699 lines
82 KiB
XML
699 lines
82 KiB
XML
<document id="C23B0D5442AA2984AEED5F9D4246FC77" ID-DOI="10.1649/0010-065x-64.4.347" ID-ISSN="1938-4394" ID-Zenodo-Dep="10102342" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630154024321" checkinUser="diego" docAuthor="Neita-Moreno, Jhon César & Ratcliffe, Brett C." docDate="2010" docId="03FFC037FFC1FFD7FF0DFCC5FB7084C1" docLanguage="en" docName="0010-065X-64.4.347.pdf.imf" docOrigin="The Coleopterists Bulletin 64 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-64.4.347" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Hemiphileurus elbitae Neita and Ratcliffe, new species" docType="treatment" docVersion="3" lastPageNumber="354" masterDocId="FFC6B84FFFC0FFD0FFB9FFA1FFE4875E" masterDocTitle="A New Species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) from Colombia, with Description of Its Larva and Pupa" masterLastPageNumber="355" masterPageNumber="347" pageNumber="348" updateTime="1699620528731" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5A25D83F00389C21D93AD153889EB3C8">A New Species of Hemiphileurus Kolbe (Coleoptera: Scarabaeidae: Dynastinae: Phileurini) from Colombia, with Description of Its Larva and Pupa</mods:title>
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<mods:namePart id="1BDF55E584A63DCBA16038370F90A815">Neita-Moreno, Jhon César</mods:namePart>
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<mods:namePart id="FAA757DC997AB2B352D54931C1619F5D">Ratcliffe, Brett C.</mods:namePart>
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<mods:title id="10E8870AB2FA46456F228AA1C8A0C56B">The Coleopterists Bulletin</mods:title>
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<mods:part id="517EEBF0702B7E5969B2CB4399D31D97">
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<mods:date id="32896AA32805FE05F8433737987228A3">2010</mods:date>
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<mods:number id="A64B0DC0C3E8ED9CD01359C288C6914C">2010-12-31</mods:number>
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<mods:identifier id="B3F6A3E42E7D657747F7CA5807E5D317" type="DOI">10.1649/0010-065x-64.4.347</mods:identifier>
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<paragraph id="8BE97121FFC1FFD1FF0DFCC5FE2184CA" blockId="1.[180,609,868,943]" pageId="1" pageNumber="348">
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<taxonomicName id="4C560AA2FFC1FFD1FF0DFCC5FDB88424" authority="Neita and Ratcliffe" authorityName="Neita and Ratcliffe" box="[180,604,868,890]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="elbitae" status="sp. nov.">
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<emphasis id="B922AD33FFC1FFD1FF0DFCC5FE6B8424" bold="true" box="[180,399,868,890]" italics="true" pageId="1" pageNumber="348">Hemiphileurus elbitae</emphasis>
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Neita and Ratcliffe
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2111048FFC1FFD1FEF6FCDEFE2184CA" box="[335,453,895,916]" pageId="1" pageNumber="348" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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(
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)
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</paragraph>
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<subSubSection id="C34C22AAFFC1FFD7FF16FC65FEE68487" lastPageId="7" lastPageNumber="354" pageId="1" pageNumber="348" type="materials_examined">
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<emphasis id="B922AD33FFC1FFD1FF16FC65FEA88487" bold="true" box="[175,332,964,985]" pageId="1" pageNumber="348">Type Material.</emphasis>
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<materialsCitation id="3B3E7B7CFFC1FFD1FEE0FC65FE0C83CA" collectorName="J. C. Neita Leg." country="Colombia" county="Tronco" elevation="10" latitude="8.624698" location="Choco" longLatPrecision="1" longitude="-77.34936" municipality="Serrania del Darien. En" pageId="1" pageNumber="348" specimenCount="3" typeStatus="paratype">
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H o l o t y p e m a l e l a b e l e d “
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<collectingCountry id="F34131B1FFC1FFD1FF1BFC7EFECC84AA" box="[162,296,991,1012]" name="Colombia" pageId="1" pageNumber="348">COLOMBIA</collectingCountry>
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:
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<location id="8E8927FAFFC1FFD1FE82FC7EFE9984AA" LSID="urn:lsid:plazi:treatment:03FFC037FFC1FFD7FF0DFCC5FB7084C1:8E8927FAFFC1FFD1FE82FC7EFE9984AA" box="[315,381,991,1012]" country="Colombia" county="Tronco" latitude="8.624698" longLatPrecision="1" longitude="-77.34936" municipality="Serrania del Darien. En" name="Choco" pageId="1" pageNumber="348">Chocó</location>
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,
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<location id="8E8927FAFFC1FFD1FE2BFC7EFD9E84AA" LSID="urn:lsid:plazi:treatment:03FFC037FFC1FFD7FF0DFCC5FB7084C1:8E8927FAFFC1FFD1FE2BFC7EFD9E84AA" box="[402,634,991,1012]" country="Colombia" county="Tronco" latitude="8.624698" longLatPrecision="1" longitude="-77.34936" municipality="Serrania del Darien. En" name="Municipio de Acandi" pageId="1" pageNumber="348">Municipio de Acandí</location>
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,
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<location id="8E8927FAFFC1FFD1FF2EFC5BFEE08351" LSID="urn:lsid:plazi:treatment:03FFC037FFC1FFD7FF0DFCC5FB7084C1:8E8927FAFFC1FFD1FF2EFC5BFEE08351" box="[151,260,1018,1039]" country="Colombia" county="Tronco" latitude="8.624698" longLatPrecision="1" longitude="-77.34936" municipality="Serrania del Darien. En" name="Capurgana" pageId="1" pageNumber="348">Capurganá</location>
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,
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<collectingMunicipality id="6B8DEB5BFFC1FFD1FEA0FC5BFDC08351" box="[281,548,1018,1039]" pageId="1" pageNumber="348">Serranía del Darién. En</collectingMunicipality>
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:
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<collectingCounty id="628809ADFFC1FFD1FD8EFC5BFD998351" box="[567,637,1018,1039]" pageId="1" pageNumber="348">Tronco</collectingCounty>
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descompuesto en pie de
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<taxonomicName id="4C560AA2FFC1FFD1FE23FBB5FEA5831A" authority="(Nees) Mez" authorityName="Mez" baseAuthorityName="Nees" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="1" pageNumber="348" phylum="Tracheophyta" rank="species" species="cernua">
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<emphasis id="B922AD33FFC1FFD1FE23FBB5FDD58377" box="[410,561,1044,1065]" italics="true" pageId="1" pageNumber="348">Ocotea cernua</emphasis>
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(Nees) Mez (Lauraceae)
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</taxonomicName>
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.
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<geoCoordinate id="EE6217E6FFC1FFD1FEF4FB8EFE31831A" box="[333,469,1070,1092]" degrees="8" direction="north" minutes="37" orientation="latitude" pageId="1" pageNumber="348" precision="1" seconds="28.91" value="8.624698">8°37′28.91″N</geoCoordinate>
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;
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.
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<quantity id="4CAEDCC4FFC1FFD1FF2EFBE8FF2D8300" box="[151,201,1097,1118]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="348" unit="m" value="10.0">
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<elevation id="007B9612FFC1FFD1FF2EFBE8FF2D8300" box="[151,201,1097,1118]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="348" unit="m" value="10.0">10 m</elevation>
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</quantity>
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alt. 13 ene de 2010.
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<collectorName id="26A314F7FFC1FFD1FE21FBE8FDD58300" box="[408,561,1097,1118]" pageId="1" pageNumber="348">J. C. Neita Leg.</collectorName>
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” Single adult male
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, one third instar
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<typeStatus id="54EDCF83FFC1FFD1FE44FBC5FDAB8327" box="[509,591,1124,1145]" pageId="1" pageNumber="348" type="paratype">paratype</typeStatus>
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, and one pupal
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<typeStatus id="54EDCF83FFC1FFD1FF44FBDEFEA983CA" box="[253,333,1151,1172]" pageId="1" pageNumber="348" type="paratype">paratype</typeStatus>
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with same data
|
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</materialsCitation>
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.
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</paragraph>
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<caption id="DF2921A9FFC1FFD1FD7CFC7CFB6A84AE" ID-DOI="http://doi.org/10.5281/zenodo.10102344" ID-Zenodo-Dep="10102344" box="[709,1166,989,1008]" httpUri="https://zenodo.org/record/10102344/files/figure.png" pageId="1" pageNumber="348" startId="1.[709,741,989,1008]" targetBox="[718,1125,209,948]" targetPageId="1">
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<emphasis id="B922AD33FFC1FFD1FD7CFC7CFCE584AE" bold="true" box="[709,769,989,1008]" pageId="1" pageNumber="348">Fig. 1.</emphasis>
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<taxonomicName id="4C560AA2FFC1FFD1FCAEFC7CFC3084AE" box="[791,980,989,1008]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="348" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC1FFD1FCAEFC7CFC3084AE" box="[791,980,989,1008]" italics="true" pageId="1" pageNumber="348">Hemiphileurus elbitae</emphasis>
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</taxonomicName>
|
||
, habitus of holotype.
|
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</paragraph>
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</caption>
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<paragraph id="8BE97121FFC1FFD1FF16FB38FE3B826D" blockId="1.[151,639,964,1729]" pageId="1" pageNumber="348">
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<materialsCitation id="3B3E7B7CFFC1FFD1FF16FB38FE3B826D" collectorName="Lincoln, NE" country="Colombia" county="Universidad Nacional" location="University of Nebraska State Museum" municipality="Museo Entomologico" pageId="1" pageNumber="348" specimenCount="3" stateProvince="Cesar" typeStatus="holotype">
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||
<typeStatus id="54EDCF83FFC1FFD1FF16FB38FEE283F0" box="[175,262,1177,1198]" pageId="1" pageNumber="348" type="holotype">Holotype</typeStatus>
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deposited at the
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(
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<collectorName id="26A314F7FFC1FFD1FE90FB15FE458397" box="[297,417,1204,1225]" pageId="1" pageNumber="348">Lincoln, NE</collectorName>
|
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). Adult
|
||
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|
||
deposited in the collection Jhon
|
||
<collectingRegion id="4992BFC3FFC1FFD1FE12FB6FFE0583BD" box="[427,481,1230,1251]" country="Colombia" name="Cesar" pageId="1" pageNumber="348">César</collectingRegion>
|
||
Neita (
|
||
<collectingRegion id="4992BFC3FFC1FFD1FD88FB6FFD9E83BD" box="[561,634,1230,1251]" country="Colombia" name="Bogota" pageId="1" pageNumber="348">Bogotá</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34131B1FFC1FFD1FF2EFB48FF1D83A0" box="[151,249,1257,1278]" name="Colombia" pageId="1" pageNumber="348">Colombia</collectingCountry>
|
||
). Larval and pupal
|
||
<typeStatus id="54EDCF83FFC1FFD1FE06FB48FDFD83A0" box="[447,537,1257,1278]" pageId="1" pageNumber="348" type="paratype">paratypes</typeStatus>
|
||
deposited in the
|
||
<collectingMunicipality id="6B8DEB5BFFC1FFD1FF6FFAA2FE408246" box="[214,420,1283,1304]" pageId="1" pageNumber="348">Museo Entomológico</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="628809ADFFC1FFD1FE09FAA2FD9A8246" box="[432,638,1283,1304]" pageId="1" pageNumber="348">Universidad Nacional</collectingCounty>
|
||
de
|
||
<collectingCountry id="F34131B1FFC1FFD1FF0FFABFFEF6826D" box="[182,274,1310,1331]" name="Colombia" pageId="1" pageNumber="348">Colombia</collectingCountry>
|
||
(
|
||
<collectingRegion id="4992BFC3FFC1FFD1FE9BFABFFE8D826D" box="[290,361,1310,1331]" country="Colombia" name="Bogota" pageId="1" pageNumber="348">Bogotá</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34131B1FFC1FFD1FECCFABFFE31826D" box="[373,469,1310,1331]" name="Colombia" pageId="1" pageNumber="348">Colombia</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC1FFD1FF16FA99FB92817C" blockId="1.[151,639,964,1729]" lastBlockId="1.[685,1172,1071,1729]" pageId="1" pageNumber="348">
|
||
<emphasis id="B922AD33FFC1FFD1FF16FA99FE6B8213" bold="true" box="[175,399,1336,1357]" pageId="1" pageNumber="348">
|
||
<typeStatus id="54EDCF83FFC1FFD1FF16FA99FEEF8213" box="[175,267,1336,1357]" pageId="1" pageNumber="348" type="holotype">Holotype</typeStatus>
|
||
Description.
|
||
</emphasis>
|
||
Length
|
||
<quantity id="4CAEDCC4FFC1FFD1FE5FFA99FDDA8213" box="[486,574,1336,1357]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" pageId="1" pageNumber="348" unit="mm" value="21.8">21.8 mm</quantity>
|
||
; width
|
||
<quantity id="4CAEDCC4FFC1FFD1FF2EFAF2FF118236" box="[151,245,1363,1384]" metricMagnitude="-2" metricUnit="m" metricValue="1.01" pageId="1" pageNumber="348" unit="mm" value="10.1">10.1 mm</quantity>
|
||
. Color black.
|
||
<emphasis id="B922AD33FFC1FFD1FE30FAF2FE248236" bold="true" box="[393,448,1363,1384]" pageId="1" pageNumber="348">Head</emphasis>
|
||
: Frons and vertex longitudinally concave, rugopunctate. Frons with distinct, longitudinal tubercle anteromedial of each eye. Clypeus triangular, apex strongly reflexed into acute spine, a small carina extending from apex to base of each tubercle; surface lateral of each carina nearly smooth. Sides of clypeus declivous. Interocular width equals 3.3 transverse eye diameters. Antenna 10-segmented, club subequal in length to segments 2–7. Mandibles arcuate on lateral edge, apices acute, reflexed.
|
||
<emphasis id="B922AD33FFC1FFD1FE66F9FCFDA2812C" bold="true" box="[479,582,1629,1650]" pageId="1" pageNumber="348">Pronotum</emphasis>
|
||
: Surface mostly densely punctate, a little less so on basal half; anterior angles rugopunctate. Disc at center with weak, longitudinal furrow in basal half only. Base with marginal line.
|
||
<emphasis id="B922AD33FFC1FFD1FC4DFB8EFBD3831A" bold="true" box="[1012,1079,1071,1092]" pageId="1" pageNumber="348">Elytra</emphasis>
|
||
: Surface between suture and humerus with 6 rows of punctures in weak furrows; punctures large, deep, ocellate-umbilicate. Intervals weakly convex, all of equal height, minutely shagreened. Sides with 5 rows of similar punctures.
|
||
<emphasis id="B922AD33FFC1FFD1FC6EFB12FBDD8396" bold="true" box="[983,1081,1203,1224]" pageId="1" pageNumber="348">Pygidium</emphasis>
|
||
: Surface densely punctate; punctures large, deep, setigerous; setae minute, pale. In lateral view, surface regularly convex.
|
||
<emphasis id="B922AD33FFC1FFD1FCB8FAA2FCD68246" bold="true" box="[769,818,1282,1304]" pageId="1" pageNumber="348">
|
||
Leg
|
||
<emphasis id="B922AD33FFC1FFD1FC90FAA3FCD68246" bold="true" box="[809,818,1282,1304]" italics="true" pageId="1" pageNumber="348">s</emphasis>
|
||
</emphasis>
|
||
: Protibia tridentate, teeth long, subequally spaced. Apex of metatibia with upper angle prolonged into sharp tooth with 7 stout spinules below tooth on left leg, 5 spinules below tooth on right leg. Apex of first metatarsomere prolonged into long spine.
|
||
<emphasis id="B922AD33FFC1FFD1FCE8FA29FC7782C3" bold="true" box="[849,915,1416,1437]" pageId="1" pageNumber="348">Venter</emphasis>
|
||
: Prosternal process erect, stout, subrectangular, apex nearly flat on top, posterior face at base with distinct ledge. Last sternite with transverse band of punctures at base and with a few sparse punctures elsewere.
|
||
<emphasis id="B922AD33FFC1FFD1FB9EFA53FB778159" bold="true" box="[1063,1171,1522,1543]" pageId="1" pageNumber="348">Parameres</emphasis>
|
||
(
|
||
<figureCitation id="136D6DA4FFC1FFD1FD0DF9ACFCF2817C" box="[692,790,1549,1570]" captionStart="Figs" captionStartId="2.[197,240,952,971]" captionTargetBox="[187,638,195,912]" captionTargetId="figure-303@2.[173,660,183,928]" captionTargetPageId="2" captionText="Figs. 2–3. Parameres of Hemiphileurus elbitae." figureDoi="http://doi.org/10.5281/zenodo.10102348" httpUri="https://zenodo.org/record/10102348/files/figure.png" pageId="1" pageNumber="348">Figs. 2, 3</figureCitation>
|
||
): Distinctly hollowed in the center.
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC1FFD1FD7CF986FCEC81D2" blockId="1.[685,1172,1071,1729]" pageId="1" pageNumber="348">
|
||
<emphasis id="B922AD33FFC1FFD1FD7CF986FC888162" bold="true" box="[709,876,1575,1596]" pageId="1" pageNumber="348">
|
||
Adult
|
||
<typeStatus id="54EDCF83FFC1FFD1FCB0F986FC8C8162" box="[777,872,1575,1596]" pageId="1" pageNumber="348" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male. Length
|
||
<quantity id="4CAEDCC4FFC1FFD1FC47F986FBB18162" box="[1022,1109,1575,1596]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="1" pageNumber="348" unit="mm" value="22.9">22.9 mm</quantity>
|
||
; width
|
||
<quantity id="4CAEDCC4FFC1FFD1FD14F9E3FD1A8109" box="[685,766,1602,1623]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="348" unit="mm" value="9.5">9.5 mm</quantity>
|
||
. As
|
||
<typeStatus id="54EDCF83FFC1FFD1FC89F9E3FC608109" box="[816,900,1602,1623]" pageId="1" pageNumber="348" type="holotype">holotype</typeStatus>
|
||
except apex of meso- and metatibiae with 4–6 spinules below tooth on right leg.
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC1FFD1FD7CF933FB70819F" blockId="1.[685,1172,1071,1729]" pageId="1" pageNumber="348">
|
||
<emphasis id="B922AD33FFC1FFD1FD7CF933FBC781F9" bold="true" box="[709,1059,1682,1703]" pageId="1" pageNumber="348">
|
||
Third Instar
|
||
<typeStatus id="54EDCF83FFC1FFD1FCF0F933FC4081F9" box="[841,932,1682,1703]" pageId="1" pageNumber="348" type="paratype">Paratype</typeStatus>
|
||
Description.
|
||
</emphasis>
|
||
One larva, 3 exuviae.
|
||
<emphasis id="B922AD33FFC1FFD1FC9EF90DFC63819F" bold="true" box="[807,903,1708,1729]" pageId="1" pageNumber="348">Cranium</emphasis>
|
||
(
|
||
<figureCitation id="136D6DA4FFC1FFD1FC25F90DFC04819F" box="[924,992,1708,1729]" captionStart="Fig" captionStartId="2.[731,763,750,769]" captionTargetBox="[726,1176,217,706]" captionTargetId="figure-320@2.[707,1194,183,727]" captionTargetPageId="2" captionText="Fig. 4. Cranium of Hemiphileurus elbitae, third instar." figureDoi="http://doi.org/10.5281/zenodo.10102346" httpUri="https://zenodo.org/record/10102346/files/figure.png" pageId="1" pageNumber="348">Fig. 4</figureCitation>
|
||
): Width of head
|
||
</paragraph>
|
||
<caption id="DF2921A9FFC2FFD2FD62FD4FFB4E845F" ID-DOI="http://doi.org/10.5281/zenodo.10102346" ID-Zenodo-Dep="10102346" box="[731,1194,750,769]" httpUri="https://zenodo.org/record/10102346/files/figure.png" pageId="2" pageNumber="349" startId="2.[731,763,750,769]" targetBox="[726,1176,217,706]" targetPageId="2">
|
||
<paragraph id="8BE97121FFC2FFD2FD62FD4FFB4E845F" blockId="2.[731,1194,750,769]" box="[731,1194,750,769]" pageId="2" pageNumber="349">
|
||
<emphasis id="B922AD33FFC2FFD2FD62FD4FFCF3845F" bold="true" box="[731,791,750,769]" pageId="2" pageNumber="349">Fig. 4.</emphasis>
|
||
Cranium of
|
||
<taxonomicName id="4C560AA2FFC2FFD2FC37FD4FFBA7845F" box="[910,1091,750,769]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC2FFD2FC37FD4FFBA7845F" box="[910,1091,750,769]" italics="true" pageId="2" pageNumber="349">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
, third instar.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2921A9FFC2FFD2FF7CFC19FD778495" ID-DOI="http://doi.org/10.5281/zenodo.10102348" ID-Zenodo-Dep="10102348" box="[197,659,951,971]" httpUri="https://zenodo.org/record/10102348/files/figure.png" pageId="2" pageNumber="349" startId="2.[197,240,952,971]" targetBox="[187,638,195,912]" targetPageId="2">
|
||
<paragraph id="8BE97121FFC2FFD2FF7CFC19FD778495" blockId="2.[173,659,951,995]" box="[197,659,951,971]" pageId="2" pageNumber="349">
|
||
<emphasis id="B922AD33FFC2FFD2FF7CFC19FEC38495" bold="true" box="[197,295,951,971]" pageId="2" pageNumber="349">Figs. 2–3.</emphasis>
|
||
Parameres of
|
||
<taxonomicName id="4C560AA2FFC2FFD2FE79FC16FD698494" box="[448,653,951,970]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC2FFD2FE79FC16FD698494" box="[448,653,951,970]" italics="true" pageId="2" pageNumber="349">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE97121FFC2FFD2FF14FC71FE2F84BD" blockId="2.[173,659,951,995]" box="[173,459,976,995]" pageId="2" pageNumber="349">
|
||
<emphasis id="B922AD33FFC2FFD2FF14FC71FF2484BD" bold="true" box="[173,192,976,995]" pageId="2" pageNumber="349">2)</emphasis>
|
||
Caudal view;
|
||
<emphasis id="B922AD33FFC2FFD2FEF9FC71FEB784BD" bold="true" box="[320,339,976,995]" pageId="2" pageNumber="349">3)</emphasis>
|
||
Lateral view.
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC2FFD2FF14FBB5FB4D83FB" blockId="2.[173,660,1043,1729]" lastBlockId="2.[707,1193,823,1189]" pageId="2" pageNumber="349">
|
||
capsule 6.07–7.00 mm. Color reddish brown. Surface strongly punctate. Frontal suture and clypeofrontal suture distinct.
|
||
<emphasis id="B922AD33FFC2FFD2FE75FBE8FDE38300" bold="true" box="[460,519,1097,1118]" pageId="2" pageNumber="349">Frons</emphasis>
|
||
(
|
||
<figureCitation id="136D6DA4FFC2FFD2FDAEFBE8FDB28300" box="[535,598,1097,1118]" captionStart="Fig" captionStartId="2.[731,763,750,769]" captionTargetBox="[726,1176,217,706]" captionTargetId="figure-320@2.[707,1194,183,727]" captionTargetPageId="2" captionText="Fig. 4. Cranium of Hemiphileurus elbitae, third instar." figureDoi="http://doi.org/10.5281/zenodo.10102346" httpUri="https://zenodo.org/record/10102346/files/figure.png" pageId="2" pageNumber="349">Fig. 4</figureCitation>
|
||
): Surface with 1 exterior frontal seta and 1 posterior frontal seta on each side; each anterior angle with 2 setae; remaining cranial surface with 3 dorsoepicranial setae, 10 epicranial setae distributed irregularly, and 7–8 paraocellar setae on each side. Anterior frontal setae absent. Ocellus present.
|
||
<emphasis id="B922AD33FFC2FFD2FF14FAA2FF1B8246" bold="true" box="[173,255,1283,1304]" pageId="2" pageNumber="349">Clypeus</emphasis>
|
||
: Form trapezoidal. Surface of postclypeus reddish-brown, well sclerotized, densely punctuate; surface of preclypeus light brown, with 2 central setae and 2 lateral seta on each side.
|
||
<emphasis id="B922AD33FFC2FFD2FD81FAF2FD688236" bold="true" box="[568,652,1363,1384]" pageId="2" pageNumber="349">Labrum</emphasis>
|
||
: Surface rugopunctate, slightly asymmetrical, with 6 posterior setae, 3 lateral setae on each side and 4 central setae.
|
||
<emphasis id="B922AD33FFC2FFD2FE84FA03FE5082E9" bold="true" box="[317,436,1442,1463]" pageId="2" pageNumber="349">Epipharynx</emphasis>
|
||
(
|
||
<figureCitation id="136D6DA4FFC2FFD2FE78FA03FE1E82E9" box="[449,506,1442,1463]" captionStart="Fig" captionStartId="2.[731,763,1686,1705]" captionTargetBox="[723,1183,1242,1655]" captionTargetId="figure-331@2.[707,1193,1230,1663]" captionTargetPageId="2" captionText="Fig. 5. Epipharynx of Hemiphileurus elbitae, third instar." figureDoi="http://doi.org/10.5281/zenodo.10102350" httpUri="https://zenodo.org/record/10102350/files/figure.png" pageId="2" pageNumber="349">Fig. 5</figureCitation>
|
||
): Form suboval, asymmetrical, lateral edges angulate. Haptomeral process prominent, entire. Right chaetoparia with 57 setae; left chaetoparia with 43 setae, with some sensillae between the setae. Acroparia with 15 straight, long, thick setae; corypha with 5 slender, long setae; right acanthoparia with 8 short, curved, spine-like setae; left acanthoparia with 9 short, curved, spine-like setae. Pedium longer than wide, without setae. Dexiotorma narrow, elongate; laeotorma slightly shorter than dexiotorma; epitorma curved onto disc of pedium, pternotorma rounded. Dexiophoba absent; laephoba poorly developed; haptolochus and inner side of laeotorma with 13 small, slender setae. Sclerotized plate of right nesium elongate, truncate at apex; sense cone on left nesium represented by longitudinal, wellsclerotized plate, apex with 3 sensilla. Crepis poorly defined.
|
||
<emphasis id="B922AD33FFC2FFD2FCA3FC50FC588358" bold="true" box="[794,956,1009,1030]" pageId="2" pageNumber="349">Right mandible</emphasis>
|
||
(
|
||
<figureCitation id="136D6DA4FFC2FFD2FC75FC50FBD38358" box="[972,1079,1008,1030]" captionStart="Figs" captionStartId="3.[175,215,1362,1381]" captionTargetBox="[156,1131,220,1311]" captionTargetId="figure-187@3.[151,1171,183,1338]" captionTargetPageId="3" captionText="Figs. 6–7. Hemiphileurus elbitae, third instar. 6a) Right mandible, dorsal view; 6b) Right mandible, ventral view;" figureDoi="http://doi.org/10.5281/zenodo.10102352" httpUri="https://zenodo.org/record/10102352/files/figure.png" pageId="2" pageNumber="349">Figs. 6a–b</figureCitation>
|
||
): Scissorial area with blade-like, apical tooth (S
|
||
<subScript id="17D27364FFC2FFD2FB9FFBB4FBCB837D" attach="left" box="[1062,1071,1045,1059]" fontSize="6" pageId="2" pageNumber="349">1</subScript>
|
||
+ S
|
||
<subScript id="17D27364FFC2FFD2FBE2FBB4FB80837D" attach="left" box="[1115,1124,1045,1059]" fontSize="6" pageId="2" pageNumber="349">2</subScript>
|
||
fused) and 1 small, rounded tooth (S
|
||
<subScript id="17D27364FFC2FFD2FC55FB8EFC118363" attach="right" box="[1004,1013,1071,1085]" fontSize="6" pageId="2" pageNumber="349">3</subScript>
|
||
) behind scissorial notch. Scrobe with 17 slender, long setae. Dorsal surface with line of 12 slender, long setae. Ventral surface with elongate-oval, stridulatory area formed by 37 narrowly separated ridges. Ventral process
|
||
</paragraph>
|
||
<caption id="DF2921A9FFC2FFD2FD62F937FD13819F" ID-DOI="http://doi.org/10.5281/zenodo.10102350" ID-Zenodo-Dep="10102350" httpUri="https://zenodo.org/record/10102350/files/figure.png" pageId="2" pageNumber="349" startId="2.[731,763,1686,1705]" targetBox="[723,1183,1242,1655]" targetPageId="2">
|
||
<paragraph id="8BE97121FFC2FFD2FD62F937FD13819F" blockId="2.[707,1193,1686,1729]" pageId="2" pageNumber="349">
|
||
<emphasis id="B922AD33FFC2FFD2FD62F937FCFC81F7" bold="true" box="[731,792,1686,1705]" pageId="2" pageNumber="349">Fig. 5.</emphasis>
|
||
Epipharynx of
|
||
<taxonomicName id="4C560AA2FFC2FFD2FC0AF937FB9081F7" box="[947,1140,1686,1705]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="349" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC2FFD2FC0AF937FB9081F7" box="[947,1140,1686,1705]" italics="true" pageId="2" pageNumber="349">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
, third instar.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2921A9FFC3FFD3FF16FAF3FB70823B" ID-DOI="http://doi.org/10.5281/zenodo.10102352" ID-Zenodo-Dep="10102352" box="[175,1172,1361,1381]" httpUri="https://zenodo.org/record/10102352/files/figure.png" pageId="3" pageNumber="350" startId="3.[175,215,1362,1381]" targetBox="[156,1131,220,1311]" targetPageId="3">
|
||
<paragraph id="8BE97121FFC3FFD3FF16FAF3FB70823B" blockId="3.[151,1172,1361,1404]" box="[175,1172,1361,1381]" pageId="3" pageNumber="350">
|
||
<emphasis id="B922AD33FFC3FFD3FF16FAF3FEEE823B" bold="true" box="[175,266,1361,1381]" pageId="3" pageNumber="350">Figs. 6–7.</emphasis>
|
||
<taxonomicName id="4C560AA2FFC3FFD3FEA6FAF0FE38823A" box="[287,476,1361,1380]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="350" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC3FFD3FEA6FAF0FE38823A" box="[287,476,1361,1380]" italics="true" pageId="3" pageNumber="350">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
, third instar.
|
||
<emphasis id="B922AD33FFC3FFD3FDF7FAF3FD89823B" bold="true" box="[590,621,1362,1381]" pageId="3" pageNumber="350">6a)</emphasis>
|
||
Right mandible, dorsal view;
|
||
<emphasis id="B922AD33FFC3FFD3FCC9FAF3FC74823B" bold="true" box="[880,912,1362,1381]" pageId="3" pageNumber="350">6b)</emphasis>
|
||
Right mandible, ventral view;
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE97121FFC3FFD3FF2EFAC8FD288222" blockId="3.[151,1172,1361,1404]" box="[151,716,1385,1404]" pageId="3" pageNumber="350">
|
||
<emphasis id="B922AD33FFC3FFD3FF2EFAC8FF528222" bold="true" box="[151,182,1385,1404]" pageId="3" pageNumber="350">7a)</emphasis>
|
||
Left mandible, dorsal view;
|
||
<emphasis id="B922AD33FFC3FFD3FE08FAC8FE358222" bold="true" box="[433,465,1385,1404]" pageId="3" pageNumber="350">7b)</emphasis>
|
||
Left mandible, ventral view.
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC3FFD5FF2EFA1CFB708302" blockId="3.[151,639,1469,1729]" lastBlockId="5.[685,1173,802,1116]" lastPageId="5" lastPageNumber="352" pageId="3" pageNumber="350">
|
||
well-developed, rounded, with many asperites. Brustia with 8 stout, long setae. Calyx large, with 6 basolateral setae. Molar area with 3 wide, convex, ridged lobes (M
|
||
<subScript id="17D27364FFC3FFD3FE8AF9B7FED8817A" attach="left" box="[307,316,1558,1572]" fontSize="6" pageId="3" pageNumber="350">1</subScript>
|
||
–
|
||
<subScript id="17D27364FFC3FFD3FEF1F9B7FEB5817A" attach="right" box="[328,337,1558,1572]" fontSize="6" pageId="3" pageNumber="350">3</subScript>
|
||
).
|
||
<emphasis id="B922AD33FFC3FFD3FED1F9ACFE13817C" bold="true" box="[360,503,1549,1570]" pageId="3" pageNumber="350">Left mandible</emphasis>
|
||
(
|
||
<figureCitation id="136D6DA4FFC3FFD3FDBEF9ACFD96817C" box="[519,626,1549,1570]" captionStart="Figs" captionStartId="3.[175,215,1362,1381]" captionTargetBox="[156,1131,220,1311]" captionTargetId="figure-187@3.[151,1171,183,1338]" captionTargetPageId="3" captionText="Figs. 6–7. Hemiphileurus elbitae, third instar. 6a) Right mandible, dorsal view; 6b) Right mandible, ventral view;" figureDoi="http://doi.org/10.5281/zenodo.10102352" httpUri="https://zenodo.org/record/10102352/files/figure.png" pageId="3" pageNumber="350">Figs. 7a, b</figureCitation>
|
||
): Form falcate. Scissorial region with 4 teeth, basal tooth separated from teeth S
|
||
<subScript id="17D27364FFC3FFD3FE10F9EAFE568107" attach="left" box="[425,434,1611,1625]" fontSize="6" pageId="3" pageNumber="350">1</subScript>
|
||
–S
|
||
<subScript id="17D27364FFC3FFD3FE75F9EAFE318107" attach="left" box="[460,469,1611,1625]" fontSize="6" pageId="3" pageNumber="350">3</subScript>
|
||
; S
|
||
<subScript id="17D27364FFC3FFD3FE48F9EAFE1E8107" attach="left" box="[497,506,1611,1625]" fontSize="6" pageId="3" pageNumber="350">1</subScript>
|
||
and S
|
||
<subScript id="17D27364FFC3FFD3FD82F9EAFDA08107" attach="left" box="[571,580,1611,1625]" fontSize="6" pageId="3" pageNumber="350">2</subScript>
|
||
separated from S
|
||
<subScript id="17D27364FFC3FFD3FEA5F9C7FEC1812A" attach="left" box="[284,293,1638,1652]" fontSize="6" pageId="3" pageNumber="350">3</subScript>
|
||
by scissorial notch. Scrobe with 15 slender, long setae. Dorsal surface with line of 8 slender, moderately long setae; acia welldeveloped, sharp, with 6 basolateral setae; dorso- molar area with row of 13 stout, slender, moderately long setae. Ventral surface with elongate-oval stridulatory area formed by 40 narrowly separated ridges; ventral process well-developed, rounded, with many asperites. Brustia with 14 stout, long setae arranged in form of a “U”. Molar area with 3 lobes, first molar lobe (M
|
||
<subScript id="17D27364FFC3FFD3FC8FF9C7FCDB812A" attach="left" box="[822,831,1638,1652]" fontSize="6" pageId="3" pageNumber="350">1</subScript>
|
||
) large.
|
||
<emphasis id="B922AD33FFC3FFD3FC31F9FCFC31812C" bold="true" box="[904,981,1629,1650]" pageId="3" pageNumber="350">Maxilla</emphasis>
|
||
(
|
||
<figureCitation id="136D6DA4FFC3FFD3FC5BF9FCFBAD812C" box="[994,1097,1628,1650]" captionStart-0="Fig" captionStart-1="Figs" captionStartId-0="4.[197,229,709,728]" captionStartId-1="4.[197,239,1662,1681]" captionTargetBox-0="[209,618,198,653]" captionTargetBox-1="[205,639,1227,1619]" captionTargetId-0="figure-396@4.[173,660,183,686]" captionTargetId-1="figure-413@4.[173,660,1205,1639]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionText-0="Fig. 8. Hemiphileurus elbitae, third instar. 8a) Hypopharynx, dorsal view; 8b) Maxilla, dorsal view." captionText-1="Figs. 9–10. Hemiphileurus elbitae, third instar. 9) Maxilla, inner view showing unci; 10) Stridulatory teeth of maxilla." figureDoi-0="http://doi.org/10.5281/zenodo.10102354" figureDoi-1="http://doi.org/10.5281/zenodo.10102356" httpUri-0="https://zenodo.org/record/10102354/files/figure.png" httpUri-1="https://zenodo.org/record/10102356/files/figure.png" pageId="3" pageNumber="350">Figs. 8–10</figureCitation>
|
||
). Cardo subrectangular. Galea with 6 stout setae and 1 well developed uncus at apex. Lacinia with many stout setae and 3 unci fused at their bases, 1
|
||
<superScript id="7C23DC69FFC3FFD3FBB2F906FBF381EB" attach="left" box="[1035,1047,1703,1717]" fontSize="6" pageId="3" pageNumber="350">st</superScript>
|
||
and 2
|
||
<superScript id="7C23DC69FFC3FFD3FBEBF906FB8081EB" attach="left" box="[1106,1124,1703,1717]" fontSize="6" pageId="3" pageNumber="350">nd</superScript>
|
||
unci separated from third. Maxillary palpus 4-segmented, segment 4 twice as long as segment 2. Stridulatory area (
|
||
<figureCitation id="136D6DA4FFC4FFD4FF67FCC0FEC48428" box="[222,288,865,886]" captionStart="Figs" captionStartId="4.[197,239,1662,1681]" captionTargetBox="[205,639,1227,1619]" captionTargetId="figure-413@4.[173,660,1205,1639]" captionTargetPageId="4" captionText="Figs. 9–10. Hemiphileurus elbitae, third instar. 9) Maxilla, inner view showing unci; 10) Stridulatory teeth of maxilla." figureDoi="http://doi.org/10.5281/zenodo.10102356" httpUri="https://zenodo.org/record/10102356/files/figure.png" pageId="4" pageNumber="351">Fig. 10</figureCitation>
|
||
) with 6–7 blunt, truncate ridges, anterior process truncate.
|
||
<emphasis id="B922AD33FFC4FFD4FEE5FCDAFE0F84CE" bold="true" box="[348,491,891,912]" pageId="4" pageNumber="351">Hypopharynx</emphasis>
|
||
(
|
||
<figureCitation id="136D6DA4FFC4FFD4FE45FCDAFDDF84CE" box="[508,571,891,912]" captionStart="Fig" captionStartId="4.[197,229,709,728]" captionTargetBox="[209,618,198,653]" captionTargetId="figure-396@4.[173,660,183,686]" captionTargetPageId="4" captionText="Fig. 8. Hemiphileurus elbitae, third instar. 8a) Hypopharynx, dorsal view; 8b) Maxilla, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10102354" httpUri="https://zenodo.org/record/10102354/files/figure.png" pageId="4" pageNumber="351">Fig. 8</figureCitation>
|
||
): Glossa with 34 slender, long setae and 56 stout, short setae. Hypopharyngeal sclerome asymmetrical, concave medially, with acute dorsal process. Left lateral lobe with 22 slender, moderately long setae; right lateral lobe with 18 slender, moderately long setae. Left margin with row of 22 stout, moderately long setae directed toward center of sclerome and 4 setae near base of sclerome.
|
||
<emphasis id="B922AD33FFC4FFD4FEE6FBF1FE52833B" bold="true" box="[351,438,1104,1125]" pageId="4" pageNumber="351">Antenna</emphasis>
|
||
: Four-segmented, segments 1–2 subequal in length, each slightly longer than segment 3; terminal segment about half length of segment 2. Apical segment on dorsal surface with 5 sensory spots (
|
||
<figureCitation id="136D6DA4FFC4FFD4FC2CFF7CFC3E87AC" box="[917,986,221,242]" captionStart="Figs" captionStartId="4.[731,775,1662,1681]" captionTargetBox="[768,1146,1129,1603]" captionTargetId="figure-435@4.[707,1194,1098,1639]" captionTargetPageId="4" captionText="Figs. 11–12. Last antennomere of third instar Hemiphileurus elbitae, showing sensory spots. 11) Dorsal view; 12) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10102358" httpUri="https://zenodo.org/record/10102358/files/figure.png" pageId="4" pageNumber="351">Fig. 11</figureCitation>
|
||
), ventral surface with 7 sensory spots (
|
||
<figureCitation id="136D6DA4FFC4FFD4FCDBFF56FC4D8652" box="[866,937,247,268]" captionStart="Figs" captionStartId="4.[731,775,1662,1681]" captionTargetBox="[768,1146,1129,1603]" captionTargetId="figure-435@4.[707,1194,1098,1639]" captionTargetPageId="4" captionText="Figs. 11–12. Last antennomere of third instar Hemiphileurus elbitae, showing sensory spots. 11) Dorsal view; 12) Ventral view." figureDoi="http://doi.org/10.5281/zenodo.10102358" httpUri="https://zenodo.org/record/10102358/files/figure.png" pageId="4" pageNumber="351">Fig. 12</figureCitation>
|
||
).
|
||
<emphasis id="B922AD33FFC4FFD4FC02FF56FBE18652" bold="true" box="[955,1029,247,268]" pageId="4" pageNumber="351">Thorax</emphasis>
|
||
: Pronotum wide, irregularly and weakly sclerotized, with 6 slender, long setae. Prothoracic spiracle (
|
||
<figureCitation id="136D6DA4FFC4FFD4FC4CFE8DFBDE861F" box="[1013,1082,300,321]" captionStart="Figs" captionStartId="5.[175,217,1170,1189]" captionTargetBox="[194,604,219,1121]" captionTargetId="figure-237@5.[151,638,191,1153]" captionTargetPageId="5" captionText="Figs. 13–14. Hemiphileurus elbitae, third instar." figureDoi="http://doi.org/10.5281/zenodo.10102362" httpUri="https://zenodo.org/record/10102362/files/figure.png" pageId="4" pageNumber="351">Fig. 13</figureCitation>
|
||
)
|
||
<quantity id="4CAEDCC4FFC4FFD4FBF1FE8DFB4D861F" box="[1096,1193,300,321]" metricMagnitude="-4" metricUnit="m" metricValue="2.37" pageId="4" pageNumber="351" unit="mm" value="0.237">0.237 mm</quantity>
|
||
long,
|
||
<quantity id="4CAEDCC4FFC4FFD4FD40FEE6FCBD8602" box="[761,857,327,348]" metricMagnitude="-4" metricUnit="m" metricValue="1.58" pageId="4" pageNumber="351" unit="mm" value="0.158">0.158 mm</quantity>
|
||
wide; respiratory plate dark brown, regularly shaped as a closed “C”, spiracular bulla rounded, slightly prominent; distance between respiratory lobes less than diameter of bulla; plate with 25 holes across diameter at middle, holes with irregular edges (
|
||
<figureCitation id="136D6DA4FFC4FFD4FCE2FE6DFC4586BF" box="[859,929,460,481]" captionStart="Figs" captionStartId="5.[175,217,1170,1189]" captionTargetBox="[194,604,219,1121]" captionTargetId="figure-237@5.[151,638,191,1153]" captionTargetPageId="5" captionText="Figs. 13–14. Hemiphileurus elbitae, third instar." figureDoi="http://doi.org/10.5281/zenodo.10102362" httpUri="https://zenodo.org/record/10102362/files/figure.png" pageId="4" pageNumber="351">Fig. 14</figureCitation>
|
||
). Dorsum of prothorax with transverse row of 6 long, slender setae and 2 spinelike setae. Mesoprescutum with transverse, irregular row of 14 long, slender setae; mesoscutellum with transverse row of 10 long, slender setae and 33 spine-like setae. Metaprescutum without setae; metascutellum with 6 long, slender setae and 22 stout, spine-like setae.
|
||
<emphasis id="B922AD33FFC4FFD4FC25FD27FC2F85C5" bold="true" box="[924,971,646,667]" pageId="4" pageNumber="351">Legs</emphasis>
|
||
: Tarsal claws (
|
||
<figureCitation id="136D6DA4FFC4FFD4FBE5FD27FB4685C5" box="[1116,1186,646,667]" captionStart="Fig" captionStartId="5.[709,744,706,725]" captionTargetBox="[724,1137,189,649]" captionTargetId="figure-253@5.[685,1172,183,682]" captionTargetPageId="5" captionText="Fig. 15. Lateral view of metatarsal claw of Hemiphileurus elbitae, third instar." figureDoi="http://doi.org/10.5281/zenodo.10102360" httpUri="https://zenodo.org/record/10102360/files/figure.png" pageId="4" pageNumber="351">Fig. 15</figureCitation>
|
||
) with enlarged apical process, 1 basoexternal seta, and 1 internal, preapical seta. Tarsal claw on pro- and mesothoracic legs longer than those of metathoracic legs. Coxa, trochanter, and tibiotarsus of all legs with many fine setae.
|
||
<emphasis id="B922AD33FFC4FFD4FC04FCABFBC48441" bold="true" box="[957,1056,778,799]" pageId="4" pageNumber="351">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="136D6DA4FFC4FFD4FB94FCABFB978441" box="[1069,1139,778,799]" captionStart="Fig" captionStartId="5.[709,742,1686,1705]" captionTargetBox="[697,1163,1163,1650]" captionTargetId="figure-267@5.[685,1172,1154,1663]" captionTargetPageId="5" captionText="Fig. 16. Habitus of third instar of Hemiphileurus elbitae." figureDoi="http://doi.org/10.5281/zenodo.10102364" httpUri="https://zenodo.org/record/10102364/files/figure.png" pageId="4" pageNumber="351">Fig. 16</figureCitation>
|
||
): Abdominal spiracle I
|
||
<quantity id="4CAEDCC4FFC4FFD4FCD7FC84FC268464" box="[878,962,805,826]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="351" unit="mm" value="0.23">0.23 mm</quantity>
|
||
long and
|
||
<quantity id="4CAEDCC4FFC4FFD4FBA4FC84FB958464" box="[1053,1137,805,826]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="351" unit="mm" value="0.16">0.16 mm</quantity>
|
||
wide, shorter than spiracles
|
||
<collectionCode id="ED47E9E4FFC4FFD4FC28FCE1FC44840B" box="[913,928,832,853]" pageId="4" pageNumber="351">II</collectionCode>
|
||
–VII. Spiracles on segments
|
||
<collectionCode id="ED47E9E4FFC4FFD4FD7AFCFBFD368431" box="[707,722,858,879]" pageId="4" pageNumber="351">II</collectionCode>
|
||
and
|
||
<collectionCode id="ED47E9E4FFC4FFD4FCBBFCFBFCF78431" box="[770,787,858,879]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="351" type="Museum">V</collectionCode>
|
||
equal in size (
|
||
<quantity id="4CAEDCC4FFC4FFD4FC24FCFBFC178431" box="[925,1011,858,879]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="4" pageNumber="351" unit="mm" value="0.26">0.26 mm</quantity>
|
||
long and
|
||
<quantity id="4CAEDCC4FFC4FFD4FBEDFCFBFB4D8431" box="[1108,1193,858,879]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="351" unit="mm" value="0.17">0.17 mm</quantity>
|
||
wide), slightly larger than spiracles on segments
|
||
<collectionCode id="ED47E9E4FFC4FFD4FD7AFC2EFD3F84FA" box="[707,731,911,932]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="4" pageNumber="351">VI</collectionCode>
|
||
–VII (
|
||
<quantity id="4CAEDCC4FFC4FFD4FCACFC2EFC8984FA" box="[789,877,911,932]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="4" pageNumber="351" unit="mm" value="0.23">0.23 mm</quantity>
|
||
long and
|
||
<quantity id="4CAEDCC4FFC4FFD4FC69FC2EFBC284FA" box="[976,1062,911,932]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="351" unit="mm" value="0.17">0.17 mm</quantity>
|
||
wide). Spiracle on segment VIII slightly larger than those on segments I–VII (
|
||
<quantity id="4CAEDCC4FFC4FFD4FCD9FC65FC518487" box="[864,949,964,985]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="351" unit="mm" value="0.21">0.21 mm</quantity>
|
||
long and
|
||
<quantity id="4CAEDCC4FFC4FFD4FBADFC65FB8C8487" box="[1044,1128,964,985]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="4" pageNumber="351" unit="mm" value="0.17">0.17 mm</quantity>
|
||
wide). Abdominal segment I with 4 long, slender setae and 18 short, spine-like setae on prescutum; subscutum with 4 long, slender setae and without short subscutum with 4 long, slender setae and without short, spine-like setae; scutum with 15 long, slen- der setae and 54 short, spine-like setae; scutellum with 2 long, slender setae and 46 short, spine-like setae. Abdominal segment
|
||
<collectionCode id="ED47E9E4FFC5FFD5FC0CFC2CFC2284FC" box="[949,966,909,930]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="352" type="Museum">V</collectionCode>
|
||
with 6 long, slender setae and 60 short, spine-like setae on prescutum; subscutum with 4 long, slender setae and without short, spine-like setae; scutum with 14 long, slender setae and 55 short, spine-like setae; scutellum with 2 long, slender setae and 48 short, spine-like setae. Abdominal segment
|
||
<collectionCode id="ED47E9E4FFC5FFD5FCD2FB8DFC67831F" box="[875,899,1068,1089]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="352">VI</collectionCode>
|
||
with 6 long, slender setae and 58 short, spine-like setae on prescutum; subscutum
|
||
</paragraph>
|
||
<caption id="DF2921A9FFC4FFD4FF7CFD64FDDD85AE" ID-DOI="http://doi.org/10.5281/zenodo.10102354" ID-Zenodo-Dep="10102354" httpUri="https://zenodo.org/record/10102354/files/figure.png" pageId="4" pageNumber="351" startId="4.[197,229,709,728]" targetBox="[209,618,198,653]" targetPageId="4">
|
||
<paragraph id="8BE97121FFC4FFD4FF7CFD64FDDD85AE" blockId="4.[173,659,709,752]" pageId="4" pageNumber="351">
|
||
<emphasis id="B922AD33FFC4FFD4FF7CFD64FEE58586" bold="true" box="[197,257,709,728]" pageId="4" pageNumber="351">Fig. 8.</emphasis>
|
||
<taxonomicName id="4C560AA2FFC4FFD4FEAFFD64FE298586" box="[278,461,709,728]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC4FFD4FEAFFD64FE298586" box="[278,461,709,728]" italics="true" pageId="4" pageNumber="351">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
, third instar.
|
||
<emphasis id="B922AD33FFC4FFD4FD82FD64FDBD8586" bold="true" box="[571,601,709,728]" pageId="4" pageNumber="351">8a)</emphasis>
|
||
Hypopharynx, dorsal view;
|
||
<emphasis id="B922AD33FFC4FFD4FEDFFD7CFE6185AE" bold="true" box="[358,389,733,752]" pageId="4" pageNumber="351">8b)</emphasis>
|
||
Maxilla, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2921A9FFC4FFD4FF7CF9DFFEEE819F" ID-DOI="http://doi.org/10.5281/zenodo.10102356" ID-Zenodo-Dep="10102356" httpUri="https://zenodo.org/record/10102356/files/figure.png" pageId="4" pageNumber="351" startId="4.[197,239,1662,1681]" targetBox="[205,639,1227,1619]" targetPageId="4">
|
||
<paragraph id="8BE97121FFC4FFD4FF7CF9DFFEEE819F" blockId="4.[173,660,1662,1729]" pageId="4" pageNumber="351">
|
||
<emphasis id="B922AD33FFC4FFD4FF7CF9DFFED681CF" bold="true" box="[197,306,1662,1681]" pageId="4" pageNumber="351">Figs. 9–10.</emphasis>
|
||
<taxonomicName id="4C560AA2FFC4FFD4FEF1F9DFFDF181CF" box="[328,533,1662,1681]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC4FFD4FEF1F9DFFDF181CF" box="[328,533,1662,1681]" italics="true" pageId="4" pageNumber="351">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
, third instar.
|
||
<emphasis id="B922AD33FFC4FFD4FF14F937FF2481F7" bold="true" box="[173,192,1686,1705]" pageId="4" pageNumber="351">9)</emphasis>
|
||
Maxilla, inner view showing unci;
|
||
<emphasis id="B922AD33FFC4FFD4FE5DF937FDE681F7" bold="true" box="[484,514,1686,1705]" pageId="4" pageNumber="351">10)</emphasis>
|
||
Stridulatory teeth of maxilla.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2921A9FFC4FFD4FD62F9DFFC50819F" ID-DOI="http://doi.org/10.5281/zenodo.10102358" ID-Zenodo-Dep="10102358" httpUri="https://zenodo.org/record/10102358/files/figure.png" pageId="4" pageNumber="351" startId="4.[731,775,1662,1681]" targetBox="[768,1146,1129,1603]" targetPageId="4">
|
||
<paragraph id="8BE97121FFC4FFD4FD62F9DFFC50819F" blockId="4.[707,1194,1662,1729]" pageId="4" pageNumber="351">
|
||
<emphasis id="B922AD33FFC4FFD4FD62F9DFFCB181CF" bold="true" box="[731,853,1662,1681]" pageId="4" pageNumber="351">Figs. 11–12.</emphasis>
|
||
Last antennomere of third instar
|
||
<taxonomicName id="4C560AA2FFC4FFD4FD7AF937FC6581F7" box="[707,897,1686,1705]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="351" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC4FFD4FD7AF937FC6581F7" box="[707,897,1686,1705]" italics="true" pageId="4" pageNumber="351">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
, showing sensory spots.
|
||
<emphasis id="B922AD33FFC4FFD4FBE7F937FB9D81F7" bold="true" box="[1118,1145,1686,1705]" pageId="4" pageNumber="351">11)</emphasis>
|
||
Dorsal view;
|
||
<emphasis id="B922AD33FFC4FFD4FCA3F90FFCDC819F" bold="true" box="[794,824,1710,1729]" pageId="4" pageNumber="351">12)</emphasis>
|
||
Ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2921A9FFC5FFD5FD7CFD63FC0D85B3" ID-DOI="http://doi.org/10.5281/zenodo.10102360" ID-Zenodo-Dep="10102360" httpUri="https://zenodo.org/record/10102360/files/figure.png" pageId="5" pageNumber="352" startId="5.[709,744,706,725]" targetBox="[724,1137,189,649]" targetPageId="5">
|
||
<paragraph id="8BE97121FFC5FFD5FD7CFD63FB77858B" blockId="5.[685,1171,706,749]" box="[709,1171,706,725]" pageId="5" pageNumber="352">
|
||
<emphasis id="B922AD33FFC5FFD5FD7CFD63FCF1858B" bold="true" box="[709,789,706,725]" pageId="5" pageNumber="352">Fig. 15.</emphasis>
|
||
Lateral view of metatarsal claw of
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC5FFD5FD14FD78FC0D85B3" blockId="5.[685,1171,706,749]" box="[685,1001,729,749]" pageId="5" pageNumber="352">
|
||
<taxonomicName id="4C560AA2FFC5FFD5FD14FD78FC9485B2" box="[685,880,729,748]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC5FFD5FD14FD78FC9485B2" box="[685,880,729,748]" italics="true" pageId="5" pageNumber="352">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
, third instar.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2921A9FFC5FFD5FF16FB33FD9B83FB" ID-DOI="http://doi.org/10.5281/zenodo.10102362" ID-Zenodo-Dep="10102362" box="[175,639,1170,1189]" httpUri="https://zenodo.org/record/10102362/files/figure.png" pageId="5" pageNumber="352" startId="5.[175,217,1170,1189]" targetBox="[194,604,219,1121]" targetPageId="5">
|
||
<paragraph id="8BE97121FFC5FFD5FF16FB33FD9B83FB" blockId="5.[151,639,1170,1213]" box="[175,639,1170,1189]" pageId="5" pageNumber="352">
|
||
<emphasis id="B922AD33FFC5FFD5FF16FB33FEC383FB" bold="true" box="[175,295,1170,1189]" pageId="5" pageNumber="352">Figs. 13–14.</emphasis>
|
||
<taxonomicName id="4C560AA2FFC5FFD5FE85FB33FDE183FB" box="[316,517,1170,1189]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC5FFD5FE85FB33FDE183FB" box="[316,517,1170,1189]" italics="true" pageId="5" pageNumber="352">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
, third instar.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE97121FFC5FFD5FF2EFB0BFDF283E3" blockId="5.[151,639,1170,1213]" box="[151,534,1194,1213]" pageId="5" pageNumber="352">
|
||
<emphasis id="B922AD33FFC5FFD5FF2EFB0BFF5283E3" bold="true" box="[151,182,1194,1213]" pageId="5" pageNumber="352">13)</emphasis>
|
||
Thoracic spiracle;
|
||
<emphasis id="B922AD33FFC5FFD5FEE6FB0BFE9983E3" bold="true" box="[351,381,1194,1213]" pageId="5" pageNumber="352">14)</emphasis>
|
||
Spiracular holes.
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC5FFD6FF2EFAA2FC9886F2" blockId="5.[151,638,1283,1729]" lastBlockId="6.[707,1194,193,507]" lastPageId="6" lastPageNumber="353" pageId="5" pageNumber="352">
|
||
setae; scutum with about 5 short, spine-like setae and 1 long, slender seta; scutellum with about 37 short, spine-like setae and 13 long, slender setae. Abdominal segment
|
||
<collectionCode id="ED47E9E4FFC5FFD5FE99FAF2FED48236" box="[288,304,1363,1384]" pageId="5" pageNumber="352">II</collectionCode>
|
||
with 4 long, slender setae and 42 short, spine-like setae on prescutum; subscutum with 4 long setae and without short, spine-like setae; scutum with 2 long, slender setae and 36 short, spine-like setae; scutellum with 15 long, slender setae and 50 short, spine-like setae. Abdominal segment III with 4 long, slender setae and 44 short, spine-like setae on prescutum; subscutum with 4 long, slender seta and without short, spine-like setae; scutum with 2 long, slender setae and 44 short, spine-like setae; scutellum with 12 long, slender setae and 46 short, spine-like setae. Abdominal segment IV with 4 long, slender setae and 57 short, spine-like setae on prescutum; with 4 long, slender setae and without short, spinelike setae; scutum with 10 long, slender setae and 48 short, spine-like setae; scutellum with 2 long, slender setae and 44 short, spine-like setae. Abdominal segment VII with 2 rows of 16 slender, long setae and 54 short, spine-like setae mixed. Abdominal segment VIII with 2 rows of 12 slender, long setae each, and 39 short, spine-like setae. Abdominal segment IX with 9 slender, long setae distributed irregularly and 36 short, spine-like setae mixed. Abdominal segment X with approximately 60 moderate to long, slender setae and 22 short, spine-like setae mixed. Pleural lobes with 6–12 long, slender setae and short, spine-like setae absent. All spiracular areas with 6 long, slender setae and 2 short, spine-like setae.
|
||
<emphasis id="B922AD33FFC6FFD6FC02FEB3FC1B8679" bold="true" box="[955,1023,274,295]" pageId="6" pageNumber="353">Raster</emphasis>
|
||
: Surface without palidia; campus with 4 slender, long setae; teges with 55–62 short setae that project toward superior anal lobe; barbula with 33 long, slender setae. Anal slit transverse. Dorsal body length approximately
|
||
<quantity id="4CAEDCC4FFC6FFD6FD7AFE36FCFD86F2" box="[707,793,407,428]" metricMagnitude="-2" metricUnit="m" metricValue="7.459999999999999" pageId="6" pageNumber="353" unit="mm" value="74.6">74.6 mm</quantity>
|
||
(
|
||
<figureCitation id="136D6DA4FFC6FFD6FC90FE36FC9586F2" box="[809,881,407,428]" captionStart="Figs" captionStartId="5.[175,217,1170,1189]" captionTargetBox="[194,604,219,1121]" captionTargetId="figure-237@5.[151,638,191,1153]" captionTargetPageId="5" captionText="Figs. 13–14. Hemiphileurus elbitae, third instar." figureDoi="http://doi.org/10.5281/zenodo.10102362" httpUri="https://zenodo.org/record/10102362/files/figure.png" pageId="6" pageNumber="353">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF2921A9FFC5FFD5FD7CF937FD08819F" ID-DOI="http://doi.org/10.5281/zenodo.10102364" ID-Zenodo-Dep="10102364" httpUri="https://zenodo.org/record/10102364/files/figure.png" pageId="5" pageNumber="352" startId="5.[709,742,1686,1705]" targetBox="[697,1163,1163,1650]" targetPageId="5">
|
||
<paragraph id="8BE97121FFC5FFD5FD7CF937FD08819F" blockId="5.[685,1172,1686,1729]" pageId="5" pageNumber="352">
|
||
<emphasis id="B922AD33FFC5FFD5FD7CF937FCEA81F7" bold="true" box="[709,782,1686,1705]" pageId="5" pageNumber="352">Fig. 16.</emphasis>
|
||
Habitus of third instar of
|
||
<taxonomicName id="4C560AA2FFC5FFD5FBADF937FD02819F" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="352" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC5FFD5FBADF937FD02819F" italics="true" pageId="5" pageNumber="352">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE97121FFC6FFD7FD62FE10FEE68487" blockId="6.[707,1194,193,507]" lastBlockId="7.[151,639,194,1729]" lastPageId="7" lastPageNumber="354" pageId="6" pageNumber="353">
|
||
<emphasis id="B922AD33FFC6FFD6FD62FE10FCF08698" bold="true" box="[731,788,433,454]" pageId="6" pageNumber="353">Pupal</emphasis>
|
||
<materialsCitation id="3B3E7B7CFFC6FFD7FCA3FE10FF1A8487" collectionCode="II, V, VI" collectorName="Description. Female & Length & Body & Color & Entire & Head & Surface & Compound & Thorax & Pronotal & Form & Elytral & Protibia & Abdomen & Segments & Segment & Segment X & Segments I - X & Pleural & Spiracle I & Abdominal" lastPageId="7" lastPageNumber="354" pageId="6" pageNumber="353" specimenCount="1" typeStatus="paratype">
|
||
<emphasis id="B922AD33FFC6FFD6FCA3FE10FC148698" bold="true" box="[794,1008,433,454]" pageId="6" pageNumber="353">
|
||
<typeStatus id="54EDCF83FFC6FFD6FCA3FE10FC978698" box="[794,883,433,454]" pageId="6" pageNumber="353" type="paratype">Paratype</typeStatus>
|
||
<collectorName id="26A314F7FFC6FFD6FCC0FE10FC148698" box="[889,1008,433,454]" pageId="6" pageNumber="353">Description.</collectorName>
|
||
</emphasis>
|
||
Female (
|
||
<figureCitation id="136D6DA4FFC6FFD6FBE9FE10FD3A86BF" captionStart="Figs" captionStartId="6.[197,237,1710,1729]" captionTargetBox="[225,1136,587,1674]" captionTargetId="figure-195@6.[173,1193,565,1687]" captionTargetPageId="6" captionText="Figs. 17–19. Pupa of Hemiphileurus elbitae. 17) Dorsal view; 18) Ventral view; 19) Lateral view." figureDoi="http://doi.org/10.5281/zenodo.10102366" httpUri="https://zenodo.org/record/10102366/files/figure.png" pageId="6" pageNumber="353">Figs. 17– 19</figureCitation>
|
||
).
|
||
<collectorName id="26A314F7FFC6FFD6FD4BFE6DFCDC86BF" box="[754,824,460,481]" pageId="6" pageNumber="353">Length</collectorName>
|
||
<quantity id="4CAEDCC4FFC6FFD6FCF9FE6DFC4D86BF" box="[832,937,460,481]" metricMagnitude="-2" metricUnit="m" metricValue="2.613" pageId="6" pageNumber="353" unit="mm" value="26.13">26.13 mm</quantity>
|
||
; widest width
|
||
<quantity id="4CAEDCC4FFC6FFD6FB87FE6DFB4286BF" box="[1086,1190,460,481]" metricMagnitude="-2" metricUnit="m" metricValue="1.175" pageId="6" pageNumber="353" unit="mm" value="11.75">11.75 mm</quantity>
|
||
.
|
||
<collectorName id="26A314F7FFC6FFD6FD7AFE47FD1286A5" box="[707,758,486,507]" pageId="6" pageNumber="353">Body</collectorName>
|
||
shape elongate, oval, stout, exarate.
|
||
<collectorName id="26A314F7FFC6FFD6FBFCFE47FB9E86A5" box="[1093,1146,486,507]" pageId="6" pageNumber="353">Color</collectorName>
|
||
dark reddish orange.
|
||
<collectorName id="26A314F7FFC7FFD7FE89FF63FE8D8789" box="[304,361,194,215]" pageId="7" pageNumber="354">Entire</collectorName>
|
||
body with fine, velvety-gold vestiture.
|
||
<emphasis id="B922AD33FFC7FFD7FF44FF7CFED787AC" bold="true" box="[253,307,221,242]" pageId="7" pageNumber="354">
|
||
<collectorName id="26A314F7FFC7FFD7FF44FF7CFED787AC" box="[253,307,221,242]" pageId="7" pageNumber="354">Head</collectorName>
|
||
</emphasis>
|
||
:
|
||
<collectorName id="26A314F7FFC7FFD7FEFFFF7CFE6B87AC" box="[326,399,221,242]" pageId="7" pageNumber="354">Surface</collectorName>
|
||
glabrous, bent sharply beneath thorax, mouthparts directed ventrally; antenna, labrum, mandibles, maxillae, and palps discernible; antennal tecae expanded, stout, with apices rounded.
|
||
<collectorName id="26A314F7FFC7FFD7FF4BFEE6FEBF8602" box="[242,347,327,348]" pageId="7" pageNumber="354">Compound</collectorName>
|
||
eyes sunken, scarcely visible, with 2 tubercles between eyes.
|
||
<emphasis id="B922AD33FFC7FFD7FE69FEC3FDF98629" bold="true" box="[464,541,354,375]" pageId="7" pageNumber="354">
|
||
<collectorName id="26A314F7FFC7FFD7FE69FEC3FDF98629" box="[464,541,354,375]" pageId="7" pageNumber="354">Thorax</collectorName>
|
||
</emphasis>
|
||
:
|
||
<collectorName id="26A314F7FFC7FFD7FD95FEC3FD9A8629" box="[556,638,354,375]" pageId="7" pageNumber="354">Pronotal</collectorName>
|
||
<emphasis id="B922AD33FFC7FFD7FF2EFEDAFF4486CE" box="[151,160,379,400]" italics="true" pageId="7" pageNumber="354">s</emphasis>
|
||
urface glabrous.
|
||
<collectorName id="26A314F7FFC7FFD7FEECFEDDFE6E86CF" box="[341,394,380,401]" pageId="7" pageNumber="354">Form</collectorName>
|
||
transverse, with wide, anteromedial, round apex, tubercle slightly defined. Meso- and metanotum well differentiated.
|
||
<collectorName id="26A314F7FFC7FFD7FF2EFE6DFF3286BF" box="[151,214,460,481]" pageId="7" pageNumber="354">Elytral</collectorName>
|
||
and posterior wing tecae closely appressed, curved ventrally around body; elytral tecae extending to middle of abdominal segment
|
||
<collectionCode id="ED47E9E4FFC7FFD7FDB2FDA0FDF98548" box="[523,541,513,534]" pageId="7" pageNumber="354">II</collectionCode>
|
||
; posterior wing tecae extending to middle of abdominal segment III.
|
||
<collectorName id="26A314F7FFC7FFD7FF4CFD97FEDA8515" box="[245,318,566,587]" pageId="7" pageNumber="354">Protibia</collectorName>
|
||
with 3 distinct teeth on external edge. Meso- and metatibiae with inner and external spines well developed at apices.
|
||
<emphasis id="B922AD33FFC7FFD7FDABFDCAFD9385DE" bold="true" box="[530,631,619,640]" pageId="7" pageNumber="354">
|
||
<collectorName id="26A314F7FFC7FFD7FDABFDCAFD9385DE" box="[530,631,619,640]" pageId="7" pageNumber="354">Abdomen</collectorName>
|
||
</emphasis>
|
||
:
|
||
<collectorName id="26A314F7FFC7FFD7FF2EFD27FF1685C5" box="[151,242,646,667]" pageId="7" pageNumber="354">Segments</collectorName>
|
||
III–X (ventral view) well-defined.
|
||
<collectorName id="26A314F7FFC7FFD7FDEAFD27FF2385EB" pageId="7" pageNumber="354">Segment</collectorName>
|
||
VII slightly long than preceding and VIII 0.25 times longer than segment VII; segments VIII and IX fused.
|
||
<collectorName id="26A314F7FFC7FFD7FEA2FD74FE6685B4" box="[283,386,725,746]" pageId="7" pageNumber="354">Segment X</collectorName>
|
||
with small genital ampulla (female).
|
||
<collectorName id="26A314F7FFC7FFD7FF4EFD51FE66845B" box="[247,386,751,773]" pageId="7" pageNumber="354">Segments I–X</collectorName>
|
||
(dorsal view) with welldefined, dioneiform organs between segments I–
|
||
<collectionCode id="ED47E9E4FFC7FFD7FDD0FCABFD9E8441" box="[617,634,778,799]" pageId="7" pageNumber="354">II</collectionCode>
|
||
,
|
||
<collectionCode id="ED47E9E4FFC7FFD7FF2EFC84FF438464" box="[151,167,805,826]" pageId="7" pageNumber="354">II</collectionCode>
|
||
–III, III–IV, IV–
|
||
<collectionCode id="ED47E9E4FFC7FFD7FEF3FC84FEBF8464" box="[330,347,805,826]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="354" type="Museum">V</collectionCode>
|
||
,
|
||
<collectionCode id="ED47E9E4FFC7FFD7FED1FC84FE9D8464" box="[360,377,805,826]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="354" type="Museum">V</collectionCode>
|
||
–
|
||
<collectionCode id="ED47E9E4FFC7FFD7FE3FFC84FE468464" box="[390,418,805,826]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="354">VI</collectionCode>
|
||
, and
|
||
<collectionCode id="ED47E9E4FFC7FFD7FE63FC84FE178464" box="[474,499,805,826]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="354">VI</collectionCode>
|
||
–VII, the first strongly sclerotized.
|
||
<collectorName id="26A314F7FFC7FFD7FEDBFCE1FE40840B" box="[354,420,832,853]" pageId="7" pageNumber="354">Pleural</collectorName>
|
||
lobes rounded.
|
||
<collectorName id="26A314F7FFC7FFD7FDFCFCE1FF238431" pageId="7" pageNumber="354">Spiracle I</collectorName>
|
||
elongate, with fine peritreme, covered by wing thecae; spiracles
|
||
<collectionCode id="ED47E9E4FFC7FFD7FE33FCD4FE7E84D4" box="[394,410,885,906]" pageId="7" pageNumber="354">II</collectionCode>
|
||
–IV ovate, prominent, each with strongly sclerotized peritreme; spiracles
|
||
<collectionCode id="ED47E9E4FFC7FFD7FF2EFC0BFF4C84E1" box="[151,168,938,959]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="354" type="Museum">V</collectionCode>
|
||
–VIII closed.
|
||
<collectorName id="26A314F7FFC7FFD7FE93FC0BFE7784E1" box="[298,403,938,959]" pageId="7" pageNumber="354">Abdominal</collectorName>
|
||
apex rounded, with fine, short setae
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF2921A9FFC6FFD6FF7CF90FFBC9819F" ID-DOI="http://doi.org/10.5281/zenodo.10102366" ID-Zenodo-Dep="10102366" box="[197,1069,1709,1729]" httpUri="https://zenodo.org/record/10102366/files/figure.png" pageId="6" pageNumber="353" startId="6.[197,237,1710,1729]" targetBox="[225,1136,587,1674]" targetPageId="6">
|
||
<paragraph id="8BE97121FFC6FFD6FF7CF90FFBC9819F" blockId="6.[197,1069,1709,1729]" box="[197,1069,1709,1729]" pageId="6" pageNumber="353">
|
||
<emphasis id="B922AD33FFC6FFD6FF7CF90FFED1819F" bold="true" box="[197,309,1709,1729]" pageId="6" pageNumber="353">Figs. 17–19.</emphasis>
|
||
Pupa of
|
||
<taxonomicName id="4C560AA2FFC6FFD6FE2CF90CFDB6819E" box="[405,594,1709,1728]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="353" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC6FFD6FE2CF90CFDB6819E" box="[405,594,1709,1728]" italics="true" pageId="6" pageNumber="353">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B922AD33FFC6FFD6FDE6F90FFD99819F" bold="true" box="[607,637,1710,1729]" pageId="6" pageNumber="353">17)</emphasis>
|
||
Dorsal view;
|
||
<emphasis id="B922AD33FFC6FFD6FD40F90FFCF3819F" bold="true" box="[761,791,1710,1729]" pageId="6" pageNumber="353">18)</emphasis>
|
||
Ventral view;
|
||
<emphasis id="B922AD33FFC6FFD6FC2EF90FFC52819F" bold="true" box="[919,950,1710,1729]" pageId="6" pageNumber="353">19)</emphasis>
|
||
Lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C34C22AAFFC7FFD7FF16FC7EFF0283F0" pageId="7" pageNumber="354" type="distribution">
|
||
<paragraph id="8BE97121FFC7FFD7FF16FC7EFEFC8377" blockId="7.[151,639,194,1729]" pageId="7" pageNumber="354">
|
||
<emphasis id="B922AD33FFC7FFD7FF16FC7EFED684AA" bold="true" box="[175,306,991,1012]" pageId="7" pageNumber="354">Distribution.</emphasis>
|
||
<taxonomicName id="4C560AA2FFC7FFD7FE86FC7EFDFD84AA" box="[319,537,991,1012]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="elbitae">
|
||
<emphasis id="B922AD33FFC7FFD7FE86FC7EFDFD84AA" box="[319,537,991,1012]" italics="true" pageId="7" pageNumber="354">Hemiphileurus elbitae</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="54EDCF83FFC7FFD7FE95FC5BFEB28351" box="[300,342,1018,1039]" pageId="7" pageNumber="354">type</typeStatus>
|
||
locality in the Chocó region of
|
||
<collectingCountry id="F34131B1FFC7FFD7FF0AFBB5FEF08377" box="[179,276,1044,1065]" name="Colombia" pageId="7" pageNumber="354">Colombia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC7FFD7FF16FB8EFE4C8327" blockId="7.[151,639,194,1729]" pageId="7" pageNumber="354">
|
||
<emphasis id="B922AD33FFC7FFD7FF16FB8EFE80831A" bold="true" box="[175,356,1071,1092]" pageId="7" pageNumber="354">Locality Records.</emphasis>
|
||
(
|
||
<specimenCount id="9D50BAA8FFC7FFD7FEC0FB8EFE21831A" box="[377,453,1071,1092]" pageId="7" pageNumber="354" type="adult">2 adults</specimenCount>
|
||
,
|
||
<specimenCount id="9D50BAA8FFC7FFD7FE74FB8EFDEB831A" box="[461,527,1071,1092]" pageId="7" pageNumber="354" type="larva">1 larva</specimenCount>
|
||
, and
|
||
<specimenCount id="9D50BAA8FFC7FFD7FD86FB8EFD99831A" box="[575,637,1071,1092]" pageId="7" pageNumber="354" type="pupa">1 pupa</specimenCount>
|
||
examined).
|
||
<materialsCitation id="3B3E7B7CFFC7FFD7FEB7FBE8FE408327" country="Colombia" location="Municipio de Acandi" municipality="Serrania del Darien" pageId="7" pageNumber="354" specimenCount="1">
|
||
<emphasis id="B922AD33FFC7FFD7FEB7FBE8FE7E8300" bold="true" box="[270,410,1097,1118]" pageId="7" pageNumber="354">
|
||
<collectingCountry id="F34131B1FFC7FFD7FEB7FBE8FE7E8300" box="[270,410,1097,1118]" name="Colombia" pageId="7" pageNumber="354">COLOMBIA</collectingCountry>
|
||
</emphasis>
|
||
: Chocó,
|
||
<location id="8E8927FAFFC7FFD7FE41FBE8FF3A8327" LSID="urn:lsid:plazi:treatment:03FFC037FFC1FFD7FF0DFCC5FB7084C1:8E8927FAFFC7FFD7FE41FBE8FF3A8327" country="Colombia" municipality="Serrania del Darien" name="Municipio de Acandi" pageId="7" pageNumber="354">Municipio de Acandí</location>
|
||
,
|
||
<collectingMunicipality id="6B8DEB5BFFC7FFD7FF51FBC5FE408327" box="[232,420,1124,1145]" pageId="7" pageNumber="354">Serranía del Darién</collectingMunicipality>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC7FFD7FF16FBDEFF0283F0" blockId="7.[151,639,194,1729]" pageId="7" pageNumber="354">
|
||
<emphasis id="B922AD33FFC7FFD7FF16FBDEFE6B83CA" bold="true" box="[175,399,1151,1172]" pageId="7" pageNumber="354">Temporal Distribution.</emphasis>
|
||
January (
|
||
<specimenCount id="9D50BAA8FFC7FFD7FE54FBDEFDD183CA" box="[493,565,1151,1172]" pageId="7" pageNumber="354" type="adult">2 adults</specimenCount>
|
||
,
|
||
<specimenCount id="9D50BAA8FFC7FFD7FD85FBDEFD9E83CA" box="[572,634,1151,1172]" pageId="7" pageNumber="354" type="larva">1 larva</specimenCount>
|
||
,
|
||
<specimenCount id="9D50BAA8FFC7FFD7FF2EFB38FF3F83F0" box="[151,219,1177,1198]" pageId="7" pageNumber="354" type="pupa">1 pupa</specimenCount>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C22AAFFC7FFD7FF16FB15FE0E819F" pageId="7" pageNumber="354" type="diagnosis">
|
||
<paragraph id="8BE97121FFC7FFD7FF16FB15FEA882E6" blockId="7.[151,639,194,1729]" pageId="7" pageNumber="354">
|
||
<emphasis id="B922AD33FFC7FFD7FF16FB15FEFC8397" bold="true" box="[175,280,1204,1225]" pageId="7" pageNumber="354">Diagnosis.</emphasis>
|
||
The males of this species are most easily recognized by the unique form of the parameres which, in caudal view, are distinctly hollowed in the center of each paramere (
|
||
<figureCitation id="136D6DA4FFC7FFD7FF26FABFFF39826D" box="[159,221,1310,1331]" captionStart="Figs" captionStartId="2.[197,240,952,971]" captionTargetBox="[187,638,195,912]" captionTargetId="figure-303@2.[173,660,183,928]" captionTargetPageId="2" captionText="Figs. 2–3. Parameres of Hemiphileurus elbitae." figureDoi="http://doi.org/10.5281/zenodo.10102348" httpUri="https://zenodo.org/record/10102348/files/figure.png" pageId="7" pageNumber="354">Fig. 2</figureCitation>
|
||
). In addition, this species is characterized by the presence of frontoclypeal tubercles (not horns), pronotum strongly punctate everywhere, elytral intervals smooth and of equal height, last sternite densely punctate only across base, and tridentate protibia.
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC7FFD7FF16FA1CFE0E819F" blockId="7.[151,639,194,1729]" pageId="7" pageNumber="354">
|
||
The third instar of
|
||
<taxonomicName id="4C560AA2FFC7FFD7FED6FA1CFDAF828C" box="[367,587,1469,1490]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B922AD33FFC7FFD7FED6FA1CFE1A828C" box="[367,510,1469,1490]" italics="true" pageId="7" pageNumber="354">Hemiphileurus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
may be distinguished, tentatively, from other phileurine larvae by the angulate margin on both sides of the labrum, absence of anterior frontal setae, and an entire haptomeral process (
|
||
<bibRefCitation id="EFC70CD0FFC7FFD7FE26F986FDC48162" author="Ritcher" box="[415,544,1575,1596]" pageId="7" pageNumber="354" refId="ref4619" refString="Ritcher, P. O. 1966. White Grubs and Their Allies. Oregon State University Press, Corvallis, OR." type="book" year="1966">Ritcher 1966</bibRefCitation>
|
||
; Ocampo and Morón 2004). However, our knowledge of phileurine immatures is still so fragmentary that many more larvae must be known in order to study characters at the species level and arrive at accurate diagnoses for the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C22AAFFC7FFD7FD7CFF63FCC08652" pageId="7" pageNumber="354" type="etymology">
|
||
<paragraph id="8BE97121FFC7FFD7FD7CFF63FCC08652" blockId="7.[685,1172,194,401]" pageId="7" pageNumber="354">
|
||
<emphasis id="B922AD33FFC7FFD7FD7CFF63FCDC8789" bold="true" box="[709,824,194,215]" pageId="7" pageNumber="354">Etymology.</emphasis>
|
||
The species is dedicated to Lic. Elba Moreno de Neita, mother of the first author, to honor her memory.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C22AAFFC7FFD7FD7CFEB3FBE286CF" pageId="7" pageNumber="354" type="biology_ecology">
|
||
<paragraph id="8BE97121FFC7FFD7FD7CFEB3FBE286CF" blockId="7.[685,1172,194,401]" pageId="7" pageNumber="354">
|
||
<emphasis id="B922AD33FFC7FFD7FD7CFEB3FCF38679" bold="true" box="[709,791,274,295]" pageId="7" pageNumber="354">Biology.</emphasis>
|
||
The larva and pupa were found in a rotted tree trunk of
|
||
<taxonomicName id="4C560AA2FFC7FFD7FCC4FE8DFCCC8602" authority="(Nees) Mez" authorityName="Mez" baseAuthorityName="Nees" class="Magnoliopsida" family="Lauraceae" genus="Ocotea" kingdom="Plantae" order="Laurales" pageId="7" pageNumber="354" phylum="Tracheophyta" rank="species" species="cernua">
|
||
<emphasis id="B922AD33FFC7FFD7FCC4FE8DFBF7861F" box="[893,1043,300,321]" italics="true" pageId="7" pageNumber="354">Ocotea cernua</emphasis>
|
||
(Nees) Mez (Lauraceae)
|
||
</taxonomicName>
|
||
. The adults were observed preying on the larvae of a species of
|
||
<taxonomicName id="4C560AA2FFC7FFD7FC49FEC0FB708629" authority="Kirby" authorityName="Kirby" authorityYear="1828" box="[1008,1172,353,375]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B922AD33FFC7FFD7FC49FEC0FBA98628" box="[1008,1101,353,374]" italics="true" pageId="7" pageNumber="354">Chrysina</emphasis>
|
||
Kirby
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C560AA2FFC7FFD7FD0DFEDDFCD186CF" authorityName="Latreille" authorityYear="1802" box="[692,821,380,401]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="family">Scarabaeidae</taxonomicName>
|
||
: Rutelinae: Rutelini).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C22AAFFC7FFD7FD58FE14FB7084C1" pageId="7" pageNumber="354" type="key">
|
||
<paragraph id="8BE97121FFC7FFD7FD58FE14FBE786B6" blockId="7.[737,1120,437,489]" pageId="7" pageNumber="354">
|
||
<emphasis id="B922AD33FFC7FFD7FD58FE14FCBC86B9" bold="true" pageId="7" pageNumber="354">KEY TO THE KNOWN THIRD INSTARS OF</emphasis>
|
||
<taxonomicName id="4C560AA2FFC7FFD7FCD8FE73FBE786B6" box="[865,1027,466,489]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B922AD33FFC7FFD7FCD8FE73FBE786B6" bold="true" box="[865,1027,466,489]" italics="true" pageId="7" pageNumber="354">HEMIPHILEURUS</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<key id="FFC72610FFC7FFD7FD14FE5FFB7084C1" pageId="7" pageNumber="354">
|
||
<keyStep id="30A26984FFC7FFD7FD14FE5FFB7785E5" pageId="7" pageNumber="354">
|
||
<paragraph id="8BE97121FFC7FFD7FD14FE5FFB77853D" blockId="7.[685,1172,510,928]" pageId="7" pageNumber="354">
|
||
<keyLead id="30A7D214FFC7FFD7FD14FE5FFB77853D" pageId="7" pageNumber="354">
|
||
1. Abdominal spiracle I shorter than spiracles II–VII. Last segment of antenna with 5 dorsal sensory spots ..................................................
|
||
<emphasis id="B922AD33FFC7FFD7FD5AFDECFB77853D" bold="true" box="[739,1171,589,611]" pageId="7" pageNumber="354">
|
||
<taxonomicName id="4C560AA2FFC7FFD7FD5AFDECFBF4853D" authority="Neita and Ratcliffe" authorityName="Neita and Ratcliffe" box="[739,1040,589,611]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="elbitae" status="sp. nov.">
|
||
<emphasis id="B922AD33FFC7FFD7FD5AFDECFCA7853D" bold="true" box="[739,835,589,611]" italics="true" pageId="7" pageNumber="354">H. elbitae</emphasis>
|
||
Neita and Ratcliffe
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2111048FFC7FFD7FBA4FDEFFB77853D" box="[1053,1171,590,611]" pageId="7" pageNumber="354" rank="species">new species</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC7FFD7FD14FDD0FB7785E5" blockId="7.[685,1172,510,928]" pageId="7" pageNumber="354">
|
||
<keyLead id="30A7D214FFC7FFD7FD14FDD0FB7785E5" pageId="7" pageNumber="354">1′. Abdominal spiracles similar in size. Last segment of antenna with less than 5 dorsal sensory spots.....................................................2</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30A26984FFC7FFD7FD14FD69FB7084C1" pageId="7" pageNumber="354">
|
||
<paragraph id="8BE97121FFC7FFD7FD14FD69FB778473" blockId="7.[685,1172,510,928]" pageId="7" pageNumber="354">
|
||
<keyLead id="30A7D214FFC7FFD7FD14FD69FB778473" pageId="7" pageNumber="354">
|
||
2. Last segment of antenna with 2 dorsal sensory spots. Inner margin of left mandible, distad of molar area, with a short, rounded tooth ....................................
|
||
<taxonomicName id="4C560AA2FFC7FFD7FC4BFCB6FB778473" authority="Kolbe" authorityName="Kolbe" box="[1010,1171,791,813]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="dispar">
|
||
<emphasis id="B922AD33FFC7FFD7FC4BFCB6FB778473" bold="true" box="[1010,1171,791,813]" pageId="7" pageNumber="354">
|
||
<emphasis id="B922AD33FFC7FFD7FC4BFCB6FBAB8473" bold="true" box="[1010,1103,791,813]" italics="true" pageId="7" pageNumber="354">H. dispar</emphasis>
|
||
Kolbe
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BE97121FFC7FFD7FD14FC9AFB7084C1" blockId="7.[685,1172,510,928]" pageId="7" pageNumber="354">
|
||
<keyLead id="30A7D214FFC7FFD7FD14FC9AFB7084C1" pageId="7" pageNumber="354">
|
||
2′. Last segment of antenna with 3–4 dorsal sensory spots. Inner margin of left mandible, distad of molar area, with a prominent, triangular tooth ......................
|
||
<taxonomicName id="4C560AA2FFC7FFD7FC61FC2BFB7084C1" authority="LeConte" authorityName="LeConte" box="[984,1172,906,928]" class="Insecta" family="Dynastidae" genus="Hemiphileurus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="354" phylum="Arthropoda" rank="species" species="illatus">
|
||
<emphasis id="B922AD33FFC7FFD7FC61FC2BFB7084C1" bold="true" box="[984,1172,906,928]" pageId="7" pageNumber="354">
|
||
<emphasis id="B922AD33FFC7FFD7FC61FC2BFBD084FE" bold="true" box="[984,1076,906,928]" italics="true" pageId="7" pageNumber="354">H. illatus</emphasis>
|
||
LeConte
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |