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<mods:title id="CE015678E77E28C85C5EB24E9959D7E4">Deep-water, offshore, and new records of Schindlers fishes, Schindleria (Teleostei, Gobiidae), from the Indo-west Pacific collected during the Dana-Expedition, 1928 - 1930</mods:title>
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<subSubSection id="168C655A417DFFACFF76F9667597A5C9" box="[151,484,1757,1784]" pageId="1" pageNumber="452" type="nomenclature">
<paragraph id="5E2936D1417DFFACFF76F9667597A5C9" blockId="1.[151,1437,1757,2036]" box="[151,484,1757,1784]" pageId="1" pageNumber="452">
<heading id="056181BD417DFFACFF76F9667597A5C9" bold="true" box="[151,484,1757,1784]" fontSize="11" level="1" pageId="1" pageNumber="452" reason="1">
<emphasis id="6CE2EAC3417DFFACFF76F9667597A5C9" bold="true" box="[151,484,1757,1784]" pageId="1" pageNumber="452">
Classification of
<taxonomicName id="99964D52417DFFACFEBFF9667597A5C9" ID-CoL="7DF7" ID-ENA="1229173" authorityName="Giltay" authorityYear="1934" box="[350,484,1757,1784]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="452" phylum="Chordata" rank="genus">
<emphasis id="6CE2EAC3417DFFACFEBFF9667597A5C9" bold="true" box="[350,484,1757,1784]" italics="true" pageId="1" pageNumber="452">Schindleria</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="168C655A417DFFAFFF76F8B97632A00D" lastPageId="2" lastPageNumber="453" pageId="1" pageNumber="452" type="description">
<paragraph id="5E2936D1417DFFACFF76F8B9772FA471" blockId="1.[151,1437,1757,2036]" pageId="1" pageNumber="452">
In the classification of
<taxonomicName id="99964D52417DFFACFE7BF8BA764FA42A" authorityName="Giltay" authorityYear="1934" box="[410,572,1793,1819]" class="Actinopterygii" family="Schindleriidae" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="452" phylum="Chordata" rank="family">Schindleriidae</taxonomicName>
as a junior synonym of
<taxonomicName id="99964D52417DFFACFCAFF8BA77C6A42A" box="[846,949,1793,1819]" class="Actinopterygii" family="Gobiidae" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="452" phylum="Chordata" rank="family">Gobiidae</taxonomicName>
we follow
<bibRefCitation id="3A074B20417DFFACFBD4F8BA7096A42D" author="Thacker, C. E." box="[1077,1253,1793,1820]" pageId="1" pageNumber="452" pagination="93 - 104" refId="ref13786" refString="Thacker, C. E. (2009) Phylogeny of Gobioidei and placement within Acanthomorpha, with a new classification and investigation of diversification and character evolution. Copeia, 2009, 93 - 104. https: // doi. org / 10.1643 / CI- 08 - 004" type="journal article" year="2009">Thacker (2009)</bibRefCitation>
, a view adopted by several authors (
<bibRefCitation id="3A074B20417DFFACFE93F89E7631A471" author="Gill, A. C. &amp; Mooi, R. D." box="[370,578,1829,1856]" pageId="1" pageNumber="452" pagination="51 - 59" refId="ref12108" refString="Gill, A. C. &amp; Mooi, R. D. (2010) Character evidence for the monophyly of Microdesminae, with comments on relationships to Schindleria (Teleostei: Gobioidei: Gobiidae). Zootaxa, 2442 (1), 51 - 59. https: // doi. org / 10.11646 / zootaxa. 2442.1.4" type="journal article" year="2010">Gill &amp; Mooi 2010</bibRefCitation>
;
<bibRefCitation id="3A074B20417DFFACFDACF89D7723A471" author="Betancur-R, R. &amp; Wiley, E. O. &amp; Arratia, G. &amp; Acero, A. &amp; Bailly, N. &amp; Miya, M. &amp; Lecointre, G. &amp; Orti, G." box="[589,848,1829,1856]" pageId="1" pageNumber="452" pagination="162" refId="ref11523" refString="Betancur-R, R., Wiley, E. O., Arratia, G., Acero, A., Bailly, N., Miya, M., Lecointre, G. &amp; Orti, G. (2017) Phylogenetic classification of bony fishes. BMC Evolutionary Biology, 17, 162. https: // doi. org / 10.1186 / s 12862 - 017 - 0958 - 3" type="journal article" year="2017">
Betancur-R
<emphasis id="6CE2EAC3417DFFACFD34F89D777EA40E" box="[725,781,1829,1855]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="5E2936D1417DFFAFFF26F8F276ADA272" blockId="1.[151,1437,1757,2036]" lastBlockId="2.[151,1436,153,828]" lastPageId="2" lastPageNumber="453" pageId="1" pageNumber="452">
Six nominal species of Schindlers fishes are documented, four from the Pacific and two from the Indian Ocean (Red Sea) (
<bibRefCitation id="3A074B20417DFFACFEFBF8D67649A4B9" author="Ahnelt, H. &amp; Sauberer, M." box="[282,570,1901,1928]" pageId="1" pageNumber="452" pagination="95 - 108" refId="ref11356" refString="Ahnelt, H. &amp; Sauberer, M. (2018) A new species of Schindler's fish (Teleostei: Gobiidae: Schindleria) from the Malay Archipelago (Southeast Asia), with notes on the caudal fin complex of Schindleria. Zootaxa, 4531 (1), 95 - 108. https: // doi. org / 10.11646 / zootaxa. 4531.1.4" type="journal article" year="2018">Ahnelt &amp; Sauberer 2018</bibRefCitation>
). But the number of species is obviously underestimated.
<bibRefCitation id="3A074B20417DFFACFB37F8D571EBA4B9" author="Kon, T. &amp; Yoshino, T. &amp; Mukai, T. &amp; Nishida M." box="[1238,1432,1901,1928]" pageId="1" pageNumber="452" pagination="53 - 62" refId="ref12605" refString="Kon, T., Yoshino, T., Mukai, T. &amp; Nishida M. (2007) DNA sequences identify numerous cryptic species of the vertebrate: A lesson from the gobioid fish Schindleria. Molecular Phylogenetics and Evolution, 44, 53 - 62. https: // doi. org / 10.1016 / j. ympev. 2006.12.007" type="journal article" year="2007">
Kon
<emphasis id="6CE2EAC3417DFFACFAF1F8D57138A4B6" box="[1296,1355,1901,1927]" italics="true" pageId="1" pageNumber="452">et al.</emphasis>
(2007
</bibRefCitation>
,
<bibRefCitation id="3A074B20417DFFACFF76F82974A3A49D" author="Kon, T. &amp; Yoshino, T. &amp; Nishida, M." box="[151,208,1938,1964]" pageId="1" pageNumber="452" pagination="62 - 66" refId="ref12669" refString="Kon, T., Yoshino, T. &amp; Nishida, M. (2011) Cryptic species of the gobioid paedomorphic genus Schindleria from Palau, western Pacific Ocean. Ichthyological Research, 58, 62 - 66. https: // doi. org / 10.1007 / s 10228 - 010 - 0178 - y" type="journal article" year="2011">2011</bibRefCitation>
) found more than 30 cryptic species just in the western Pacific. Separation of these extremely paedomorphic species based on morphological characters is challenging (
<bibRefCitation id="3A074B20417DFFACFCC4F80D77ABA4E1" author="Kon, T. &amp; Yoshino, T. &amp; Mukai, T. &amp; Nishida M." box="[805,984,1973,2000]" pageId="1" pageNumber="452" pagination="53 - 62" refId="ref12605" refString="Kon, T., Yoshino, T., Mukai, T. &amp; Nishida M. (2007) DNA sequences identify numerous cryptic species of the vertebrate: A lesson from the gobioid fish Schindleria. Molecular Phylogenetics and Evolution, 44, 53 - 62. https: // doi. org / 10.1016 / j. ympev. 2006.12.007" type="journal article" year="2007">
Kon
<emphasis id="6CE2EAC3417DFFACFCBFF80D77FCA4FE" box="[862,911,1973,1999]" italics="true" pageId="1" pageNumber="452">et al</emphasis>
. 2007
</bibRefCitation>
;
<bibRefCitation id="3A074B20417DFFACFC03F80E7084A4E1" author="Ahnelt, H. &amp; Sauberer, M." box="[994,1271,1973,2000]" pageId="1" pageNumber="452" pagination="95 - 108" refId="ref11356" refString="Ahnelt, H. &amp; Sauberer, M. (2018) A new species of Schindler's fish (Teleostei: Gobiidae: Schindleria) from the Malay Archipelago (Southeast Asia), with notes on the caudal fin complex of Schindleria. Zootaxa, 4531 (1), 95 - 108. https: // doi. org / 10.11646 / zootaxa. 4531.1.4" type="journal article" year="2018">Ahnelt &amp; Sauberer 2018</bibRefCitation>
;
<bibRefCitation id="3A074B20417DFFACFAE0F80E71E2A4E1" author="Ahnelt, H." box="[1281,1425,1973,2000]" pageId="1" pageNumber="452" pagination="450 - 456" refId="ref11305" refString="Ahnelt, H. (2019) Redescription of the paedomorphic goby Schindleria nigropunctata Fricke &amp; El-Regal 2017 (Teleostei: Gobiidae) from the Red Sea. Zootaxa, 4615 (3), 450 - 456. https: // doi. org / 10.11646 / zootaxa. 4615.3.2" type="journal article" year="2019">Ahnelt 2019</bibRefCitation>
). The most striking difference between the first described species of
<taxonomicName id="99964D52417DFFACFC9FF862778CA4C2" authorityName="Giltay" authorityYear="1934" box="[894,1023,2009,2035]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="452" phylum="Chordata" rank="genus">
<emphasis id="6CE2EAC3417DFFACFC9FF862778CA4C2" box="[894,1023,2009,2035]" italics="true" pageId="1" pageNumber="452">Schindleria</emphasis>
</taxonomicName>
,
<taxonomicName id="99964D52417DFFACFBEDF861711DA4C5" authority="(Schindler 1930)" baseAuthorityName="Schindler" baseAuthorityYear="1930" box="[1036,1390,2009,2036]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="452" phylum="Chordata" rank="species" species="praematura">
<emphasis id="6CE2EAC3417DFFACFBEDF86170D8A4C2" box="[1036,1195,2010,2036]" italics="true" pageId="1" pageNumber="452">S. praematura</emphasis>
(
<bibRefCitation id="3A074B20417DFFACFB58F8627116A4C5" author="Schindler, O." box="[1209,1381,2009,2036]" pageId="1" pageNumber="452" pagination="79 - 80" refId="ref13610" refString="Schindler, O. (1930) Ein neuer Hemirhamphus aus dem Pazifischen Ozean. Anzeiger der Akademie der Wissenschaften Wien, 67, 79 - 80." type="journal article" year="1930">Schindler 1930</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="99964D52417EFFAFFF76FF22767DA382" authority="(Schindler 1931)" baseAuthorityName="Schindler" baseAuthorityYear="1931" box="[151,526,153,179]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="pietschmanni">
<emphasis id="6CE2EAC3417EFFAFFF76FF22753BA382" box="[151,328,153,179]" italics="true" pageId="2" pageNumber="453">S. pietschmanni</emphasis>
(
<bibRefCitation id="3A074B20417EFFAFFEB6FF227675A382" author="Schindler, O." box="[343,518,153,179]" pageId="2" pageNumber="453" pagination="2 - 3" refId="ref13638" refString="Schindler, O. (1931) Ein neuer Hemirhamphus aus dem Pazifischen Ozean. Anzeiger der Akademie der Wissenschaften Wien, 68, 2 - 3." type="journal article" year="1931">Schindler 1931</bibRefCitation>
)
</taxonomicName>
, was the relative positions of the dorsal and anal fins: in
<taxonomicName id="99964D52417EFFAFFB68FF22715AA382" baseAuthorityName="Schindler" baseAuthorityYear="1930" box="[1161,1321,153,179]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="praematura">
<emphasis id="6CE2EAC3417EFFAFFB68FF22715AA382" box="[1161,1321,153,179]" italics="true" pageId="2" pageNumber="453">S. praematura</emphasis>
</taxonomicName>
the origin of the dorsal fin is well in advance of the origin of the anal fin, while in
<taxonomicName id="99964D52417EFFAFFC3AFF0670FEA3E6" baseAuthorityName="Schindler" baseAuthorityYear="1931" box="[987,1165,189,215]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="pietschmanni">
<emphasis id="6CE2EAC3417EFFAFFC3AFF0670FEA3E6" box="[987,1165,189,215]" italics="true" pageId="2" pageNumber="453">S. pietschmanni</emphasis>
</taxonomicName>
the origin of the dorsal fin is just above the origin of the anal fin (
<bibRefCitation id="3A074B20417EFFAFFD60FF5A7741A3CA" author="Schindler, O." box="[641,818,225,251]" pageId="2" pageNumber="453" pagination="79 - 80" refId="ref13610" refString="Schindler, O. (1930) Ein neuer Hemirhamphus aus dem Pazifischen Ozean. Anzeiger der Akademie der Wissenschaften Wien, 67, 79 - 80." type="journal article" year="1930">Schindler 1930</bibRefCitation>
;
<bibRefCitation id="3A074B20417EFFAFFCA1FF5A7708A3CA" author="Schindler, O." box="[832,891,225,251]" pageId="2" pageNumber="453" pagination="2 - 3" refId="ref13638" refString="Schindler, O. (1931) Ein neuer Hemirhamphus aus dem Pazifischen Ozean. Anzeiger der Akademie der Wissenschaften Wien, 68, 2 - 3." type="journal article" year="1931">1931</bibRefCitation>
;
<bibRefCitation id="3A074B20417EFFAFFC69FF5A70C2A3CA" author="Johnson, G. D. &amp; Brothers, E. B." box="[904,1201,225,251]" pageId="2" pageNumber="453" pagination="441 - 471" refId="ref12430" refString="Johnson, G. D. &amp; Brothers, E. B. (1993) Schindleria: a paedomorphic goby (Teleostei: Gobioidei). Bulletin of Marine Science, 52, 441 - 471." type="journal article" year="1993">Johnson &amp; Brothers 1993</bibRefCitation>
). Subsequently most authors followed this character and consequently named specimens from their samples either
<taxonomicName id="99964D52417EFFAFFB5FFEBE712CA22E" baseAuthorityName="Schindler" baseAuthorityYear="1930" box="[1214,1375,261,287]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="praematura">
<emphasis id="6CE2EAC3417EFFAFFB5FFEBE712CA22E" box="[1214,1375,261,287]" italics="true" pageId="2" pageNumber="453">S. praematura</emphasis>
</taxonomicName>
or
<taxonomicName id="99964D52417EFFAFFA66FEBE755FA272" baseAuthorityName="Schindler" baseAuthorityYear="1931" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="pietschmanni">
<emphasis id="6CE2EAC3417EFFAFFA66FEBE755FA272" italics="true" pageId="2" pageNumber="453">S. pietschmanni</emphasis>
</taxonomicName>
depending on the relative fin position.
</paragraph>
<paragraph id="5E2936D1417EFFAFFF26FEF6749DA152" blockId="2.[151,1436,153,828]" pageId="2" pageNumber="453">
We checked 49 publications (72 records) which reported findings of Schindlers fishes (
<tableCitation id="1314036A417EFFAFFB58FEF6717CA256" box="[1209,1295,333,359]" captionStart="TABLE 1" captionStartId="2.[151,239,874,898]" captionTargetBox="[166,1313,982,1990]" captionText="TABLE 1. 72 records from 49 publications, including the publications of the six formally described species of Schindleria (highlighted in grey). * refers to the type specimens of S. nigropunctata, actually a LDF type, and therefore listed twice. Nominal species highlighted in grey. List of references see below." httpUri="http://table.plazi.org/id/0AE96659417EFFAFFF76FCD177EEA0FA" pageId="2" pageNumber="453" tableUuid="0AE96659417EFFAFFF76FCD177EEA0FA">Table 1</tableCitation>
). Excluding those publications which formally described a species (n = 6), the remaining 43 studies comprised 65 records of
<taxonomicName id="99964D52417EFFAFFF76FE2E756BA29E" authorityName="Giltay" authorityYear="1934" box="[151,280,405,431]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="genus">
<emphasis id="6CE2EAC3417EFFAFFF76FE2E756BA29E" box="[151,280,405,431]" italics="true" pageId="2" pageNumber="453">Schindleria</emphasis>
</taxonomicName>
. In 41 of these records the specimens were identified as
<taxonomicName id="99964D52417EFFAFFC45FE2E7035A29E" baseAuthorityName="Schindler" baseAuthorityYear="1930" box="[932,1094,405,431]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="praematura">
<emphasis id="6CE2EAC3417EFFAFFC45FE2E7035A29E" box="[932,1094,405,431]" italics="true" pageId="2" pageNumber="453">S. praematura</emphasis>
</taxonomicName>
, in 14 as
<taxonomicName id="99964D52417EFFAFFB59FE2E7118A29E" baseAuthorityName="Schindler" baseAuthorityYear="1931" box="[1208,1387,405,431]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="pietschmanni">
<emphasis id="6CE2EAC3417EFFAFFB59FE2E7118A29E" box="[1208,1387,405,431]" italics="true" pageId="2" pageNumber="453">S. pietschmanni</emphasis>
</taxonomicName>
and in two as
<taxonomicName id="99964D52417EFFAFFEEBFE0275C0A2E2" authorityName="Watson &amp; Walker" authorityYear="2004" box="[266,435,441,467]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="brevipinguis">
<emphasis id="6CE2EAC3417EFFAFFEEBFE0275C0A2E2" box="[266,435,441,467]" italics="true" pageId="2" pageNumber="453">S. brevipinguis</emphasis>
</taxonomicName>
. Eight of the records were not assigned to a species and were identified as
<taxonomicName id="99964D52417EFFAFFAFAFE0271EFA2E2" authorityName="Giltay" authorityYear="1934" box="[1307,1436,441,467]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="genus">
<emphasis id="6CE2EAC3417EFFAFFAFAFE0271EFA2E2" box="[1307,1436,441,467]" italics="true" pageId="2" pageNumber="453">Schindleria</emphasis>
</taxonomicName>
sp. or
<taxonomicName id="99964D52417EFFAFFF3FFE6675F7A2C6" authorityName="Giltay" authorityYear="1934" box="[222,388,477,503]" class="Actinopterygii" family="Schindleriidae" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="family">Schindleriidae</taxonomicName>
. In the light of high endemism and a large number of cryptic species (
<bibRefCitation id="3A074B20417EFFAFFB7DFE667121A2C6" author="Kon, T. &amp; Yoshino, T. &amp; Mukai, T. &amp; Nishida M." box="[1180,1362,477,503]" pageId="2" pageNumber="453" pagination="53 - 62" refId="ref12605" refString="Kon, T., Yoshino, T., Mukai, T. &amp; Nishida M. (2007) DNA sequences identify numerous cryptic species of the vertebrate: A lesson from the gobioid fish Schindleria. Molecular Phylogenetics and Evolution, 44, 53 - 62. https: // doi. org / 10.1016 / j. ympev. 2006.12.007" type="journal article" year="2007">
Kon
<emphasis id="6CE2EAC3417EFFAFFB37FE65717CA2C6" box="[1238,1295,477,503]" italics="true" pageId="2" pageNumber="453">et al.</emphasis>
2007
</bibRefCitation>
,
<bibRefCitation id="3A074B20417EFFAFFABFFE6671EBA2C6" author="Kon, T. &amp; Yoshino, T. &amp; Nishida, M." box="[1374,1432,477,503]" pageId="2" pageNumber="453" pagination="62 - 66" refId="ref12669" refString="Kon, T., Yoshino, T. &amp; Nishida, M. (2011) Cryptic species of the gobioid paedomorphic genus Schindleria from Palau, western Pacific Ocean. Ichthyological Research, 58, 62 - 66. https: // doi. org / 10.1007 / s 10228 - 010 - 0178 - y" type="journal article" year="2011">2011</bibRefCitation>
;
<bibRefCitation id="3A074B20417EFFAFFF76FDBA75AFA12A" author="Bogorodsky, S. &amp; Randall, J. E." box="[151,476,513,539]" pageId="2" pageNumber="453" pagination="239 - 265" refId="ref11598" refString="Bogorodsky, S. &amp; Randall, J. E. (2019) Endemic fishes of the Red Sea. In: Rasul, N. M. A. &amp; Stewart, I. C. F. (Eds.), Oceanographic and Biological Aspects of the Red Sea. Springer Oceanography, pp. 239 - 265. https: // doi. org / 10.1007 / 978 - 3 - 319 - 99417 - 8 _ 14" type="book chapter" year="2019">Bogorodsky &amp; Randall 2019</bibRefCitation>
) it is unlikely that all these documented records of Schindlers fishes belong to one of these two species.
<bibRefCitation id="3A074B20417EFFAFFE89FD9E7663A10E" author="Randall, J. E." box="[360,528,549,575]" pageId="2" pageNumber="453" refId="ref13515" refString="Randall, J. E. (2007) Reef and shore fishes of the Hawaiian Islands. University of Hawai'i Sea Grant College Program, Hono- lulu, 546 pp." type="book" year="2007">Randall (2007)</bibRefCitation>
concluded that
<taxonomicName id="99964D52417EFFAFFD23FD9E7711A10E" baseAuthorityName="Schindler" baseAuthorityYear="1930" box="[706,866,549,575]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="praematura">
<emphasis id="6CE2EAC3417EFFAFFD23FD9E7711A10E" box="[706,866,549,575]" italics="true" pageId="2" pageNumber="453">S. praematura</emphasis>
</taxonomicName>
and
<taxonomicName id="99964D52417EFFAFFC78FD9E7039A10E" baseAuthorityName="Schindler" baseAuthorityYear="1931" box="[921,1098,549,575]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="pietschmanni">
<emphasis id="6CE2EAC3417EFFAFFC78FD9E7039A10E" box="[921,1098,549,575]" italics="true" pageId="2" pageNumber="453">S. pietschmanni</emphasis>
</taxonomicName>
are endemic for the Hawaiian Islands.
</paragraph>
<paragraph id="5E2936D1417EFFAFFF26FDD67632A00D" blockId="2.[151,1436,153,828]" pageId="2" pageNumber="453">
It will require a series of integrative studies including ecology, genetics and morphology just to get an overview of the number of species of
<emphasis id="6CE2EAC3417EFFAFFE31FD2A762BA19A" box="[464,600,657,683]" italics="true" pageId="2" pageNumber="453">
<taxonomicName id="99964D52417EFFAFFE31FD2A7627A19A" authorityName="Giltay" authorityYear="1934" box="[464,596,657,683]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="genus">Schindleria</taxonomicName>
,
</emphasis>
which possibly number several dozen. Species with very distinct morphological characters like
<taxonomicName id="99964D52417EFFAFFE76FD0E764CA1FE" authorityName="Watson &amp; Walker" authorityYear="2004" box="[407,575,693,719]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="brevipinguis">
<emphasis id="6CE2EAC3417EFFAFFE76FD0E764CA1FE" box="[407,575,693,719]" italics="true" pageId="2" pageNumber="453">S. brevipinguis</emphasis>
</taxonomicName>
(tiny and toothless) or
<taxonomicName id="99964D52417EFFAFFCA9FD0E7073A1FE" authorityName="Ahnelt &amp; Sauberer" authorityYear="2018" box="[840,1024,693,719]" class="Actinopterygii" family="Schindleriidae" genus="Schindleria" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="453" phylum="Chordata" rank="species" species="macrodentata">
<emphasis id="6CE2EAC3417EFFAFFCA9FD0E7073A1FE" box="[840,1024,693,719]" italics="true" pageId="2" pageNumber="453">S. macrodentata</emphasis>
</taxonomicName>
(relatively large, with few, but large teeth, last procurrent spine with extra spiny process) may be formally distinguished and may well prove to be valid species. Therefore, we preliminarily attribute all published records (
<tableCitation id="1314036A417EFFAFFC75FD46779AA026" box="[916,1001,765,791]" captionStart="TABLE 1" captionStartId="2.[151,239,874,898]" captionTargetBox="[166,1313,982,1990]" captionText="TABLE 1. 72 records from 49 publications, including the publications of the six formally described species of Schindleria (highlighted in grey). * refers to the type specimens of S. nigropunctata, actually a LDF type, and therefore listed twice. Nominal species highlighted in grey. List of references see below." httpUri="http://table.plazi.org/id/0AE96659417EFFAFFF76FCD177EEA0FA" pageId="2" pageNumber="453" tableUuid="0AE96659417EFFAFFF76FCD177EEA0FA">Table 1</tableCitation>
), including those of the Dana-Expedition, to two morphotypes (see below).
</paragraph>
</subSubSection>
</treatment>
</document>