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<document id="CDC980FD7593306A15AD28AF9202F0FB" ID-DOI="10.11646/zootaxa.4093.3.1" ID-GBIF-Dataset="85f537f5-faf4-4a89-839b-8f56feadb0e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="264106" ID-ZooBank="565068DE-7042-483D-99FD-50BD2FA86BC1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1458719684729" checkinUser="plazi" docAuthor="Pollock, Darren A." docDate="2016" docId="038A6816711943382EDCFB4F369EFE0C" docLanguage="en" docName="zootaxa.4093.3.1.pdf.imf" docOrigin="Zootaxa 4093 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thisiomorphus Pic" docType="treatment" docVersion="9" lastPageNumber="304" masterDocId="FFB3106E711B433C2E4BFF88367EFFDC" masterDocTitle="Revision of Thisiomorphus Pic (Coleoptera: Mycteridae: Eurypinae) with descriptions of eleven new species from Central and South America and a key to genera of Neotropical Eurypinae" masterLastPageNumber="322" masterPageNumber="301" pageNumber="303" updateTime="1698185977904" updateUser="plazi">
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<mods:title id="75488E5928D4DFA1DD146AD425ACAB31">Revision of Thisiomorphus Pic (Coleoptera: Mycteridae: Eurypinae) with descriptions of eleven new species from Central and South America and a key to genera of Neotropical Eurypinae</mods:title>
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<mods:namePart id="6C553996ACAA5FCB41261C2CF801A4F2">Pollock, Darren A.</mods:namePart>
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<mods:date id="402987E98B81EA9D2B50722099641EB6">2016</mods:date>
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<treatment id="038A6816711943382EDCFB4F369EFE0C" ID-DOI="http://doi.org/10.5281/zenodo.6063828" ID-GBIF-Taxon="119257837" ID-Zenodo-Dep="6063828" LSID="urn:lsid:plazi:treatment:038A6816711943382EDCFB4F369EFE0C" httpUri="http://treatment.plazi.org/id/038A6816711943382EDCFB4F369EFE0C" lastPageId="4" lastPageNumber="304" pageId="2" pageNumber="303">
<subSubSection id="C3398A8B7119433E2EDCFB4F3703FB3D" box="[151,381,1223,1249]" pageId="2" pageNumber="303" type="nomenclature">
<paragraph id="8B9CD9007119433E2EDCFB4F3703FB3D" blockId="2.[151,381,1223,1249]" box="[151,381,1223,1249]" pageId="2" pageNumber="303">
<heading id="D0D46E6C7119433E2EDCFB4F3703FB3D" bold="true" box="[151,381,1223,1249]" fontSize="11" level="1" pageId="2" pageNumber="303" reason="1">
<taxonomicName id="4C23A2837119433E2EDCFB4F3703FB3D" authority="Pic" authorityName="Pic" box="[151,381,1223,1249]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2EDCFB4F3703FB3D" bold="true" box="[151,381,1223,1249]" pageId="2" pageNumber="303">
<emphasis id="B95705127119433E2EDCFB4F3731FB3D" bold="true" box="[151,335,1223,1249]" italics="true" pageId="2" pageNumber="303">Thisiomorphus</emphasis>
Pic
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3398A8B7119433E2EDCFA863682F9BC" pageId="2" pageNumber="303" type="reference_group">
<paragraph id="8B9CD9007119433E2EDCFA8634C6FA9F" blockId="2.[151,1150,1294,1347]" pageId="2" pageNumber="303">
<taxonomicName id="4C23A2837119433E2EDCFA8637C6FAF8" authority="Pic 1931: 31" authorityName="Pic" authorityPageNumber="31" authorityYear="1931" box="[151,440,1294,1316]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2EDCFA863751FAF8" box="[151,303,1294,1316]" italics="true" pageId="2" pageNumber="303">Thisiomorphus</emphasis>
Pic 1931: 31
</taxonomicName>
; Blackwelder 1945: 493; Pollock 2010: 693; Ivie &amp; Pollock 2012: 10.
<emphasis id="B95705127119433E2EDCFAA53762FA9F" bold="true" box="[151,284,1325,1347]" pageId="2" pageNumber="303">
<typeStatus id="549867A27119433E2EDCFAA536B4FA9F" box="[151,202,1325,1347]" pageId="2" pageNumber="303">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C23A2837119433E2F62FAA5345CFA9F" authority="Pic" authorityName="Pic" box="[297,546,1325,1347]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B95705127119433E2F62FAA53786FA9F" box="[297,504,1325,1347]" italics="true" pageId="2" pageNumber="303">Thisiomorphus latus</emphasis>
Pic
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="8B9CD9007119433E2EDCFAE73682F9BC" blockId="2.[151,1437,1391,1920]" pageId="2" pageNumber="303">
Original description (translated from Pic 1931: 31 by the author): “
<taxonomicName id="4C23A2837119433E2DEAFAE73233FA54" box="[929,1101,1391,1416]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus" status="gen. nov.">Thisiomorphus</taxonomicName>
<taxonomicNameLabel id="A264B8697119433E2A1CFAF832D8FA54" box="[1111,1190,1392,1416]" pageId="2" pageNumber="303" rank="genus">n. gen.</taxonomicNameLabel>
(fam.
<taxonomicName id="4C23A2837119433E2ABEFAE733EFFA54" box="[1269,1425,1391,1416]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="subFamily" subFamily="Lacconotinae">Lacconotinae</taxonomicName>
). This genus, near
<taxonomicName id="4C23A2837119433E2F12FA1B3794FA70" authority="Chp." authorityName="Chp." box="[345,490,1427,1452]" class="Insecta" family="Mycteridae" genus="Thisias" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2F12FA1B37D4FA70" box="[345,426,1427,1452]" italics="true" pageId="2" pageNumber="303">Thisias</emphasis>
Chp.
</taxonomicName>
, which can be distinguished at first glance by the structure of the prothorax and the pygidium protruding posterior of the elytra, presents the following principal characters: fore coxae approximate and median coxae with trochantin, legs robust, tarsal claws appendiculate; antennae short and slender, with club of four articles; prothorax short and transverse, with weak lateral bead, vanishing anteriorly, and posteriorly bordered; body robust, very slightly pubescent; pygidium prominent. This genus is established for the following new species.”
</paragraph>
</subSubSection>
<subSubSection id="C3398A8B711943382E8CF9E3369EFE0C" lastPageId="4" lastPageNumber="305" pageId="2" pageNumber="303" type="description">
<paragraph id="8B9CD9007119433E2E8CF9E33785F917" blockId="2.[151,1437,1391,1920]" pageId="2" pageNumber="303">
<emphasis id="B95705127119433E2E8CF9E33443F958" bold="true" box="[199,573,1643,1668]" pageId="2" pageNumber="303">Redescription of genus. Body</emphasis>
subparallel-sided, elongate oval or broadly oval (TL/GEW 2.023.05), flattened to slightly convex dorsally. Pubescence short, inconspicuous (most pronounced in
<taxonomicName id="4C23A2837119433E2AEAF9193365F974" box="[1185,1307,1680,1704]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="B95705127119433E2AEAF91932D0F974" box="[1185,1198,1681,1704]" italics="true" pageId="2" pageNumber="303">T</emphasis>
.
<emphasis id="B95705127119433E2AF7F9193365F974" box="[1212,1307,1681,1704]" italics="true" pageId="2" pageNumber="303">osaensis</emphasis>
</taxonomicName>
). TL
<quantity id="4CDB74E57119433E2B17F9183680F917" metricMagnitude="-3" metricUnit="m" metricValue="6.01" metricValueMax="7.9" metricValueMin="4.12" pageId="2" pageNumber="303" unit="mm" value="6.01" valueMax="7.9" valueMin="4.12">4.12 7.90 mm</quantity>
; GEW
<quantity id="4CDB74E57119433E2F1BF93B3789F917" box="[336,503,1715,1740]" metricMagnitude="-3" metricUnit="m" metricValue="2.52" metricValueMax="3.52" metricValueMin="1.52" pageId="2" pageNumber="303" unit="mm" value="2.52" valueMax="3.52" valueMin="1.52">1.523.52 mm</quantity>
.
</paragraph>
<paragraph id="8B9CD9007119433E2E8CF95F3308F8A3" blockId="2.[151,1437,1391,1920]" pageId="2" pageNumber="303">
<emphasis id="B95705127119433E2E8CF95F3779F92C" bold="true" box="[199,263,1751,1776]" pageId="2" pageNumber="303">Head</emphasis>
relatively short, slightly to moderately narrowed posterior of eyes; distance between posterior extent of eye and anterior margin of pronotum much less than length of one eye; eyes moderately large, hemispherical in some species, finely faceted, without intrafacetal setae; frontoclypeal suture not indicated; frontal furrows present, of variable length and depth, parallel or slightly convergent anteriorly; frons slightly elevated over very slightly concealed antennal insertions; labrum distinctly transverse, anterior margin slightly convex to slightly concave.
</paragraph>
<paragraph id="8B9CD9007119433E2EDCF834370AF82D" blockId="2.[151,1436,1980,2033]" pageId="2" pageNumber="303">
1. It is doubtful that all species of
<taxonomicName id="4C23A2837119433E2C41F8343401F80E" box="[522,639,1980,2002]" class="Insecta" family="Mycteridae" genus="Loboglossa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127119433E2C41F8343401F80E" box="[522,639,1980,2002]" italics="true" pageId="2" pageNumber="303">Loboglossa</emphasis>
</taxonomicName>
described by Pic and Champion are congeneric with
<taxonomicName id="4C23A2837119433E2AEBF835330CF80E" authority="Solier" authorityName="Solier" box="[1184,1394,1980,2002]" class="Insecta" family="Mycteridae" genus="Loboglossa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="303" phylum="Arthropoda" rank="species" species="variipennis">
<emphasis id="B95705127119433E2AEBF83532D0F80E" box="[1184,1198,1981,2002]" italics="true" pageId="2" pageNumber="303">L</emphasis>
.
<emphasis id="B95705127119433E2AF7F8353350F80E" box="[1212,1326,1981,2002]" italics="true" pageId="2" pageNumber="303">variipennis</emphasis>
Solier
</taxonomicName>
, the
<typeStatus id="549867A27119433E2E8CF854368DF82D" box="[199,243,2012,2033]" pageId="2" pageNumber="303">type</typeStatus>
of the genus
</paragraph>
<paragraph id="8B9CD9007118433F2E8CFF1F343CFE9C" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CFF1F3744FF6C" bold="true" box="[199,314,151,176]" pageId="3" pageNumber="304">Antennae</emphasis>
relatively short, extending posteriorly to near midlength of pronotum (in
<taxonomicName id="4C23A2837118433F2AE4FF113335FF6C" box="[1199,1355,152,176]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="curticornis" status="sp. nov.">
<emphasis id="B95705127118433F2AE4FF1132C3FF6C" box="[1199,1213,153,176]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2A9BFF113335FF6C" box="[1232,1355,153,176]" italics="true" pageId="3" pageNumber="304">curticornis</emphasis>
</taxonomicName>
<emphasis id="B95705127118433F2B1CFF1F33E2FF6C" bold="true" box="[1367,1436,151,176]" pageId="3" pageNumber="304">
<taxonomicNameLabel id="A264B8697118433F2B1CFF1F33E2FF6C" box="[1367,1436,151,176]" pageId="3" pageNumber="304" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
slightly past anterior margin), not exhibiting obvious sexual dimorphism; scape and pedicel short, submoniliform; antennomeres 410 variable, from submoniliform (e.g.
<taxonomicName id="4C23A2837118433F2D5BFF6935F2FF24" box="[784,908,224,248]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="B95705127118433F2D5BFF693563FF24" box="[784,797,225,248]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2D66FF6935F2FF24" box="[813,908,225,248]" italics="true" pageId="3" pageNumber="304">osaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C23A2837118433F2DD7FF69324AFF24" box="[924,1076,224,248]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="curticornis">
<emphasis id="B95705127118433F2DD7FF6935D7FF24" box="[924,937,225,248]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2DF2FF69324AFF24" box="[953,1076,225,248]" italics="true" pageId="3" pageNumber="304">curticornis</emphasis>
</taxonomicName>
) to filiform, some species with vaguely indicated, subserrate distal club of 4 antennomeres; sensilla obvious on distal surfaces of antennomeres
<quantity id="4CDB74E57118433F2BCBFE8C36AEFE9C" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.54" metricValueMin="1.27" pageId="3" pageNumber="304" unit="in" value="7.5" valueMax="10.0" valueMin="5.0">5 10 in</quantity>
some species (e.g.
<taxonomicName id="4C23A2837118433F2FE2FEA1344CFE9C" box="[425,562,295,320]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="davidsoni">
<emphasis id="B95705127118433F2FE2FEA137C8FE9C" box="[425,438,297,320]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2F8EFEAF344CFE9C" box="[453,562,295,320]" italics="true" pageId="3" pageNumber="304">davidsoni</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B9CD9007118433F2E8CFEC43520FDC4" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CFEC4373BFEB9" bold="true" box="[199,325,332,357]" pageId="3" pageNumber="304">Mandibles</emphasis>
relatively short, slightly asymmetrical, apically shallowly bidentate; inner, terebral teeth absent, represented by indentations or crenulations, especially on right mandible; left and right mandibular molae subequal in size, approximately subquadrate; prostheca distinct on both mandibles, nearly half the length of mandible;
<emphasis id="B95705127118433F2EDCFE3F3682FE0C" bold="true" box="[151,252,439,464]" pageId="3" pageNumber="304">maxillae</emphasis>
with inner lacinia slightly shorter than outer galea, both bluntly rounded and densely setose; maxillae with palpomeres elongate, 3-segmented, terminal palpomere variously securiform to slightly cultriform;
<emphasis id="B95705127118433F2B73FE5433E5FE29" bold="true" box="[1336,1435,476,501]" pageId="3" pageNumber="304">mentum</emphasis>
relatively short,
<emphasis id="B95705127118433F2F06FE7737AAFDC4" bold="true" box="[333,468,511,536]" pageId="3" pageNumber="304">labial palpi</emphasis>
with terminal palpomere fusiform.
</paragraph>
<paragraph id="8B9CD9007118433F2E8CFDAC3215FCC9" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CFDAC373CFDE1" bold="true" box="[199,322,548,573]" pageId="3" pageNumber="304">Prothorax</emphasis>
subquadrate to distinctly transverse (GPW/PL 1.141.73); lateral pronotal margins parallel-sided (e.g.
<taxonomicName id="4C23A2837118433F2E98FDC13750FDBC" box="[211,302,583,608]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="solisi">
<emphasis id="B95705127118433F2E98FDC1369EFDBC" box="[211,224,585,608]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2EB8FDCF3750FDBC" box="[243,302,583,608]" italics="true" pageId="3" pageNumber="304">solisi</emphasis>
</taxonomicName>
), convergent anteriorly (e.g.
<taxonomicName id="4C23A2837118433F2CCFFDC1355FFDBC" box="[644,801,584,608]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="convergens">
<emphasis id="B95705127118433F2CCFFDC134EFFDBC" box="[644,657,585,608]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2CE8FDC1355FFDBC" box="[675,801,585,608]" italics="true" pageId="3" pageNumber="304">convergens</emphasis>
</taxonomicName>
), to evenly arcuate (e.g.
<taxonomicName id="4C23A2837118433F2A01FDC13295FDBC" box="[1098,1259,583,608]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B95705127118433F2A01FDC13226FDBC" box="[1098,1112,585,608]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2A21FDCF3295FDBC" box="[1130,1259,583,608]" italics="true" pageId="3" pageNumber="304">brasiliensis</emphasis>
</taxonomicName>
); pronotal disc indistinctly to distinctly convex, with (e.g.
<taxonomicName id="4C23A2837118433F2CD8FDE5354AFD59" box="[659,820,620,645]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="brasiliensis">
<emphasis id="B95705127118433F2CD8FDE534DEFD58" box="[659,672,621,644]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2CF8FDE4354AFD59" box="[691,820,620,645]" italics="true" pageId="3" pageNumber="304">brasiliensis</emphasis>
</taxonomicName>
) or without broad, paired depressions; lateral carina absent, or at most vaguely impressed, posteriorly only; posterior margin of pronotum with distinct, relatively narrow but deep groove, ending laterally in slightly widened pits; prosternum anterad procoxae relatively short, intercoxal process narrow, elongate, extending to at least one-half length of procoxae, not broadened distally; procoxae rounded to slightly elongate; cavities broadly open, protrochantins concealed.
</paragraph>
<paragraph id="8B9CD9007118433F2E8CFC9732DFFC31" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CFC973728FCE4" bold="true" box="[199,342,799,824]" pageId="3" pageNumber="304">Mesothorax</emphasis>
. Elytra moderately ovate, ovate-elongate, to distinctly elongate; disc flat to moderately convex, upper surface evenly punctate/setose, setae mostly decumbent, but with a few scattered, erect setae; apical elytral patch present, conspicuous or inconspicuous dorsally (in intact specimens); each epipleuron narrow through most of its length, but distinct to very nearly the elytral apex; mesosternum with posterior intercoxal process elongate, subparallel-sided, extended posteriorly to past one-half length of mesocoxae; mesocoxae completely separated, trochantins exposed; coxal cavities partially closed laterally by inner margin of mesepimera.
</paragraph>
<paragraph id="8B9CD9007118433F2E8CFC7F3680FBA1" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CFC7F372AFBCC" bold="true" box="[199,340,1015,1040]" pageId="3" pageNumber="304">Metathorax</emphasis>
. Metasternum large, convex, with distinct discrimen; intercoxal process distinct, in contact with, or slightly overlapping mesosternal process; metendosternite with relatively wide stalk; anterior tendons attached to anterior surface of metendosternite body; laminae (lateral extensions) relatively short, rounded to somewhat angulate.
</paragraph>
<paragraph id="8B9CD9007118433F2E8CFB0F3569FB34" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CFB0F3776FB7C" bold="true" box="[199,264,1159,1184]" pageId="3" pageNumber="304">Wing</emphasis>
fully developed, with wing membrane beyond closed radial cell moderately short; wing lightly to moderately darkly pigmented; 3 MP veins reaching hind wing margin, proximal to CuA + AA; pigmented flecks present, near junction of
<collectingCountry id="F33499907118433F2FE5FB5937AEFB34" box="[430,464,1233,1256]" name="Poland" pageId="3" pageNumber="304">RP</collectingCountry>
and MP, and near radial cell.
</paragraph>
<paragraph id="8B9CD9007118433F2E8CFB7C36A6FAA4" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CFB7C3681FAD1" bold="true" box="[199,255,1268,1293]" pageId="3" pageNumber="304">Legs</emphasis>
well developed, similar in relative size and shape on all thoracic segments; legs relatively slender, femora slightly deepened at midlength; tibiae straight, subequal in length to femora; tibial spurs very short, inconspicuous; tarsi 5-5-4, simple ventrally except penultimate tarsomere distinctly broadened; tarsal claws with large, blunt basal tooth.
</paragraph>
<paragraph id="8B9CD9007118433F2E8CFA0C33F9FA1C" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CFA0C3742FA41" bold="true" box="[199,316,1412,1437]" pageId="3" pageNumber="304">Abdomen</emphasis>
with all ventrites freely articulated, uniformly punctate and setose; male sex patch present on second ventrite, consisting of relatively small, longitudinally ovate patch of thickened, posteriorly directed setae.
</paragraph>
<paragraph id="8B9CD9007118433F2E8CFA4434E6F944" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CFA44370DFA39" bold="true" box="[199,371,1484,1509]" pageId="3" pageNumber="304">
<collectingCountry id="F33499907118433F2E8CFA44377AFA39" box="[199,260,1484,1509]" name="Malta" pageId="3" pageNumber="304">Male</collectingCountry>
genitalia
</emphasis>
(
<figureCitation id="1318C5857118433F2FC9FA44347AFA38" box="[386,516,1484,1509]" captionStart="FIGURES 13 20" captionStartId="7.[151,264,1862,1884]" captionTargetBox="[211,1373,353,1833]" captionTargetId="figure@7.[203,1384,339,1841]" captionTargetPageId="7" captionText="FIGURES 13 20. Male genitalia of Thisiomorphus species. 13. T. andrewsi, n. sp.; 14. T. brasiliensis, n. sp.; 15. T. caeruleus, n. sp.; 16. T. curticornis, n. sp.; 17. T. festivus, n. sp.; 18. T. inaequalis, n. sp.; 19. T. osaensis, n. sp.; 20. T. solisi, n. sp.." httpUri="https://zenodo.org/record/264109/files/figure.png" pageId="3" pageNumber="304">Figs 1320</figureCitation>
) relatively short, stocky, median lobe oriented dorsad tegmen; sternite 9 forming ring-like sclerite; tegmen of two distinct parts; basale broad, deeply emarginate proximally; apicale divided into two separate parameres, shorter than basale (along lateral margin); inner margins of parameres relatively straight or slightly emarginate/sinuate distally, apices distinctly hooked medially; penis relatively stout, distinctly longer than tegmen; ventral surface emarginate; dorsal side subquadrate basally, sides somewhat explanate or “flanged”; apex of penis bluntly triangular to slightly bulbose.
</paragraph>
<paragraph id="8B9CD9007118433F2E8CF92C375DF891" blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CF92C37EEF961" bold="true" box="[199,400,1700,1725]" pageId="3" pageNumber="304">Female genitalia</emphasis>
with relatively short, lightly sclerotized ovipositor; coxites short, multi-segmented, with short setae; distal segment darker, more sclerotized than remainder of coxite; valviferal and proctigeral baculi well developed; coxital baculi present, situated distinctly distad ovipositor midlength; spiculum distinct, long, exceeding length of segment 8; bursa copulatrix small to moderately large, separated from vagina by narrow constriction.
</paragraph>
<paragraph id="8B9CD900711843382E8CF8DF369EFE0C" blockId="3.[151,1437,151,2013]" lastBlockId="4.[151,1437,151,464]" lastPageId="4" lastPageNumber="305" pageId="3" pageNumber="304">
<emphasis id="B95705127118433F2E8CF8DF3701F8AC" bold="true" box="[199,383,1879,1904]" pageId="3" pageNumber="304">Natural history</emphasis>
. No biocontextual information has been published for any species of
<taxonomicName id="4C23A2837118433F2AD4F8DF3339F8AC" box="[1183,1351,1879,1904]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127118433F2AD4F8DF3339F8AC" box="[1183,1351,1879,1904]" italics="true" pageId="3" pageNumber="304">Thisiomorphus</emphasis>
</taxonomicName>
; this is not surprising, as there was only one previously described species, based on a single specimen. The only data available have been gleaned from specimen labels, and even that is rather scant, regarding natural history of adult
<taxonomicName id="4C23A2837118433F2EDCF84C3741F801" box="[151,319,1988,2013]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus">
<emphasis id="B95705127118433F2EDCF84C3741F801" box="[151,319,1988,2013]" italics="true" pageId="3" pageNumber="304">Thisiomorphus</emphasis>
</taxonomicName>
. For several species (
<taxonomicName id="4C23A2837118433F2C76F84D34C5F800" box="[573,699,1989,2012]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="osaensis">
<emphasis id="B95705127118433F2C76F84D3434F800" box="[573,586,1989,2012]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2C17F84D34C5F800" box="[604,699,1989,2012]" italics="true" pageId="3" pageNumber="304">osaensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C23A2837118433F2C80F84D3520F801" box="[715,862,1988,2013]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="inaequalis">
<emphasis id="B95705127118433F2C80F84D34A6F800" box="[715,728,1989,2012]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2CA2F84C3520F801" box="[745,862,1988,2013]" italics="true" pageId="3" pageNumber="304">inaequalis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C23A2837118433F2DEAF84D3252F801" box="[929,1068,1988,2013]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="caeruleus">
<emphasis id="B95705127118433F2DEAF84D35D0F800" box="[929,942,1989,2012]" italics="true" pageId="3" pageNumber="304">T</emphasis>
.
<emphasis id="B95705127118433F2DF4F84C3252F801" box="[959,1068,1988,2013]" italics="true" pageId="3" pageNumber="304">caeruleus</emphasis>
</taxonomicName>
) the only relevant data indicate capture in a Malaise trap. Specimens of
<taxonomicName id="4C23A283711F43382C11FF1134CFFF6C" box="[602,689,151,176]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="solisi">
<emphasis id="B9570512711F43382C11FF113419FF6C" box="[602,615,153,176]" italics="true" pageId="4" pageNumber="305">T</emphasis>
.
<emphasis id="B9570512711F43382C3DFF1F34CFFF6C" box="[630,689,151,176]" italics="true" pageId="4" pageNumber="305">solisi</emphasis>
</taxonomicName>
have been captured in both Malaise and flight intercept traps. The greatest detail in relevant natural history data is in that of
<taxonomicName id="4C23A283711F43382D50FF3535F5FF08" box="[795,907,188,212]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="B9570512711F43382D50FF353557FF08" box="[795,809,189,212]" italics="true" pageId="4" pageNumber="305">T</emphasis>
.
<emphasis id="B9570512711F43382D7CFF3435F5FF08" box="[823,907,188,212]" italics="true" pageId="4" pageNumber="305">festivus</emphasis>
</taxonomicName>
. In addition to being collected by Malaise trap, and in primary and secondary forest, there are three associations with plants. There is a generic “palm” association on one of the specimen labels; two other species are also indicated:
<taxonomicName id="4C23A283711F43382DC3FE8C32B6FEC1" authority="Loomis" authorityName="Loomis" box="[904,1224,260,285]" class="Liliopsida" family="Arecaceae" genus="Astrocaryum" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="305" phylum="Tracheophyta" rank="species" species="alatum">
<emphasis id="B9570512711F43382DC3FE8C3212FEC1" box="[904,1132,260,285]" italics="true" pageId="4" pageNumber="305">Astrocaryum alatum</emphasis>
Loomis
</taxonomicName>
(underside of leaf) and
<taxonomicName id="4C23A283711F43382E82FEAF344EFEE3" authority="(Mart.) Mart." authorityName="(Mart.) Mart." box="[201,560,295,320]" class="Insecta" family="Noctuidae" genus="Raphia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="taedigera">
<emphasis id="B9570512711F43382E82FEAF37F0FE9C" box="[201,398,295,320]" italics="true" pageId="4" pageNumber="305">Raphia taedigera</emphasis>
(Mart.) Mart.
</taxonomicName>
(on frond). Both species are members of the palm family (
<taxonomicName id="4C23A283711F43382A96FEA03326FE9C" box="[1245,1368,296,320]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="4" pageNumber="305" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
), and, although there are only two records, they may indicate an association of
<taxonomicName id="4C23A283711F43382D8FFEC5324AFEB8" box="[964,1076,332,356]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="festivus">
<emphasis id="B9570512711F43382D8FFEC535AFFEB8" box="[964,977,333,356]" italics="true" pageId="4" pageNumber="305">T</emphasis>
.
<emphasis id="B9570512711F43382DABFEC4324AFEB8" box="[992,1076,332,356]" italics="true" pageId="4" pageNumber="305">festivus</emphasis>
</taxonomicName>
with palms. Immature stages of
<taxonomicName id="4C23A283711F43382EDCFEE73741FE54" box="[151,319,367,392]" class="Insecta" family="Mycteridae" genus="Thisiomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="genus">
<emphasis id="B9570512711F43382EDCFEE73741FE54" box="[151,319,367,392]" italics="true" pageId="4" pageNumber="305">Thisiomorphus</emphasis>
</taxonomicName>
remain unknown; however, based on known larvae of other genera of
<taxonomicName id="4C23A283711F43382A39FEE73296FE54" box="[1138,1256,367,392]" class="Insecta" family="Mycteridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="subFamily" subFamily="Eurypinae">Eurypinae</taxonomicName>
, it seems likely that larvae are subcortical inhabitants of dead trees or within dead vegetation (see Pollock 1995; Pollock et al. 2000).
</paragraph>
</subSubSection>
</treatment>
</document>