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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1098.73223" ID-Pensoft-Pub="1313-2970-1098-1" ID-Pensoft-UUID="B22184E75B1A59C9B985C0DD9F092196" ID-ZooBank="B4F4033F296E43CCBE54B9413BC19268" LSID="urn:lsid:zoobank.org:act:BDE5DB12-AFBD-41F7-95F5-966AF8E790F3" ModsDocID="1313-2970-1098-1" checkinTime="1651618226429" checkinUser="pensoft" docAuthor="Rakotonirina, Jean Claude &amp; Fisher, Brian L." docDate="2022" docId="E59160DA8FA65AB8B878429A200E7150" docLanguage="en" docName="ZooKeys 1098: 1-180" docOrigin="ZooKeys 1098" docPubDate="2022-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1098.73223" docTitle="Camponotus kelimaso Rakotonirina &amp; Fisher 2022, sp. nov." docType="treatment" docUuid="BDE5DB12-AFBD-41F7-95F5-966AF8E790F3" docUuidSource="ZooBank" docVersion="1" id="B22184E75B1A59C9B985C0DD9F092196" lastPageNumber="1" masterDocId="B22184E75B1A59C9B985C0DD9F092196" masterDocTitle="Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="180" masterPageNumber="1" pageId="0" pageNumber="1" updateTime="1651618226429" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rakotonirina, Jean Claude</mods:namePart>
<mods:affiliation>Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
<mods:nameIdentifier type="email">jcrakoto25@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
<mods:affiliation>Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-05-03</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1098</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>180</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1098.73223</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1098-1</mods:identifier>
<mods:identifier type="ZooBank">B4F4033F296E43CCBE54B9413BC19268</mods:identifier>
<mods:identifier type="Pensoft-UUID">B22184E75B1A59C9B985C0DD9F092196</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:BDE5DB12-AFBD-41F7-95F5-966AF8E790F3" httpUri="http://treatment.plazi.org/id/E59160DA8FA65AB8B878429A200E7150" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/BDE5DB12-AFBD-41F7-95F5-966AF8E790F3" authority="Rakotonirina &amp; Fisher, 2022" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus kelimaso" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kelimaso" status="sp. nov.">Camponotus kelimaso</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Head in full-face view A C. liandia (CASENT 0491145) B C. kelimaso (CASENT 0487718)." figureDoi="10.3897/zookeys.1098.73223.figure34" httpUri="https://binary.pensoft.net/fig/678093" pageId="0" pageNumber="1">Figs 34B</figureCitation>
<figureCitation captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Second and third abdominal tergites in dorsal view A C. kelimaso (CASENT 0487718) B C. quadrimaculatus (CASENT 0096044) C C. gibber (CASENT 0491497)." figureDoi="10.3897/zookeys.1098.73223.figure35" httpUri="https://binary.pensoft.net/fig/678094" pageId="0" pageNumber="1">, 35A</figureCitation>
<figureCitation captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Head in full-face view A C. kelimaso (CASENT 0487718) B C. lubbocki (CASENT 0486998)." figureDoi="10.3897/zookeys.1098.73223.figure37" httpUri="https://binary.pensoft.net/fig/678096" pageId="0" pageNumber="1">, 37A</figureCitation>
<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Camponotus kelimaso A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0487718 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure63" httpUri="https://binary.pensoft.net/fig/678122" pageId="0" pageNumber="1">, 63</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="holotype worker">
<paragraph pageId="0" pageNumber="1">Holotype worker.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
: Province
<emphasis bold="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
: PN Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, -14.435, 49.76, 775 m, rainforest, ex rotten log, 16 Nov 2003 (B.L. Fisher et al.) collection code: BLF09010, specimen code: CASENT0487718 (CAS).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="paratypes">
<paragraph pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="1">3 minor workers of same data as holotype but specimens coded as: CASENT0837637, CASENT0837636, CASENT0837635 (NHMUK, PBZT, CAS).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="B. L. Fisher, PN Marojejy" country="Madagascar" county="Makirovana forest" elevation="715" latitude="-14.10295" location="Manantenina River" longLatPrecision="1" longitude="50.01984" municipality="Makirovana forest" specimenCount="1" stateProvince="Antsiranana">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingCountry name="Madagascar">Madagascar</collectingCountry>
:
<collectingRegion country="Madagascar" name="Antsiranana">Antsiranana</collectingRegion>
</emphasis>
:
<collectingCounty>Makirovana forest</collectingCounty>
, -14.16666, 49.95,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.15" unit="m" value="715.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.15" unit="m" value="715.0">715 m</elevation>
</quantity>
, rainforest (
<collectorName>B.L. Fisher</collectorName>
et al.) (CAS);
<collectingMunicipality>Makirovana forest</collectingMunicipality>
,
<geoCoordinate degrees="14.10295" direction="south" orientation="latitude" precision="1" value="-14.10295">-14.10295</geoCoordinate>
,
<geoCoordinate degrees="50.01984" direction="east" orientation="longitude" precision="1" value="50.01984">50.01984</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.9" unit="m" value="390.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.9" unit="m" value="390.0">390 m</elevation>
</quantity>
, rainforest (
<collectorName>B.L. Fisher</collectorName>
et al.) (CAS);
<collectorName>PN Marojejy</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:E59160DA8FA65AB8B878429A200E7150:501C61627AC43920AADFF347C729287D" country="Madagascar" county="Makirovana forest" latitude="-14.10295" longLatPrecision="1" longitude="50.01984" municipality="Makirovana forest" name="Manantenina River" stateProvince="Antsiranana">Manantenina River</location>
,
<locationDeviation location="Andapa">
<quantity metricMagnitude="4" metricUnit="m" metricValue="2.76" unit="km" value="27.6">27.6 km</quantity>
35° NE Andapa
</locationDeviation>
,
<locationDeviation location="Manantenina">
<quantity metricMagnitude="3" metricUnit="m" metricValue="9.6" unit="km" value="9.6">9.6 km</quantity>
327° NNW Manantenina
</locationDeviation>
, -14.435, 49.76,
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.75" unit="m" value="775.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.75" unit="m" value="775.0">775 m</elevation>
</quantity>
, rainforest (
<collectorName>B.L. Fisher</collectorName>
) (CAS)
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="R. Harin'Hala, M. E. Irwin" country="Madagascar" county="Foret d'Ambalagoavy Nord" elevation="625" latitude="-21.857067" location="Ambatombe" longLatPrecision="1" longitude="47.37849" municipality="Ikongo" specimenCount="1" stateProvince="Fianarantsoa">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Madagascar" name="Fianarantsoa">Fianarantsoa</collectingRegion>
</emphasis>
:
<collectingCounty>
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAmbalagoavy">d'Ambalagoavy</normalizedToken>
Nord
</collectingCounty>
,
<collectingMunicipality>Ikongo</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:E59160DA8FA65AB8B878429A200E7150:7A70946CC7D53B4A670D4B8D2504B5BD" country="Madagascar" county="Foret d'Ambalagoavy Nord" latitude="-21.857067" longLatPrecision="1" longitude="47.37849" municipality="Ikongo" name="Ambatombe" stateProvince="Fianarantsoa">Ambatombe</location>
,
<geoCoordinate degrees="21.857068" direction="south" orientation="latitude" precision="1" value="-21.857067">-21.857068</geoCoordinate>
,
<geoCoordinate degrees="47.37849" direction="east" orientation="longitude" precision="1" value="47.37849">47.37849</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.25" unit="m" value="625.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.25" unit="m" value="625.0">625 m</elevation>
</quantity>
(
<collectorName>
R.
<normalizedToken originalValue="HarinHala">Harin'Hala</normalizedToken>
</collectorName>
&amp;
<collectorName>M.E. Irwin</collectorName>
) (CAS)
</materialsCitation>
.
<materialsCitation collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="RS Ambatovaky, B. L. Fisher" country="Madagascar" elevation="430" latitude="-16.7702" location="Sandrangato river" longLatPrecision="5" longitude="49.26638" municipality="Sandrangato river" specimenCount="1" stateProvince="Toamasina">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="Madagascar" name="Toamasina">Toamasina</collectingRegion>
</emphasis>
:
<collectorName>RS Ambatovaky</collectorName>
,
<collectingMunicipality>Sandrangato river</collectingMunicipality>
,
<geoCoordinate degrees="16.7755" direction="south" orientation="latitude" precision="5" value="-16.7755">-16.7755</geoCoordinate>
,
<geoCoordinate degrees="49.26427" direction="east" orientation="longitude" precision="1" value="49.26427">49.26427</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.3" unit="m" value="430.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.3" unit="m" value="430.0">430 m</elevation>
</quantity>
, rainforest (
<collectorName>B.L. Fisher</collectorName>
et al.) (CAS)
<collectorName>RS Ambatovaky</collectorName>
,
<location LSID="urn:lsid:plazi:treatment:E59160DA8FA65AB8B878429A200E7150:EA68BE2298368E5E25948A460584FC24" country="Madagascar" latitude="-16.7702" longLatPrecision="5" longitude="49.26638" municipality="Sandrangato river" name="Sandrangato river" stateProvince="Toamasina">Sandrangato river</location>
,
<geoCoordinate degrees="16.7702" direction="south" orientation="latitude" precision="5" value="-16.7702">-16.7702</geoCoordinate>
,
<geoCoordinate degrees="49.26638" direction="east" orientation="longitude" precision="1" value="49.26638">49.26638</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</elevation>
</quantity>
, rainforest (
<collectorName>B.L. Fisher</collectorName>
et al.) (CAS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">In full-face view, lateral margins of head anterior to eye level diverging posteriorly; anterior clypeal margin truncate; dorsum of second and third abdominal tergites lacking white spot; propodeal dorsum straight; body color dark brown; antennal scape without erect hairs.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, head sides diverging towards anterior to level of eye, evenly rounding to approximately straight posterior margin; head widest at eye level; eyes almost flattened and small (EL/CS: 0.19
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.16-0.21), not breaking lateral cephalic margin, level of posterior margin located approximately at posterior 1/3 of head (PoOc/CL: 0.28
<normalizedToken originalValue="±">+/-</normalizedToken>
0.01; 0.27-0.29); frontal carinae more or less wide (FR/CS: 0.31
<normalizedToken originalValue="±">+/-</normalizedToken>
0.02; 0.28-0.34), posteriorly diverging, distance between them smaller than their smallest distance to eye; clypeus with anterolateral angle and approximately straight anteromedian margin; mandible with two apical teeth distantly spaced; antennal scape relatively short (SL/CS: 1.19
<normalizedToken originalValue="±">+/-</normalizedToken>
0.05; 1.11-1.27). Promesonotum weakly convex; mesonotum with posterior portion flat immediately anterior to weakly visible metanotal groove; propodeal dorsum almost straight, its angle to declivity widely rounded; propodeal dorsum ca. 2
<normalizedToken originalValue="×">x</normalizedToken>
as long as declivity. Petiolar node flattened anteroposteriorly, without obvious dorsal margin; anterior face almost the same height as posterior face; femur of hind leg rounded axially, not twisted basally.
</paragraph>
<paragraph pageId="0" pageNumber="1">First and second gastral tergites without a pair of white spots; lateral cephalic margin with erect hairs anterior to level of eyes; no erect hairs on lateral margin of head posterior to eye level; two erect hairs present near posterior margin; antennal scape only covered with appressed hairs; pronotum with few erect hairs; mesonotum with a pair of erect hairs; two erect hairs present on anterior to posterodorsal corner of propodeum. Body color reddish brown to pale brown.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
Differing from minor worker in the following characters: enlarged head (CS: 2.83
<normalizedToken originalValue="±">+/-</normalizedToken>
0.25; 2.35-3.15; CWb/CL: 0.99
<normalizedToken originalValue="±">+/-</normalizedToken>
0.04; 0.92-1.04) with broadly concave posterior margin; apical 1/4 of antennal scape surpassing posterior cephalic margin; robust mesosoma with separate convexity of promesonotum, lower position of metanotum and propodeum, propodeal dorsum approximately the same length as declivity surface and with rounded junction; petiolar node more flattened anteroposteriorly.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The distribution of
<taxonomicName genus="C." lsidName="C. kelimaso" pageId="0" pageNumber="1" rank="species" species="kelimaso">
<emphasis italics="true" pageId="0" pageNumber="1">C. kelimaso</emphasis>
</taxonomicName>
is limited to the eastern lowland rainforest of Madagascar between the Makirovana forest in the north and the
<normalizedToken originalValue="Forêt">Foret</normalizedToken>
<normalizedToken originalValue="dAmbalagoavy">d'Ambalagoavy</normalizedToken>
Nord in the south (Fig.
<figureCitation captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Camponotus kelimaso A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0487718 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure63" httpUri="https://binary.pensoft.net/fig/678122" pageId="0" pageNumber="1">63D</figureCitation>
). It nests mainly in rotten logs and rarely in root mat layers in the ground. Foraging is carried out on the ground and through leaf litter.
</paragraph>
<caption doi="10.3897/zookeys.1098.73223.figure63" httpUri="https://binary.pensoft.net/fig/678122" pageId="0" pageNumber="1" start="Figure 63" startId="F63">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 63.</emphasis>
<taxonomicName authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus kelimaso" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kelimaso">
<emphasis italics="true" pageId="0" pageNumber="1">Camponotus kelimaso</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral view
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
head in full-face view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
dorsal view of holotype minor worker CASENT0487718
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
distribution map.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus kelimaso" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kelimaso">
<emphasis italics="true" pageId="0" pageNumber="1">Camponotus kelimaso</emphasis>
</taxonomicName>
can be separated from
<taxonomicName genus="C." lsidName="C. immaculatus" pageId="0" pageNumber="1" rank="species" species="immaculatus">
<emphasis italics="true" pageId="0" pageNumber="1">C. immaculatus</emphasis>
</taxonomicName>
by its straight propodeal dorsum. It can be differentiated from
<taxonomicName genus="C." lsidName="C. lubbocki" pageId="0" pageNumber="1" rank="species" species="lubbocki">
<emphasis italics="true" pageId="0" pageNumber="1">C. lubbocki</emphasis>
</taxonomicName>
by the approximately straight posterior margin of the head and its small compound eyes.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Based on the information provided by the NC-clustering method, the grouping of
<taxonomicName genus="C." lsidName="C. kelimaso" pageId="0" pageNumber="1" rank="species" species="kelimaso">
<emphasis italics="true" pageId="0" pageNumber="1">C. kelimaso</emphasis>
</taxonomicName>
supports the distinction of the species by conventional qualitative taxonomy. The confirmatory LDA identified the samples successfully at 100%.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The species name
<emphasis italics="true" pageId="0" pageNumber="1">kelimaso</emphasis>
is a non-latin word derived from the Malagasy word for &quot;small eye&quot;. It refers to the fact that its compound eyes are small compared to those of other species in the subgenus
<taxonomicName class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus (Myrmosaga)" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Myrmosaga">Camponotus Myrmosaga</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>