D81A1753F9FF" box="[955,1100,1609,1632]" italics="true" pageId="2" pageNumber="283">C. p. jomuda</emphasis>
</taxonomicName>
, central lobe is narrowed and rounded).
</paragraph>
</subSubSection>
<subSubSection id="C32A6527FFA2FF92FF4DD8DD1075F957" pageId="2" pageNumber="283" type="distribution">
<paragraph id="8B8F36ACFFA2FF92FF4DD8DD1075F957" blockId="2.[151,1436,1331,1874]" pageId="2" pageNumber="283">
<emphasis id="B944EABEFFA2FF92FF4DD8DD1118F93A" bold="true" box="[199,519,1678,1701]" pageId="2" pageNumber="283">Distribution and bionomics.</emphasis>
The new species is known only from the
<typeStatus id="548B880EFFA2FF92FC46D8DD10E5F93A" box="[972,1018,1678,1701]" pageId="2" pageNumber="283">type</typeStatus>
locality so far (plate 3). Imagoes were found in the beginning of June being slightly worn. See Discussion.
</paragraph>
</subSubSection>
<subSubSection id="C32A6527FFA2FF94FF4DD880105CFC3F" lastPageId="4" lastPageNumber="285" pageId="2" pageNumber="283" type="etymology">
<paragraph id="8B8F36ACFFA2FF92FF4DD88017A2F8CD" blockId="2.[151,1436,1331,1874]" pageId="2" pageNumber="283">
<emphasis id="B944EABEFFA2FF92FF4DD8801258F975" bold="true" box="[199,327,1747,1770]" pageId="2" pageNumber="283">Etymology.</emphasis>
<taxonomicName id="4C304D2FFFA2FF92FEC4D88012B6F975" box="[334,425,1747,1770]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="285" phylum="Arthropoda" rank="species" species="succuba" status="sp. nov.">Succuba</taxonomicName>
(paramour, Latin)—a folkloric demon that appears in dreams and takes a form of a woman in order to seduce men. The name refers to the serrated submarginal line unusual for the genus members, and wavy outer margin of a hindwing—in the Medieval culture succubae were often pictured and sculptured as a beautiful young girl possessing demonic features such as serrated wings, bird claws or serpentine tails (
<bibRefCitation id="EFA14B5DFFA2FF92FB83D96817ADF8CD" author="Davidson" box="[1033,1202,1851,1874]" pageId="2" pageNumber="283" refString="Davidson, J. P. (2012) Early modern supernatural: the dark side of European culture, 1400 - 1700. Santa Barbara, Praeger, 248 pp." type="book chapter" year="2012">Davidson 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8B8F36ACFFA3FF93FF1DD941126DF86B" blockId="3.[151,1436,1809,2036]" pageId="3" pageNumber="284">
<emphasis id="B944EABEFFA3FF93FF1DD94113E0F8B8" bold="true" box="[151,255,1810,1831]" pageId="3" pageNumber="284">PLATE 2.</emphasis>
<taxonomicName id="4C304D2FFFA3FF93FE8CD942126EF8B8" box="[262,369,1809,1831]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="284" phylum="Arthropoda" rank="genus">
<emphasis id="B944EABEFFA3FF93FE8CD942126EF8B8" box="[262,369,1809,1831]" italics="true" pageId="3" pageNumber="284">Callophrys</emphasis>
</taxonomicName>
spp., genitalia. Scale bar equals 1,0 mm. 1—
<taxonomicName id="4C304D2FFFA3FF93FCA4D94210F1F8B8" box="[814,1006,1809,1831]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="284" phylum="Arthropoda" rank="species" species="succuba" status="sp. nov.">
<emphasis id="B944EABEFFA3FF93FCA4D94210F1F8B8" box="[814,1006,1809,1831]" italics="true" pageId="3" pageNumber="284">Callophrys succuba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757C5FFA3FF93FC7CD9411731F8B8" box="[1014,1070,1810,1831]" pageId="3" pageNumber="284" rank="species">
<emphasis id="B944EABEFFA3FF93FC7CD9411731F8B8" bold="true" box="[1014,1070,1810,1831]" pageId="3" pageNumber="284">sp. n.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="548B880EFFA3FF93FBB1D941178FF8B8" box="[1083,1168,1810,1831]" pageId="3" pageNumber="284" type="paratype">paratype</typeStatus>
, male genitalia (a—general view from ventral side, b—aedeagus, lateral view),
<collectingCountry id="F327763CFFA3FF93FD2BD97C1006F8DB" box="[673,793,1839,1860]" name="Afghanistan" pageId="3" pageNumber="284">Afghanistan</collectingCountry>
, Bamyan Province, Band-e Amir,
<geoCoordinate id="EE04506BFFA3FF93FBFED97C17CCF8DB" box="[1140,1235,1839,1860]" direction="north" orientation="latitude" pageId="3" pageNumber="284" precision="925" value="34.8">34°48’ N</geoCoordinate>
,
<geoCoordinate id="EE04506BFFA3FF93FB6AD97C1623F8DB" box="[1248,1340,1839,1860]" direction="east" orientation="longitude" pageId="3" pageNumber="284" precision="925" value="67.183334">67°11’ E</geoCoordinate>
,
<quantity id="4CC89B49FFA3FF93FAC3D97C1687F8DB" box="[1353,1432,1839,1860]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="284" unit="m" value="3100.0">3100 m</quantity>
,
<date id="FF8E106CFFA3FF93FF1DD91E1219F8FD" box="[151,262,1869,1890]" pageId="3" pageNumber="284" value="2012-06-01">
01.
<bibRefCitation id="EFA14B5DFFA3FF93FF38D91E1219F8FD" author="VI" box="[178,262,1869,1890]" pageId="3" pageNumber="284" refString="Davidson, J. P. (2012) Early modern supernatural: the dark side of European culture, 1400 - 1700. Santa Barbara, Praeger, 248 pp." type="book chapter" year="2012">VI.2012</bibRefCitation>
</date>
, I.G. Pljushtch leg. (AKM); 2—
<emphasis id="B944EABEFFA3FF93FDC9D91F11D4F8FD" box="[579,715,1868,1890]" italics="true" pageId="3" pageNumber="284">
С.
<taxonomicName id="4C304D2FFFA3FF93FDE3D91E11D4F8FD" box="[617,715,1868,1890]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="284" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">p. jomuda</taxonomicName>
</emphasis>
, male genitalia, N
<collectingCountry id="F327763CFFA3FF93FC03D91E10ADF8FD" box="[905,946,1869,1890]" name="Iran" pageId="3" pageNumber="284">Iran</collectingCountry>
, Golestan, Shahkuh Mts., 50 SSE Gorgan, vic. of Tahar,
<quantity id="4CC89B49FFA3FF93FF50D939127DF8E0" box="[218,354,1898,1919]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="3" pageNumber="284" unit="m" value="2200.0" valueMax="2300.0" valueMin="2100.0">2100–2300 m</quantity>
,
<date id="FF8E106CFFA3FF93FEE7D93912CEF8E0" box="[365,465,1898,1919]" pageId="3" pageNumber="284" value="2009-05-07">07.V.2009</date>
, A.L. Devyatkin leg. (AKM); 3—
<emphasis id="B944EABEFFA3FF93FCAAD93A10B9F8E0" box="[800,934,1897,1919]" italics="true" pageId="3" pageNumber="284">
С.
<taxonomicName id="4C304D2FFFA3FF93FCCED93910B9F8E0" box="[836,934,1897,1919]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="284" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">p. jomuda</taxonomicName>
</emphasis>
, male genitalia, W.–
<collectingCountry id="F327763CFFA3FF93FBE5D93917F8F8E0" box="[1135,1255,1898,1919]" name="Afghanistan" pageId="3" pageNumber="284">Afghanistan</collectingCountry>
, Heart—Kala Nao [road], Kashka–Pass,
<quantity id="4CC89B49FFA3FF93FEE4D9D412A3F803" box="[366,444,1927,1948]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="284" unit="m" value="2300.0">2300 m</quantity>
, Polsterwuchstformat.[ion], 18.5.[19]56, Amsel leg. (ZSM); 4—
<taxonomicName id="4C304D2FFFA3FF93FBB0D9D417B7F802" box="[1082,1192,1927,1949]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="284" phylum="Arthropoda" rank="species" species="succuba" status="sp. nov.">
<emphasis id="B944EABEFFA3FF93FBB0D9D417B7F802" box="[1082,1192,1927,1949]" italics="true" pageId="3" pageNumber="284">C. succuba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757C5FFA3FF93FB3BD9D417F5F803" box="[1201,1258,1927,1948]" pageId="3" pageNumber="284" rank="species">
<emphasis id="B944EABEFFA3FF93FB3BD9D417F5F803" bold="true" box="[1201,1258,1927,1948]" pageId="3" pageNumber="284">sp. n.</emphasis>
</taxonomicNameLabel>
,
<typeStatus id="548B880EFFA3FF93FB73D9D41651F803" box="[1273,1358,1927,1948]" pageId="3" pageNumber="284" type="paratype">paratype</typeStatus>
, female genitalia (dorsal view),
<collectingCountry id="F327763CFFA3FF93FE0AD9F612E7F825" box="[384,504,1957,1978]" name="Afghanistan" pageId="3" pageNumber="284">Afghanistan</collectingCountry>
, Bamyan Province, Band-e Amir,
<geoCoordinate id="EE04506BFFA3FF93FCC1D9F610B7F825" box="[843,936,1957,1978]" direction="north" orientation="latitude" pageId="3" pageNumber="284" precision="925" value="34.8">34°48’ N</geoCoordinate>
,
<geoCoordinate id="EE04506BFFA3FF93FC3ED9F61712F825" box="[948,1037,1957,1978]" direction="east" orientation="longitude" pageId="3" pageNumber="284" precision="925" value="67.183334">67°11’ E</geoCoordinate>
,
<quantity id="4CC89B49FFA3FF93FB93D9F61779F825" box="[1049,1126,1957,1978]" metricMagnitude="3" metricUnit="m" metricValue="3.1" pageId="3" pageNumber="284" unit="m" value="3100.0">3100 m</quantity>
,
<date id="FF8E106CFFA3FF93FBF8D9F617FEF825" box="[1138,1249,1957,1978]" pageId="3" pageNumber="284" value="2012-06-01">
01.
<bibRefCitation id="EFA14B5DFFA3FF93FB07D9F617FEF825" author="VI" box="[1165,1249,1957,1978]" pageId="3" pageNumber="284" refString="Davidson, J. P. (2012) Early modern supernatural: the dark side of European culture, 1400 - 1700. Santa Barbara, Praeger, 248 pp." type="book chapter" year="2012">VI.2012</bibRefCitation>
</date>
, I.G. Pljushtch leg. (AKM); 5—
<emphasis id="B944EABEFFA3FF93FE9ED9921284F848" box="[276,411,1985,2007]" italics="true" pageId="3" pageNumber="284">
С.
<taxonomicName id="4C304D2FFFA3FF93FEBCD9911284F848" box="[310,411,1985,2007]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="284" phylum="Arthropoda" rank="subSpecies" species="paulae" subSpecies="jomuda">p. jomuda</taxonomicName>
</emphasis>
, female genitalia, N
<collectingCountry id="F327763CFFA3FF93FDF3D99111BDF848" box="[633,674,1986,2007]" name="Iran" pageId="3" pageNumber="284">Iran</collectingCountry>
, Golestan, Shahkuh Mts., 35 S Gorgan, plateau,
<quantity id="4CC89B49FFA3FF93FB17D99117F2F848" box="[1181,1261,1986,2007]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="3" pageNumber="284" unit="m" value="2300.0">2300 m</quantity>
,
<date id="FF8E106CFFA3FF93FB76D991167FF848" box="[1276,1376,1986,2007]" pageId="3" pageNumber="284" value="2009-05-10">10.V.2009</date>
, A.L. Devyatkin leg. (AKM)
</paragraph>
<paragraph id="8B8F36ACFFA4FF94FF1DDD03105CFC3F" blockId="4.[151,1436,848,928]" pageId="4" pageNumber="285">
<emphasis id="B944EABEFFA4FF94FF1DDD0313E2FCFA" bold="true" box="[151,253,848,869]" pageId="4" pageNumber="285">PLATE 3.</emphasis>
Distribution of the
<taxonomicName id="4C304D2FFFA4FF94FE31DD031173FCF9" box="[443,620,848,870]" class="Insecta" family="Lycaenidae" genus="Callophrys" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="285" phylum="Arthropoda" rank="species" species="paulae">
<emphasis id="B944EABEFFA4FF94FE31DD031173FCF9" box="[443,620,848,870]" italics="true" pageId="4" pageNumber="285">Callophrys paulae</emphasis>
</taxonomicName>
species group based on original data and reliable data from literature (
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Hesselbarth
<emphasis id="B944EABEFFA4FF94FA01DD0213B5FC1C" italics="true" pageId="4" pageNumber="285">et al</emphasis>
. 1995
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;
<bibRefCitation id="EFA14B5DFFA4FF94FF7FDD3D126FFC1C" author="Nazari" box="[245,368,878,899]" pageId="4" pageNumber="285" refString="Nazari, V. (2003) Butterflies of Iran. Tehran, Dayereh-Sabz, National Museum of Natural History of the Islamic Republic of Iran, 568 pp. [in Persian]" type="book" year="2003">Nazari 2003</bibRefCitation>
; ten Hagen &amp; Miller 2010;
<bibRefCitation id="EFA14B5DFFA4FF94FD09DD3D10D4FC1C" author="Tshikolovets" box="[643,971,878,899]" pageId="4" pageNumber="285" refString="Tshikolovets, V. &amp; Nekrutenko, Y. (2012) The butterflies of Caucasus and Transcaucasia. Pardubice, Tshikolovets Publications, 423 pp." type="book" year="2012">Tshikolovets &amp; Nekrutenko 2012</bibRefCitation>
;
<bibRefCitation id="EFA14B5DFFA4FF94FC5DDD3D17D9FC1C" author="Tshikolovets" box="[983,1222,877,899]" pageId="4" pageNumber="285" refString="Tshikolovets, V., Naderi, A. &amp; Eckweiler, W. (2014) The butterflies of Iran and Iraq. Pardubice, Tshikolovets Publications, 440 pp." type="book" year="2014">
Tshikolovets
<emphasis id="B944EABEFFA4FF94FBD3DD3D179AFC1C" box="[1113,1157,877,899]" italics="true" pageId="4" pageNumber="285">et al</emphasis>
. 2014
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). Triangles mark
<typeStatus id="548B880EFFA4FF94FAF8DD3D1683FC1C" box="[1394,1436,878,899]" pageId="4" pageNumber="285">type</typeStatus>
localities. Question-mark indicates a lack of material from the territory.
</paragraph>
</subSubSection>
</treatment>
</document>
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