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<mods:title id="3D6CA41520E935F36E6B376499DB90E4">Provisional revision of American snapping shrimp allied to Alpheus floridanus Kingsley, 1878 (Crustacea: Decapoda: Alpheidae) with notes on A. floridanus africanus</mods:title>
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<treatment id="03979151FFEBFFEDFF06F11AFA2B5490" ID-DOI="http://doi.org/10.5281/zenodo.5686427" ID-GBIF-Taxon="119598574" ID-Zenodo-Dep="5686427" LSID="urn:lsid:plazi:treatment:03979151FFEBFFEDFF06F11AFA2B5490" httpUri="http://treatment.plazi.org/id/03979151FFEBFFEDFF06F11AFA2B5490" lastPageId="40" lastPageNumber="491" pageId="39" pageNumber="490">
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<paragraph id="8B812047FFEBFFE2FF06F11AFCC150DC" blockId="39.[151,886,1660,1686]" box="[151,886,1660,1686]" pageId="39" pageNumber="490">
<heading id="D0C9972BFFEBFFE2FF06F11AFCC150DC" bold="true" box="[151,886,1660,1686]" fontSize="11" level="1" pageId="39" pageNumber="490" reason="1">
<emphasis id="B94AFC55FFEBFFE2FF06F11AFCC150DC" bold="true" box="[151,886,1660,1686]" pageId="39" pageNumber="490">
Key to American species of the
<taxonomicName id="4C3E5BC4FFEBFFE2FD8EF11AFCB050DC" ID-CoL="84W7Q" box="[543,775,1660,1686]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="490" phylum="Arthropoda" rank="species" species="floridanus">
<emphasis id="B94AFC55FFEBFFE2FD8EF11AFCB050DC" bold="true" box="[543,775,1660,1686]" italics="true" pageId="39" pageNumber="490">Alpheus floridanus</emphasis>
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complex
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<subSubSection id="C32473CCFFEBFFEDFF06F1A7FA2B5490" lastPageId="40" lastPageNumber="491" pageId="39" pageNumber="490" type="key">
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<paragraph id="8B812047FFEBFFE2FF06F1A7FA2B5107" blockId="39.[151,1436,1729,2015]" pageId="39" pageNumber="490">
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1. Antennal scale (scaphocerite) lateral margin sinuous (
<figureCitation id="13053CC2FFEBFFE2FD73F1A7FCAD509D" box="[738,794,1729,1751]" captionStart="FIGURE 6" captionStartId="11.[151,250,1727,1749]" captionTargetBox="[152,1422,204,1695]" captionTargetId="figure@11.[151,1436,193,1706]" captionTargetPageId="11" captionText="FIGURE 6. Alpheus roblesi sp. nov.: a g, holotype male (USNM 1265091 = tissue ULLZ 9141); h, paratype ovigerous female (ULLZ 7197 B): a, habitus, lateral; b, anterior carapace, left side, lateral; c, anterior carapace, dorsal; d, left major cheliped, lateral; e, right minor cheliped, lateral; f, right minor cheliped, propodus and dactylus, mesial (balaeniceps setae shown); g, right second pleopod, appendix interna and appendix masculina (serrated setae shown only in bracket); h, sixth abdominal somite, telson and uropods, dorsal. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/229147/files/figure.png" pageId="39" pageNumber="490">Fig. 6</figureCitation>
c), medial bend (toward body midline) originating near midlength; pereopod 5 propodus bearing 17 movable spines (rarely lacking, often conspicuous) on ventromesial surface (
<figureCitation id="13053CC2FFEBFFE2FA90F1B9FA8E50BF" box="[1281,1337,1759,1781]" captionStart="FIGURE 8" captionStartId="13.[151,250,1764,1786]" captionTargetBox="[151,1436,391,1731]" captionTargetId="figure@13.[151,1436,373,1743]" captionTargetPageId="13" captionText="FIGURE 8. Alpheus roblesi sp. nov.: a e, holotype male (USNM 1265091 = tissue ULLZ 9141); f, paratype ovigerous female (ULLZ 7197 B): a, left second pereopod, lateral; b, left third pereopod, lateral; c, left fourth pereopod, lateral; d, left fifth pereopod, lateral; e, left fifth pereopod, propodus and dactylus, mesial; f, abdomen, telson, uropods, pleopods, and egg, lateral. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/229149/files/figure.png" pageId="39" pageNumber="490">Fig. 8</figureCitation>
e); in life, weakly pigmented except for irregular orange banding on chelae, body translucence overlain by diffuse dorsal aggregations of small orange chromatophores [inner continental shelf, southern Gulf of
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, southeastern Florida and Caribbean; eastern Atlantic, Africa]...................................................................
<taxonomicName id="4C3E5BC4FFEBFFE2FB25F051FAFE5107" ID-CoL="85LNS" box="[1204,1353,1847,1869]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="39" pageNumber="490" phylum="Arthropoda" rank="species" species="roblesi" status="sp. nov.">
<emphasis id="B94AFC55FFEBFFE2FB25F051FAFE5107" box="[1204,1353,1847,1869]" italics="true" pageId="39" pageNumber="490">Alpheus roblesi</emphasis>
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<emphasis id="B94AFC55FFEBFFE2FADEF051FA2B5107" bold="true" box="[1359,1436,1847,1869]" pageId="39" pageNumber="490">sp. nov.</emphasis>
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</paragraph>
<paragraph id="8B812047FFEBFFE2FF06F033FA2C51EF" blockId="39.[151,1436,1729,2015]" pageId="39" pageNumber="490">
<keyLead id="30CF8372FFEBFFE2FF06F033FA2C51EF" pageId="39" pageNumber="490">
- Antennal scale lateral margin concave (
<figureCitation id="13053CC2FFEBFFE2FDCBF032FD2C5120" box="[602,667,1876,1898]" captionStart="FIGURE 3" captionStartId="5.[151,250,1803,1825]" captionTargetBox="[170,1415,205,1737]" captionTargetId="figure@5.[156,1431,193,1749]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus floridanus Kingsley, 1878: a o, lectotype female (MCZ 4987): a, anterior carapace, dorsal (right antennal scale broken); b, anterior carapace, left lateral view; c, left mandible, mesial; d, left maxillule, lateral; e, left second maxilla, lateral; f, left first maxilliped, lateral; g, left second maxilliped, lateral; h, right third maxilliped, lateral; i, right major chela, dactylus and propodus, lateral; j, left minor cheliped, ishium, merus, carpus, propodus, and dactylus, mesial; k, right major cheliped, coxa, basis, ishium, merus, and carpus, mesial; l, right third pereopod, lateral; m, left fourth pereopod, lateral; n, abdomen, telson (broken) and uropods, lateral; o, telson and uropods, lateral. Scale bars = 1.0 mm (d g), 2.0 mm (a c, h o)." httpUri="https://zenodo.org/record/229144/files/figure.png" pageId="39" pageNumber="490">Figs. 3</figureCitation>
a, 4 b, 9 b, 12 b, 15 d), medial bend (if any) restricted to spine at tip; pereopod 5 propodus lacking movable spines along ventromesial surface; in life, body pigmentation varied, sometimes including strongly contrasting dorsal patterns of red, orange, green, brown, or yellow............................................... 2
</keyLead>
</paragraph>
<footnote id="E8253C49FFEBFFE2FF06F0CAFA2C5195" pageId="39" pageNumber="490">
<paragraph id="8B812047FFEBFFE2FF06F0CAFA2C5195" blockId="39.[151,1436,1729,2015]" pageId="39" pageNumber="490">2. Antennal scale (scaphocerite) blade and its distolateral tooth distinctly over-reaching distal end of antennular peduncle (Figs. 3 a, 4 b, 9 b), distolateral tooth distinctly over-reaching blade, often by&gt;1/10 blade length............................. 3</paragraph>
</footnote>
<paragraph id="8B812047FFE4FFEDFF06F7F1FA2C5680" blockId="40.[151,1436,151,730]" pageId="40" pageNumber="491">
<keyLead id="30CF8372FFE4FFEDFF06F7F1FA2C5680" pageId="40" pageNumber="491">- Antennal scale blade and its distolateral tooth reaching about to distal end of antennular peduncle, rarely with both slightly over-reaching it, distolateral tooth exceeding blade by &lt;1/10 blade length, if at all.................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30CA38E2FFE4FFEDFF06F7B7FA2B5798" pageId="40" pageNumber="491">
<paragraph id="8B812047FFE4FFEDFF06F7B7FA2B570A" blockId="40.[151,1436,151,730]" pageId="40" pageNumber="491">
<keyLead id="30CF8372FFE4FFEDFF06F7B7FA2B570A" pageId="40" pageNumber="491">
3. Pereopod 3 and 4 propodus bearing 12 movable spines on proximal half of ventrolateral margin (
<figureCitation id="13053CC2FFE4FFEDFB32F7B7FB5F56AD" box="[1187,1256,209,231]" captionStart="FIGURE 11" captionStartId="18.[151,250,1339,1361]" captionTargetBox="[162,1436,447,1308]" captionTargetId="figure@18.[151,1436,445,1318]" captionTargetPageId="18" captionText="FIGURE 11. Alpheus ulalae sp. nov.: a f, holotype ovig. female (USNM 1265092 = tissue / sequence ULLZ 6815): a, left second pereopod, lateral; b, left third pereopod, lateral (epipod not illustrated); c, dactylus of left third pereopod, dorsal; d, left third pereopod, carpus, propodus, dactylus, mesial; e, left fourth pereopod, lateral; f, left fifth pereopod, lateral. Scale bars = 2.0 mm." httpUri="https://zenodo.org/record/229152/files/figure.png" pageId="40" pageNumber="491">Fig. 11</figureCitation>
b, d, e); mandible incisor with strong dentition on &lt;1/2 of cutting edge (
<figureCitation id="13053CC2FFE4FFEDFD77F789FC9C574F" box="[742,811,239,261]" captionStart="FIGURE 10" captionStartId="17.[151,250,1505,1527]" captionTargetBox="[151,1427,200,1471]" captionTargetId="figure@17.[151,1436,193,1484]" captionTargetPageId="17" captionText="FIGURE 10. Alpheus ulalae sp. nov.: a f, holotype ovig. female (USNM 1265092 = tissue / sequence ULLZ 6815): a, left mandible, mesial; b, left maxillule, lateral; c, left second maxilla, lateral; d, left first maxilliped, lateral; e, left second maxilliped, lateral; f, left third maxilliped, lateral. Scale bars = 1.0 mm (b e), 2.0 mm (a, f)." httpUri="https://zenodo.org/record/229151/files/figure.png" pageId="40" pageNumber="491">Fig. 10</figureCitation>
a); in life, body translucence overlain by diffuse dorsal aggregations of small orange chromatophores, lacking distinct longitudinal bars on carapace and abdomen (
<figureCitation id="13053CC2FFE4FFEDFB09F66AFB675768" box="[1176,1232,268,290]" captionStart="FIGURE 2" captionStartId="4.[151,250,1738,1760]" captionTargetBox="[260,1321,193,1708]" captionTargetId="figure@4.[260,1321,193,1709]" captionTargetPageId="4" captionText="FIGURE 2. Color patterns of species allied to Alpheus floridanus Kingsley, 1878. Alpheus roblesi sp. nov.: a, b, dorsal and lateral views of male, cl 8.0 mm, Campeche Banks, Mexico (ULLZ 7197 A); c, lateral view of ovigerous female, cl 7.5 mm, Campeche Banks, Mexico (ULLZ 7197 B); d, dorsal view of female, cl 4.9 mm, Campeche Banks, Mexcio (ULLZ 7440). Alpheus ulalae sp. nov.: e, f, lateral view of holotype ovigerous female, cl 7.5 mm, northeastern Gulf of Mexico (USNM 1265092 = tissue / sequence ULLZ 6815). Alpheus hephaestus sp. nov.: g, lateral view of male, cl 9.1 mm, Pacific coast of Costa Rica (ULLZ 6204); h, dorsal view of male, cl 7.1 mm, Pacific coast of Panama (ULLZ 8004); i, dorsal view of male, cl 8.9 mm, Pacific coast of Panama (ULLZ 9508 A). Alpheus platycheirus Boone, 1927: j, dorsal view of damaged female, cl 7.9 mm, northern Gulf of Mexico (ULLZ 6421); k, dorsal view of ovigerous female, cl 9.0 mm, northern Gulf of Mexico (ULLZ 5858); l, dorsal view of male, cl 9.8 mm, northern Gulf of Mexico (ULLZ 5952)." httpUri="https://zenodo.org/record/229143/files/figure.png" pageId="40" pageNumber="491">Fig. 2</figureCitation>
ef) [inner continental shelf, Gulf of
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and Carribean]......................................................
<taxonomicName id="4C3E5BC4FFE4FFEDFB63F64FFAFF5775" ID-CoL="85LQP" box="[1266,1352,297,319]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="491" phylum="Arthropoda" rank="species" species="ulalae" status="sp. nov.">
<emphasis id="B94AFC55FFE4FFEDFB63F64FFAFF5775" box="[1266,1352,297,319]" italics="true" pageId="40" pageNumber="491">A. ulalae</emphasis>
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<emphasis id="B94AFC55FFE4FFEDFADEF64CFA2B570A" bold="true" box="[1359,1436,298,320]" pageId="40" pageNumber="491">sp. nov.</emphasis>
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</paragraph>
<paragraph id="8B812047FFE4FFEDFF06F621FA2B5798" blockId="40.[151,1436,151,730]" pageId="40" pageNumber="491">
<keyLead id="30CF8372FFE4FFEDFF06F621FA2B5798" pageId="40" pageNumber="491">
- Pereopod 3 and 4 propodus bearing 3 or more movable spines along ventrolateral margin, often extending onto distal half (
<figureCitation id="13053CC2FFE4FFEDFF79F602FE9B5730" box="[232,300,356,378]" captionStart="FIGURE 3" captionStartId="5.[151,250,1803,1825]" captionTargetBox="[170,1415,205,1737]" captionTargetId="figure@5.[156,1431,193,1749]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus floridanus Kingsley, 1878: a o, lectotype female (MCZ 4987): a, anterior carapace, dorsal (right antennal scale broken); b, anterior carapace, left lateral view; c, left mandible, mesial; d, left maxillule, lateral; e, left second maxilla, lateral; f, left first maxilliped, lateral; g, left second maxilliped, lateral; h, right third maxilliped, lateral; i, right major chela, dactylus and propodus, lateral; j, left minor cheliped, ishium, merus, carpus, propodus, and dactylus, mesial; k, right major cheliped, coxa, basis, ishium, merus, and carpus, mesial; l, right third pereopod, lateral; m, left fourth pereopod, lateral; n, abdomen, telson (broken) and uropods, lateral; o, telson and uropods, lateral. Scale bars = 1.0 mm (d g), 2.0 mm (a c, h o)." httpUri="https://zenodo.org/record/229144/files/figure.png" pageId="40" pageNumber="491">Figs. 3</figureCitation>
l, m; 5 b, c); mandible incisor with strong dentition along entire margin (
<figureCitation id="13053CC2FFE4FFEDFB90F602FB8B5730" box="[1025,1084,356,378]" captionStart="FIGURE 3" captionStartId="5.[151,250,1803,1825]" captionTargetBox="[170,1415,205,1737]" captionTargetId="figure@5.[156,1431,193,1749]" captionTargetPageId="5" captionText="FIGURE 3. Alpheus floridanus Kingsley, 1878: a o, lectotype female (MCZ 4987): a, anterior carapace, dorsal (right antennal scale broken); b, anterior carapace, left lateral view; c, left mandible, mesial; d, left maxillule, lateral; e, left second maxilla, lateral; f, left first maxilliped, lateral; g, left second maxilliped, lateral; h, right third maxilliped, lateral; i, right major chela, dactylus and propodus, lateral; j, left minor cheliped, ishium, merus, carpus, propodus, and dactylus, mesial; k, right major cheliped, coxa, basis, ishium, merus, and carpus, mesial; l, right third pereopod, lateral; m, left fourth pereopod, lateral; n, abdomen, telson (broken) and uropods, lateral; o, telson and uropods, lateral. Scale bars = 1.0 mm (d g), 2.0 mm (a c, h o)." httpUri="https://zenodo.org/record/229144/files/figure.png" pageId="40" pageNumber="491">Fig. 3</figureCitation>
c); in life, chromatophores forming strong pattern of brownish-maroon to orange longitudinal bars on off-white to pale green background of carapace and abdomen (
<figureCitation id="13053CC2FFE4FFEDFE88F6F9FEE457FF" box="[281,339,415,437]" captionStart="FIGURE 1" captionStartId="3.[151,250,1891,1913]" captionTargetBox="[208,1377,193,1862]" captionTargetId="figure@3.[208,1377,193,1863]" captionTargetPageId="3" captionText="FIGURE 1. Color patterns in Alpheus floridanus Kingsley, 1878: a, dorsal view of male, cl 12.1 mm., Fort Pierce, Florida (ULLZ 10569 A); b, dorsal view of ovigerous female, cl 12.4 mm, Fort Pierce, Florida (ULLZ 10569 B); c, dorsal view of ovigerous female, cl 11.8 mm, Carrie Bow Cay, Belize (ULLZ 11128); d, lateral view of ovig. female, cl 10.3 mm, Fort Pierce, Florida (ULLZ 5631); e, lateral view of ovigerous female, cl 11.6 mm, Fort Pierce, Florida (ULLZ 6207); f, g, dorsal and lateral views of male, cl 11.0 mm, Campeche Banks, Mexico (ULLZ 6203)." httpUri="https://zenodo.org/record/229142/files/figure.png" pageId="40" pageNumber="491">Fig. 1</figureCitation>
ag) [nearshore and shallow western Atlantic, southern Gulf of
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,
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, southeastern Florida and Caribbean]..................................................................................
<taxonomicName id="4C3E5BC4FFE4FFEDFB6BF6DAFAC05798" ID-CoL="84W7Q" box="[1274,1399,444,466]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="491" phylum="Arthropoda" rank="species" sensu="stricto" species="floridanus">
<emphasis id="B94AFC55FFE4FFEDFB6BF6DAFAC05798" box="[1274,1399,444,466]" italics="true" pageId="40" pageNumber="491">A. floridanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A279412EFFE4FFEDFAECF6DBFA2B5798" box="[1405,1436,445,466]" pageId="40" pageNumber="491" sensu="stricto">s.s.</taxonomicNameLabel>
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4. In life, chromatophores typically rust-orange to red, loosely distributed into interconnected spots and ill-defined lateral bars or crescents on abdominal somites, sixth somite not dorsally more pigmented than preceding somites or proximal half of uropodal endopods (
<figureCitation id="13053CC2FFE4FFEDFEDAF572FE325460" box="[331,389,532,554]" captionStart="FIGURE 2" captionStartId="4.[151,250,1738,1760]" captionTargetBox="[260,1321,193,1708]" captionTargetId="figure@4.[260,1321,193,1709]" captionTargetPageId="4" captionText="FIGURE 2. Color patterns of species allied to Alpheus floridanus Kingsley, 1878. Alpheus roblesi sp. nov.: a, b, dorsal and lateral views of male, cl 8.0 mm, Campeche Banks, Mexico (ULLZ 7197 A); c, lateral view of ovigerous female, cl 7.5 mm, Campeche Banks, Mexico (ULLZ 7197 B); d, dorsal view of female, cl 4.9 mm, Campeche Banks, Mexcio (ULLZ 7440). Alpheus ulalae sp. nov.: e, f, lateral view of holotype ovigerous female, cl 7.5 mm, northeastern Gulf of Mexico (USNM 1265092 = tissue / sequence ULLZ 6815). Alpheus hephaestus sp. nov.: g, lateral view of male, cl 9.1 mm, Pacific coast of Costa Rica (ULLZ 6204); h, dorsal view of male, cl 7.1 mm, Pacific coast of Panama (ULLZ 8004); i, dorsal view of male, cl 8.9 mm, Pacific coast of Panama (ULLZ 9508 A). Alpheus platycheirus Boone, 1927: j, dorsal view of damaged female, cl 7.9 mm, northern Gulf of Mexico (ULLZ 6421); k, dorsal view of ovigerous female, cl 9.0 mm, northern Gulf of Mexico (ULLZ 5858); l, dorsal view of male, cl 9.8 mm, northern Gulf of Mexico (ULLZ 5952)." httpUri="https://zenodo.org/record/229143/files/figure.png" pageId="40" pageNumber="491">Fig. 2</figureCitation>
jl); mature specimens with major cheliped ischium lateral longitudinal furrow deeply defined; pereopod 4 propodus bearing 14 movable spines on ventrolateral surface [muddy middle and inner continental shelf, northern Gulf of
<collectingCountry id="F32960D7FFE4FFEDFF70F529FE9D542F" box="[225,298,591,613]" name="Mexico" pageId="40" pageNumber="491">Mexico</collectingCountry>
through Caribbean and
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<taxonomicName id="4C3E5BC4FFE4FFEDFB40F529FA2D542F" ID-CoL="85LMZ" box="[1233,1434,591,613]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="491" phylum="Arthropoda" rank="species" species="platycheirus">
<emphasis id="B94AFC55FFE4FFEDFB40F529FA2D542F" box="[1233,1434,591,613]" italics="true" pageId="40" pageNumber="491">Alpheus platycheirus</emphasis>
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- In life, chromatophores typically deep fire-red to reddish brown, densely distributed overall, sixth abdominal somite more densely pigmented than fifth or proximal half of uropodal endopods (
<figureCitation id="13053CC2FFE4FFEDFC0FF5EFFC6D54D5" box="[926,986,649,671]" captionStart="FIGURE 2" captionStartId="4.[151,250,1738,1760]" captionTargetBox="[260,1321,193,1708]" captionTargetId="figure@4.[260,1321,193,1709]" captionTargetPageId="4" captionText="FIGURE 2. Color patterns of species allied to Alpheus floridanus Kingsley, 1878. Alpheus roblesi sp. nov.: a, b, dorsal and lateral views of male, cl 8.0 mm, Campeche Banks, Mexico (ULLZ 7197 A); c, lateral view of ovigerous female, cl 7.5 mm, Campeche Banks, Mexico (ULLZ 7197 B); d, dorsal view of female, cl 4.9 mm, Campeche Banks, Mexcio (ULLZ 7440). Alpheus ulalae sp. nov.: e, f, lateral view of holotype ovigerous female, cl 7.5 mm, northeastern Gulf of Mexico (USNM 1265092 = tissue / sequence ULLZ 6815). Alpheus hephaestus sp. nov.: g, lateral view of male, cl 9.1 mm, Pacific coast of Costa Rica (ULLZ 6204); h, dorsal view of male, cl 7.1 mm, Pacific coast of Panama (ULLZ 8004); i, dorsal view of male, cl 8.9 mm, Pacific coast of Panama (ULLZ 9508 A). Alpheus platycheirus Boone, 1927: j, dorsal view of damaged female, cl 7.9 mm, northern Gulf of Mexico (ULLZ 6421); k, dorsal view of ovigerous female, cl 9.0 mm, northern Gulf of Mexico (ULLZ 5858); l, dorsal view of male, cl 9.8 mm, northern Gulf of Mexico (ULLZ 5952)." httpUri="https://zenodo.org/record/229143/files/figure.png" pageId="40" pageNumber="491">
Fig.
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gi); mature specimens with major cheliped ischium lateral longitudinal furrow broad, shallow; pereopod 4 propodus bearing 34 movable spines on ventrolateral surface [muddy middle and inner continental margins, eastern Pacific, Gulf of California to
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]...
<taxonomicName id="4C3E5BC4FFE4FFEDFB1AF5A2FAFE5490" ID-CoL="85LJF" box="[1163,1353,708,730]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="491" phylum="Arthropoda" rank="species" species="hephaestus" status="sp. nov.">
<emphasis id="B94AFC55FFE4FFEDFB1AF5A2FAFE5490" box="[1163,1353,708,730]" italics="true" pageId="40" pageNumber="491">Alpheus hephaestus</emphasis>
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