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<document id="DF32A9F88E5F3A1533C4956694460E00" ID-CLB-Dataset="20038" ID-DOI="10.5281/zenodo.5318986" ID-GBIF-Dataset="3b197df1-d7f8-4513-9a9b-e5c3654415e4" ID-ISSN="0374-1036" ID-Zenodo-Dep="5318986" ID-ZooBank="A159251E-8C8E-4509-8C0C-34FBD0A79279M" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630107039701" checkinUser="marcus" docAuthor="Löbl, Ivan" docDate="2015" docId="03998786FF845F21FE75FD960B4AFD48" docLanguage="en" docName="ActaEntMusNatPra.2015.55.1.129-144.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 55 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Scaphisoma punctatum" docType="treatment" docVersion="8" lastPageNumber="138" masterDocId="FFA0FFFEFF8C5F28FFCBFFED082FFFFB" masterDocTitle="Notes on Scaphisoma of Kalimantan (Coleoptera: Staphylinidae: Scaphidiinae)" masterLastPageNumber="144" masterPageNumber="129" pageNumber="137" updateTime="1699156701922" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="EB657D28550FD26D8E4E3060048CEF85">Notes on Scaphisoma of Kalimantan (Coleoptera: Staphylinidae: Scaphidiinae)</mods:title>
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<mods:namePart id="6CFB3A4CD3732388AC9C10AC65072458">Löbl, Ivan</mods:namePart>
<mods:affiliation id="DD1C4D3E43851FF371F306470A4E7EF3">Muséum dhistoire naturelle, Route de Malagnou 1, CH- 1208 Genève, Switzerland</mods:affiliation>
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<treatment id="03998786FF845F21FE75FD960B4AFD48" ID-DOI="http://doi.org/10.5281/zenodo.5332399" ID-GBIF-Taxon="194318107" ID-Zenodo-Dep="5332399" LSID="urn:lsid:plazi:treatment:03998786FF845F21FE75FD960B4AFD48" httpUri="http://treatment.plazi.org/id/03998786FF845F21FE75FD960B4AFD48" lastPageId="9" lastPageNumber="138" pageId="8" pageNumber="137">
<subSubSection id="C32A651BFF845F20FE75FD960B10FD68" box="[446,831,635,659]" pageId="8" pageNumber="137" type="nomenclature">
<paragraph id="8B8F3690FF845F20FE75FD960B10FD68" blockId="8.[446,831,635,659]" box="[446,831,635,659]" pageId="8" pageNumber="137">
<heading id="D0C781FCFF845F20FE75FD960B10FD68" bold="true" box="[446,831,635,659]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="137" reason="3">
<taxonomicName id="4C304D13FF845F20FE75FD960B10FD68" ID-CoL="79SVN" authority="(Pic, 1915)" baseAuthorityName="Pic" baseAuthorityYear="1915" box="[446,831,635,659]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="punctatum">
<emphasis id="B944EA82FF845F20FE75FD960B10FD68" bold="true" box="[446,831,635,659]" pageId="8" pageNumber="137">
<emphasis id="B944EA82FF845F20FE75FD960AEEFD68" bold="true" box="[446,705,635,659]" italics="true" pageId="8" pageNumber="137">Scaphisoma punctatum</emphasis>
(
<bibRefCitation id="EFA14B61FF845F20FD04FD910B17FD68" author="PIC M." box="[719,824,636,659]" pageId="8" pageNumber="137" pagination="30 - 32" refId="ref8700" refString="PIC M. 1915: Diagnoses de nouveaux genres et nouvelles especes de Scaphidiides. L'Echange, Revue Linneenne 31: 30 - 32." type="journal article" year="1915">Pic, 1915</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A651BFF845F20FF5CFD4C0A33FD4E" box="[151,540,673,693]" pageId="8" pageNumber="137" type="reference_group">
<paragraph id="8B8F3690FF845F20FF5CFD4C0A33FD4E" blockId="8.[151,540,673,693]" box="[151,540,673,693]" pageId="8" pageNumber="137">
<treatmentCitationGroup id="AB2011BEFF845F20FF5CFD4C0A33FD4E" box="[151,540,673,693]" pageId="8" pageNumber="137">
<treatmentCitation id="0A911081FF845F20FF5CFD4C0A37FD4E" author="PIC M." box="[151,536,673,693]" page="31" pageId="8" pageNumber="137" year="1915">
<taxonomicName id="4C304D13FF845F20FF5CFD4C0A37FD4E" ID-CoL="78GWH" authority="Pic, 1915: 31" authorityName="Pic" authorityPageNumber="31" authorityYear="1915" box="[151,536,673,693]" class="Insecta" family="Staphylinidae" genus="Pseudoscaphosoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="punctatum">
<emphasis id="B944EA82FF845F20FF5CFD4C09B0FD4F" box="[151,415,673,692]" italics="true" pageId="8" pageNumber="137">Pseudoscaphosoma punctatum</emphasis>
<bibRefCitation id="EFA14B61FF845F20FE6FFD4F0A37FD4E" author="PIC M." box="[420,536,674,693]" pageId="8" pageNumber="137" pagination="30 - 32" refId="ref8700" refString="PIC M. 1915: Diagnoses de nouveaux genres et nouvelles especes de Scaphidiides. L'Echange, Revue Linneenne 31: 30 - 32." type="journal article" year="1915">Pic, 1915: 31</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32A651BFF845F20FF5CFD210B88FC84" pageId="8" pageNumber="137" type="materials_examined">
<paragraph id="8B8F3690FF845F20FF5CFD210A6EFD24" blockId="8.[151,1127,716,895]" box="[151,577,716,735]" pageId="8" pageNumber="137">
<emphasis id="B944EA82FF845F20FF5CFD21093FFD24" bold="true" box="[151,272,716,735]" pageId="8" pageNumber="137">
<typeStatus id="548B8832FF845F20FF5CFD2108EBFD24" box="[151,196,716,735]" pageId="8" pageNumber="137">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3277600FF845F20FEDEFD210945FD24" box="[277,362,716,735]" name="Indonesia" pageId="8" pageNumber="137">Indonesia</collectingCountry>
,
<collectingRegion id="49F4F872FF845F20FEB8FD2109F6FD24" box="[371,473,716,735]" country="Indonesia" name="Kalimantan Utara" pageId="8" pageNumber="137">Kalimantan</collectingRegion>
, Martapura.
</paragraph>
<paragraph id="8B8F3690FF845F20FF5CFD0B0A4FFC9F" blockId="8.[151,1127,716,895]" pageId="8" pageNumber="137">
<emphasis id="B944EA82FF845F20FF5CFD0B0957FD02" bold="true" box="[151,376,742,761]" pageId="8" pageNumber="137">Type material examined.</emphasis>
<materialsCitation id="3B583CCDFF845F20FEB7FD0B098CFCB2" ID-GBIF-Occurrence="3744299311" collectionCode="MNHN" collectorName="Doherty" location="Martapura" municipality="Martapura" pageId="8" pageNumber="137" specimenCount="1" specimenCount-female="1" stateProvince="S. E. Borneo" typeStatus="lectotype">
<typeStatus id="548B8832FF845F20FEB7FD0B09FAFD03" box="[380,469,742,761]" pageId="8" pageNumber="137" type="lectotype">LECTOTYPE</typeStatus>
(by present designation):
<specimenCount id="9D36FD19FF845F20FD7AFD0D0AEEFD02" box="[689,705,736,761]" pageId="8" pageNumber="137" type="female"></specimenCount>
,
<collectingMunicipality id="6BEBACEAFF845F20FD05FD0B0B04FD02" box="[718,811,742,761]" pageId="8" pageNumber="137">Martapura</collectingMunicipality>
<collectingRegion id="49F4F872FF845F20FCFBFD0B0BBDFD02" box="[816,914,742,761]" pageId="8" pageNumber="137">S.E.Borneo</collectingRegion>
<collectorName id="26C55346FF845F20FC5CFD0B0BF2FD02" box="[919,989,742,761]" pageId="8" pageNumber="137">Doherty</collectorName>
1891 [printed] / Type [handwritten by Pic] / TYPE [red, printed] /
<taxonomicName id="4C304D13FF845F20FDF0FCEC0B70FCEF" authority="Pic" authorityName="Pic" authorityYear="1915" box="[571,863,769,788]" class="Insecta" family="Staphylinidae" genus="Pseudoscaphosoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="punctatum">Pseudoscaphosoma punctatum Pic</taxonomicName>
[handwritten by Pic] /
<taxonomicName id="4C304D13FF845F20FBEAFCEC08EBFCD4" class="Insecta" family="Staphylinidae" genus="Scaphosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="genus">Scaphosoma</taxonomicName>
probabl [handwritten by Pic] /
<taxonomicName id="4C304D13FF845F20FE03FCF10A94FCD4" authority="(Pic)" baseAuthorityName="Pic" baseAuthorityYear="1915" box="[456,699,796,815]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="punctatum">Scaphisoma punctatum (Pic)</taxonomicName>
det. Löbl 1976 / Lectotype
<taxonomicName id="4C304D13FF845F20FC6AFCF108EEFCB2" authority="(Pic)" baseAuthorityName="Pic" baseAuthorityYear="1915" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="punctatum">Scaphisoma punctatum (Pic)</taxonomicName>
des. Löbl 2014 (
<collectionCode id="ED21AE55FF845F20FE98FCDB09B6FCB2" box="[339,409,822,841]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="8" pageNumber="137">MNHN</collectionCode>
).
</materialsCitation>
The
<typeStatus id="548B8832FF845F20FE00FCDB0A36FCB2" box="[459,537,822,841]" pageId="8" pageNumber="137" type="lectotype">lectotype</typeStatus>
has the left antenna mounted on celluloid slide, the antennomeres
<collectionCode id="ED21AE55FF845F20FB82FCDB0C48FCB2" box="[1097,1127,822,841]" pageId="8" pageNumber="137">VII</collectionCode>
to XI of the right antenna are broken off and missing.
</paragraph>
<paragraph id="8B8F3690FF845F20FF5CFC810B88FC84" blockId="8.[151,1127,716,895]" box="[151,935,869,895]" pageId="8" pageNumber="137">
<emphasis id="B944EA82FF845F20FF5CFC810982FC84" bold="true" box="[151,429,876,895]" pageId="8" pageNumber="137">Additional material examined.</emphasis>
<materialsCitation id="3B583CCDFF845F20FE79FC810B88FC84" ID-GBIF-Occurrence="3744299309" box="[434,935,869,895]" collectingDate="1891" collectionCode="MHNG" collectorName="Doherty" location="Martapura" municipality="Martapura" pageId="8" pageNumber="137" specimenCount="1" specimenCount-female="1" stateProvince="S. E. Borneo" typeStatus="lectotype">
<specimenCount id="9D36FD19FF845F20FE79FC8109FDFC85" box="[434,466,869,895]" pageId="8" pageNumber="137" type="female">1 ♀</specimenCount>
, with the same locality data as the
<typeStatus id="548B8832FF845F20FD36FC810B64FC84" box="[765,843,876,895]" pageId="8" pageNumber="137" type="lectotype">lectotype</typeStatus>
(
<collectionCode id="ED21AE55FF845F20FC9CFC810BB2FC84" box="[855,925,876,895]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="137" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32A651BFF845F21FF5CFC7A0BB6FDE8" lastPageId="9" lastPageNumber="138" pageId="8" pageNumber="137" type="description">
<paragraph id="8B8F3690FF845F21FF5CFC7A0BB6FDE8" blockId="8.[151,1127,919,1582]" lastBlockId="9.[113,1089,220,691]" lastPageId="9" lastPageNumber="138" pageId="8" pageNumber="137">
<emphasis id="B944EA82FF845F20FF5CFC7A0912FC55" bold="true" box="[151,317,919,942]" pageId="8" pageNumber="137">Redescription.</emphasis>
Length
<quantity id="4CC89B75FF845F20FE5DFC7A09D7FC55" box="[406,504,919,942]" metricMagnitude="-3" metricUnit="m" metricValue="1.37" pageId="8" pageNumber="137" unit="mm" value="1.37">1.37 mm</quantity>
, width
<quantity id="4CC89B75FF845F20FD8CFC7A0A85FC55" box="[583,682,919,942]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="8" pageNumber="137" unit="mm" value="0.92">0.92 mm</quantity>
. Head and most of body very dark brown with somewhat reddish lustre, apical sixth of elytra, apical abdominal segments, tarsi and antennae yellowish or very light brown, femora and tibiae also lighter than most of body, but not as light as tarsi. Pronotum and elytra without microsculpture. Antennae with length/ width ratio of antennomeres as follows: III 12/5: IV 20/5: V 30/6: VI 30/7: VII 42/10: VIII 28/8: IX 41/10: X 42/10: XI 50/14. Pronotum with lateral margins rounded, lateral margin carinae not visible in dorsal view, discal punctation fine, dense, well delimited, near base distinctly coarser than on anterior half; pubescence indistinct. Apical part of scutellum exposed. Elytra with lateral margin carinae entirely exposed in dorsal view, apical margins slightly rounded, inner apical angle not prominent, situated posterior level of outer angles, sutural margin raised, sutural striae deep, not curved at base and not extended along basal margins, gradually converging apically, adsutural areas flat at sutural margin, combined
<quantity id="4CC89B75FF845F20FBF3FB1A08EFFAD5" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="8" pageNumber="137" unit="mm" value="0.18">0.18 mm</quantity>
wide at elytral base, each with two distinct puncture rows reaching mid-length, single puncture row in apical halves. Elytral punctation conspicuous, each elytron with 5 or 6 oblique puncture rows consisting of large and well delimited punctures, mostly much larger than puncture intervals. Inner puncture rows reaching apical third of disc, latter very finely and sparsely punctate. Humeral areas very finely punctate, areas between puncture rows and lateral margins impunctate. Inner puncture rows long, starting near base, outer puncture rows short, limited onto middle third of each elytron. Punctation on surfaces between anterior parts of inner puncture rows and sutural striae distinct, similar to that on basal part of pronotum. Hypomera not microsculptured, appearing impunctate. Mesepimeres as long as two thirds of interval to mesocoxae and 3 times as long as wide. Metaventrite weakly convex mesally, with very shallow apicolateral impressions, lacking microsculpture in middle and on sides, with strigulate microsculpture between mesocoxae and metacoxae, punctation mostly very fine and sparse, several large punctures in and near apicomedian impression, antecoxal puncture rows present, not impressed. Submesocoxal lines weakly convex, distinctly punctured, submesocoxal areas
<quantity id="4CC89B75FF855F21FE87FE910984FE68" box="[332,427,380,403]" metricMagnitude="-5" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="138" unit="mm" value="0.04">0.04 mm</quantity>
long, as third of interval to metacoxae. Metanepisterna flat, at inner margin impressed below level of metaventrite,
<quantity id="4CC89B75FF855F21FD6EFE710B2BFE48" box="[677,772,412,435]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="138" unit="mm" value="0.09">0.09 mm</quantity>
wide, weakly narrowed anteriad, inner suture almost straight, rounded at angles. Tibiae straight. Abdomen with strigulate microsculpture and very finely punctate. Ventrite 1 with submetaxocal areas
<quantity id="4CC89B75FF855F21FC6EFE310C2CFE08" box="[933,1027,476,499]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="138" unit="mm" value="0.08">0.08 mm</quantity>
long, strongly convex, about as two thirds of interval to apical margin of ventrite.
</paragraph>
</subSubSection>
<subSubSection id="C32A651BFF855F21FFBAFDF10A21FDC8" box="[113,526,540,563]" pageId="9" pageNumber="138" type="distribution">
<paragraph id="8B8F3690FF855F21FFBAFDF10A21FDC8" blockId="9.[113,1089,220,691]" box="[113,526,540,563]" pageId="9" pageNumber="138">
<emphasis id="B944EA82FF855F21FFBAFDF1092BFDC8" bold="true" box="[113,260,540,563]" pageId="9" pageNumber="138">Distribution.</emphasis>
<collectingCountry id="F3277600FF855F21FEC1FDF1095DFDC8" box="[266,370,540,563]" name="Indonesia" pageId="9" pageNumber="138">Indonesia</collectingCountry>
(
<collectingRegion id="49F4F872FF855F21FE4BFDF10A2CFDC8" box="[384,515,540,563]" country="Indonesia" name="Kalimantan Utara" pageId="9" pageNumber="138">Kalimantan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C32A651BFF855F21FFBAFDD10B4AFD48" pageId="9" pageNumber="138" type="discussion">
<paragraph id="8B8F3690FF855F21FFBAFDD10B4AFD48" blockId="9.[113,1089,220,691]" pageId="9" pageNumber="138">
<emphasis id="B944EA82FF855F21FFBAFDD108DBFDA8" bold="true" box="[113,244,572,595]" pageId="9" pageNumber="138">Comments.</emphasis>
This species is absent from new collections and its male characters remain unknown. The shape of the sutural striae of the elytra suggest possible relationships with members of the
<taxonomicName id="4C304D13FF855F21FED1FD9109F0FD68" authorityName="Reitter" authorityYear="1877" box="[282,479,635,659]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="haemorrhoidale">
<emphasis id="B944EA82FF855F21FED1FD910908FD68" box="[282,295,636,659]" italics="true" pageId="9" pageNumber="138">S</emphasis>
.
<emphasis id="B944EA82FF855F21FEFFFD9609F0FD68" box="[308,479,635,659]" italics="true" pageId="9" pageNumber="138">haemorrhoidale</emphasis>
</taxonomicName>
or
<taxonomicName id="4C304D13FF855F21FDCBFD910A4DFD68" authorityName="Motschulsky" authorityYear="1863" box="[512,610,636,659]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="pictum">
<emphasis id="B944EA82FF855F21FDCBFD910A22FD68" box="[512,525,636,659]" italics="true" pageId="9" pageNumber="138">S</emphasis>
.
<emphasis id="B944EA82FF855F21FDD2FD910A4DFD68" box="[537,610,636,659]" italics="true" pageId="9" pageNumber="138">pictum</emphasis>
</taxonomicName>
groups, the conspicuous pattern of the elytral punctation suggest relationships with members of the
<taxonomicName id="4C304D13FF855F21FD7AFD710B36FD48" authorityName="Pic" authorityYear="1916" box="[689,793,668,691]" class="Insecta" family="Staphylinidae" genus="Scaphisoma" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="138" phylum="Arthropoda" rank="species" species="rouyeri">
<emphasis id="B944EA82FF855F21FD7AFD710A91FD48" box="[689,702,668,691]" italics="true" pageId="9" pageNumber="138">S</emphasis>
.
<emphasis id="B944EA82FF855F21FD00FD710B36FD48" box="[715,793,668,691]" italics="true" pageId="9" pageNumber="138">rouyeri</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
</treatment>
</document>