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<document id="D03933426B47D52F467540B7C4FAD466" ID-CLB-Dataset="27502" ID-DOI="10.11646/zootaxa.4550.1.1" ID-GBIF-Dataset="291701d4-652e-440e-adab-f6b0201c0972" ID-ISSN="1175-5326" ID-Zenodo-Dep="2625030" ID-ZooBank="2F7D1EC4-D4ED-4FAE-B227-CF7B79EAE833" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554274879827" checkinUser="plazi" docAuthor="Huber, Bernhard A." docDate="2019" docId="CA3B104CFF8EFF86FF3DF991FF41E357" docLanguage="en" docName="zootaxa.4550.1.1.pdf" docOrigin="Zootaxa 4550 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tissahamia maturata" docType="treatment" docVersion="15" lastPageNumber="30" masterDocId="36026834FF95FF9BFFAAFFA1FFB6E16A" masterDocTitle="The pholcid spiders of Sri Lanka (Araneae: Pholcidae)" masterLastPageNumber="57" masterPageNumber="1" pageNumber="28" updateTime="1698716853299" updateUser="ExternalLinkService">
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<mods:title id="DE7839523DE8F6B097A2017CDF0846B1">The pholcid spiders of Sri Lanka (Araneae: Pholcidae)</mods:title>
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<treatment id="CA3B104CFF8EFF86FF3DF991FF41E357" ID-DOI="http://doi.org/10.5281/zenodo.4581674" ID-GBIF-Taxon="154534131" ID-Zenodo-Dep="4581674" LSID="urn:lsid:plazi:treatment:CA3B104CFF8EFF86FF3DF991FF41E357" httpUri="http://treatment.plazi.org/id/CA3B104CFF8EFF86FF3DF991FF41E357" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28">
<subSubSection id="0A88F2D1FF8EFF80FF3DF991FDFEE720" box="[151,584,1584,1610]" pageId="27" pageNumber="28" type="nomenclature">
<paragraph id="422DA15AFF8EFF80FF3DF991FDFEE720" blockId="27.[151,584,1584,1643]" box="[151,584,1584,1610]" pageId="27" pageNumber="28">
<heading id="19651636FF8EFF80FF3DF991FDFEE720" bold="true" box="[151,584,1584,1610]" fontSize="11" level="1" pageId="27" pageNumber="28" reason="1">
<taxonomicName id="8592DAD9FF8EFF80FF3DF991FDFEE720" ID-CoL="577S5" authority="(Huber, 2011)" baseAuthorityName="Huber" baseAuthorityYear="2011" box="[151,584,1584,1610]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF8EFF80FF3DF991FDFEE720" bold="true" box="[151,584,1584,1610]" pageId="27" pageNumber="28">
<emphasis id="70E67D48FF8EFF80FF3DF991FE21E720" bold="true" box="[151,407,1584,1610]" italics="true" pageId="27" pageNumber="28">Tissahamia maturata</emphasis>
(
<bibRefCitation id="2603DCABFF8EFF80FE0DF991FDF7E720" author="Huber, B. A." box="[423,577,1584,1610]" pageId="27" pageNumber="28" pagination="1 - 509" refId="ref27101" refString="Huber, B. A. (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien, 58, 1 - 509." type="journal article" year="2011">Huber, 2011</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF8EFF80FF3DF9F3FD81E701" box="[151,567,1618,1643]" pageId="27" pageNumber="28" type="description">
<paragraph id="422DA15AFF8EFF80FF3DF9F3FD81E701" blockId="27.[151,584,1584,1643]" box="[151,567,1618,1643]" pageId="27" pageNumber="28">
<figureCitation id="DAA9BDDFFF8EFF80FF3DF9F3FE92E701" box="[151,292,1618,1643]" captionStart="FIGURES 93102" captionStartId="25.[151,264,1918,1941]" captionTargetBox="[151,1436,418,1859]" captionTargetId="figure@25.[151,1436,418,1859]" captionTargetPageId="25" captionText="FIGURES 93102. Tissahamia, live specimens. 9396. T. ethagala (Huber, 2011), male from Ethagala, female with egg-sac from Kandalama, and females from Dematagala. 9798. T. karuna sp. n., male and female with egg-sac from Labugama. 99 100. T. maturata (Huber, 2011), male with mite and female with egg-sac from Hakgala. 101102. T. kottawagamaensis (Yao &amp; Li, 2016), male from Kottawa, female from Viharekele." httpUri="https://zenodo.org/record/2625062/files/figure.png" pageId="27" pageNumber="28">Figs 99100</figureCitation>
,
<figureCitation id="DAA9BDDFFF8EFF80FE9AF9F2FE25E701" box="[304,403,1619,1643]" captionStart="FIGURES 110115" captionStartId="28.[151,264,1879,1902]" captionTargetBox="[202,1418,342,1855]" captionTargetId="figure@28.[164,1427,332,1861]" captionTargetPageId="28" captionText="FIGURES 110115. Left procursi in prolateral (upper row) and retrolateral views, all images at same scale. 110111. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20066); asterisk marks area where males from Mitirigala have a small conical process. 112113. T. maturata (Huber, 2011), from Hakgala (ZFMK Ar 20069); arrow points at distinctive prolateral process. 114115. T. kottawagamaensis (Yao &amp; Li, 2016), from Kottawa (ZFMK Ar 20072). Abbreviations: bp, bifid process; hs, hinged sclerite; m, membrane; po, pocket; r, ridge; ta, tarsus. Scale line: 0.5 mm." httpUri="https://zenodo.org/record/2625066/files/figure.png" pageId="27" pageNumber="28">112113</figureCitation>
,
<figureCitation id="DAA9BDDFFF8EFF80FE34F9F2FE7FE701" box="[414,457,1619,1643]" captionStart="FIGURES 116119" captionStartId="29.[151,264,1920,1943]" captionTargetBox="[155,1435,1202,1900]" captionTargetId="figure@29.[151,1436,1193,1908]" captionTargetPageId="29" captionText="FIGURES 116119. 116117. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20066), male chelicerae, frontal view; and processes of left genital bulb, prolateral view. 118119. T. maturata (Huber, 2011) from Hakgala (ZFMK Ar 20069) and T. kottawagamaensis (Yao &amp; Li, 2016) from Kottawa (ZFMK Ar 20072), processes of left genital bulbs in prolateral views (all genital bulbs at same scale). Abbreviations: a, appendix (main bulbal process); e, embolus. Scale lines: 0.3 mm." httpUri="https://zenodo.org/record/2625070/files/figure.png" pageId="27" pageNumber="28">118</figureCitation>
,
<figureCitation id="DAA9BDDFFF8EFF80FE7FF9F2FD81E701" box="[469,567,1619,1643]" captionStart="FIGURES 120125" captionStartId="30.[151,264,1133,1156]" captionTargetBox="[151,1436,193,1113]" captionTargetId="figure@30.[151,1436,193,1113]" captionTargetPageId="30" captionText="FIGURES 120125. Epigyna in ventral views (upper row) and cleared female genitalia in dorsal views. 120121. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20067); arrow points at internal arc (valve). 122123. T. maturata (Huber, 2011), from Hakgala (ZFMK Ar 20070). 124125. T. kottawagamaensis (Yao &amp; Li, 2016), from Kottawa (ZFMK Ar 20073). Abbreviation: pp, pore plate. At various scales." httpUri="https://zenodo.org/record/2625072/files/figure.png" pageId="27" pageNumber="28">122123</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF8EFF80FF3DF93AFC4AE7A5" pageId="27" pageNumber="28" type="reference_group">
<paragraph id="422DA15AFF8EFF80FF3DF93AFBA4E7DB" blockId="27.[151,1042,1691,1743]" box="[151,1042,1691,1713]" pageId="27" pageNumber="28">
<treatmentCitationGroup id="62828674FF8EFF80FF3DF93AFBA4E7DB" box="[151,1042,1691,1713]" pageId="27" pageNumber="28">
<taxonomicName id="8592DAD9FF8EFF80FF3DF93AFDB2E7DB" ID-CoL="94739" authority="Huber, 2011: 177" authorityName="Huber" authorityPageNumber="177" authorityYear="2011" box="[151,516,1691,1713]" class="Arachnida" family="Pholcidae" genus="Pholcus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF8EFF80FF3DF93AFEFAE7DB" box="[151,332,1691,1713]" italics="true" pageId="27" pageNumber="28">Pholcus maturata</emphasis>
<treatmentCitation id="C333874BFF8EFF80FEF8F93AFDB2E7DB" author="Huber, B. A." box="[338,516,1691,1713]" page="177" pageId="27" pageNumber="28" year="2011">
<bibRefCitation id="2603DCABFF8EFF80FEF8F93AFE64E7DB" author="Huber, B. A." box="[338,466,1691,1713]" pageId="27" pageNumber="28" pagination="1 - 509" refId="ref27101" refString="Huber, B. A. (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien, 58, 1 - 509." type="journal article" year="2011">Huber, 2011</bibRefCitation>
: 177
</treatmentCitation>
</taxonomicName>
, figs 745748, 777778, 805808 (
<specimenCount id="54946AD3FF8EFF80FCC7F93AFC32E7DB" box="[877,900,1691,1713]" pageId="27" pageNumber="28" type="male"></specimenCount>
<specimenCount id="54946AD3FF8EFF80FC2EF93AFC22E7DB" box="[900,916,1691,1713]" pageId="27" pageNumber="28" type="female"></specimenCount>
,
<collectingCountry id="3A85E1CAFF8EFF80FC35F93AFBB1E7DB" box="[927,1031,1691,1713]" name="Sri Lanka" pageId="27" pageNumber="28">Sri Lanka</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="422DA15AFF8EFF80FF3DF918FC4AE7A5" blockId="27.[151,1042,1691,1743]" box="[151,1020,1721,1743]" pageId="27" pageNumber="28">
<taxonomicName id="8592DAD9FF8EFF80FF3DF918FEDBE7A5" ID-CoL="577S5" baseAuthorityName="Huber" baseAuthorityYear="2011" box="[151,365,1721,1743]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF8EFF80FF3DF918FEDBE7A5" box="[151,365,1721,1743]" italics="true" pageId="27" pageNumber="28">Tissahamia maturata</emphasis>
</taxonomicName>
<bibRefCitation id="2603DCABFF8EFF80FE2DF918FD8DE7A5" author="Eberle, J. &amp; Dimitrov, D. &amp; Valdez-Mondragon, A. &amp; Huber, B. A." box="[391,571,1721,1743]" pageId="27" pageNumber="28" pagination="141" refId="ref26630" refString="Eberle, J., Dimitrov, D., Valdez-Mondragon, A. &amp; Huber, B. A. (2018) Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology, 18, 141. https: // doi. org / 10.1186 / s 12862 - 018 - 1244 - 8" type="journal article" year="2018">
Eberle
<emphasis id="70E67D48FF8EFF80FE65F91BFDB4E7A5" box="[463,514,1721,1743]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2018
</bibRefCitation>
(molecular data).
<bibRefCitation id="2603DCABFF8EFF80FD5EF918FC1FE7A5" author="Huber, B. A. &amp; Eberle, J. &amp; Dimitrov, D." box="[756,937,1721,1743]" pageId="27" pageNumber="28" pagination="51 - 101" refId="ref27545" refString="Huber, B. A., Eberle, J. &amp; Dimitrov, D. (2018) The phylogeny of pholcid spiders (Araneae, Pholcidae): a critical evaluation of relationships suggested by molecular data. ZooKeys, 789, 51 - 101. https: // doi. org / 10.3897 / zookeys. 789.22781" type="journal article" year="2018">
Huber
<emphasis id="70E67D48FF8EFF80FC90F91BFCDBE7A5" box="[826,877,1721,1743]" italics="true" pageId="27" pageNumber="28">et al.</emphasis>
2018
</bibRefCitation>
: fig. 10.
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF8EFF80FF3DF95DFC9BE6A3" pageId="27" pageNumber="28" type="diagnosis">
<paragraph id="422DA15AFF8EFF80FF3DF95DFC9BE6A3" blockId="27.[151,1437,1788,2029]" pageId="27" pageNumber="28">
<emphasis id="70E67D48FF8EFF80FF3DF95DFEBCE67F" bold="true" box="[151,266,1788,1813]" pageId="27" pageNumber="28">Diagnosis</emphasis>
(amendments; see
<bibRefCitation id="2603DCABFF8EFF80FE49F95DFDDBE67E" author="Huber, B. A." box="[483,621,1788,1813]" pageId="27" pageNumber="28" pagination="1 - 509" refId="ref27101" refString="Huber, B. A. (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien, 58, 1 - 509." type="journal article" year="2011">Huber 2011</bibRefCitation>
). Males are easily distinguished from
<taxonomicName id="8592DAD9FF8EFF80FBB3F95CFB34E67F" authorityName="Huber" authorityYear="2019" box="[1049,1154,1788,1813]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="karuna">
<emphasis id="70E67D48FF8EFF80FBB3F95CFB34E67F" box="[1049,1154,1788,1813]" italics="true" pageId="27" pageNumber="28">T. karuna</emphasis>
</taxonomicName>
by shape of main bulbal process (appendix) (
<figureCitation id="DAA9BDDFFF8EFF80FE27F881FE5BE652" box="[397,493,1824,1849]" captionStart="FIGURES 116119" captionStartId="29.[151,264,1920,1943]" captionTargetBox="[155,1435,1202,1900]" captionTargetId="figure@29.[151,1436,1193,1908]" captionTargetPageId="29" captionText="FIGURES 116119. 116117. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20066), male chelicerae, frontal view; and processes of left genital bulb, prolateral view. 118119. T. maturata (Huber, 2011) from Hakgala (ZFMK Ar 20069) and T. kottawagamaensis (Yao &amp; Li, 2016) from Kottawa (ZFMK Ar 20072), processes of left genital bulbs in prolateral views (all genital bulbs at same scale). Abbreviations: a, appendix (main bulbal process); e, embolus. Scale lines: 0.3 mm." httpUri="https://zenodo.org/record/2625070/files/figure.png" pageId="27" pageNumber="28">Fig. 118</figureCitation>
; strong distal spines; large prolateral process) and by details of procursus (
<figureCitation id="DAA9BDDFFF8EFF80FA86F881FF75E636" captionStart="FIGURES 110115" captionStartId="28.[151,264,1879,1902]" captionTargetBox="[202,1418,342,1855]" captionTargetId="figure@28.[164,1427,332,1861]" captionTargetPageId="28" captionText="FIGURES 110115. Left procursi in prolateral (upper row) and retrolateral views, all images at same scale. 110111. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20066); asterisk marks area where males from Mitirigala have a small conical process. 112113. T. maturata (Huber, 2011), from Hakgala (ZFMK Ar 20069); arrow points at distinctive prolateral process. 114115. T. kottawagamaensis (Yao &amp; Li, 2016), from Kottawa (ZFMK Ar 20072). Abbreviations: bp, bifid process; hs, hinged sclerite; m, membrane; po, pocket; r, ridge; ta, tarsus. Scale line: 0.5 mm." httpUri="https://zenodo.org/record/2625066/files/figure.png" pageId="27" pageNumber="28">Figs 112 113</figureCitation>
; longer but with smaller distal hinged sclerite; with sclerotized prolateral process). Females differ from
<taxonomicName id="8592DAD9FF8EFF80FA2DF8E4FED0E6EB" baseAuthorityName="Yao &amp; Li" baseAuthorityYear="2016" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="kottawagamaensis">
<emphasis id="70E67D48FF8EFF80FA2DF8E4FED0E6EB" italics="true" pageId="27" pageNumber="28">T. kottawagamaensis</emphasis>
</taxonomicName>
by trapezoidal rather than triangular epigynal plate and by shorter internal genitalia (
<figureCitation id="DAA9BDDFFF8EFF80FA80F8C9FF73E6CE" captionStart="FIGURES 120125" captionStartId="30.[151,264,1133,1156]" captionTargetBox="[151,1436,193,1113]" captionTargetId="figure@30.[151,1436,193,1113]" captionTargetPageId="30" captionText="FIGURES 120125. Epigyna in ventral views (upper row) and cleared female genitalia in dorsal views. 120121. Tissahamia karuna sp. n., from Labugama (ZFMK Ar 20067); arrow points at internal arc (valve). 122123. T. maturata (Huber, 2011), from Hakgala (ZFMK Ar 20070). 124125. T. kottawagamaensis (Yao &amp; Li, 2016), from Kottawa (ZFMK Ar 20073). Abbreviation: pp, pore plate. At various scales." httpUri="https://zenodo.org/record/2625072/files/figure.png" pageId="27" pageNumber="28">Figs 122 123</figureCitation>
); females are difficult to separate from
<taxonomicName id="8592DAD9FF8EFF80FD26F82CFD40E6CF" authorityName="Huber" authorityYear="2019" box="[652,758,1932,1957]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="karuna">
<emphasis id="70E67D48FF8EFF80FD26F82CFD40E6CF" box="[652,758,1932,1957]" italics="true" pageId="27" pageNumber="28">T. karuna</emphasis>
</taxonomicName>
(more heavily sclerotized epigynal plate; anterior epigynal margin medially straight; slightly shorter internal genitalia.
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF8EFF86FF6DF875FD72E0C6" lastPageId="29" lastPageNumber="30" pageId="27" pageNumber="28" type="description">
<paragraph id="422DA15AFF8EFF87FF6DF875FD24E077" blockId="27.[151,1437,1788,2029]" lastBlockId="28.[151,1436,151,285]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">
<emphasis id="70E67D48FF8EFF80FF6DF875FEE7E687" bold="true" box="[199,337,2004,2029]" pageId="27" pageNumber="28">Description</emphasis>
(amendments; see
<bibRefCitation id="2603DCABFF8EFF80FD97F875FD7BE686" author="Huber, B. A." box="[573,717,2004,2029]" pageId="27" pageNumber="28" pagination="1 - 509" refId="ref27101" refString="Huber, B. A. (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien, 58, 1 - 509." type="journal article" year="2011">Huber 2011</bibRefCitation>
). Most males with large, posteriorly divided, dark mark on carapace and with dark clypeus and sternum; one male with female pattern: carapace light with small U-shaped dark mark and clypeus and sternum whitish. Males from Namunukula and Mausagalla with tip of appendix slightly different (pointed processes shorter and more uniform in length). Tibia
<quantity id="856A0CBFFF89FF87FC78FF41FBB6E192" box="[978,1024,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="28" pageNumber="29" unit="in" value="1.0">1 in</quantity>
13 newly examined males: 7.29.4 (mean 8.0); in
<specimenCount id="54946AD3FF89FF87FE96FEA5FE1AE077" box="[316,428,260,285]" pageId="28" pageNumber="29" type="female">9 females</specimenCount>
: 6.07.0 (mean 6.4).
</paragraph>
<caption id="16EDF1D2FF89FF87FF3DF8F6FC5AE69F" httpUri="https://zenodo.org/record/2625066/files/figure.png" pageId="28" pageNumber="29" startId="28.[151,264,1879,1902]" targetBox="[202,1418,342,1855]" targetPageId="28">
<paragraph id="422DA15AFF89FF87FF3DF8F6FC5AE69F" blockId="28.[151,1436,1879,2037]" pageId="28" pageNumber="29">
<emphasis id="70E67D48FF89FF87FF3DF8F6FEDEE606" bold="true" box="[151,360,1879,1902]" pageId="28" pageNumber="29">FIGURES 110115</emphasis>
. Left procursi in prolateral (upper row) and retrolateral views, all images at same scale. 110111.
<taxonomicName id="8592DAD9FF89FF87FF3DF8D8FEECE6E5" authority="Huber, 2019" authorityName="Huber" authorityYear="2019" box="[151,346,1913,1935]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="karuna" status="sp. nov.">
<emphasis id="70E67D48FF89FF87FF3DF8D8FEECE6E5" box="[151,346,1913,1935]" italics="true" pageId="28" pageNumber="29">Tissahamia karuna</emphasis>
</taxonomicName>
<emphasis id="70E67D48FF89FF87FECBF8D8FE2DE6E4" bold="true" box="[353,411,1913,1934]" pageId="28" pageNumber="29">
<taxonomicNameLabel id="6BD5C033FF89FF87FECBF8D8FE2DE6E4" box="[353,411,1913,1934]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, from Labugama (ZFMK Ar 20066); asterisk marks area where males from Mitirigala have a small conical process. 112113.
<taxonomicName id="8592DAD9FF89FF87FE0CF83AFD00E6DA" authority="(Huber, 2011)" authorityName="Huber" authorityYear="2011" baseAuthorityName="Huber" baseAuthorityYear="2011" box="[422,694,1946,1968]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF89FF87FE0CF83AFDA8E6DA" box="[422,542,1947,1968]" italics="true" pageId="28" pageNumber="29">T. maturata</emphasis>
(Huber, 2011)
</taxonomicName>
, from Hakgala (ZFMK Ar 20069); arrow points at distinctive prolateral process. 114115.
<taxonomicName id="8592DAD9FF89FF87FEFEF81CFD54E6B8" authority="(Yao &amp; Li, 2016)" authorityName="Yao &amp; Li" authorityYear="2016" baseAuthorityName="Yao &amp; Li" baseAuthorityYear="2016" box="[340,738,1980,2002]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kottawagamaensis">
<emphasis id="70E67D48FF89FF87FEFEF81CFD9EE6B8" box="[340,552,1980,2002]" italics="true" pageId="28" pageNumber="29">T. kottawagamaensis</emphasis>
(Yao &amp; Li, 2016)
</taxonomicName>
, from Kottawa (ZFMK Ar 20072). Abbreviations: bp, bifid process; hs, hinged sclerite; m, membrane; po, pocket; r, ridge; ta, tarsus. Scale line: 0.5 mm.
</paragraph>
</caption>
<paragraph id="422DA15AFF88FF86FF6DFF36FD59E00F" blockId="29.[151,1437,151,573]" pageId="29" pageNumber="30">
<emphasis id="70E67D48FF88FF86FF6DFF36FEEBE1DA" bold="true" box="[199,349,151,176]" pageId="29" pageNumber="30">New records</emphasis>
.
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:
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<specimenCount id="54946AD3FF88FF86FD9DFF39FDECE1DA" box="[567,602,152,176]" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
,
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</materialsCitation>
,
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<specimenCount id="54946AD3FF88FF86FD69FF39FD45E1DA" box="[707,755,152,176]" pageId="29" pageNumber="30" type="male">11♂</specimenCount>
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<specimenCount id="54946AD3FF88FF86FC98FF39FCC5E1DA" box="[818,883,151,176]" pageId="29" pageNumber="30" type="generic">1 juv.</specimenCount>
,
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(
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2006970)
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and
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<specimenCount id="54946AD3FF88FF86FB1AFF39FB65E1DA" box="[1200,1235,152,176]" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
<specimenCount id="54946AD3FF88FF86FB70FF39FB4AE1DA" box="[1242,1276,152,176]" pageId="29" pageNumber="30" type="female">3♀</specimenCount>
2 juvs in pure ethanol,
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(
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),
<collectingRegion id="80566FB8FF88FF86FE16FF1DFDC8E1BF" box="[444,638,188,213]" country="Sri Lanka" name="Central" pageId="29" pageNumber="30">Central Province</collectingRegion>
,
<location id="474DF781FF88FF86FD21FF1DFC81E1BE" LSID="urn:lsid:plazi:treatment:CA3B104CFF8EFF86FF3DF991FF41E357:474DF781FF88FF86FD21FF1DFC81E1BE" box="[651,823,188,213]" country="Sri Lanka" latitude="6.93" longLatPrecision="78" longitude="80.814" name="Hakgala Forest" pageId="29" pageNumber="30" stateProvince="Central">Hakgala Forest</location>
(
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,
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),
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<quantity id="856A0CBFFF88FF86FB93FF1DFB26E1BE" box="[1081,1168,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.7899999999999998" pageId="29" pageNumber="30" unit="m" value="1790.0">1790 m</quantity>
a.s.l.
</elevation>
,
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<collectingDate id="26687E72FF88FF86FB76FF1DFAE3E1BF" box="[1244,1365,188,213]" pageId="29" pageNumber="30" value="2017-03-14">14.iii.2017</collectingDate>
</date>
(
<collectorName id="EF67C48CFF88FF86FACDFF1DFF54E192" pageId="29" pageNumber="30">B.A. Huber</collectorName>
)
</materialsCitation>
.
<materialsCitation id="F2FAAB07FF88FF86FF5FFF41FD91E192" ID-GBIF-Occurrence="2234226217" box="[245,551,224,248]" collectionCode="ZFMK" collectorName="Ar" country="Sri Lanka" pageId="29" pageNumber="30" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="54946AD3FF88FF86FF5FFF41FEA1E192" box="[245,279,224,248]" pageId="29" pageNumber="30" type="male">2♂</specimenCount>
<specimenCount id="54946AD3FF88FF86FEB5FF41FEF4E192" box="[287,322,224,248]" pageId="29" pageNumber="30" type="female">1♀</specimenCount>
,
<collectionCode id="2483399FFF88FF86FEFBFF41FE29E19D" box="[337,415,224,247]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="29" pageNumber="30" type="Museum">ZFMK</collectionCode>
(
<collectorName id="EF67C48CFF88FF86FE04FF41FE7BE192" box="[430,461,224,248]" pageId="29" pageNumber="30">Ar</collectorName>
20071)
</materialsCitation>
,
<materialsCitation id="F2FAAB07FF88FF86FD99FF41FB03E077" ID-GBIF-Occurrence="2234226203" collectingDate="2014-01-23" collectorName="S. P. Benjamin &amp; N. Athukorala" country="Sri Lanka" county="Badulla District" elevation="2000" latitude="6.867" location="Namunukula Peak" longLatPrecision="78" longitude="81.117" municipality="Passara-Ella" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Uva">
<collectingRegion id="80566FB8FF88FF86FD99FF41FD79E192" box="[563,719,223,248]" country="Sri Lanka" name="Uva" pageId="29" pageNumber="30">Uva Province</collectingRegion>
,
<collectingCounty id="AB4CD9D6FF88FF86FD76FF7EFC27E192" box="[732,913,223,248]" pageId="29" pageNumber="30">Badulla District</collectingCounty>
, along
<collectingMunicipality id="A2493B20FF88FF86FC49FF7EFBDBE192" box="[995,1133,223,248]" pageId="29" pageNumber="30">Passara-Ella</collectingMunicipality>
road (B113),
<location id="474DF781FF88FF86FAA1FF7EFF7BE077" LSID="urn:lsid:plazi:treatment:CA3B104CFF8EFF86FF3DF991FF41E357:474DF781FF88FF86FAA1FF7EFF7BE077" country="Sri Lanka" county="Badulla District" latitude="6.867" longLatPrecision="78" longitude="81.117" municipality="Passara-Ella" name="Namunukula Peak" pageId="29" pageNumber="30" stateProvince="Uva">Namunukula Peak</location>
(
<geoCoordinate id="27A6C79DFF88FF86FF77FEA5FEF6E077" box="[221,320,260,285]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="55" value="6.867">6.867°N</geoCoordinate>
,
<geoCoordinate id="27A6C79DFF88FF86FEE6FEA5FE0EE076" box="[332,440,260,284]" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="55" value="81.117">81.117°E</geoCoordinate>
), ~
<elevation id="C9BF4669FF88FF86FE70FEA5FDD9E077" box="[474,623,260,285]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="m" value="2000.0">
<quantity id="856A0CBFFF88FF86FE70FEA5FD84E076" box="[474,562,260,284]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="29" pageNumber="30" unit="m" value="2000.0">2000 m</quantity>
a.s.l.
</elevation>
, beating,
<date id="362C879AFF88FF86FD74FEA5FCF1E077" box="[734,839,260,285]" pageId="29" pageNumber="30" value="2014-01-23">
<collectingDate id="26687E72FF88FF86FD74FEA5FCF1E077" box="[734,839,260,285]" pageId="29" pageNumber="30" value="2014-01-23">23.i.2014</collectingDate>
</date>
(
<collectorName id="EF67C48CFF88FF86FCF3FEA5FC4DE077" box="[857,1019,260,285]" pageId="29" pageNumber="30">S.P. Benjamin</collectorName>
,
<collectorName id="EF67C48CFF88FF86FBADFEA4FB1BE077" box="[1031,1197,260,285]" pageId="29" pageNumber="30">N. Athukorala</collectorName>
)
</materialsCitation>
.
<materialsCitation id="F2FAAB07FF88FF86FB6AFEA5FEE4E02A" ID-GBIF-Occurrence="2234226161" collectionCode="ZFMK" collectorName="Benj" country="Sri Lanka" pageId="29" pageNumber="30" specimenCount="1" specimenCount-male="1">
<specimenCount id="54946AD3FF88FF86FB6AFEA5FB55E076" box="[1216,1251,260,284]" pageId="29" pageNumber="30" type="male">1♂</specimenCount>
in pure ethanol,
<collectionCode id="2483399FFF88FF86FF3DFE89FF53E055" box="[151,229,296,319]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="29" pageNumber="30" type="Museum">ZFMK</collectionCode>
(
<collectorName id="EF67C48CFF88FF86FF5EFE86FE9BE02A" box="[244,301,295,320]" pageId="29" pageNumber="30">Benj</collectorName>
49)
</materialsCitation>
,
<materialsCitation id="F2FAAB07FF88FF86FEF4FE89FD5DE00F" ID-GBIF-Occurrence="2234226209" collectingDate="2014-01-22" collectorName="S. P. Benjamin &amp; N. Athukorala" country="Sri Lanka" county="Badulla District" latitude="6.909" location="Mausagalla" longLatPrecision="78" longitude="81.132" municipality="Passara-Ella" pageId="29" pageNumber="30" specimenCount="1" stateProvince="Uva">
<collectingRegion id="80566FB8FF88FF86FEF4FE89FE4CE02A" box="[350,506,295,320]" country="Sri Lanka" name="Uva" pageId="29" pageNumber="30">Uva Province</collectingRegion>
,
<collectingCounty id="AB4CD9D6FF88FF86FDAFFE86FD0DE02A" box="[517,699,295,320]" pageId="29" pageNumber="30">Badulla District</collectingCounty>
,
<collectingMunicipality id="A2493B20FF88FF86FD6CFE86FCE7E02A" box="[710,849,295,320]" pageId="29" pageNumber="30">Passara-Ella</collectingMunicipality>
road (B113),
<location id="474DF781FF88FF86FC44FE86FBD9E02A" LSID="urn:lsid:plazi:treatment:CA3B104CFF8EFF86FF3DF991FF41E357:474DF781FF88FF86FC44FE86FBD9E02A" box="[1006,1135,295,320]" country="Sri Lanka" county="Badulla District" latitude="6.909" longLatPrecision="78" longitude="81.132" municipality="Passara-Ella" name="Mausagalla" pageId="29" pageNumber="30" stateProvince="Uva">Mausagalla</location>
[=Maussagolla] (
<geoCoordinate id="27A6C79DFF88FF86FA9FFE86FA2EE02A" box="[1333,1432,295,320]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="55" value="6.909">6.909°N</geoCoordinate>
,
<geoCoordinate id="27A6C79DFF88FF86FF3DFEEDFEB2E00E" box="[151,260,332,356]" direction="east" orientation="longitude" pageId="29" pageNumber="30" precision="55" value="81.132">81.132°E</geoCoordinate>
),
<date id="362C879AFF88FF86FEBDFEEDFE37E00F" box="[279,385,332,357]" pageId="29" pageNumber="30" value="2014-01-22">
<collectingDate id="26687E72FF88FF86FEBDFEEDFE37E00F" box="[279,385,332,357]" pageId="29" pageNumber="30" value="2014-01-22">22.i.2014</collectingDate>
</date>
(
<collectorName id="EF67C48CFF88FF86FE3BFEEDFD85E00F" box="[401,563,332,357]" pageId="29" pageNumber="30">S.P. Benjamin</collectorName>
,
<collectorName id="EF67C48CFF88FF86FD94FEECFD55E00F" box="[574,739,332,357]" pageId="29" pageNumber="30">N. Athukorala</collectorName>
)
</materialsCitation>
.
</paragraph>
<paragraph id="422DA15AFF88FF86FF6DFECEFD72E0C6" blockId="29.[151,1437,151,573]" pageId="29" pageNumber="30">
<emphasis id="70E67D48FF88FF86FF6DFECEFEC9E0E2" bold="true" box="[199,383,367,392]" pageId="29" pageNumber="30">Natural history</emphasis>
. This species was found under large live leaves along a brook in the forest. The webs were very indistinct, in contrast to those of
<taxonomicName id="8592DAD9FF88FF86FD95FE34FD0BE0C7" baseAuthorityName="Huber" baseAuthorityYear="2011" box="[575,701,404,429]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ethagala">
<emphasis id="70E67D48FF88FF86FD95FE34FD0BE0C7" box="[575,701,404,429]" italics="true" pageId="29" pageNumber="30">T. ethagala</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="0A88F2D1FF88FF86FF6DFE16FF41E357" pageId="29" pageNumber="30" type="distribution">
<paragraph id="422DA15AFF88FF86FF6DFE16FF41E357" blockId="29.[151,1437,151,573]" pageId="29" pageNumber="30">
<emphasis id="70E67D48FF88FF86FF6DFE16FEEFE0BA" bold="true" box="[199,345,439,464]" pageId="29" pageNumber="30">Distribution</emphasis>
. Known from several high elevation localities in central
<collectingCountry id="3A85E1CAFF88FF86FC77FE16FBFDE0BA" box="[989,1099,439,464]" name="Sri Lanka" pageId="29" pageNumber="30">Sri Lanka</collectingCountry>
(
<figureCitation id="DAA9BDDFFF88FF86FBF1FE16FB08E0BA" box="[1115,1214,439,464]" captionStart="FIGURES 224225" captionStartId="52.[151,264,1873,1896]" captionTargetBox="[151,1436,1159,1861]" captionTargetId="figure@52.[151,1436,1159,1861]" captionTargetPageId="52" captionText="FIGURES 224225. Known distribution of Tissahamia in Sri Lanka; “?”, female with uncertain identity from Eastern Sinharaja." httpUri="https://zenodo.org/record/2625102/files/figure.png" pageId="29" pageNumber="30">Fig. 225</figureCitation>
). The single female from Avissawella (ZFMK, Ar 5088) assigned to
<taxonomicName id="8592DAD9FF88FF86FD66FE7CFBA2E09E" authority="in Huber (2011)" authorityName="in Huber" authorityYear="2011" box="[716,1044,476,501]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF88FF86FD66FE7CFCE7E09E" box="[716,849,477,500]" italics="true" pageId="29" pageNumber="30">T. maturata</emphasis>
in
<bibRefCitation id="2603DCABFF88FF86FCD1FE7DFBA2E09E" author="Huber, B. A." box="[891,1044,476,501]" pageId="29" pageNumber="30" pagination="1 - 509" refId="ref27101" refString="Huber, B. A. (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien, 58, 1 - 509." type="journal article" year="2011">Huber (2011)</bibRefCitation>
</taxonomicName>
is here considered to represent
<taxonomicName id="8592DAD9FF88FF86FA2DFE7CFF50E372" authorityName="Huber" authorityYear="2019" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="karuna">
<emphasis id="70E67D48FF88FF86FA2DFE7CFF50E372" italics="true" pageId="29" pageNumber="30">T. karuna</emphasis>
</taxonomicName>
. The identity of the single female from Eastern Sinharaja assigned to
<taxonomicName id="8592DAD9FF88FF86FB84FDA0FAC8E372" authority="in Huber (2011)" authorityName="in Huber" authorityYear="2011" box="[1070,1406,511,536]" class="Arachnida" family="Pholcidae" genus="Tissahamia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="maturata">
<emphasis id="70E67D48FF88FF86FB84FDA0FB03E372" box="[1070,1205,513,536]" italics="true" pageId="29" pageNumber="30">T. maturata</emphasis>
in
<bibRefCitation id="2603DCABFF88FF86FB48FE5EFAC8E372" author="Huber, B. A." box="[1250,1406,511,536]" pageId="29" pageNumber="30" pagination="1 - 509" refId="ref27101" refString="Huber, B. A. (2011) Revision and cladistic analysis of Pholcus and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien, 58, 1 - 509." type="journal article" year="2011">Huber (2011)</bibRefCitation>
</taxonomicName>
is dubious.
</paragraph>
</subSubSection>
</treatment>
</document>