treatments-xml/data/03/8A/35/038A3565BC0C1768FCA60F3BDEE0CFD9.xml

<document id="D0A07E9EE3089063C9AEF91C72E0A8D2" ID-CLB-Dataset="20463" ID-DOI="10.15553/c2016v712a10" ID-GBIF-Dataset="a90e897b-d6db-4a44-bc59-3fa9ece479d2" ID-ISSN="2235-3658" ID-Zenodo-Dep="5721713" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637687381177" checkinUser="carolina" docAuthor="Callmander, Martin W. &amp; Buerki, Sven" docDate="2016" docId="038A3565BC0C1768FCA60F3BDEE0CFD9" docLanguage="en" docName="Candollea.71.2.257-263.pdf" docOrigin="Candollea 71 (2)" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Benstonea fortuita Callm. &amp; Buerki 2016, spec. nova" docType="treatment" docVersion="6" lastPageNumber="259" masterDocId="FFB34D1DBC0D176AFF820E01DC58C844" masterDocTitle="Two new threatened species of Benstonea Callm. &amp; Buerki (Pandanaceae) from Sabah (Borneo, Malaysia)" masterLastPageNumber="263" masterPageNumber="257" pageNumber="258" updateTime="1699469569050" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="FD54D24F0736592862D1EE8D95D358E3">Two new threatened species of Benstonea Callm. &amp; Buerki (Pandanaceae) from Sabah (Borneo, Malaysia)</mods:title>
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<mods:name id="A11F915EAE0805699DA611A6D3E2FAB0" type="personal">
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<mods:namePart id="C0E9ACEEB21B754EE2757BB4A51752E7">Callmander, Martin W.</mods:namePart>
<mods:affiliation id="A7663C228130BF1048F4074B0AB005D9">Conservatoire et Jardin botaniques de la Ville de Genève, C. P. 60, 1292 Chambésy, Genève, Switzerland</mods:affiliation>
<mods:nameIdentifier id="75734B2CC7788ED4A7BD32E17E2BD88D" type="email">martin.callmander@ville-ge.ch</mods:nameIdentifier>
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<mods:namePart id="B13BE40214AAA8D280A77A2CE4AD97CD">Buerki, Sven</mods:namePart>
<mods:affiliation id="A4A9AA5653F15FAFA00592B54AE5C2CE">Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, U. K.</mods:affiliation>
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<mods:title id="D1FB3A0A687E7CF056F6214CF8F010F6">Candollea</mods:title>
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<mods:date id="91E86367AF27795EA70027AF03889BDB">2016</mods:date>
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<taxonomicName id="4C23FFF0BC0C176BFCA60F3BD8F8C916" ID-CoL="8WRPB" authority="Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" authorityYear="2016" box="[804,1184,314,338]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="fortuita" status="sp. nov.">
<emphasis id="B9575861BC0C176BFCA60F3BDFBEC916" bold="true" box="[804,998,314,338]" italics="true" pageId="1" pageNumber="258">Benstonea fortuita</emphasis>
Callm. &amp; Buerki
</taxonomicName>
, 
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<taxonomicNameLabel id="A264E51ABC0C176BFB280F3BD945C916" box="[1194,1309,314,338]" pageId="1" pageNumber="258" rank="species">spec. nova</taxonomicNameLabel>
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(
<figureCitation id="131898F6BC0C176BFAA90F3BD934C916" box="[1323,1388,314,338]" captionStart="Fig" captionStartId="3.[116,143,1860,1879]" captionTargetBox="[116,1454,279,1814]" captionTargetId="figure-45@3.[116,1454,276,1127]" captionTargetPageId="3" captionText="Fig. 2. – Benstonea fortuita Callm. &amp; Buerki. A. Habit; B. Infructescence; C. Close up of styles and stigmas. [Callmander et al.1197] [Photos: M. Callmander]" figureDoi="http://doi.org/10.5281/zenodo.5721720" httpUri="https://zenodo.org/record/5721720/files/figure.png" pageId="1" pageNumber="258">Fig. 2</figureCitation>
).
</heading>
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<paragraph id="8B9C8473BC0C176BFCC80F64D9F7C999" blockId="1.[842,1455,356,477]" pageId="1" pageNumber="258">
<emphasis id="B9575861BC0C176BFCC80F64DFCEC939" bold="true" box="[842,918,357,381]" pageId="1" pageNumber="258">Typus:</emphasis>
<materialsCitation id="3B4B8E2EBC0C176BFC190F64D8C7C999" ID-GBIF-Occurrence="3499009303" collectingDate="2014-10-09" collectionCode="G" collectorName="Callmander &amp; al." country="Malaysia" elevation="620" latitude="5.006111" location="Tenom" longLatPrecision="21" longitude="115.86472" municipality="Tenom Dist" pageId="1" pageNumber="258" specimenCode="G00341180" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<emphasis id="B9575861BC0C176BFC190F64D944C938" bold="true" box="[923,1308,356,381]" pageId="1" pageNumber="258">
<collectingCountry id="F334C4E3BC0C176BFC190F64D854C93F" box="[923,1036,357,381]" name="Malaysia" pageId="1" pageNumber="258">MALAYSIA</collectingCountry>
[BORNEO]. State 
<collectingRegion id="49E74A91BC0C176BFB530F65D94FC938" box="[1233,1303,356,380]" country="Malaysia" name="Sabah" pageId="1" pageNumber="258">Sabah</collectingRegion>
:
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., 
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from 
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on road to 
<location id="8EFCD2A8BC0C176BFB290F84D94BC9D9" LSID="urn:lsid:plazi:treatment:038A3565BC0C1768FCA60F3BDEE0CFD9:8EFCD2A8BC0C176BFB290F84D94BC9D9" box="[1195,1299,389,413]" country="Malaysia" latitude="5.006111" longLatPrecision="21" longitude="115.86472" municipality="Tenom Dist" name="Sipittang" pageId="1" pageNumber="258" stateProvince="Sabah">Sipittang</location>
, 
<location id="8EFCD2A8BC0C176BFAA20F84D925C9D9" LSID="urn:lsid:plazi:treatment:038A3565BC0C1768FCA60F3BDEE0CFD9:8EFCD2A8BC0C176BFAA20F84D925C9D9" box="[1312,1405,389,413]" country="Malaysia" latitude="5.006111" longLatPrecision="21" longitude="115.86472" municipality="Tenom Dist" name="Lumako" pageId="1" pageNumber="258" stateProvince="Sabah">Lumako</location>
FR, 
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<geoCoordinate id="EE17E2B4BC0C176BFC3E0FA5D867C9F8" box="[956,1087,420,444]" degrees="115" direction="east" minutes="51" orientation="longitude" pageId="1" pageNumber="258" precision="15" seconds="53" value="115.86472">115°51’53”E</geoCoordinate>
, 
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<elevation id="000E6340BC0C176BFBC80FA4D8D5C9F8" box="[1098,1165,421,445]" metricMagnitude="2" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="258" unit="m" value="620.0">620 m</elevation>
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, 
<date id="FF9DA2B3BC0C176BFB150FA4D8AFC9F9" box="[1175,1271,421,445]" pageId="1" pageNumber="258" value="2014-10-09">
<collectingDate id="EFD95B5BBC0C176BFB150FA4D8AFC9F9" box="[1175,1271,421,445]" pageId="1" pageNumber="258" value="2014-10-09">9.X.2014</collectingDate>
</date>
, fr., 
<emphasis id="B9575861BC0C176BFAA00FA4DFC3C999" italics="true" pageId="1" pageNumber="258">
<collectorName id="26D6E1A5BC0C176BFAA00FA4D9C0C9F9" box="[1314,1432,421,445]" pageId="1" pageNumber="258">Callmander</collectorName>
et 
<collectorName id="26D6E1A5BC0C176BFCC80FC4DF3BC999" box="[842,867,453,477]" pageId="1" pageNumber="258">al.</collectorName>
1197
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(
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: 
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[
<specimenCode id="DB852C08BC0C176BFB8C0FC4D8CDC999" box="[1038,1173,453,477]" pageId="1" pageNumber="258">G00341180</specimenCode>
]!
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; 
<materialsCitation id="3B4B8E2EBC0C176BFB2A0FC4D9FBC999" ID-GBIF-Occurrence="3499009301" box="[1192,1443,453,477]" collectingDate="2014-10-09" collectionCode="BM, SING, SAN" collectorName="Callmander &amp; al." country="Malaysia" elevation="620" latitude="5.006111" location="Tenom" longLatPrecision="21" longitude="115.86472" municipality="Tenom Dist" pageId="1" pageNumber="258" specimenCount="1" stateProvince="Sabah" typeStatus="isotype">
<typeStatus id="54983AD1BC0C176BFB2A0FC4D88AC999" box="[1192,1234,453,477]" pageId="1" pageNumber="258" type="isotype">iso-</typeStatus>
: 
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!, 
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!, 
<collectionCode id="ED321CB6BC0C176BFAEB0FC4D9FBC999" box="[1385,1443,453,477]" collectionName="SAN" pageId="1" pageNumber="258">SAN</collectionCode>
</materialsCitation>
!)
</paragraph>
</subSubSection>
<subSubSection id="C339D7F8BC0C176BFCC80FF1D959CDC3" pageId="1" pageNumber="258" type="description">
<paragraph id="8B9C8473BC0C176BFCC80FF1D8A2CAC3" blockId="1.[842,1455,495,647]" pageId="1" pageNumber="258">
<emphasis id="B9575861BC0C176BFCC80FF1D8A2CAC3" italics="true" pageId="1" pageNumber="258">
<taxonomicName id="4C23FFF0BC0C176BFCC80FF1D8E6CA43" authority="Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" box="[842,1214,495,519]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="fortuita">Benstonea fortuita Callm. &amp; Buerki</taxonomicName>
is distinguished among the genus by its subacaulescent habit, its leaf abruptly attenuate at the apex with a short apicule (0.5-1 cm), its ventral pleats armed, its stout acuminate style (3-4 mm), slightly curved proximally, and its solitary globose syncarp.
</emphasis>
</paragraph>
<paragraph id="8B9C8473BC0C176BFCC80CAED959CDC3" blockId="1.[804,1455,687,1415]" pageId="1" pageNumber="258">
Subacaulescent 
<emphasis id="B9575861BC0C176BFC7D0CAED861CA83" box="[1023,1081,687,711]" italics="true" pageId="1" pageNumber="258">shrub</emphasis>
&lt;1m tall. 
<emphasis id="B9575861BC0C176BFB2D0CB1D8A0CA83" box="[1199,1272,688,711]" italics="true" pageId="1" pageNumber="258">Leaves</emphasis>
glaucous, white below, 100-120 cm long, 4-5 cm wide in the middle, 3.5 cm wide near the sheath, abruptly attenuate in the upper part, shortly apiculate (c. 0.5-1 cm long), narrowing towards the base, subcoriaceous; marginal prickles beginning at 6-9 cm above the base and extending to the apex, antrorse, c. 2 mm in the lower third, 2-6 mm apart, to c. 1 mm in the mid third, 5-10 mm apart; minute in the distal third, c. 2 mm apart; ventral pleats armed in the last 25-28 cm of the leaf with minute prickles; sheath not auriculate, whitish. 
<emphasis id="B9575861BC0C176BFCA60DEEDFEECC43" box="[804,950,1007,1031]" italics="true" pageId="1" pageNumber="258">Infructescence</emphasis>
terminal, monosyncarpic, erect at maturity; syncarp in the middle of the clump of leaves, pale green, subglobose, c. 6 cm in diam.; peduncle straight, 10 cm long, 0.7 cm wide at apex, trigonous, white 
<emphasis id="B9575861BC0C176BFB4E0A51D947CC23" box="[1228,1311,1104,1127]" italics="true" pageId="1" pageNumber="258">in vivo,</emphasis>
longitudinal veins visible, covered with 4-5 bracts scars. 
<emphasis id="B9575861BC0C176BFB6D0A71D960CCC3" box="[1263,1336,1136,1159]" italics="true" pageId="1" pageNumber="258">Drupes</emphasis>
c. 150, 23- 25 mm high, 4-6 mm wide, c. 6 mm in depth, 5-6-angled; pileus pyramidal, pale to dark green 
<emphasis id="B9575861BC0C176BFB140AB1D8BACC83" box="[1174,1250,1200,1223]" italics="true" pageId="1" pageNumber="258">in vivo,</emphasis>
shrunk when dried, 8-10 mm high (excluding style). 
<emphasis id="B9575861BC0C176BFBF80ACED8F2CCA3" box="[1146,1194,1231,1255]" italics="true" pageId="1" pageNumber="258">Style</emphasis>
unique, stout acuminate, 3-4 mm long, slightly curved proximally; stigmatic groove brown 
<emphasis id="B9575861BC0C176BFCF20B11DF99CD63" box="[880,961,1296,1319]" italics="true" pageId="1" pageNumber="258">in vivo,</emphasis>
linear, positioned on abaxial side; endocarp c. 6 mm long, walls c. 0.4 mm thick; seed locule elliptic, 4 × 3 mm; apical mesocarp fibrous, discrete, c. 4 mm long; lower mesocarp fibrous. 
<emphasis id="B9575861BC0C176BFC6A0B71D8D5CDC3" box="[1000,1165,1391,1415]" italics="true" pageId="1" pageNumber="258">Staminate plant</emphasis>
unknown.
</paragraph>
</subSubSection>
<subSubSection id="C339D7F8BC0C176BFCC80BAED812CE03" pageId="1" pageNumber="258" type="etymology">
<paragraph id="8B9C8473BC0C176BFCC80BAED812CE03" blockId="1.[804,1455,1455,1607]" pageId="1" pageNumber="258">
<emphasis id="B9575861BC0C176BFCC80BAEDF9BCD83" box="[842,963,1455,1479]" italics="true" pageId="1" pageNumber="258">Etymology.</emphasis>
– This species is only known from a single population where only one fruiting individual was observed. We were very fortunate to come across this rare and discrete taxon and the species epithet reflects our feeling when we discovered this new 
<taxonomicName id="4C23FFF0BC0C176BFC5F0831D81ACE03" box="[989,1090,1584,1607]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="genus">
<emphasis id="B9575861BC0C176BFC5F0831D81ACE03" box="[989,1090,1584,1607]" italics="true" pageId="1" pageNumber="258">Benstonea</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C339D7F8BC0C176BFCC8086ED966CE83" pageId="1" pageNumber="258" type="distribution">
<paragraph id="8B9C8473BC0C176BFCC8086ED966CE83" blockId="1.[804,1455,1647,1735]" pageId="1" pageNumber="258">
<emphasis id="B9575861BC0C176BFCC8086ED94DCEC3" box="[842,1301,1647,1671]" italics="true" pageId="1" pageNumber="258">
Distribution and ecology. – 
<taxonomicName id="4C23FFF0BC0C176BFBDC0871D94DCEC3" box="[1118,1301,1647,1671]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="1" pageNumber="258" phylum="Tracheophyta" rank="species" species="fortuita">Benstonea fortuita</taxonomicName>
</emphasis>
is only known by few individuals in southwestern Sabah in the lowland Dipterocarp forest at an altitude of 600 m (
<figureCitation id="131898F6BC0C176BFB7008AED977CE83" box="[1266,1327,1711,1735]" captionStart="Fig" captionStartId="2.[140,167,1084,1103]" captionTargetBox="[159,787,297,980]" captionTargetId="figure-721@2.[157,789,294,982]" captionTargetPageId="2" captionText="Fig. 1. – Distribution of Benstonea fortuita Callm. &amp; Buerki (white star) and B. serpentinica Callm. &amp; Buerki (white circle) in Borneo. Abbreviations: 1. Crocker Range National Park; 2. Kinabalu Park." figureDoi="http://doi.org/10.5281/zenodo.5721718" httpUri="https://zenodo.org/record/5721718/files/figure.png" pageId="1" pageNumber="258">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C339D7F8BC0C1768FCC808F1DF51CD59" lastPageId="2" lastPageNumber="259" pageId="1" pageNumber="258" type="conservation">
<paragraph id="8B9C8473BC0C1768FCC808F1DF51CD59" blockId="1.[804,1454,1775,1927]" lastBlockId="2.[140,789,1221,1309]" lastPageId="2" lastPageNumber="259" pageId="1" pageNumber="258">
<emphasis id="B9575861BC0C176BFCC808F1D873CF43" box="[842,1067,1776,1799]" italics="true" pageId="1" pageNumber="258">Conservation status. –</emphasis>
Despite being known from a Forest Reserve [FR], this species is considered rare and seriously threatened. The only known population is located in the southern tip of the Lumako FR in a secondary Dipterocarp forest where few dominant trees still protect the understorey due to past logging. 
<taxonomicName id="4C23FFF0BC0F1768FEBC0AC4DDA5CC99" box="[318,509,1221,1245]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="fortuita">
<emphasis id="B9575861BC0F1768FEBC0AC4DDA5CC99" box="[318,509,1221,1245]" italics="true" pageId="2" pageNumber="259">Benstonea fortuita</emphasis>
</taxonomicName>
is assigned a preliminary status of “Critically Endangered” [CR B1ab(i,iii,iv,v)] following IUCN Red List Categories and Criteria (
<bibRefCitation id="EFB2F982BC0F1768FDF30B04DEA5CD59" author="IUCN" box="[625,765,1285,1309]" firstAuthor="IUCN" pageId="2" pageNumber="259" refId="ref3963" refString="IUCN (2012). IUCN Red List Categories and Criteria: Version 3.1. 2 nd ed. IUCN Species Survival Commission, Gland &amp; Cambridge." type="book" year="2012">IUCN, 2012</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5CD4FBBC0F1768FF0E0A3DDE90CCC6" ID-DOI="http://doi.org/10.5281/zenodo.5721718" ID-Zenodo-Dep="5721718" httpUri="https://zenodo.org/record/5721718/files/figure.png" pageId="2" pageNumber="259" startId="2.[140,167,1084,1103]" targetBox="[159,787,297,980]" targetPageId="2">
<paragraph id="8B9C8473BC0F1768FF0E0A3DDE28CC2D" blockId="2.[140,786,1084,1154]" pageId="2" pageNumber="259">
<emphasis id="B9575861BC0F1768FF0E0A3DDC93CC0B" bold="true" box="[140,203,1084,1103]" pageId="2" pageNumber="259">Fig. 1. –</emphasis>
Distribution of 
<taxonomicName id="4C23FFF0BC0F1768FED10A3CDEDCCC14" authority="Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" box="[339,644,1085,1104]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="fortuita">
<emphasis id="B9575861BC0F1768FED10A3CDDA0CC14" box="[339,504,1085,1104]" italics="true" pageId="2" pageNumber="259">Benstonea fortuita</emphasis>
Callm. &amp; Buerki
</taxonomicName>
(white star) and 
<taxonomicName id="4C23FFF0BC0F1768FF0E0A57DDC4CC2D" authority="Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" box="[140,412,1110,1129]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="serpentinica">
<emphasis id="B9575861BC0F1768FF0E0A57DD48CC2D" box="[140,272,1110,1129]" italics="true" pageId="2" pageNumber="259">B. serpentinica</emphasis>
Callm. &amp; Buerki
</taxonomicName>
(white circle) in Borneo.
</paragraph>
<paragraph id="8B9C8473BC0F1768FF0E0A6EDE90CCC6" blockId="2.[140,786,1084,1154]" box="[140,712,1135,1154]" pageId="2" pageNumber="259">Abbreviations: 1. Crocker Range National Park; 2. Kinabalu Park.</paragraph>
</caption>
<subSubSection id="C339D7F8BC0F1768FF330B44DEE0CFD9" pageId="2" pageNumber="259" type="discussion">
<paragraph id="8B9C8473BC0F1768FF330B44DEDECE59" blockId="2.[139,791,1349,1949]" pageId="2" pageNumber="259">
<emphasis id="B9575861BC0F1768FF330B44DD59CD18" box="[177,257,1349,1372]" italics="true" pageId="2" pageNumber="259">Notes. –</emphasis>
The overall morphology of the new species can be compared to: 
<taxonomicName id="4C23FFF0BC0F1768FE9F0B64DEECCD39" authority="(Ridl.) Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" baseAuthorityName="Ridl." box="[285,692,1381,1405]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="glaucophylla">
<emphasis id="B9575861BC0F1768FE9F0B64DDEACD39" box="[285,434,1381,1405]" italics="true" pageId="2" pageNumber="259">B. glaucophylla</emphasis>
(Ridl.) Callm. &amp; Buerki
</taxonomicName>
, 
<taxonomicName id="4C23FFF0BC0F1768FD3C0B64DDCACDD9" authority="(Ridl.) Callm. &amp; Buerki" authorityName="Callm. &amp; Buerki" baseAuthorityName="Ridl." class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="parva">
<emphasis id="B9575861BC0F1768FD3C0B64DF4ECD38" box="[702,790,1381,1404]" italics="true" pageId="2" pageNumber="259">B. parva</emphasis>
(Ridl.) Callm. &amp; Buerki
</taxonomicName>
, and 
<taxonomicName id="4C23FFF0BC0F1768FE4E0B84DD59CDF9" authority="(Ridl.) Callm. &amp; Buerki. All" authorityName="Callm. &amp; Buerki. All" baseAuthorityName="Ridl." class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="undulifolia">
<emphasis id="B9575861BC0F1768FE4E0B84DE02CDD9" box="[460,602,1413,1437]" italics="true" pageId="2" pageNumber="259">B. undulifolia</emphasis>
(Ridl.) Callm. &amp; Buerki. All
</taxonomicName>
three species share wide, abruptly attenuate leaves and a solitary syncarp with 
<taxonomicName id="4C23FFF0BC0F1768FE360BC4DE47CD99" box="[436,543,1477,1501]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="fortuita">
<emphasis id="B9575861BC0F1768FE360BC4DE47CD99" box="[436,543,1477,1501]" italics="true" pageId="2" pageNumber="259">B. fortuita</emphasis>
</taxonomicName>
. Nevertheless, the new species can easily be discriminated from these species by the combination of characters presented in 
<tableCitation id="C6A1B1C8BC0F1768FDAD0804DE27CE59" box="[559,639,1541,1565]" captionStart="Table 1" captionStartId="4.[139,191,286,305]" captionTargetBox="[147,1473,363,702]" captionTargetPageId="4" captionText="Table 1. – Key morphological differences between Benstonea fortuita Callm &amp; Buerki, B. glaucophylla (Ridl.) Callm. &amp; Buerki, B. parva (Ridl.) Callm. &amp; Buerki, and B. undulifolia (Ridl.) Callm. &amp; Buerki." httpUri="http://table.plazi.org/id/DF5CD4FBBC09176EFF090F1FDF4CC902" pageId="2" pageNumber="259" tableUuid="DF5CD4FBBC09176EFF090F1FDF4CC902">Table 1</tableCitation>
.
</paragraph>
<paragraph id="8B9C8473BC0F1768FF330824DEE0CFD9" blockId="2.[139,791,1349,1949]" pageId="2" pageNumber="259">
In their phylogeny, 
<bibRefCitation id="EFB2F982BC0F1768FE060824DE39CE79" author="BUERKI" box="[388,609,1573,1597]" etAl="et al." firstAuthor="BUERKI" pageId="2" pageNumber="259" pagination="211 - 223" refId="ref3548" refString="BUERKI, S., T. GALLAHER, T. BOOTH, G. BREWER, F. FOREST, J. T. PEREIRA &amp; M. W. CALLMANDER (2016). Biogeography and evolution of the screw-pine genus Benstonea Callm &amp; Buerki (Pandanaceae). Candollea 71: 211 - 223. DOI: http: // dx. doi. org / 10.15553 / c 2016 v 712 a 7" type="journal article" year="2016">BUERKI et al. (2016)</bibRefCitation>
showed that the new species 
<taxonomicName id="4C23FFF0BC0F1768FE960844DDDACE19" box="[276,386,1605,1629]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="fortuita">
<emphasis id="B9575861BC0F1768FE960844DDDACE19" box="[276,386,1605,1629]" italics="true" pageId="2" pageNumber="259">B. fortuita</emphasis>
</taxonomicName>
belongs to clade I together with the endemic Bornean 
<taxonomicName id="4C23FFF0BC0F1768FED30864DCA9CED9" authority="(Martelli) Callm. &amp; Buerki. Although" authorityName="Callm. &amp; Buerki. Although" baseAuthorityName="Martelli" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="platystigma">
<emphasis id="B9575861BC0F1768FED30864DDBCCE39" box="[337,484,1637,1661]" italics="true" pageId="2" pageNumber="259">B. platystigma</emphasis>
(Martelli) Callm. &amp; Buerki. Although
</taxonomicName>
both species have the same ecology (lowland humid forest), 
<taxonomicName id="4C23FFF0BC0F1768FF5D08A4DD13CEF9" box="[223,331,1701,1725]" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="fortuita">
<emphasis id="B9575861BC0F1768FF5D08A4DD13CEF9" box="[223,331,1701,1725]" italics="true" pageId="2" pageNumber="259">B. fortuita</emphasis>
</taxonomicName>
can easily be distinguished from 
<taxonomicName id="4C23FFF0BC0F1768FD3B08A4DC89CE98" class="Liliopsida" family="Pandanaceae" genus="Benstonea" kingdom="Plantae" order="Pandanales" pageId="2" pageNumber="259" phylum="Tracheophyta" rank="species" species="platystigma">
<emphasis id="B9575861BC0F1768FD3B08A4DC89CE98" italics="true" pageId="2" pageNumber="259">B. platystigma</emphasis>
</taxonomicName>
by its habit (subacaulescent shrub &lt;1 m tall vs. low shrub), leaves (wide, 4.5 cm and apex abruptly attenuate vs. narrow, 1.5 cm and apex gradually attenuate), style and stigmatic groove (stout acuminate, slightly curved proximally with a linear stigmatic groove vs. antrorsely bent down, flattened proximally with a subreniform groove) and fruits (pale green syncarp when ripe vs. bright red syncarp when ripe).
</paragraph>
</subSubSection>
</treatment>
</document>