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<document id="0F1CE9C3DFDEAC5B79613042C7FC6289" ID-DOI="10.11646/zootaxa.3838.2.9" ID-GBIF-Dataset="4fb0d9e1-07d3-4d05-868e-1e9a264b12a6" ID-ISSN="1175-5326" ID-Zenodo-Dep="224625" ID-ZooBank="485D8CBC-75BE-4C41-B85E-EA695B96E83A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460459308428" checkinUser="plazi" docAuthor="Kury, Adriano B. &amp; Quintero, Diomedes" docDate="2014" docId="E778A2756F13FFA2FF5FF958FCE2F3CC" docLanguage="en" docName="zt03838p246.pdf" docOrigin="Zootaxa 3838 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hernandarioides plana F.O. Pickard-Cambridge 1905" docType="treatment" docVersion="8" lastPageNumber="245" masterDocId="1B41DA0D6F12FFA1FFC8FF8AFFEBF50B" masterDocTitle="Characterization of the genus Hernandarioides F. O. Pickard-Cambridge, 1905 (Opiliones, Gonyleptidae, Ampycinae)" masterLastPageNumber="246" masterPageNumber="242" pageNumber="243" updateTime="1698303343701" updateUser="plazi">
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<mods:title id="D7ED2DEEB458FBE24ACFF4B49D5BE4E7">Characterization of the genus Hernandarioides F. O. Pickard-Cambridge, 1905 (Opiliones, Gonyleptidae, Ampycinae)</mods:title>
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<mods:date id="D2FA22DBD4798BBD0E2603CD99274A62">2014</mods:date>
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<treatment id="E778A2756F13FFA2FF5FF958FCE2F3CC" ID-DOI="http://doi.org/10.5281/zenodo.6144333" ID-GBIF-Taxon="119512015" ID-Zenodo-Dep="6144333" LSID="urn:lsid:plazi:treatment:E778A2756F13FFA2FF5FF958FCE2F3CC" httpUri="http://treatment.plazi.org/id/E778A2756F13FFA2FF5FF958FCE2F3CC" lastPageId="3" lastPageNumber="245" pageId="1" pageNumber="243">
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<paragraph id="6F6E13636F13FFA0FF5FF958FD12F3E2" blockId="1.[151,761,1746,1770]" box="[151,761,1746,1770]" pageId="1" pageNumber="243">
<heading id="3426A40F6F13FFA0FF5FF958FD12F3E2" bold="true" box="[151,761,1746,1770]" fontSize="10" level="2" pageId="1" pageNumber="243" reason="2">
<taxonomicName id="A8D168E06F13FFA0FF5FF958FD12F3E2" ID-CoL="7WL5B" authority="F.O. Pickard-Cambridge, 1905" authorityName="F.O. Pickard-Cambridge" authorityYear="1905" box="[151,761,1746,1770]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="243" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="5DA5CF716F13FFA0FF5FF958FD12F3E2" bold="true" box="[151,761,1746,1770]" pageId="1" pageNumber="243">
<emphasis id="5DA5CF716F13FFA0FF5FF958FE7DF3E1" bold="true" box="[151,406,1746,1770]" italics="true" pageId="1" pageNumber="243">Hernandarioides plana</emphasis>
<bibRefCitation id="0B406E926F13FFA0FE54F958FD12F3E2" author="Pickard-Cambridge" box="[412,761,1746,1769]" pageId="1" pageNumber="243" refString="Pickard-Cambridge, F. O. (1905) Order Opiliones [2 nd part]. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia centrali- Americana. Vol. 2. Arachnida. Araneidea and Opiliones. R. H. Porter / Dulau &amp; Co., London, pp. 561 - 610, pls. 53, 54." type="journal article" year="1905">F.O. Pickard-Cambridge, 1905</bibRefCitation>
</emphasis>
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<subSubSection id="27CB40E86F13FFA0FF5FF892FC6CF284" pageId="1" pageNumber="243" type="reference_group">
<paragraph id="6F6E13636F13FFA0FF5FF892FC8DF244" blockId="1.[151,1434,1816,1935]" pageId="1" pageNumber="243">
<treatmentCitationGroup id="4FC1344D6F13FFA0FF5FF892FC8DF244" pageId="1" pageNumber="243">
<taxonomicName id="A8D168E06F13FFA0FF5FF892FCFBF224" ID-CoL="7WL5B" authority="F.O. Pickard-Cambridge 1905: 574" authorityName="F.O. Pickard-Cambridge" authorityPageNumber="574" authorityYear="1905" box="[151,784,1816,1839]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="243" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="5DA5CF716F13FFA0FF5FF892FE7BF224" box="[151,400,1816,1839]" italics="true" pageId="1" pageNumber="243">Hernandarioides plana</emphasis>
<treatmentCitation id="EE7035726F13FFA0FE5FF892FCFBF224" author="Pickard-Cambridge" box="[407,784,1816,1839]" page="574" pageId="1" pageNumber="243" year="1905">
<bibRefCitation id="0B406E926F13FFA0FE5FF892FD30F224" author="Pickard-Cambridge" box="[407,731,1816,1839]" pageId="1" pageNumber="243" refString="Pickard-Cambridge, F. O. (1905) Order Opiliones [2 nd part]. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia centrali- Americana. Vol. 2. Arachnida. Araneidea and Opiliones. R. H. Porter / Dulau &amp; Co., London, pp. 561 - 610, pls. 53, 54." type="journal article" year="1905">F.O. Pickard-Cambridge 1905</bibRefCitation>
: 574
</treatmentCitation>
</taxonomicName>
, pl. 54 figs. 1–1a;
<treatmentCitation id="EE7035726F13FFA0FC1DF892FAA6F224" author="Kury" box="[981,1357,1816,1839]" page="53" pageId="1" pageNumber="243" year="2011">
<bibRefCitation id="0B406E926F13FFA0FC1DF892FACFF224" author="Kury" box="[981,1316,1816,1839]" pageId="1" pageNumber="243" refString="Kury, A. B. &amp; Alonso-Zarazaga, M. - A. (2011) Addenda and corrigenda to the &quot; Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones) &quot;. Zootaxa, 3034, 47 - 68." type="journal article" year="2011">Kury &amp; Alonso-Zarazaga 2011</bibRefCitation>
: 53
</treatmentCitation>
.
<taxonomicName id="A8D168E06F13FFA0FF5FF8B2FE71F244" box="[151,410,1848,1871]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="243" phylum="Arthropoda" rank="species" species="planus">
<emphasis id="5DA5CF716F13FFA0FF5FF8B2FE71F244" box="[151,410,1848,1871]" italics="true" pageId="1" pageNumber="243">Hernandarioides planus</emphasis>
</taxonomicName>
[unjustified emendation]:
<treatmentCitation id="EE7035726F13FFA0FD7EF8B2FC89F244" author="Kury" box="[694,866,1848,1871]" page="105" pageId="1" pageNumber="243" year="2003">
<bibRefCitation id="0B406E926F13FFA0FD7EF8B2FCC7F244" author="Kury" box="[694,812,1848,1871]" pageId="1" pageNumber="243" refString="Kury, A. B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. especial monografico, 1, 1 - 337." type="journal article" year="2003">Kury 2003</bibRefCitation>
: 105
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6F6E13636F13FFA0FF5FF8D2FC6CF284" blockId="1.[151,1434,1816,1935]" pageId="1" pageNumber="243">
<treatmentCitationGroup id="4FC1344D6F13FFA0FF5FF8D2FC6CF284" pageId="1" pageNumber="243">
<taxonomicName id="A8D168E06F13FFA0FF5FF8D2FD2DF264" authority="Goodnight &amp; Goodnight 1942: 18" authorityName="Goodnight &amp; Goodnight" authorityPageNumber="18" authorityYear="1942" box="[151,710,1880,1903]" class="Arachnida" family="Gonyleptidae" genus="Kaluga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="243" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="5DA5CF716F13FFA0FF5FF8D2FEA1F264" box="[151,330,1880,1903]" italics="true" pageId="1" pageNumber="243">Kaluga elongata</emphasis>
<treatmentCitation id="EE7035726F13FFA0FE9BF8D2FD2DF264" author="Goodnight" box="[339,710,1880,1903]" page="18" pageId="1" pageNumber="243" year="1942">
<bibRefCitation id="0B406E926F13FFA0FE9BF8D2FD76F264" author="Goodnight" box="[339,669,1880,1903]" pageId="1" pageNumber="243" refString="Goodnight, C. J. &amp; Goodnight, M. L. (1942) Phalangids from Central America and the West Indies. American Museum Novitates, 1184, 1 - 23." type="journal article" year="1942">Goodnight &amp; Goodnight 1942</bibRefCitation>
: 18
</treatmentCitation>
</taxonomicName>
, fig. 33 [junior subjective synonym of
<taxonomicName id="A8D168E06F13FFA0FBA7F8D2FE38F284" ID-CoL="7WL5B" authority="F.O. Pickard-Cambridge 1905" authorityName="F.O. Pickard-Cambridge" authorityYear="1905" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="243" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="5DA5CF716F13FFA0FBA7F8D2FA81F264" box="[1135,1386,1880,1903]" italics="true" pageId="1" pageNumber="243">Hernandarioides plana</emphasis>
<bibRefCitation id="0B406E926F13FFA0FABAF8D2FE38F284" author="Pickard-Cambridge" pageId="1" pageNumber="243" refString="Pickard-Cambridge, F. O. (1905) Order Opiliones [2 nd part]. In: Godman, F. D. &amp; Salvin, O. (Eds.), Biologia centrali- Americana. Vol. 2. Arachnida. Araneidea and Opiliones. R. H. Porter / Dulau &amp; Co., London, pp. 561 - 610, pls. 53, 54." type="journal article" year="1905">F.O. Pickard-Cambridge 1905</bibRefCitation>
</taxonomicName>
by
<treatmentCitation id="EE7035726F13FFA0FE33F8F2FC90F284" author="Goodnight" box="[507,891,1912,1935]" page="15" pageId="1" pageNumber="243" year="1947">
<bibRefCitation id="0B406E926F13FFA0FE33F8F2FC90F284" author="Goodnight" box="[507,891,1912,1935]" pageId="1" pageNumber="243" refString="Goodnight, C. J. &amp; Goodnight, M. L. (1947) Studies on the phalangid fauna of Central America. American Museum Novitates, 1340, 1 - 21." type="journal article" year="1947">Goodnight &amp; Goodnight (1947: 15)</bibRefCitation>
</treatmentCitation>
].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="27CB40E86F13FFA2FF5FF830FAD2F5DA" lastPageId="3" lastPageNumber="245" pageId="1" pageNumber="243" type="materials_examined">
<paragraph id="6F6E13636F13FFA0FF5FF830FCA4F2FF" blockId="1.[151,1436,1978,2036]" pageId="1" pageNumber="243">
<emphasis id="5DA5CF716F13FFA0FF5FF830FEFFF2DA" bold="true" box="[151,276,1978,2001]" pageId="1" pageNumber="243">
<typeStatus id="B06AADC16F13FFA0FF5FF830FF24F2DA" box="[151,207,1978,2001]" pageId="1" pageNumber="243">Type</typeStatus>
data.
</emphasis>
<taxonomicName id="A8D168E06F13FFA0FEE9F830FDCBF2DA" box="[289,544,1978,2001]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="243" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="5DA5CF716F13FFA0FEE9F830FDCBF2DA" box="[289,544,1978,2001]" italics="true" pageId="1" pageNumber="243">Hernandarioides plana</emphasis>
</taxonomicName>
: ♀
<typeStatus id="B06AADC16F13FFA0FD9DF830FD59F2DA" box="[597,690,1978,2001]" pageId="1" pageNumber="243" type="holotype">holotype</typeStatus>
(
<collectionCode id="09C08BA66F13FFA0FD0EF830FCF1F2DA" box="[710,794,1978,2001]" pageId="1" pageNumber="243">BMNH</collectionCode>
, examined), from
<collectingCountry id="17C653F36F13FFA0FC27F830FBADF2DA" box="[1007,1094,1978,2001]" name="Panama" pageId="1" pageNumber="243">Panama</collectingCountry>
, Bugaba.
<taxonomicName id="A8D168E06F13FFA0FB74F830FA9FF2DA" box="[1212,1396,1978,2001]" class="Arachnida" family="Gonyleptidae" genus="Kaluga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="1" pageNumber="243" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="5DA5CF716F13FFA0FB74F830FA9FF2DA" box="[1212,1396,1978,2001]" italics="true" pageId="1" pageNumber="243">Kaluga elongata</emphasis>
</taxonomicName>
: ♀
<typeStatus id="B06AADC16F13FFA0FF5FF857FF1FF2FF" box="[151,244,2013,2036]" pageId="1" pageNumber="243" type="holotype">holotype</typeStatus>
(
<collectionCode id="09C08BA66F13FFA0FECAF857FEB3F2FF" LSID="urn:lsid:biocol.org:col:34925" box="[258,344,2013,2036]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="1" pageNumber="243">AMNH</collectionCode>
, not examined),
<collectingCountry id="17C653F36F13FFA0FDC0F857FDB4F2FF" box="[520,607,2013,2036]" name="Panama" pageId="1" pageNumber="243">Panama</collectingCountry>
, Chiriqui. Casita Alta.
</paragraph>
<caption id="3BAE43EB6F10FFA3FF5FF8CDFCADF2CD" httpUri="https://zenodo.org/record/224627/files/figure.png" pageId="2" pageNumber="244" targetBox="[165,1422,194,1838]" targetPageId="2">
<paragraph id="6F6E13636F10FFA3FF5FF8CDFCADF2CD" blockId="2.[151,1436,1863,1990]" pageId="2" pageNumber="244">
<emphasis id="5DA5CF716F10FFA3FF5FF8CDFEBDF255" bold="true" box="[151,342,1863,1886]" pageId="2" pageNumber="244">FIGURES 8–13.</emphasis>
<taxonomicName id="A8D168E06F10FFA3FEABF8CDFC2CF255" authority="F.O. Pickard-Cambridge, 1905" authorityName="F.O. Pickard-Cambridge" authorityYear="1905" box="[355,967,1863,1886]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="244" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="5DA5CF716F10FFA3FEABF8CDFD88F255" box="[355,611,1863,1886]" italics="true" pageId="2" pageNumber="244">Hernandarioides plana</emphasis>
F.O. Pickard-Cambridge, 1905
</taxonomicName>
. ♂ (MIUP AK 004), from Jurutungo. 6. Habitus, dorsal view; 7. Habitus, lateral view; 8. Left Mt-Ta IV, prolateral view; 9. Left Pa-Ti IV, prolateral view. Penis, distal part: 10. Dorsal view; 11. Lateral view; 12. Ventral view; 13. Western Panama, showing the three localities of the specimens of
<taxonomicName id="A8D168E06F10FFA3FEE1F825FE6CF2CD" box="[297,391,1967,1990]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="2" pageNumber="244" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="5DA5CF716F10FFA3FEE1F825FE6CF2CD" box="[297,391,1967,1990]" italics="true" pageId="2" pageNumber="244">H. plana</emphasis>
</taxonomicName>
cited in the literature. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="6F6E13636F11FFA2FF0FFF12FAD2F5DA" blockId="3.[151,1436,152,1735]" pageId="3" pageNumber="245">
<emphasis id="5DA5CF716F11FFA2FF0FFF12FE05F5A4" bold="true" box="[199,494,152,175]" pageId="3" pageNumber="245">Other material examined.</emphasis>
<specimenCount id="79D7D8EA6F11FFA2FE3DFF12FDF5F5A4" box="[501,542,152,175]" pageId="3" pageNumber="245" type="generic">1 ♂</specimenCount>
(
<collectionCode id="09C08BA66F11FFA2FDE5FF12FD9AF5A4" box="[557,625,152,175]" pageId="3" pageNumber="245">MIUP</collectionCode>
AK 004) from
<collectingCountry id="17C653F36F11FFA2FCD1FF12FC9BF5A4" box="[793,880,152,175]" name="Panama" pageId="3" pageNumber="245">Panama</collectingCountry>
, Prov. Chiriquí: Jurutungo, Parque Internacional La Amistad (
<collectionCode id="09C08BA66F11FFA2FF37FF30FEABF5DA" box="[255,320,186,209]" pageId="3" pageNumber="245">PILA</collectionCode>
), Distrito Renacimiento,
<quantity id="A829BE866F11FFA2FD83FF30FD0BF5DA" box="[587,736,186,209]" metricMagnitude="3" metricUnit="m" metricValue="1.955" metricValueMax="2.11" metricValueMin="1.8" pageId="3" pageNumber="245" unit="m" value="1955.0" valueMax="2110.0" valueMin="1800.0">1800–2110 m</quantity>
,
<date id="1B6F35A36F11FFA2FD23FF30FC53F5DA" box="[747,952,186,209]" pageId="3" pageNumber="245" value="2006-03-13" valueMax="2006-03-16" valueMin="2006-03-13">13–16 March 2006</date>
, R.Miranda leg. See map in
<figureCitation id="F7EA0FE66F11FFA2FB2FFF30FADEF5DA" box="[1255,1333,186,209]" captionStart="FIGURES 8 – 13" captionStartId="2.[151,269,1863,1886]" captionTargetBox="[165,1422,194,1838]" captionTargetId="figure@2.[165,1422,194,1839]" captionTargetPageId="2" captionText="FIGURES 8 – 13. Hernandarioides plana F. O. Pickard-Cambridge, 1905. ♂ (MIUP AK 004), from Jurutungo. 6. Habitus, dorsal view; 7. Habitus, lateral view; 8. Left Mt-Ta IV, prolateral view; 9. Left Pa-Ti IV, prolateral view. Penis, distal part: 10. Dorsal view; 11. Lateral view; 12. Ventral view; 13. Western Panama, showing the three localities of the specimens of H. plana cited in the literature. Scale bars = 1 mm." httpUri="https://zenodo.org/record/224627/files/figure.png" pageId="3" pageNumber="245">Fig. 13</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="27CB40E86F11FFA2FF0FFF57FCE2F3CC" pageId="3" pageNumber="245" type="description">
<paragraph id="6F6E13636F11FFA2FF0FFF57FA77F172" blockId="3.[151,1436,152,1735]" pageId="3" pageNumber="245">
<emphasis id="5DA5CF716F11FFA2FF0FFF57FEA0F5FF" bold="true" box="[199,331,221,244]" pageId="3" pageNumber="245">Description</emphasis>
, male (MIUP AK 004).
<emphasis id="5DA5CF716F11FFA2FD98FF57FD15F5FF" bold="true" box="[592,766,221,244]" pageId="3" pageNumber="245">Measurements.</emphasis>
CL = 2.1, CW = 3.0, AL = 3.0, AW = 4.7. Appendages: Fe I = 1.8, Ti I = 1.6, Fe II = 3.1, Ti II = 2.0, Fe III = 2.9, Ti III = 2.1, Fe IV = 4.4, Ti IV = 2.4.
<emphasis id="5DA5CF716F11FFA2FBF3FE8AFB75F41C" bold="true" box="[1083,1182,256,279]" pageId="3" pageNumber="245">Dorsum.</emphasis>
Dorsal scutum gammashaped, widest at area II (
<figureCitation id="F7EA0FE66F11FFA2FE60FEA8FE19F432" box="[424,498,290,313]" captionStart="FIGURES 1 – 5" captionStartId="1.[151,269,1584,1607]" captionTargetBox="[151,1436,194,1559]" captionTargetId="figure@1.[151,1436,194,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 5. Hernandarioides plana F. O. Pickard-Cambridge, 1905. ♂ (MIUP AK 004), from Jurutungo. 1. Habitus, dorsal view; 2. Body, lateral view; 3. Free tergites and sternites, posterior view; 4. Left leg IV, dorso-retrolateral view; 5. Habitus, frontal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/224626/files/figure.png" pageId="3" pageNumber="245">Figs. 1</figureCitation>
, 6). Mesotergum divided into 4 ill-defined areas, areas 3–4 partially fused to each other (Fig. 6). Areas with transverse rows of small setiferous tubercles (Figs. 6, 7). Free tergites I–III each with a row of acuminate setiferous tubercles, free tergite II with central spine much larger than the others (Figs. 6, 7).
<emphasis id="5DA5CF716F11FFA2FADBFEE2FA8AF474" bold="true" box="[1299,1377,360,383]" pageId="3" pageNumber="245">Venter.</emphasis>
Free sternites, stigmatic area and coxae II–IV irregularly covered with minute granules. Coxa I with 1 row of robust setiferous tubercles plus some scantily placed. Stigmatic area attenuate Y-shaped, stigmata small. Coxae I–III oriented transversely, coxa IV oblique.
<emphasis id="5DA5CF716F11FFA2FE9FFE5AFE37F4EC" bold="true" box="[343,476,464,487]" pageId="3" pageNumber="245">Pedipalpus.</emphasis>
Articles only moderately elongate. Trochanter with 1 ventral setiferous tubercle; femur with a ventral row of 3 small setiferous tubercles; patella unarmed. Tibia-Tarsus flattened ventrally, each with 2 rows of spines: ventro-mesal and ventro-ectal IiIi.
<emphasis id="5DA5CF716F11FFA2FDACFD9FFD4AF727" bold="true" box="[612,673,533,556]" pageId="3" pageNumber="245">Legs.</emphasis>
Femora I–II straight, III–IV arched (
<figureCitation id="F7EA0FE66F11FFA2FBFEFD9FFB9CF727" box="[1078,1143,533,556]" captionStart="FIGURES 1 – 5" captionStartId="1.[151,269,1584,1607]" captionTargetBox="[151,1436,194,1559]" captionTargetId="figure@1.[151,1436,194,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 5. Hernandarioides plana F. O. Pickard-Cambridge, 1905. ♂ (MIUP AK 004), from Jurutungo. 1. Habitus, dorsal view; 2. Body, lateral view; 3. Free tergites and sternites, posterior view; 4. Left leg IV, dorso-retrolateral view; 5. Habitus, frontal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/224626/files/figure.png" pageId="3" pageNumber="245">Fig. 1</figureCitation>
). Podomeres I–IV without significant armature. Metatarsus I with calcaneus thickened. Femora III and IV (latter larger) arched, each with 2 rows of pointed tubercles, ventro-ectal and ventro-mesal. Tibia III with 2 ventral rows of setiferous tubercles. Coxa IV with dorso-lateral uniramous spiniform apophysis, pointed backwards. Trochanter IV short, with short retroapical spiniform apophysis. Tibia IV slightly incrassate at distal half, with 6 longitudinal rows of setiferous tubercles (
<figureCitation id="F7EA0FE66F11FFA2FB19FD2AFAF9F7BC" box="[1233,1298,672,695]" captionStart="FIGURES 8 – 13" captionStartId="2.[151,269,1863,1886]" captionTargetBox="[165,1422,194,1838]" captionTargetId="figure@2.[165,1422,194,1839]" captionTargetPageId="2" captionText="FIGURES 8 – 13. Hernandarioides plana F. O. Pickard-Cambridge, 1905. ♂ (MIUP AK 004), from Jurutungo. 6. Habitus, dorsal view; 7. Habitus, lateral view; 8. Left Mt-Ta IV, prolateral view; 9. Left Pa-Ti IV, prolateral view. Penis, distal part: 10. Dorsal view; 11. Lateral view; 12. Ventral view; 13. Western Panama, showing the three localities of the specimens of H. plana cited in the literature. Scale bars = 1 mm." httpUri="https://zenodo.org/record/224627/files/figure.png" pageId="3" pageNumber="245">Fig. 8</figureCitation>
). Metatarsus IV incrassate, densely covered with setiferous tubercles, loosely arranged into rows (
<figureCitation id="F7EA0FE66F11FFA2FBF1FD48FB96F7D2" box="[1081,1149,706,729]" captionStart="FIGURES 8 – 13" captionStartId="2.[151,269,1863,1886]" captionTargetBox="[165,1422,194,1838]" captionTargetId="figure@2.[165,1422,194,1839]" captionTargetPageId="2" captionText="FIGURES 8 – 13. Hernandarioides plana F. O. Pickard-Cambridge, 1905. ♂ (MIUP AK 004), from Jurutungo. 6. Habitus, dorsal view; 7. Habitus, lateral view; 8. Left Mt-Ta IV, prolateral view; 9. Left Pa-Ti IV, prolateral view. Penis, distal part: 10. Dorsal view; 11. Lateral view; 12. Ventral view; 13. Western Panama, showing the three localities of the specimens of H. plana cited in the literature. Scale bars = 1 mm." httpUri="https://zenodo.org/record/224627/files/figure.png" pageId="3" pageNumber="245">Fig. 9</figureCitation>
). Tarsal counts: 6(3)-6(3)/ 8(3)-8(3)/6-6/6-6.
<emphasis id="5DA5CF716F11FFA2FE94FD6FFDC5F7F7" bold="true" box="[348,558,741,764]" pageId="3" pageNumber="245">Color (in alcohol):</emphasis>
(see also
<figureCitation id="F7EA0FE66F11FFA2FD50FD6FFD14F7F7" box="[664,767,741,764]" captionStart="FIGURES 1 – 5" captionStartId="1.[151,269,1584,1607]" captionTargetBox="[151,1436,194,1559]" captionTargetId="figure@1.[151,1436,194,1559]" captionTargetPageId="1" captionText="FIGURES 1 – 5. Hernandarioides plana F. O. Pickard-Cambridge, 1905. ♂ (MIUP AK 004), from Jurutungo. 1. Habitus, dorsal view; 2. Body, lateral view; 3. Free tergites and sternites, posterior view; 4. Left leg IV, dorso-retrolateral view; 5. Habitus, frontal view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/224626/files/figure.png" pageId="3" pageNumber="245">Figs. 1–5</figureCitation>
). Body, coxae I–IV and Tr IV Strong Orange (50). Mesotergal areas lighter – Brilliant Orange (49). Body with some black mottling, strongest on free tergites I–III. Legs I–III Dark Greenish Yellow (103). Leg IV Deep Reddish Brown (41).
<emphasis id="5DA5CF716F11FFA2FCE3FCA0FC75F64A" bold="true" box="[811,926,810,833]" pageId="3" pageNumber="245">Genitalia:</emphasis>
(
<figureCitation id="F7EA0FE66F11FFA2FC67FCA0FBDFF64A" box="[943,1076,810,833]" captionStart="FIGURES 8 – 13" captionStartId="2.[151,269,1863,1886]" captionTargetBox="[165,1422,194,1838]" captionTargetId="figure@2.[165,1422,194,1839]" captionTargetPageId="2" captionText="FIGURES 8 – 13. Hernandarioides plana F. O. Pickard-Cambridge, 1905. ♂ (MIUP AK 004), from Jurutungo. 6. Habitus, dorsal view; 7. Habitus, lateral view; 8. Left Mt-Ta IV, prolateral view; 9. Left Pa-Ti IV, prolateral view. Penis, distal part: 10. Dorsal view; 11. Lateral view; 12. Ventral view; 13. Western Panama, showing the three localities of the specimens of H. plana cited in the literature. Scale bars = 1 mm." httpUri="https://zenodo.org/record/224627/files/figure.png" pageId="3" pageNumber="245">Figs. 10–12</figureCitation>
). Ventral plate elongate, roughly subrectangular, but with proximal constriction and narrowing distally. Ventral plate also apical cleft creating two lobes and strongly projected dorsally in basal half. Distribution of macrosetae asymmetrical in number and/or insertion. Latero-proximal = two pairs, with apex slightly spatulate; Latero-distal = one or two pairs, short, straight, laterosubdistal; Dorso-lateral = one pair, near the insertion of glans in the follis; Ventral = two pairs, minute, positioned as a rectangle on ventral surface of VP. Follis roughly cubic, well-developed. Glans elongate, with stylus sinuous, inserted at its ventro-apical region.
<emphasis id="5DA5CF716F11FFA2FE61FC70FD7FF11A" bold="true" box="[425,660,1018,1041]" pageId="3" pageNumber="245">Sexual dimorphism:</emphasis>
armature and proportions of body and tergites similar in both sexes. Spiniform processes of coxa to tibia IV more strongly developed in male. Femur IV slightly incrassate distally in male. Metatarsus IV clavate only in male.
<emphasis id="5DA5CF716F11FFA2FDEAFBCAFD63F15C" bold="true" box="[546,648,1088,1111]" pageId="3" pageNumber="245">Remark:</emphasis>
The male MIUP AK 004 is here assigned to
<taxonomicName id="A8D168E06F11FFA2FBBEFBCAFB3EF15C" box="[1142,1237,1088,1111]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="species" species="plana">
<emphasis id="5DA5CF716F11FFA2FBBEFBCAFB3EF15C" box="[1142,1237,1088,1111]" italics="true" pageId="3" pageNumber="245">H. plana</emphasis>
</taxonomicName>
because all subtle details in granulation and shape of posterior margin of dorsal scutum and free tergites match exactly the female
<typeStatus id="B06AADC16F11FFA2FAFFFBE8FA73F172" box="[1335,1432,1122,1145]" pageId="3" pageNumber="245" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="6F6E13636F11FFA2FF0FFB0FFBA9F337" blockId="3.[151,1436,152,1735]" pageId="3" pageNumber="245">
Most genera of Ampycinae are poorly known, but
<taxonomicName id="A8D168E06F11FFA2FD29FB0FFC7CF197" box="[737,919,1157,1180]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FD29FB0FFC7CF197" box="[737,919,1157,1180]" italics="true" pageId="3" pageNumber="245">Hernandarioides</emphasis>
</taxonomicName>
does not seem to be especially closely related to any other genus.
<taxonomicName id="A8D168E06F11FFA2FE86FB22FDEFF1B4" box="[334,516,1192,1215]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FE86FB22FDEFF1B4" box="[334,516,1192,1215]" italics="true" pageId="3" pageNumber="245">Hernandarioides</emphasis>
</taxonomicName>
differs from the complex
<taxonomicName id="A8D168E06F11FFA2FCD6FB22FC21F1B4" authority="Simon" authorityName="Simon" box="[798,970,1192,1215]" class="Arachnida" family="Gonyleptidae" genus="Ampycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FCD6FB22FC95F1B4" box="[798,894,1192,1215]" italics="true" pageId="3" pageNumber="245">Ampycus</emphasis>
Simon
</taxonomicName>
/
<taxonomicName id="A8D168E06F11FFA2FC11FB22FB40F1B4" authority="Roewer" authorityName="Roewer" box="[985,1195,1192,1215]" class="Arachnida" family="Gonyleptidae" genus="Hexabunus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FC11FB22FBBAF1B4" box="[985,1105,1192,1215]" italics="true" pageId="3" pageNumber="245">Hexabunus</emphasis>
Roewer
</taxonomicName>
/
<taxonomicName id="A8D168E06F11FFA2FB72FB22FA73F1B4" authority="Roewer" authorityName="Roewer" box="[1210,1432,1192,1215]" class="Arachnida" family="Gonyleptidae" genus="Pirunipygus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FB72FB22FAD7F1B4" box="[1210,1340,1192,1215]" italics="true" pageId="3" pageNumber="245">Pirunipygus</emphasis>
Roewer
</taxonomicName>
, because those genera have diamond–shaped ventral plate, canoe-shaped erect stylus and four pairs of latero-distal macrosetae.
<taxonomicName id="A8D168E06F11FFA2FED4FB67FDD5F00F" authority="Soares &amp; Soares" authorityName="Soares &amp; Soares" box="[284,574,1261,1284]" class="Arachnida" family="Gonyleptidae" genus="Hutamaia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FED4FB67FE6CF00F" box="[284,391,1261,1284]" italics="true" pageId="3" pageNumber="245">Hutamaia</emphasis>
Soares &amp; Soares
</taxonomicName>
differs from
<taxonomicName id="A8D168E06F11FFA2FD04FB67FC69F00F" box="[716,898,1261,1284]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FD04FB67FC69F00F" box="[716,898,1261,1284]" italics="true" pageId="3" pageNumber="245">Hernandarioides</emphasis>
</taxonomicName>
in having free tergite III armed instead of II, stout hooked dorsal apophysis on Tr II, strong retrolateral armature on Tr IV, pyriform ventral plate, oblique in relation to truncus, with the macrosetae displaced distally, very thin follis and stylus erect and much elongate.
<taxonomicName id="A8D168E06F11FFA2FB1DFAB8FA73F042" authority="Roewer" authorityName="Roewer" box="[1237,1432,1330,1353]" class="Arachnida" family="Gonyleptidae" genus="Glysterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FB1DFAB8FAD0F042" box="[1237,1339,1330,1353]" italics="true" pageId="3" pageNumber="245">Glysterus</emphasis>
Roewer
</taxonomicName>
, possesses free tergite III in males with stout armature (which may be bi- or trifurcate), trochanter IV with inner armature, metatarsus IV sinuous and armed. Penis (undescribed) is significantly different in pattern of macrosetae and shape of stylus and ventral plate.
<taxonomicName id="A8D168E06F11FFA2FE68FA10FDBDF0BA" box="[416,598,1434,1457]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FE68FA10FDBDF0BA" box="[416,598,1434,1457]" italics="true" pageId="3" pageNumber="245">Hernandarioides</emphasis>
</taxonomicName>
looks similar to
<taxonomicName id="A8D168E06F11FFA2FCC7FA10FBC7F0BA" authority="Chamberlin" authorityName="Chamberlin" box="[783,1068,1434,1457]" class="Arachnida" family="Gonyleptidae" genus="Nesopachylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FCC7FA10FC48F0BA" box="[783,931,1434,1457]" italics="true" pageId="3" pageNumber="245">Nesopachylus</emphasis>
Chamberlin
</taxonomicName>
, (only one species from
<collectingCountry id="17C653F36F11FFA2FAF0FA10FA7AF0BA" box="[1336,1425,1434,1457]" name="Panama" pageId="3" pageNumber="245">Panama</collectingCountry>
), both with unarmed scutum, prominent medial apophysis on free tergite II, leg IV roughly granular, but without further conspicuous armature, and metatarsus IV incrassate.
<taxonomicName id="A8D168E06F11FFA2FD17FA6AFC98F0FC" box="[735,883,1504,1527]" class="Arachnida" family="Gonyleptidae" genus="Nesopachylus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FD17FA6AFC98F0FC" box="[735,883,1504,1527]" italics="true" pageId="3" pageNumber="245">Nesopachylus</emphasis>
</taxonomicName>
differs by having long dorsal and latero-proximal macrosetae, a pyriform ventral plate and a canoe-shaped stylus, much elongate and not nearly as sinuous as in
<taxonomicName id="A8D168E06F11FFA2FF5FF9AFFEA6F337" box="[151,333,1573,1596]" class="Arachnida" family="Gonyleptidae" genus="Hernandarioides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="245" phylum="Arthropoda" rank="genus">
<emphasis id="5DA5CF716F11FFA2FF5FF9AFFEA6F337" box="[151,333,1573,1596]" italics="true" pageId="3" pageNumber="245">Hernandarioides</emphasis>
</taxonomicName>
, with marginal serrations, and also follis much narrower than truncus).
</paragraph>
<paragraph id="6F6E13636F11FFA2FF0FF9C2FCE2F3CC" blockId="3.[151,1436,152,1735]" pageId="3" pageNumber="245">This study was supported by grant # 562149/2010-4 (PROTAX – OPESC project) and scholarship # 477502/2012-1 (Universal 14/2012) from CNPq, and grant E-26/111.705/2012 (APQ1) from FAPERJ to ABK. The authors wish to thank Osvaldo Villarreal for his useful suggestions and comments to the draft. Amanda Mendes and an unknown referee improved the original manuscript with insightful criticism.</paragraph>
</subSubSection>
</treatment>
</document>
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