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<document id="C0BC96E642AAC0CD338CCF1B6D53DD62" ID-DOI="10.5852/ejt.2018.396" ID-GBIF-Dataset="2b95a851-e1bf-4bdd-9cce-441ddc4135d2" ID-ISSN="2118-9773" ID-Zenodo-Dep="3787049" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1588696404769" checkinUser="valdenar" docAuthor="Camfield, Rebecca &amp; Hughes, Mark" docDate="2018" docId="0397A96F2F74FFCA004EF95CE3A12B51" docLanguage="en" docName="ejt-396_camfield_hughes.pdf.imf" docOrigin="European Journal of Taxonomy 396" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Begonia (sect. Sphenanthera) burkillii Dunn (Dunn 1920" docType="treatment" docVersion="14" lastPageNumber="30" masterDocId="FFAED1172F6DFFD7025FFFB1E00A2E5C" masterDocTitle="A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India" masterLastPageNumber="116" masterPageNumber="1" pageNumber="26" updateTime="1737391932328" updateUser="diego">
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<mods:title id="3DFFF4EED333516E6136E8F07AFE5CA5">A revision and one new species of Begonia L. (Begoniaceae, Cucurbitales) in Northeast India</mods:title>
</mods:titleInfo>
<mods:name id="A67E88D4FC2CD4841D7C60044493FE6A" type="personal">
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<mods:namePart id="06B452FD1806B72E032240CE2FDCD145">Camfield, Rebecca</mods:namePart>
<mods:affiliation id="95AB423161ACF02C25F258982E3B19C9">Royal Botanic Gardens Edinburgh, 20 a Inverleith Row, Edinburgh, EH 5 3 LR, United Kingdom.</mods:affiliation>
<mods:nameIdentifier id="08A68A041BF72800E2979CF9C9BD553B" type="email">RCamfield@rbge.org.uk</mods:nameIdentifier>
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<mods:namePart id="41965A2CABAFFCEA824523FC0701DCCC">Hughes, Mark</mods:namePart>
<mods:affiliation id="06A1E2D2B18F13EBC0FAC8525EDC6C6D">Royal Botanic Gardens Edinburgh, 20 a Inverleith Row, Edinburgh, EH 5 3 LR, United Kingdom.</mods:affiliation>
<mods:nameIdentifier id="943D3990C0B0CCAB8F5ACFE9C8809AD8" type="email">MHughes@rbge.org.uk</mods:nameIdentifier>
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<mods:title id="D4F3EADD5DB9E54259CBA9B8F4FE1A76">European Journal of Taxonomy</mods:title>
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<mods:date id="A383C3201FEE995675F5BE82EDB5EFA3">2018</mods:date>
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<mods:number id="D97DFCB601E0CEF5823324380C332129">2018-01-19</mods:number>
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<mods:number id="6CA8DBBC65414D1E80470E86EDCC92BB">396</mods:number>
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<mods:identifier id="C7E7A1E9F908F5A282E1C9C8901E6D8A" type="DOI">10.5852/ejt.2018.396</mods:identifier>
<mods:identifier id="443720B6E18C431519FB4C627432E8D7" type="GBIF-Dataset">2b95a851-e1bf-4bdd-9cce-441ddc4135d2</mods:identifier>
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<treatment id="0397A96F2F74FFCA004EF95CE3A12B51" ID-DOI="http://doi.org/10.5281/zenodo.3794337" ID-GBIF-Taxon="163963840" ID-Zenodo-Dep="3794337" LSID="urn:lsid:plazi:treatment:0397A96F2F74FFCA004EF95CE3A12B51" httpUri="http://treatment.plazi.org/id/0397A96F2F74FFCA004EF95CE3A12B51" lastPageId="29" lastPageNumber="30" pageId="25" pageNumber="26" scope_class="Equisetopsida" scope_family="Begoniaceae" scope_order="Cucurbitales" scope_subClass="Magnoliidae" scope_superOrder="Rosanae">
<subSubSection id="C3244BF22F74FFCE004EF95CE4282954" box="[529,1058,1773,1800]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="8B8118792F74FFCE004EF95CE4282954" blockId="25.[529,1058,1773,1835]" box="[529,1058,1773,1800]" pageId="25" pageNumber="26">
<heading id="D0C9AF152F74FFCE004EF95CE4282954" box="[529,1058,1773,1800]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="4C3E63FA2F74FFCE004EF95CE3222954" ID-CoL="87Y9Z" authority="Dunn" authorityName="Dunn (Dunn" authorityYear="1920" box="[529,808,1773,1800]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="species" section="Sphenanthera" species="burkillii">
<emphasis id="B94AC46B2F74FFCE004EF95CE2EA2954" bold="true" box="[529,736,1773,1800]" italics="true" pageId="25" pageNumber="26">Begonia burkillii</emphasis>
Dunn
</taxonomicName>
[sect.
<taxonomicName id="4C3E63FA2F74FFCE012AF95CE412295B" box="[885,1048,1773,1799]" class="Magnoliopsida" family="Begoniaceae" genus="Sphenanthera" higherTaxonomySource="CoL" kingdom="Plantae" order="Cucurbitales" pageId="25" pageNumber="26" phylum="Tracheophyta" rank="section">
<emphasis id="B94AC46B2F74FFCE012AF95CE412295B" box="[885,1048,1773,1799]" italics="true" pageId="25" pageNumber="26">Sphenanthera</emphasis>
</taxonomicName>
]
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F74FFCE0088F8A0E3562977" box="[727,860,1808,1835]" pageId="25" pageNumber="26" type="description">
<paragraph id="8B8118792F74FFCE0088F8A0E3562977" blockId="25.[529,1058,1773,1835]" box="[727,860,1808,1835]" pageId="25" pageNumber="26">
<figureCitation id="130504FC2F74FFCE0088F8A0E3562977" box="[727,860,1808,1835]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="26.[189,232,1994,2020]" captionStartId-1="27.[189,232,1958,1984]" captionStartId-2="28.[189,232,1924,1950]" captionTargetBox-1="[415,1173,265,1921]" captionTargetBox-2="[195,1391,269,1895]" captionTargetId-0="figure@26.[416,1172,1215,1965]" captionTargetId-1="figure@27.[415,1173,265,1921]" captionTargetId-2="figure@28.[189,1398,265,1898]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionTargetPageId-2="28" captionText-0="Fig. 16. Map showing the location of B. burkillii Dunn specimens." captionText-1="Fig. 17. Begonia burkillii Dunn. A. Leaf pattern. B. Male flowers. Photograph A courtesy of Aaron Matsumoto and photograph B courtesy of Earl I-Lan of plants in cultivation in private collections." captionText-2="Fig. 18. Illustration of B. burkillii Dunn. A. Habit of plant. B. Style. C. Female flower. D. Stamen. E. Male flower. F. Male inflorescence. Drawn by Heather Raeburn. A–C from Burkill 36720; D from Burkill 37455; E–F from Burkill 37139; all K." figureDoi-0="http://doi.org/10.5281/zenodo.14616265" figureDoi-1="http://doi.org/10.5281/zenodo.3787075" figureDoi-2="http://doi.org/10.5281/zenodo.3787077" httpUri-0="https://zenodo.org/record/14616265/files/figure.png" httpUri-1="https://zenodo.org/record/3787075/files/figure.png" httpUri-2="https://zenodo.org/record/3787077/files/figure.png" pageId="25" pageNumber="26">Figs 16–18</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F74FFCE02E2F8E3E3EA2930" box="[189,992,1874,1900]" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="8B8118792F74FFCE02E2F8E3E3EA2930" blockId="25.[189,1398,1874,1936]" box="[189,992,1874,1900]" pageId="25" pageNumber="26">
<emphasis id="B94AC46B2F74FFCE02E2F8E3E2B72930" box="[189,701,1874,1900]" italics="true" pageId="25" pageNumber="26">Bulletin of Miscellaneous Information, Kew</emphasis>
<treatmentCitation id="0A9F3E682F74FFCE009AF8E3E3D62930" box="[709,988,1874,1900]" pageId="25" pageNumber="26">
1920: 110 (
<bibRefCitation id="EFAF65882F74FFCE0115F8E3E3D92930" author="Dunn S. T." box="[842,979,1874,1900]" pageId="25" pageNumber="26" pagination="108 - 111" refId="ref53600" refString="Dunn S. T. 1920. Decades Kewensis: Plantarum Novarum in Herbario Horti Regii Conservatarum. Decas XCVI. Bulletin of Miscellaneous Information, Kew 1920: 108 - 111. Available from http: // biodiversitylibrary. org / page / 11269649 [accessed May 2009]." type="journal article" year="1920">Dunn 1920</bibRefCitation>
)
</treatmentCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F74FFCE01B7F8E2E57C29CC" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="8B8118792F74FFCE01B7F8E2E57C29CC" blockId="25.[189,1398,1874,1936]" pageId="25" pageNumber="26">
–
<typeStatus id="5485A6DB2F74FFCE01A1F8E2E4362931" box="[1022,1084,1875,1901]" pageId="25" pageNumber="26">Type</typeStatus>
:
<materialsCitation id="3B5612242F74FFCE0617F8E3E4A229D3" collectingDate="1911-11" collectingDateMax="1912-03" collectingDateMin="1911-11" collectionCode="K" collectorName="Burkill" country="India" location="Janak river" municipality="Arunachal-Pradesh" pageId="25" pageNumber="26" specimenCode="K000634633" specimenCount="1" typeStatus="lectotype">
<collectingCountry id="F32958E92F74FFCE0617F8E3E48C2930" box="[1096,1158,1874,1900]" name="India" pageId="25" pageNumber="26">India</collectingCountry>
,
<collectingMunicipality id="6BE582032F74FFCE06CEF8E3E5782930" box="[1169,1394,1874,1900]" pageId="25" pageNumber="26">Arunachal-Pradesh</collectingMunicipality>
,
<location id="8EE14EA22F74FFCE02E2F8C4E13729D3" LSID="urn:lsid:plazi:treatment:0397A96F2F74FFCA004EF95CE3A12B51:8EE14EA22F74FFCE02E2F8C4E13729D3" box="[189,317,1909,1935]" country="India" municipality="Arunachal-Pradesh" name="Janak river" pageId="25" pageNumber="26">Janak river</location>
,
<date id="FF803EB92F74FFCE0318F8C7E1B629D3" box="[327,444,1909,1936]" pageId="25" pageNumber="26" value="1911-11">
<collectingDate id="EFC4C7512F74FFCE0318F8C7E1B629D3" box="[327,444,1909,1936]" pageId="25" pageNumber="26" value="1911-11">Nov. 1911</collectingDate>
</date>
–
<date id="FF803EB92F74FFCE0391F8C7E24C29D3" box="[462,582,1909,1936]" pageId="25" pageNumber="26" value="1912-03">
<collectingDate id="EFC4C7512F74FFCE0391F8C7E24C29D3" box="[462,582,1909,1936]" pageId="25" pageNumber="26" value="1912-03">Mar. 1912</collectingDate>
</date>
,
<emphasis id="B94AC46B2F74FFCE000FF8C4E2E429D3" box="[592,750,1909,1935]" italics="true" pageId="25" pageNumber="26">
<collectorName id="26CB7DAF2F74FFCE000FF8C4E2AA29D3" box="[592,672,1909,1935]" pageId="25" pageNumber="26">Burkill</collectorName>
37139
</emphasis>
(
<typeStatus id="5485A6DB2F74FFCE00A2F8C4E34E29D3" box="[765,836,1909,1935]" pageId="25" pageNumber="26">lecto-</typeStatus>
:
<specimenCode id="DB98B0022F74FFCE0112F8C4E3EF29D3" box="[845,997,1909,1935]" pageId="25" pageNumber="26">K000634633</specimenCode>
, here designated
</materialsCitation>
;
<materialsCitation id="3B5612242F74FFCE06EDF8C4E57829CC" box="[1202,1394,1909,1936]" collectingDate="1911-11" collectingDateMax="1912-03" collectingDateMin="1911-11" collectionCode="B" collectorName="Burkill" country="India" location="Janak river" municipality="Arunachal-Pradesh" pageId="25" pageNumber="26" specimenCount="1" typeStatus="isolectotype">isolecto-: B n.v.)</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F77FFCD02E1FEBAE2302F79" box="[190,570,267,294]" pageId="26" pageNumber="27" type="multiple">
<paragraph id="8B8118792F77FFCD02E1FEBAE2302F79" blockId="26.[190,570,267,294]" box="[190,570,267,294]" pageId="26" pageNumber="27">
<heading id="D0C9AF152F77FFCD02E1FEBAE2302F79" bold="true" box="[190,570,267,294]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B94AC46B2F77FFCD02E1FEBAE2302F79" bold="true" box="[190,570,267,294]" pageId="26" pageNumber="27">Citations in other publications</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F77FFCD02E1FE8BE5392F2B" pageId="26" pageNumber="27" type="reference_group">
<paragraph id="8B8118792F77FFCD02E1FE8BE5392F2B" blockId="26.[190,1399,314,376]" pageId="26" pageNumber="27">
<treatmentCitationGroup id="AB2E3F572F77FFCD02E1FE8BE5392F2B" pageId="26" pageNumber="27">
<treatmentCitation id="0A9F3E682F77FFCD02E1FE8BE1A72F08" box="[190,429,314,340]" pageId="26" pageNumber="27">
<bibRefCitation id="EFAF65882F77FFCD02E1FE8BE1A72F08" author="Burkill I. H." box="[190,429,314,340]" pageId="26" pageNumber="27" pagination="1 - 420" refId="ref52793" refString="Burkill I. H. 1924. Botany of the Abor Expedition. Records of the Botanical Survey of India 10: 1 - 420." type="journal article" year="1924">Burkill (1924: 288)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F77FFCD03E3FE8BE38D2F08" box="[444,903,314,341]" pageId="26" pageNumber="27">
<bibRefCitation id="EFAF65882F77FFCD03E3FE8BE2902F09" author="Tebbitt M. C." box="[444,666,314,341]" pageId="26" pageNumber="27" pagination="1 - 9" refId="ref57598" refString="Tebbitt M. C. 2003 b. Notes on South Asian Begonia. Edinburgh Journal of Botan y 60: 1 - 9. https: // doi. org / 10.1017 / S 0960428603000015" type="journal article" year="2003">Tebbitt (2003b: 7)</bibRefCitation>
, Morris (2006: 88)
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F77FFCD01C8FE8BE4762F09" box="[919,1148,314,341]" pageId="26" pageNumber="27">
<bibRefCitation id="EFAF65882F77FFCD01C8FE8BE4762F09" author="Uddin A." box="[919,1148,314,341]" pageId="26" pageNumber="27" pagination="591 - 597" refId="ref57753" refString="Uddin A. 2007. Distribution and status of India Begonia species. Journal of Economic and Taxonomic Botany 31 (3): 591 - 597." type="journal article" year="2007">Uddin (2007: 593)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F77FFCD06D3FE8BE5782F08" box="[1164,1394,314,340]" pageId="26" pageNumber="27">
<bibRefCitation id="EFAF65882F77FFCD06D3FE8BE5782F08" author="Hughes M." box="[1164,1394,314,340]" pageId="26" pageNumber="27" refId="ref54915" refString="Hughes M. 2008. An annotated checklist of Southeast Asian Begonia. Royal Botanic Garden Edinburgh." type="book" year="2008">Hughes (2008: 22)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F77FFCD02E1FEECE1BA2F2B" box="[190,432,349,375]" pageId="26" pageNumber="27">
<bibRefCitation id="EFAF65882F77FFCD02E1FEECE1BA2F2B" author="Morris R." box="[190,432,349,375]" pageId="26" pageNumber="27" pagination="173" refId="ref56479" refString="Morris R. 2009 b. Announcement of 2009 - 2010 trip to India to document Begonias. The Begonian 76: 173." type="journal article" year="2009">Morris (2009b: 173)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F77FFCD03E3FEECE2772F2B" box="[444,637,349,375]" pageId="26" pageNumber="27">
<bibRefCitation id="EFAF65882F77FFCD03E3FEECE2772F2B" author="Dash S. S." box="[444,637,349,375]" pageId="26" pageNumber="27" pagination="29 - 43" refId="ref53314" refString="Dash S. S. 2010. Native species of Begonia L. (Begoniaceae) in Arunachal-Pradesh-diversity and distribution. Bulletin of Arunachal Forest Research 26: 29 - 43." type="journal article" year="2010">Dash (2010: 32)</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F77FFCD00D6FEECE3952F24" box="[649,927,349,376]" pageId="26" pageNumber="27">
Saika
<emphasis id="B94AC46B2F77FFCD008DFEEFE3072F2B" box="[722,781,349,375]" italics="true" pageId="26" pageNumber="27">et al.</emphasis>
(2011: 357)
</treatmentCitation>
,
<treatmentCitation id="0A9F3E682F77FFCD01F4FEECE5252F2B" box="[939,1327,349,375]" pageId="26" pageNumber="27">Baruah &amp; Choudbury (2014: 41)</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F77FFCD02E1FE2FE1702FE4" box="[190,378,414,440]" pageId="26" pageNumber="27" type="multiple">
<paragraph id="8B8118792F77FFCD02E1FE2FE1702FE4" blockId="26.[190,378,414,440]" box="[190,378,414,440]" pageId="26" pageNumber="27">
<heading id="D0C9AF152F77FFCD02E1FE2FE1702FE4" bold="true" box="[190,378,414,440]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B94AC46B2F77FFCD02E1FE2FE1702FE4" bold="true" box="[190,378,414,440]" pageId="26" pageNumber="27">Other material</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F77FFCD02E1FE7CE16A2D37" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8B8118792F77FFCD02E1FE7CE16A2D37" blockId="26.[190,1399,461,875]" pageId="26" pageNumber="27">
<materialsCitation id="3B5612242F77FFCD02E1FE7CE0FF2C57" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
<collectingCountry id="F32958E92F77FFCD02E1FE7CE11C2FBB" box="[190,278,461,487]" name="India" pageId="26" pageNumber="27">INDIA</collectingCountry>
:
<emphasis id="B94AC46B2F77FFCD0378FE7CE2172FBB" bold="true" box="[295,541,461,487]" pageId="26" pageNumber="27">
<collectingRegion id="49FAD69B2F77FFCD0378FE7CE2172FBB" box="[295,541,461,487]" pageId="26" pageNumber="27">Arunachal-Pradesh</collectingRegion>
</emphasis>
: Abor Hills, Jauamukh,
<date id="FF803EB92F77FFCD0108FE7FE3DF2FBB" box="[855,981,461,488]" pageId="26" pageNumber="27" value="1911-11">Nov. 1911</date>
–
<date id="FF803EB92F77FFCD01B7FE7FE4632FBB" box="[1000,1129,461,488]" pageId="26" pageNumber="27" value="1912-03">Mar. 1912</date>
,
<emphasis id="B94AC46B2F77FFCD0623FE7CE52F2FBB" box="[1148,1317,461,487]" italics="true" pageId="26" pageNumber="27">Burkill 37121</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD0764FE7CE57D2FBB" box="[1339,1399,461,487]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="26" pageNumber="27" type="Herbarium">CAL</collectionCode>
n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0359FE40E4AB2C57" ID-GBIF-Occurrence="2610421301" box="[262,1185,496,523]" pageId="26" pageNumber="27">
Bapu Mountain, Renging Camp,
<date id="FF803EB92F77FFCD00FFFE40E3172C57" box="[672,797,497,523]" pageId="26" pageNumber="27" value="1911-11">Nov. 1911</date>
–
<date id="FF803EB92F77FFCD016FFE40E3BA2C57" box="[816,944,497,523]" pageId="26" pageNumber="27" value="1912-03">Mar. 1912</date>
,
<emphasis id="B94AC46B2F77FFCD019DFE41E4622C56" box="[962,1128,496,522]" italics="true" pageId="26" pageNumber="27">Burkill 36720</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD0620FE40E4922C57" box="[1151,1176,497,523]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD06ECFE40E2542C72" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
Bapu Mountain, Rotung,
<emphasis id="B94AC46B2F77FFCD0378FDA5E1C72C72" box="[295,461,532,558]" italics="true" pageId="26" pageNumber="27">Burkill 37706</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD03BFFDA5E2162C72" box="[480,540,532,558]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="26" pageNumber="27" type="Herbarium">CAL</collectionCode>
n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0031FDA5E0F42C0D" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
Bapu Mountain, Talauma,
<date id="FF803EB92F77FFCD01ECFDA5E4242C72" box="[947,1070,532,558]" pageId="26" pageNumber="27" value="1911-11">Nov. 1911</date>
–
<date id="FF803EB92F77FFCD061EFDA5E4CA2C72" box="[1089,1216,532,558]" pageId="26" pageNumber="27" value="1912-03">Mar. 1912</date>
,
<emphasis id="B94AC46B2F77FFCD068FFDA5E57C2C72" box="[1232,1398,532,558]" italics="true" pageId="26" pageNumber="27">Burkill 36910</emphasis>
(n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0352FD86E1302C28" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
Changlang, Namdapha Biosphere Reserve, Anamica Falls,
<date id="FF803EB92F77FFCD0192FD86E4782C0D" box="[973,1138,567,593]" pageId="26" pageNumber="27" value="2004-11-30">30 Nov. 2004</date>
,
<emphasis id="B94AC46B2F77FFCD0620FD86E57C2C0D" box="[1151,1398,567,593]" italics="true" pageId="26" pageNumber="27">Amad Uddin 107934</emphasis>
(ASSAM)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0319FDEBE3852C29" ID-GBIF-Occurrence="2610421301" box="[326,911,602,629]" pageId="26" pageNumber="27">
Dihang Valley,
<date id="FF803EB92F77FFCD03A5FDEBE29F2C28" box="[506,661,602,628]" pageId="26" pageNumber="27" value="1928-02-12">12 Feb. 1928</date>
,
<emphasis id="B94AC46B2F77FFCD00FEFDEBE32B2C28" box="[673,801,602,628]" italics="true" pageId="26" pageNumber="27">Ward 7822</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD016FFDEBE3682C28" box="[816,866,602,628]" country="United Kingdom" name="Bristol Museum" pageId="26" pageNumber="27">BM</collectionCode>
,
<collectionCode id="ED2F80BC2F77FFCD0132FDEAE38C2C29" box="[877,902,603,629]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD01C4FDEAE1A52CCB" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
Igar Valley,
<date id="FF803EB92F77FFCD0677FDEBE4CF2C28" box="[1064,1221,602,629]" pageId="26" pageNumber="27" value="1911-11-29">29 Nov. 1911</date>
–
<date id="FF803EB92F77FFCD0689FDEBE57B2C28" box="[1238,1393,602,629]" pageId="26" pageNumber="27" value="1912-03-11">11 Mar. 1912</date>
,
<emphasis id="B94AC46B2F77FFCD02E1FDCCE1692CCB" box="[190,355,637,663]" italics="true" pageId="26" pageNumber="27">Burkill 16121</emphasis>
(n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD03E0FDCCE4442CCB" ID-GBIF-Occurrence="2610421301" box="[447,1102,637,664]" pageId="26" pageNumber="27">
Janak river,
<date id="FF803EB92F77FFCD000AFDCFE2DA2CCB" box="[597,720,637,664]" pageId="26" pageNumber="27" value="1911-11">Nov. 1911</date>
–
<date id="FF803EB92F77FFCD00BCFDCFE3682CCB" box="[739,866,637,664]" pageId="26" pageNumber="27" value="1912-03">Mar. 1912</date>
,
<emphasis id="B94AC46B2F77FFCD012DFDCCE41D2CCB" box="[882,1047,637,663]" italics="true" pageId="26" pageNumber="27">Burkill 37455</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD0673FDCCE44F2CCB" box="[1068,1093,637,663]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0601FDCCE1572CE7" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
Naharlagun,
<emphasis id="B94AC46B2F77FFCD06A6FDCCE57C2CCB" box="[1273,1398,637,663]" italics="true" pageId="26" pageNumber="27">Pal 70320</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD0299FD10E1112CE7" box="[198,283,673,699]" country="India" httpUri="http://grbio.org/cool/1csn-k189" name="Botanical Survey of India, Arunachal Pradesh Regional Centre" pageId="26" pageNumber="27" type="Herbarium">ARUN</collectionCode>
n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0331FD10E3F72CE7" ID-GBIF-Occurrence="2610421301" box="[366,1021,672,699]" pageId="26" pageNumber="27">
Papum Pare,
<date id="FF803EB92F77FFCD004EFD10E2AC2CE7" box="[529,678,672,699]" pageId="26" pageNumber="27" value="2005-04-03">3 Apr. 2005</date>
,
<emphasis id="B94AC46B2F77FFCD00E9FD10E34E2CE7" box="[694,836,673,699]" italics="true" pageId="26" pageNumber="27">Morris AR2</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD0107FD10E3B62CE7" box="[856,956,673,699]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12826" lsid="urn:lsid:biocol.org:col:12826" name="Clemson University" pageId="26" pageNumber="27" type="Herbarium">CLEMS</collectionCode>
n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0651FD11E1262C82" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
Pasighat,
<date id="FF803EB92F77FFCD06DBFD10E5232CE7" box="[1156,1321,672,699]" pageId="26" pageNumber="27" value="1928-02-10">10 Feb. 1928</date>
,
<emphasis id="B94AC46B2F77FFCD0766FD11E0F32C82" italics="true" pageId="26" pageNumber="27">Ward 7822</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD0355FD75E1292C82" box="[266,291,708,734]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="26" pageNumber="27" type="Herbarium">K</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0365FD75E3A12C82" ID-GBIF-Occurrence="2610421301" box="[314,939,708,734]" pageId="26" pageNumber="27">
Ponging,
<date id="FF803EB92F77FFCD03F4FD75E2292C82" box="[427,547,708,734]" pageId="26" pageNumber="27" value="1911-11">Nov. 1911</date>
–
<date id="FF803EB92F77FFCD0069FD75E2B82C82" box="[566,690,708,734]" pageId="26" pageNumber="27" value="1912-03">Mar. 1912</date>
,
<emphasis id="B94AC46B2F77FFCD00E0FD75E3682C82" box="[703,866,708,734]" italics="true" pageId="26" pageNumber="27">Burkill 37375</emphasis>
(n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD01E6FD75E1562D5D" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
Sangram to Panyu River,
<emphasis id="B94AC46B2F77FFCD06B6FD75E57C2C82" box="[1257,1398,708,734]" italics="true" pageId="26" pageNumber="27">Dash 32511</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD0299FD56E1112D5D" box="[198,283,743,769]" country="India" httpUri="http://grbio.org/cool/1csn-k189" name="Botanical Survey of India, Arunachal Pradesh Regional Centre" pageId="26" pageNumber="27" type="Herbarium">ARUN</collectionCode>
n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0333FD56E4222D5D" ID-GBIF-Occurrence="2610421301" box="[364,1064,743,769]" pageId="26" pageNumber="27">
Serpo stream,
<date id="FF803EB92F77FFCD0043FD56E29C2D5D" box="[540,662,743,769]" pageId="26" pageNumber="27" value="1911-11">Nov. 1911</date>
–
<date id="FF803EB92F77FFCD00F6FD56E3222D5D" box="[681,808,743,769]" pageId="26" pageNumber="27" value="1912-03">Mar. 1912</date>
,
<emphasis id="B94AC46B2F77FFCD0168FD56E3D62D5D" box="[823,988,743,769]" italics="true" pageId="26" pageNumber="27">Burkill 36315</emphasis>
(n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0667FD56E0F42D78" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
Shile River,
<emphasis id="B94AC46B2F77FFCD068EFD56E57C2D5D" box="[1233,1398,743,769]" italics="true" pageId="26" pageNumber="27">Burkill 37121</emphasis>
(n.v.)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0350FCBBE3C12D79" ID-GBIF-Occurrence="2610421301" box="[271,971,778,805]" pageId="26" pageNumber="27">
Subansiri District,
<date id="FF803EB92F77FFCD03ADFCBBE28C2D79" box="[498,646,778,805]" pageId="26" pageNumber="27" value="2005-04-08">8 Apr. 2005</date>
,
<emphasis id="B94AC46B2F77FFCD00CAFCBAE3042D78" box="[661,782,778,804]" italics="true" pageId="26" pageNumber="27">Morris 12</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD017DFCBBE38C2D78" box="[802,902,778,804]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12826" lsid="urn:lsid:biocol.org:col:12826" name="Clemson University" pageId="26" pageNumber="27" type="Herbarium">CLEMS</collectionCode>
n.v.).
</materialsCitation>
<materialsCitation id="3B5612242F77FFCD0189FCBBE2232D1B" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
<emphasis id="B94AC46B2F77FFCD0189FCBBE4232D78" bold="true" box="[982,1065,778,804]" pageId="26" pageNumber="27">
<collectingRegion id="49FAD69B2F77FFCD0189FCBBE4232D78" box="[982,1065,778,804]" country="India" name="Assam" pageId="26" pageNumber="27">Assam</collectingRegion>
</emphasis>
: Cacchar, Sonapur,
<date id="FF803EB92F77FFCD077FFCBBE0F02D1B" pageId="26" pageNumber="27" value="2011-01-13">13 Jan. 2011</date>
,
<emphasis id="B94AC46B2F77FFCD0357FC9FE1BC2D1B" box="[264,438,813,839]" italics="true" pageId="26" pageNumber="27">Himu Roy 143</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD0397FC9CE22B2D1B" box="[456,545,813,839]" country="India" httpUri="http://grbio.org/cool/7wsv-m0m0" name="Gauhati University" pageId="26" pageNumber="27" type="Herbarium">GUBH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5612242F77FFCD0067FC9CE16A2D37" ID-GBIF-Occurrence="2610421301" pageId="26" pageNumber="27">
Dhemaji, Silapathar, Poba Reserve Forest,
<date id="FF803EB92F77FFCD0665FC9CE4DD2D1B" box="[1082,1239,813,840]" pageId="26" pageNumber="27" value="2010-01-13">13 Jan. 2010</date>
,
<emphasis id="B94AC46B2F77FFCD06BBFC9FE0E02D37" italics="true" pageId="26" pageNumber="27">Ranjit Saika 300</emphasis>
(
<collectionCode id="ED2F80BC2F77FFCD02A5FCE0E15E2D37" box="[250,340,849,875]" country="India" httpUri="http://grbio.org/cool/7wsv-m0m0" name="Gauhati University" pageId="26" pageNumber="27" type="Herbarium">GUBH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F77FFCA02E1FC23E2102CD9" lastPageId="29" lastPageNumber="30" pageId="26" pageNumber="27" type="description">
<paragraph id="8B8118792F77FFCD02E1FC23E1452DF0" blockId="26.[190,335,914,940]" box="[190,335,914,940]" pageId="26" pageNumber="27">
<heading id="D0C9AF152F77FFCD02E1FC23E1452DF0" bold="true" box="[190,335,914,940]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="B94AC46B2F77FFCD02E1FC23E1452DF0" bold="true" box="[190,335,914,940]" pageId="26" pageNumber="27">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F77FFCA02E1FC71E2102CD9" blockId="26.[190,1399,960,1163]" lastBlockId="29.[189,1399,267,646]" lastPageId="29" lastPageNumber="30" pageId="26" pageNumber="27">
Rhizomatous, dioecious herb,
<quantity id="4CC6B59C2F77FFCD007FFC70E2902D87" box="[544,666,961,987]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="26" pageNumber="27" unit="cm" value="30.0" valueMax="40.0" valueMin="20.0">20–40 cm</quantity>
high. Rhizome: ca
<quantity id="4CC6B59C2F77FFCD0122FC70E3C52D87" box="[893,975,961,987]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="mm" value="10.0">10 mm</quantity>
wide, glabrous, internodes
<quantity id="4CC6B59C2F77FFCD074FFC70E57D2D87" box="[1296,1399,960,987]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="26" pageNumber="27" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 mm</quantity>
long. Stipules: lanceolate, 5–17 ×
<quantity id="4CC6B59C2F77FFCD0005FC55E2C92DA2" box="[602,707,996,1022]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="7.0" metricValueMin="2.0" pageId="26" pageNumber="27" unit="mm" value="4.5" valueMax="7.0" valueMin="2.0">2–7 mm</quantity>
, glabrous, persistent. Leaves: petiole 5–15(–25) cm long, glabrous; lamina ovate-lanceolate to ovate, basifixed, base cordate with lobes almost overlapping, (4–)10–20 × (3–)
<quantity id="4CC6B59C2F77FFCD03C7FB9BE1F32A19" box="[408,505,1066,1093]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" pageId="26" pageNumber="27" unit="cm" value="7.5" valueMax="9.0" valueMin="6.0">6–9 cm</quantity>
, asymmetric, upper surface variegated green and brown with a dark centre, glabrous or sparsely puberulent, underside variegated red and green, sparsely puberulent to glabrous, venation palmate-pinnate, midrib (3–)
<quantity id="4CC6B59C2F77FFCD00DAFBC0E2E42AD7" box="[645,750,1137,1163]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="26" pageNumber="27" unit="cm" value="13.0" valueMax="18.0" valueMin="8.0">8–18 cm</quantity>
long; margin entire to shallow dentate, glabrous; apex shortly acuminate. Inflorescence: cymose, axillary, several; peduncle glabrous, branching twice, male primary
<quantity id="4CC6B59C2F70FFCA037AFE9FE19F2F15" box="[293,405,302,329]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="29" pageNumber="30" unit="cm" value="10.0" valueMax="15.0" valueMin="5.0">5–15 cm</quantity>
, secondary
<quantity id="4CC6B59C2F70FFCA0078FE9EE2812F15" box="[551,651,303,329]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="29" pageNumber="30" unit="cm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 cm</quantity>
, 2–6 flowers, female primary
<quantity id="4CC6B59C2F70FFCA065AFE9FE4632F15" box="[1029,1129,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="29" pageNumber="30" unit="cm" value="5.0" valueMax="6.0" valueMin="4.0">4–6 cm</quantity>
, 1(–2) flowers; bracts lanceolate, 6–20 ×
<quantity id="4CC6B59C2F70FFCA03F8FEE0E21A2F30" box="[423,528,337,364]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="3.0" valueMax="5.0" valueMin="1.0">1–5 mm</quantity>
, deciduous. Male flower: pedicel 13–25(–35) mm long, glabrous; tepals 4; outer tepals narrowly ovate, 10–30 ×
<quantity id="4CC6B59C2F70FFCA00C1FEC4E3182FD3" box="[670,786,373,399]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.6" metricValueMin="0.7" pageId="29" pageNumber="30" unit="mm" value="11.5" valueMax="16.0" valueMin="7.0">7–16 mm</quantity>
, white to pink, glabrous, margin entire; inner tepals elliptic-obovate, 10–20 ×
<quantity id="4CC6B59C2F70FFCA03B4FE29E25A2FEF" box="[491,592,408,435]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="7.0" metricValueMin="3.0" pageId="29" pageNumber="30" unit="mm" value="5.0" valueMax="7.0" valueMin="3.0">3–7 mm</quantity>
, white to pink, glabrous; androecium with 40 –70 stamens, symmetric; filaments
<quantity id="4CC6B59C2F70FFCA036BFE0AE1942F8A" box="[308,414,443,470]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="29" pageNumber="30" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1–3 mm</quantity>
long, unequal, free; anther elliptic-obovate,
<quantity id="4CC6B59C2F70FFCA0199FE0AE4052F8A" box="[966,1039,443,470]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="mm" value="2.0">2 mm</quantity>
long, dehiscing through slits about half the length of the anther, not hooded, connective extended. Female flower: pedicel glabrous; bracteoles absent; tepals 4, similar to those of the male flower; ovary 4-locular, placentae bifid; capsule rhomboid, with 4 fleshy triangular wings/ridges; styles 4, forked once and twisted twice, deciduous. Fruit: on a stout pedicel, rhomboid; capsule oblong-ellipsoid, 20 ×
<quantity id="4CC6B59C2F70FFCA0183FDF9E4592C3F" box="[988,1107,584,611]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="11.0" metricValueMin="4.0" pageId="29" pageNumber="30" unit="mm" value="7.5" valueMax="11.0" valueMin="4.0">4–11 mm</quantity>
, puberulous to glabrous, with 4 stubby wings or horns.
</paragraph>
</subSubSection>
<caption id="DF4148F12F77FFCD02E2F87BE3C029B9" ID-DOI="http://doi.org/10.5281/zenodo.14616265" ID-Zenodo-Dep="14616265" box="[189,970,1994,2021]" httpUri="https://zenodo.org/record/14616265/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1994,2020]" targetBox="[431,1160,1226,1953]" targetPageId="26">
<paragraph id="8B8118792F77FFCD02E2F87BE3C029B9" blockId="26.[189,970,1994,2021]" box="[189,970,1994,2021]" pageId="26" pageNumber="27">
<emphasis id="B94AC46B2F77FFCD02E2F87BE11029B8" bold="true" box="[189,282,1994,2020]" pageId="26" pageNumber="27">Fig. 16.</emphasis>
Map showing the location of
<taxonomicName id="4C3E63FA2F77FFCD0026F87AE34B29B9" authority="Dunn" authorityName="Dunn (Dunn" authorityYear="1920" box="[633,833,1995,2021]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="26" pageNumber="27" phylum="Tracheophyta" rank="species" species="burkillii">
<emphasis id="B94AC46B2F77FFCD0026F87AE2F329B9" box="[633,761,1995,2021]" italics="true" pageId="26" pageNumber="27">B. burkillii</emphasis>
Dunn
</taxonomicName>
specimens.
</paragraph>
</caption>
<caption id="DF4148F12F76FFCC02E2F817E53729B8" ID-DOI="http://doi.org/10.5281/zenodo.3787075" ID-Zenodo-Dep="3787075" httpUri="https://zenodo.org/record/3787075/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1958,1984]" targetBox="[415,1173,265,1921]" targetPageId="27">
<paragraph id="8B8118792F76FFCC02E2F817E53729B8" blockId="27.[189,1398,1958,2020]" pageId="27" pageNumber="28">
<emphasis id="B94AC46B2F76FFCC02E2F817E117299D" bold="true" box="[189,285,1958,1985]" pageId="27" pageNumber="28">Fig. 17.</emphasis>
<taxonomicName id="4C3E63FA2F76FFCC0378F819E2A7299D" authority="Dunn. A. Leaf" box="[295,685,1958,1985]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="27" pageNumber="28" phylum="Tracheophyta" rank="species" species="burkillii">
<emphasis id="B94AC46B2F76FFCC0378F819E1F8299D" box="[295,498,1959,1985]" italics="true" pageId="27" pageNumber="28">Begonia burkillii</emphasis>
Dunn.
<emphasis id="B94AC46B2F76FFCC000FF817E26F299C" bold="true" box="[592,613,1958,1984]" pageId="27" pageNumber="28">A</emphasis>
. Leaf
</taxonomicName>
pattern.
<emphasis id="B94AC46B2F76FFCC0144F816E325299D" bold="true" box="[795,815,1959,1985]" pageId="27" pageNumber="28">B</emphasis>
. Male flowers. Photograph A courtesy of Aaron Matsumoto and photograph B courtesy of Earl I-Lan of plants in cultivation in private collections.
</paragraph>
</caption>
<caption id="DF4148F12F71FFCB02E2F835E2EA29BA" ID-DOI="http://doi.org/10.5281/zenodo.3787077" ID-Zenodo-Dep="3787077" httpUri="https://zenodo.org/record/3787077/files/figure.png" pageId="28" pageNumber="29" startId="28.[189,232,1924,1950]" targetBox="[195,1391,269,1895]" targetPageId="28">
<paragraph id="8B8118792F71FFCB02E2F835E2EA29BA" blockId="28.[189,1399,1924,2022]" pageId="28" pageNumber="29">
<emphasis id="B94AC46B2F71FFCB02E2F835E11729C3" bold="true" box="[189,285,1924,1951]" pageId="28" pageNumber="29">Fig. 18.</emphasis>
Illustration of
<taxonomicName id="4C3E63FA2F71FFCB0389F837E31429C3" authority="Dunn. A. Habit" box="[470,798,1924,1951]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="burkillii">
<emphasis id="B94AC46B2F71FFCB0389F837E25229C3" box="[470,600,1925,1951]" italics="true" pageId="28" pageNumber="29">B. burkillii</emphasis>
Dunn.
<emphasis id="B94AC46B2F71FFCB00E9F835E2C129C2" bold="true" box="[694,715,1924,1950]" pageId="28" pageNumber="29">A</emphasis>
. Habit
</taxonomicName>
of plant.
<emphasis id="B94AC46B2F71FFCB01C6F834E3A729C3" bold="true" box="[921,941,1925,1951]" pageId="28" pageNumber="29">B</emphasis>
. Style.
<emphasis id="B94AC46B2F71FFCB0653F835E42B29C2" bold="true" box="[1036,1057,1924,1950]" pageId="28" pageNumber="29">C</emphasis>
. Female flower.
<emphasis id="B94AC46B2F71FFCB06AFF834E50F29C3" bold="true" box="[1264,1285,1925,1951]" pageId="28" pageNumber="29">D</emphasis>
. Stamen.
<emphasis id="B94AC46B2F71FFCB02E2F819E0DB299E" bold="true" box="[189,209,1960,1986]" pageId="28" pageNumber="29">E</emphasis>
. Male flower.
<emphasis id="B94AC46B2F71FFCB0320F819E19B299E" bold="true" box="[383,401,1960,1986]" pageId="28" pageNumber="29">F</emphasis>
. Male inflorescence. Drawn by Heather Raeburn. A–C from
<emphasis id="B94AC46B2F71FFCB0634F819E504299E" box="[1131,1294,1960,1986]" italics="true" pageId="28" pageNumber="29">Burkill 36720</emphasis>
; D from
<emphasis id="B94AC46B2F71FFCB02E2F87AE15529B9" box="[189,351,1995,2021]" italics="true" pageId="28" pageNumber="29">Burkill 37455</emphasis>
; E–F from
<emphasis id="B94AC46B2F71FFCB03B2F87AE28529B9" box="[493,655,1995,2021]" italics="true" pageId="28" pageNumber="29">Burkill 37139</emphasis>
; all K.
</paragraph>
</caption>
<subSubSection id="C3244BF22F70FFCA02E2FD1DE19A2D44" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8B8118792F70FFCA02E2FD1DE21F2C9B" blockId="29.[189,533,684,711]" box="[189,533,684,711]" pageId="29" pageNumber="30">
<heading id="D0C9AF152F70FFCA02E2FD1DE21F2C9B" bold="true" box="[189,533,684,711]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B94AC46B2F70FFCA02E2FD1DE21F2C9B" bold="true" box="[189,533,684,711]" pageId="29" pageNumber="30">Distribution and phenology</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F70FFCA02E2FD6AE19A2D44" blockId="29.[189,1399,731,793]" pageId="29" pageNumber="30">
Arunachal-Pradesh and Assam; also in
<collectingCountry id="F32958E92F70FFCA00DBFD6DE2F02CAA" box="[644,762,732,758]" name="Myanmar" pageId="29" pageNumber="30">Myanmar</collectingCountry>
;
<quantity id="4CC6B59C2F70FFCA015AFD6AE3962CAA" box="[773,924,731,758]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="11.0" metricValueMin="3.0" pageId="29" pageNumber="30" unit="m" value="700.0" valueMax="1100.0" valueMin="300.0">300–1100 m</quantity>
. Flowering: February to March; fruiting: February to April.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F70FFCA02E2FCF1E4D52DF0" pageId="29" pageNumber="30" type="conservation">
<paragraph id="8B8118792F70FFCA02E2FCF1E1BF2D06" blockId="29.[189,437,832,858]" box="[189,437,832,858]" pageId="29" pageNumber="30">
<heading id="D0C9AF152F70FFCA02E2FCF1E1BF2D06" bold="true" box="[189,437,832,858]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B94AC46B2F70FFCA02E2FCF1E1BF2D06" bold="true" box="[189,437,832,858]" pageId="29" pageNumber="30">Conservation status</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F70FFCA02E2FCDEE4D52DF0" blockId="29.[189,1398,877,940]" pageId="29" pageNumber="30">
Least Concern.
<taxonomicName id="4C3E63FA2F70FFCA0321FCDEE2402DD4" box="[382,586,878,904]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="burkillii">
<emphasis id="B94AC46B2F70FFCA0321FCDEE2402DD4" box="[382,586,878,904]" italics="true" pageId="29" pageNumber="30">Begonia burkillii</emphasis>
</taxonomicName>
has an EOO of
<quantity id="4CC6B59C2F70FFCA0144FCDEE3962DD4" box="[795,924,878,905]" metricMagnitude="7" metricUnit="m" metricValue="8.34" pageId="29" pageNumber="30" unit="km" value="83400.0">83,400 km</quantity>
<superScript id="7C4BB5312F70FFCA01C2FCDCE3AC2D27" attach="left" box="[925,934,877,891]" fontSize="6" pageId="29" pageNumber="30">2</superScript>
and an AOO of
<quantity id="4CC6B59C2F70FFCA0624FCDEE4C32DD4" box="[1147,1225,878,905]" metricMagnitude="4" metricUnit="m" metricValue="7.2" pageId="29" pageNumber="30" unit="km" value="72.0">72 km</quantity>
<superScript id="7C4BB5312F70FFCA0695FCDCE4D92D27" attach="right" box="[1226,1235,877,891]" fontSize="6" pageId="29" pageNumber="30">2</superScript>
, with a wide distribution through undisturbed forest in Arunachal-Pradesh and into Northern
<collectingCountry id="F32958E92F70FFCA0639FC23E4D02DF0" box="[1126,1242,914,940]" name="Myanmar" pageId="29" pageNumber="30">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3244BF22F70FFCA02E2FC62E3A12B51" pageId="29" pageNumber="30" type="discussion">
<paragraph id="8B8118792F70FFCA02E2FC62E1252DB1" blockId="29.[189,303,979,1005]" box="[189,303,979,1005]" pageId="29" pageNumber="30">
<heading id="D0C9AF152F70FFCA02E2FC62E1252DB1" bold="true" box="[189,303,979,1005]" fontSize="11" level="3" pageId="29" pageNumber="30" reason="3">
<emphasis id="B94AC46B2F70FFCA02E2FC62E1252DB1" bold="true" box="[189,303,979,1005]" pageId="29" pageNumber="30">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B8118792F70FFCA02E2FBB0E4B72AF4" blockId="29.[189,1399,1025,1193]" pageId="29" pageNumber="30">
Specimens from the region lacked female flowers, which are described based on the protologue and photographs of cultivated plants. The leaf markings can vary in colour from red to purple-brown to almost black, but keep the same pattern (
<figureCitation id="130504FC2F70FFCA00F5FBF9E3082A3E" box="[682,770,1096,1122]" captionStart="Fig" captionStartId="27.[189,232,1958,1984]" captionTargetBox="[415,1173,265,1921]" captionTargetId="figure@27.[415,1173,265,1921]" captionTargetPageId="27" captionText="Fig. 17. Begonia burkillii Dunn. A. Leaf pattern. B. Male flowers. Photograph A courtesy of Aaron Matsumoto and photograph B courtesy of Earl I-Lan of plants in cultivation in private collections." figureDoi="http://doi.org/10.5281/zenodo.3787075" httpUri="https://zenodo.org/record/3787075/files/figure.png" pageId="29" pageNumber="30">Fig. 17</figureCitation>
). This species is perhaps most similar to
<taxonomicName id="4C3E63FA2F70FFCA06ABFBF8E57C2A3E" authorityName="Irmsch. (Irmscher" authorityYear="1921" box="[1268,1398,1096,1122]" class="Magnoliopsida" family="Begoniaceae" genus="Begonia" kingdom="Plantae" order="Cucurbitales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="handelii">
<emphasis id="B94AC46B2F70FFCA06ABFBF8E57C2A3E" box="[1268,1398,1096,1122]" italics="true" pageId="29" pageNumber="30">B. handelii</emphasis>
</taxonomicName>
in being rhizomatous and having 4-locular fruit, but that species has large rounded outer petals (not narrowly ovate), non-variegated leaves and fruit that is more pyramidal than rhomboid.
</paragraph>
<paragraph id="8B8118792F70FFCA02E2FB61E3A12B51" blockId="29.[189,1399,1232,1294]" pageId="29" pageNumber="30">
The lectotypification by
<bibRefCitation id="EFAF65882F70FFCA0382FB61E2B72AB6" author="Saikia R. &amp; Roy H. &amp; Borthakur S. K." box="[477,701,1232,1258]" pageId="29" pageNumber="30" pagination="357 - 360" refId="ref57452" refString="Saikia R., Roy H. &amp; Borthakur S. K. 2011. New report of occurrence of two species of Begonia L. (Begoniaceae) from Assam, India. Pleione 5 (2): 357 - 360." type="journal article" year="2011">
Saikia
<emphasis id="B94AC46B2F70FFCA0072FB60E2622AB6" box="[557,616,1232,1258]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
(2011)
</bibRefCitation>
list two specimens (
<emphasis id="B94AC46B2F70FFCA01F0FB61E45B2AB6" box="[943,1105,1232,1258]" italics="true" pageId="29" pageNumber="30">Burkill 37706</emphasis>
and
<emphasis id="B94AC46B2F70FFCA06D5FB61E4D92AB6" box="[1162,1235,1232,1258]" italics="true" pageId="29" pageNumber="30">37121</emphasis>
) and hence is not valid. Here
<emphasis id="B94AC46B2F70FFCA032CFB42E21F2B51" box="[371,533,1267,1293]" italics="true" pageId="29" pageNumber="30">Burkill 37139</emphasis>
(K) is designated as the
<typeStatus id="5485A6DB2F70FFCA0168FB42E3AC2B51" box="[823,934,1267,1293]" pageId="29" pageNumber="30" type="lectotype">lectotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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