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<document id="86F00A61D4708ABC9E181D43B963BA72" ID-CLB-Dataset="22458" ID-DOI="10.5852/ejt.2020.622" ID-GBIF-Dataset="9cb3ea96-0ee4-40b4-8006-19b5fed80b46" ID-Zenodo-Dep="3777282" ID-ZooBank="9CD3B2CD-F072-4994-8CA9-24145D343401" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1588217079841" checkinUser="valdenar" docAuthor="Darby, Michael" docDate="2020" docId="039987A1383FFFE02B70FD30FAEBFB6F" docLanguage="en" docName="ejt-622_darby.pdf.imf" docOrigin="European Journal of Taxonomy 622" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cissidium ishigakiense Sawada 2008" docType="treatment" docVersion="7" lastPageNumber="47" masterDocId="FFA0FFD93812FFCE2949FFEEFF9CFF98" masterDocTitle="A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species" masterLastPageNumber="188" masterPageNumber="1" pageNumber="46" updateTime="1698838121940" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A37325D4670E22B2838FA4EC1F323202">A revision of Cissidium Motschulsky (Coleoptera: Ptiliidae) with seventy seven new species</mods:title>
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<mods:namePart id="1CAB1C1CE7D0B7602EE6BE6838B69663">Darby, Michael</mods:namePart>
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<mods:title id="329C9E4AF7E03B95A31D4C79F9D5A026">European Journal of Taxonomy</mods:title>
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<mods:date id="6BAD97C4E360F64E9341DE6FE7C00644">2020</mods:date>
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<treatment id="039987A1383FFFE02B70FD30FAEBFB6F" ID-DOI="http://doi.org/10.5281/zenodo.3795578" ID-GBIF-Taxon="163877856" ID-Zenodo-Dep="3795578" LSID="urn:lsid:plazi:treatment:039987A1383FFFE02B70FD30FAEBFB6F" httpUri="http://treatment.plazi.org/id/039987A1383FFFE02B70FD30FAEBFB6F" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">
<subSubSection id="C32A653C383FFFE32B70FD30FC67FD60" box="[569,1019,734,761]" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="8B8F36B7383FFFE32B70FD30FC67FD60" blockId="45.[569,1019,734,795]" box="[569,1019,734,761]" pageId="45" pageNumber="46">
<heading id="D0C781DB383FFFE32B70FD30FC67FD60" box="[569,1019,734,761]" centered="true" fontSize="11" level="2" pageId="45" pageNumber="46" reason="2">
<taxonomicName id="4C304D34383FFFE32B70FD30FC67FD60" ID-CoL="VK8C" authority="Sawada" authorityName="Sawada" authorityYear="2008" box="[569,1019,734,761]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="ishigakiense">
<emphasis id="B944EAA5383FFFE32B70FD30FCD3FD61" bold="true" box="[569,847,734,761]" italics="true" pageId="45" pageNumber="46">Cissidium ishigakiense</emphasis>
<bibRefCitation id="EFA14B46383FFFE32A1FFD30FC67FD60" author="Sawada Y." box="[854,1019,734,760]" pageId="45" pageNumber="46" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C383FFFE32BB9FCEFFCDFFC83" box="[752,835,769,795]" pageId="45" pageNumber="46" type="description">
<paragraph id="8B8F36B7383FFFE32BB9FCEFFCDFFC83" blockId="45.[569,1019,734,795]" box="[752,835,769,795]" pageId="45" pageNumber="46">
<figureCitation id="130B2A32383FFFE32BB9FCEFFCDFFC83" box="[752,835,769,795]" captionStart="Fig" captionStartId="45.[189,232,1964,1990]" captionTargetBox="[192,1398,1087,1918]" captionTargetId="figure@45.[189,1398,1087,1922]" captionTargetPageId="45" captionText="Fig. 29. Cissidium ishigakiense Sawada, 2008. A. Habitus. B. Pronotum from slide preparation. C. Meso- and metaventrite from slide preparation." figureDoi="http://doi.org/10.5281/zenodo.3777342" httpUri="https://zenodo.org/record/3777342/files/figure.png" pageId="45" pageNumber="46">Fig. 29</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A653C383FFFE029F4FCADFB57FBFF" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="8B8F36B7383FFFE329F4FCADFE36FCC5" blockId="45.[189,426,835,861]" box="[189,426,835,861]" pageId="45" pageNumber="46">
<heading id="D0C781DB383FFFE329F4FCADFE36FCC5" bold="true" box="[189,426,835,861]" fontSize="11" level="3" pageId="45" pageNumber="46" reason="3">
<emphasis id="B944EAA5383FFFE329F4FCADFE36FCC5" bold="true" box="[189,426,835,861]" pageId="45" pageNumber="46">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7383FFFE329A3FC9CFEC0FC14" blockId="45.[189,1399,882,978]" box="[234,348,882,908]" pageId="45" pageNumber="46">
<materialsCitation id="3B583CEA383FFFE329A3FC9CFEC0FC14" ID-GBIF-Occurrence="2609493623" box="[234,348,882,908]" collectingDate="1999-05-09" collectionCode="YS" collectorName="Y. Sawada" country="Japan" location="Ishigaki Island" municipality="Komi" pageId="45" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa Pref." typeStatus="paratype">
<emphasis id="B944EAA5383FFFE329A3FC9CFEC0FC14" bold="true" box="[234,348,882,908]" pageId="45" pageNumber="46">
<typeStatus id="548B8815383FFFE329A3FC9CFEC0FC14" box="[234,348,882,908]" pageId="45" pageNumber="46" type="paratype">Paratype</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<paragraph id="8B8F36B7383FFFE329F4FC7BFD95FC4A" blockId="45.[189,1399,882,978]" pageId="45" pageNumber="46">
<collectingCountry id="F3277627383FFFE329F4FC7BFE8AFC37" box="[189,278,917,943]" name="Japan" pageId="45" pageNumber="46">JAPAN</collectingCountry>
•
<specimenCount id="9D36FD3E383FFFE32862FC7BFEC4FC37" box="[299,344,917,943]" pageId="45" pageNumber="46" type="male">1 ♂</specimenCount>
;
<collectingMunicipality id="6BEBACCD383FFFE3282DFC7AFE36FC36" box="[356,426,916,942]" pageId="45" pageNumber="46">Komi</collectingMunicipality>
,
<location id="8EEF606C383FFFE328FDFC7AFDFFFC36" LSID="urn:lsid:plazi:treatment:039987A1383FFFE02B70FD30FAEBFB6F:8EEF606C383FFFE328FDFC7AFDFFFC36" box="[436,611,916,942]" country="Japan" municipality="Komi" name="Ishigaki Island" pageId="45" pageNumber="46" stateProvince="Okinawa Pref.">Ishigaki Island</location>
,
<collectingRegion id="49F4F855383FFFE32B24FC7AFC84FC36" box="[621,792,916,942]" country="Japan" name="Okinawa" pageId="45" pageNumber="46">Okinawa Pref.</collectingRegion>
;
<date id="FF8E1077383FFFE32A6BFC7BFC2CFC37" box="[802,944,917,943]" pageId="45" pageNumber="46" value="1999-05-09">
<collectingDate id="EFCAE99F383FFFE32A6BFC7BFC2CFC37" box="[802,944,917,943]" pageId="45" pageNumber="46" value="1999-05-09">9 May 1999</collectingDate>
</date>
; pile of straw;
<collectorName id="26C55361383FFFE32D1EFC7BFB4DFC36" box="[1111,1233,916,943]" pageId="45" pageNumber="46">Y. Sawada</collectorName>
leg.; mounted as a disarticulated slide;
<collectionCode id="ED21AE72383FFFE32895FC56FD99FC4A" box="[476,517,952,978]" pageId="45" pageNumber="46">YS</collectionCode>
.
</paragraph>
<caption id="DF4F663F383FFFE329F4F842FD09F871" ID-DOI="http://doi.org/10.5281/zenodo.3777342" ID-Zenodo-Dep="3777342" httpUri="https://zenodo.org/record/3777342/files/figure.png" pageId="45" pageNumber="46" startId="45.[189,232,1964,1990]" targetBox="[192,1398,1087,1918]" targetPageId="45">
<paragraph id="8B8F36B7383FFFE329F4F842FD09F871" blockId="45.[189,1398,1964,2025]" pageId="45" pageNumber="46">
<emphasis id="B944EAA5383FFFE329F4F842FE8BF85E" bold="true" box="[189,279,1964,1990]" pageId="45" pageNumber="46">Fig. 29.</emphasis>
<taxonomicName id="4C304D34383FFFE32855F842FD4EF85E" authority="Sawada, 2008" authorityName="Sawada" authorityYear="2008" box="[284,722,1964,1990]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="ishigakiense">
<emphasis id="B944EAA5383FFFE32855F842FDBBF85E" box="[284,551,1964,1990]" italics="true" pageId="45" pageNumber="46">Cissidium ishigakiense</emphasis>
<bibRefCitation id="EFA14B46383FFFE32B65F842FD4EF85E" author="Sawada Y." box="[556,722,1964,1990]" pageId="45" pageNumber="46" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada, 2008</bibRefCitation>
</taxonomicName>
.
<emphasis id="B944EAA5383FFFE32B92F842FD6CF85E" bold="true" box="[731,752,1964,1990]" pageId="45" pageNumber="46">A</emphasis>
. Habitus.
<emphasis id="B944EAA5383FFFE32A2DF843FCE4F85F" bold="true" box="[868,888,1965,1991]" pageId="45" pageNumber="46">B</emphasis>
. Pronotum from slide preparation.
<emphasis id="B944EAA5383FFFE32C43F842FA83F85E" bold="true" box="[1290,1311,1964,1990]" pageId="45" pageNumber="46">C</emphasis>
. Meso- and metaventrite from slide preparation.
</paragraph>
</caption>
<paragraph id="8B8F36B7383CFFE029F4FEE5FD8DFEBD" blockId="46.[189,529,267,293]" box="[189,529,267,293]" pageId="46" pageNumber="47">
<heading id="D0C781DB383CFFE029F4FEE5FD8DFEBD" bold="true" box="[189,529,267,293]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B944EAA5383CFFE029F4FEE5FD8DFEBD" bold="true" box="[189,529,267,293]" pageId="46" pageNumber="47">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FED6FBFEFEED" blockId="46.[189,1398,312,373]" pageId="46" pageNumber="47">
The description below concentrates on amplifying
<bibRefCitation id="EFA14B46383CFFE02A52FED6FC51FECA" author="Sawada Y." box="[795,973,312,338]" pageId="46" pageNumber="47" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada (2008)</bibRefCitation>
with more details of the meso- and metaventral characters in order to conform with the entries in the present paper.
</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FE75FC48FE2D" blockId="46.[189,980,411,437]" box="[189,980,411,437]" pageId="46" pageNumber="47">
SIZE. Habitus (
<figureCitation id="130B2A32383CFFE0282EFE75FE53FE2D" box="[359,463,411,437]" captionStart="Fig" captionStartId="45.[189,232,1964,1990]" captionTargetBox="[192,1398,1087,1918]" captionTargetId="figure@45.[189,1398,1087,1922]" captionTargetPageId="45" captionText="Fig. 29. Cissidium ishigakiense Sawada, 2008. A. Habitus. B. Pronotum from slide preparation. C. Meso- and metaventrite from slide preparation." figureDoi="http://doi.org/10.5281/zenodo.3777342" httpUri="https://zenodo.org/record/3777342/files/figure.png" pageId="46" pageNumber="47">Fig. 29A</figureCitation>
copied from
<bibRefCitation id="EFA14B46383CFFE02B25FE75FC92FE2D" author="Sawada Y." box="[620,782,411,437]" pageId="46" pageNumber="47" pagination="101 - 126" refId="ref59928" refString="Sawada Y. 2008. A Revision of the Genus Cissidium (Coleoptera: Ptiliidae) in Japan. Special Publication 2: 101 - 126. Japan Coelopterological Society, Osaka." type="journal article" year="2008">Sawada 2008</bibRefCitation>
), length
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.
</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FE35FD48FE6D" blockId="46.[189,724,475,502]" box="[189,724,475,502]" pageId="46" pageNumber="47">COLOUR. Dark brown, antennae dusky yellow.</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FDF2FD13FDC0" blockId="46.[189,1399,540,600]" pageId="46" pageNumber="47">
HEAD. Width across eyes
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; mentum obscured; antennomeres III–XI, length
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, III–IX length
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, X–XI length
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.
</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FD91FCBEFD45" blockId="46.[189,1398,638,733]" pageId="46" pageNumber="47">
PRONOTUM.
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long,
<quantity id="4CC89B52383CFFE028A2FD90FDCDFD01" box="[491,593,638,665]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="46" pageNumber="47" unit="mm" value="0.27">0.27 mm</quantity>
wide, lateral margins rounded then almost parallel sided to rectangular hind angles, bordered, the borders not continuing along the posterior margin, medial emargination opposite the scutellum shallowly sinuate (
<figureCitation id="130B2A32383CFFE02BE0FD2CFC88FD45" box="[681,788,706,733]" captionStart="Fig" captionStartId="45.[189,232,1964,1990]" captionTargetBox="[192,1398,1087,1918]" captionTargetId="figure@45.[189,1398,1087,1922]" captionTargetPageId="45" captionText="Fig. 29. Cissidium ishigakiense Sawada, 2008. A. Habitus. B. Pronotum from slide preparation. C. Meso- and metaventrite from slide preparation." figureDoi="http://doi.org/10.5281/zenodo.3777342" httpUri="https://zenodo.org/record/3777342/files/figure.png" pageId="46" pageNumber="47">Fig. 29B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FCEDFD1FFC85" blockId="46.[189,643,771,797]" box="[189,643,771,797]" pageId="46" pageNumber="47">
ELYTRA.
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long,
<quantity id="4CC89B52383CFFE0289BFCEDFDA7FC85" box="[466,571,771,797]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="46" pageNumber="47" unit="mm" value="0.4">0.40 mm</quantity>
wide.
</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FCAAFD39FC5C" blockId="46.[189,1399,835,964]" pageId="46" pageNumber="47">
MESOVENTRITE. Medial extension of collar tapering from broad base to mid-point of mid-keel; mid-keel sides concave, hind angles not reaching mesocoxal anterior borders, raised medially before keel; keel long, narrow, terminating in a point between the mesocoxae; mesoventral lateral margins serrate in posterior half; humeri toothed (
<figureCitation id="130B2A32383CFFE02B65FC44FD04FC5C" box="[556,664,938,964]" captionStart="Fig" captionStartId="45.[189,232,1964,1990]" captionTargetBox="[192,1398,1087,1918]" captionTargetId="figure@45.[189,1398,1087,1922]" captionTargetPageId="45" captionText="Fig. 29. Cissidium ishigakiense Sawada, 2008. A. Habitus. B. Pronotum from slide preparation. C. Meso- and metaventrite from slide preparation." figureDoi="http://doi.org/10.5281/zenodo.3777342" httpUri="https://zenodo.org/record/3777342/files/figure.png" pageId="46" pageNumber="47">Fig. 29C</figureCitation>
).
</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FC05FD8AFBBF" blockId="46.[189,1398,1002,1063]" pageId="46" pageNumber="47">
METAVENTRITE. Sparsely pubescent, length
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, disc simple, width across spines
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; posterior margin of mesocoxae serrate.
</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FBA3FB57FBFF" blockId="46.[189,1227,1101,1127]" box="[189,1227,1101,1127]" pageId="46" pageNumber="47">
GENITALIA. Male aedeagus as
<figureCitation id="130B2A32383CFFE02B5CFBA3FDC4FBFF" box="[533,600,1101,1127]" captionStart="Fig" captionStartId="7.[189,232,1825,1851]" captionTargetBox="[202,1398,769,1790]" captionTargetId="figure@7.[189,1398,769,1799]" captionTargetPageId="7" captionText="Fig. 6. Figures to show genitalia types. – Spermathecae. A. Cissidium bilineatum sp. nov. B. Cissidium amboroense Darby, 2015. C. Cissidium sawadai Darby, 2015. D. Cissidium arcuatum sp. nov. E. Cissidium murphyi sp. nov. – Aedeagi. F. Cissidium petri Darby, 2015. G. Cissidium clareae sp. nov. H. Cissidium fraternum sp. nov. I. Cissidium tigrum Darby, 2015. J. Cissidium apterum sp. nov. K. Cissidium brachypterum sp. nov. L. Cissidium greensladei sp. nov. M. Cissidium kolombangaricum sp. nov. N. Cissidium mindanao sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3777294" httpUri="https://zenodo.org/record/3777294/files/figure.png" pageId="46" pageNumber="47">Fig. 6</figureCitation>
Fa–b but slightly more elongate. Females not known.
</paragraph>
</subSubSection>
<subSubSection id="C32A653C383CFFE029F4FB63FAEBFB6F" pageId="46" pageNumber="47" type="discussion">
<paragraph id="8B8F36B7383CFFE029F4FB63FEB3FB3F" blockId="46.[189,303,1165,1191]" box="[189,303,1165,1191]" pageId="46" pageNumber="47">
<heading id="D0C781DB383CFFE029F4FB63FEB3FB3F" bold="true" box="[189,303,1165,1191]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="B944EAA5383CFFE029F4FB63FEB3FB3F" bold="true" box="[189,303,1165,1191]" pageId="46" pageNumber="47">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F36B7383CFFE029F4FB54FAEBFB6F" blockId="46.[189,1399,1210,1271]" pageId="46" pageNumber="47">
This is the only Japanese species into this group apart from
<taxonomicName id="4C304D34383CFFE02AD5FB55FB6FFB4D" authority="Johnson, 2007" authorityName="Johnson" authorityYear="2007" box="[924,1267,1210,1237]" class="Insecta" family="Ptiliidae" genus="Cissidium" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="matthewsi">
<emphasis id="B944EAA5383CFFE02AD5FB55FBA5FB4D" box="[924,1081,1211,1237]" italics="true" pageId="46" pageNumber="47">C. matthewsi</emphasis>
<bibRefCitation id="EFA14B46383CFFE02D0AFB54FB6FFB4D" author="Johnson C." box="[1091,1267,1210,1237]" pageId="46" pageNumber="47" pagination="213 - 226" refId="ref59626" refString="Johnson C. 2007. Studies on Ptiliidae (Col.,) from the Solomon Islands, 1. Entomologist's Monthly Magazine 143: 213 - 226." type="journal article" year="2007">Johnson, 2007</bibRefCitation>
</taxonomicName>
. It may be distinguished from that species and all other Japanese species by the narrow almost parallel sided elytra.
</paragraph>
</subSubSection>
</treatment>
</document>
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