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<document ID-DOI="http://doi.org/10.5281/zenodo.4758613" ID-GBIF-Dataset="0f978355-d6d8-40c8-984b-d0183880c3e0" ID-GBIF-Taxon="182067881" ID-ISSN="1854-0392" ID-Zenodo-Dep="4758613" ID-ZooBank="9A7BCA7F-E6CC-4B8E-8E13-BD85A0301DC2" checkinTime="1620834534472" checkinUser="felipe" docAuthor="Mayorga, Alfredo" docDate="2016" docId="390C87B65764710AE9972C56E6BDFE52" docLanguage="en" docName="Illiesia.12.12.64-73.pdf" docOrigin="Illiesia 12 (12)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Anacroneuria quadriloba Jewett. Moreover 1958" docType="treatment" docVersion="6" lastPageId="7" lastPageNumber="71" masterDocId="C535FFCE5761710DEA052970E224FFE5" masterDocTitle="A New Species Of Anacroneuria Klapálek (Plecoptera: Perlidae) And Complementary Descriptions Of Three Additional Species From Mexico, With Comments On The Current Knowledge Of Mexican Species Of The Genus" masterLastPageNumber="73" masterPageNumber="64" pageId="5" pageNumber="69" updateTime="1643491587280" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A New Species Of Anacroneuria Klapálek (Plecoptera: Perlidae) And Complementary Descriptions Of Three Additional Species From Mexico, With Comments On The Current Knowledge Of Mexican Species Of The Genus</mods:title>
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<mods:namePart>Mayorga, Alfredo</mods:namePart>
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<mods:title>Illiesia</mods:title>
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<mods:date>2016</mods:date>
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<mods:number>12</mods:number>
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<mods:number>12</mods:number>
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<subSubSection box="[914,1363,1318,1342]" pageId="5" pageNumber="69" type="nomenclature">
<paragraph blockId="5.[914,1363,1318,1342]" box="[914,1363,1318,1342]" pageId="5" pageNumber="69">
<heading bold="true" box="[914,1363,1318,1342]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="69" reason="2">
<taxonomicName authority="Jewett, 1958" authorityName="Jewett. Moreover" authorityYear="1958" box="[914,1363,1318,1342]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="69" phylum="Arthropoda" rank="species" species="quadriloba">
<emphasis bold="true" box="[914,1363,1318,1342]" pageId="5" pageNumber="69">
<emphasis bold="true" box="[914,1213,1318,1342]" italics="true" pageId="5" pageNumber="69">Anacroneuria quadriloba</emphasis>
<bibRefCitation author="Jewett, S. G." box="[1220,1363,1318,1342]" pageId="5" pageNumber="69" pagination="159 - 175" refId="ref4306" refString="Jewett, S. G. 1958. Stoneflies of the genus Anacroneuria from Mexico and Central America. The American Midland Naturalist, 60: 159 - 175." type="journal article" year="1958">Jewett, 1958</bibRefCitation>
</emphasis>
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<paragraph blockId="5.[1068,1210,1354,1378]" box="[1068,1210,1354,1378]" pageId="5" pageNumber="69">
(
<figureCitation box="[1076,1203,1354,1378]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="5" pageNumber="69">Figs. 11-16</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="69" type="multiple">
<paragraph blockId="5.[840,1439,1429,1482]" pageId="5" pageNumber="69">
http://lsid.speciesfile.org/urn:lsid:
<taxonomicName box="[1173,1285,1429,1451]" class="Insecta" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="69" phylum="Arthropoda" rank="order">Plecoptera</taxonomicName>
.speciesfile.org: TaxonName:1771
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="69" type="reference_group">
<paragraph blockId="5.[873,1443,1533,1588]" pageId="5" pageNumber="69">
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<treatmentCitation author="Jewett, S. G." box="[873,1303,1533,1555]" page="166" pageId="5" pageNumber="69" year="1958">
<taxonomicName authority="Jewett, 1958: 166" authorityName="Jewett. Moreover" authorityPageNumber="166" authorityYear="1958" box="[873,1261,1533,1555]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="69" phylum="Arthropoda" rank="species" species="quadriloba">
<emphasis box="[873,1118,1533,1554]" italics="true" pageId="5" pageNumber="69">Anacroneuria quadriloba</emphasis>
<bibRefCitation author="Jewett, S. G." box="[1128,1303,1533,1555]" pageId="5" pageNumber="69" pagination="159 - 175" refId="ref4306" refString="Jewett, S. G. 1958. Stoneflies of the genus Anacroneuria from Mexico and Central America. The American Midland Naturalist, 60: 159 - 175." type="journal article" year="1958">Jewett, 1958:166</bibRefCitation>
</taxonomicName>
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.
<materialsCitation ID-GBIF-Occurrence="3111885304" collectionCode="FMNH" country="Mexico" location="Metlac" pageId="5" pageNumber="69" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz" typeStatus="holotype">
<typeStatus box="[1315,1415,1533,1555]" pageId="5" pageNumber="69">Holotype</typeStatus>
<specimenCount box="[1425,1443,1533,1555]" pageId="5" pageNumber="69" type="female"></specimenCount>
(
<collectionCode box="[880,960,1566,1588]" pageId="5" pageNumber="69">FMNH</collectionCode>
),
<location LSID="urn:lsid:plazi:treatment:390C87B65764710AE9972C56E6BDFE52:B47A607B57647108E9D42F6EE63FF9D1" box="[977,1051,1566,1588]" country="Mexico" name="Metlac" pageId="5" pageNumber="69" stateProvince="Veracruz">Metlac</location>
,
<collectingRegion box="[1060,1159,1566,1587]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="5" pageNumber="69">Veracruz</collectingRegion>
,
<collectingCountry box="[1169,1246,1566,1588]" name="Mexico" pageId="5" pageNumber="69">Mexico</collectingCountry>
</materialsCitation>
</treatmentCitationGroup>
</paragraph>
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<subSubSection pageId="5" pageNumber="69" type="materials_examined">
<paragraph blockId="5.[835,1443,1634,1911]" pageId="5" pageNumber="69">
<emphasis bold="true" box="[835,1071,1634,1658]" pageId="5" pageNumber="69">Material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="3111885305" collectingDate="1996-07-12" collectionCode="R" collectorName="Barba, A" country="Mexico" elevation="610" latitude="17.603462" location="Oaxaca" longLatPrecision="14" longitude="-96.59556" pageId="5" pageNumber="69" specimenCount="610" specimenCount-female="1" specimenCount-male="1" stateProvince="Oaxaca">
<emphasis bold="true" box="[1078,1291,1634,1658]" pageId="5" pageNumber="69">
<collectingCountry box="[1078,1192,1634,1658]" name="Mexico" pageId="5" pageNumber="69">MEXICO</collectingCountry>
:
<collectingRegion box="[1204,1291,1634,1658]" country="Mexico" name="Oaxaca" pageId="5" pageNumber="69">Oaxaca</collectingRegion>
</emphasis>
, km 11, carr. Cuicatlán-Sgo. Quiotepec,
<geoCoordinate box="[1228,1443,1671,1695]" degrees="17" direction="north" minutes="36" orientation="latitude" pageId="5" pageNumber="69" precision="1" seconds="12.46" value="17.603462">17°36´12.46” N</geoCoordinate>
<geoCoordinate box="[835,985,1707,1731]" degrees="96" direction="west" minutes="35" orientation="longitude" pageId="5" pageNumber="69" precision="15" seconds="44" value="-96.59556">96°35´44” W</geoCoordinate>
,
<specimenCount box="[997,1079,1707,1731]" pageId="5" pageNumber="69" type="generic">
<quantity box="[997,1075,1707,1731]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="5" pageNumber="69" unit="m" value="610.0">
<elevation box="[997,1075,1707,1731]" metricMagnitude="2" metricUnit="m" metricValue="6.1" pageId="5" pageNumber="69" unit="m" value="610.0">610 m</elevation>
</quantity>
.
</specimenCount>
a.s.l.,
<date box="[1142,1288,1707,1731]" pageId="5" pageNumber="69" value="1996-07-12">
<collectingDate box="[1142,1288,1707,1731]" pageId="5" pageNumber="69" value="1996-07-12">12 July 1996</collectingDate>
</date>
,
<collectionCode box="[1300,1321,1707,1730]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="69">R</collectionCode>
.
<collectorName box="[1332,1439,1707,1731]" pageId="5" pageNumber="69">Barba, A</collectorName>
. Rojas,
<specimenCount box="[911,952,1743,1767]" pageId="5" pageNumber="69" type="male">1 ♂</specimenCount>
,
<specimenCount box="[965,998,1743,1767]" pageId="5" pageNumber="69" type="female">1♀</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="7" lastPageNumber="71" pageId="5" pageNumber="69" type="description">
<paragraph blockId="5.[835,1443,1634,1911]" lastBlockId="7.[189,797,199,1914]" lastPageId="7" lastPageNumber="71" pageId="5" pageNumber="69">
<emphasis bold="true" box="[835,1014,1778,1802]" pageId="5" pageNumber="69">Adult habitus.</emphasis>
General body ground color brown (
<figureCitation box="[843,930,1815,1839]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="5" pageNumber="69">Fig. 11</figureCitation>
). Head brown medially; pigment pattern extends from occiput, through ocelli and anterior to M-line and lappets; M-line wide and pale, interrupted by a space and continued forward with a brown pattern V-shaped curved; lappets large, brown with small tufts of bristles at the base; brown circum-antennal ridges connect lappets to anterior margin of eyes; darker shading posterior to eyes and clothed with variously sized bristles. Pronotum with pale, narrow band at center, discs patterned with elongate, pale, C-shaped stripes; mediolateral areas of discs with dark rugosities; lateral margins pale, anterior and posterior margins with narrow, dark line (
<figureCitation box="[616,705,559,583]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="7" pageNumber="71">Fig. 12</figureCitation>
). Wing membranes amber, veins brown, costa transparent and sub-costa darker. Forelegs brown; femora grading from pale to dark banded at apex (
<figureCitation box="[695,776,667,691]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="7" pageNumber="71">Fig. 11</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.4752826" ID-Zenodo-Dep="4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="6" pageNumber="70" startId="6.[189,241,1863,1887]" targetBox="[362,1374,189,1816]" targetPageId="6">
<paragraph blockId="6.[189,1443,1863,1924]" pageId="6" pageNumber="70">
Figs. 11-16.
<taxonomicName authorityName="Jewett. Moreover" authorityYear="1958" box="[327,595,1863,1887]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="70" phylum="Arthropoda" rank="species" species="quadriloba">
<emphasis box="[327,595,1863,1887]" italics="true" pageId="6" pageNumber="70">Anacroneuria quadriloba</emphasis>
</taxonomicName>
. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate.
</paragraph>
</caption>
<paragraph blockId="7.[189,797,199,1914]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[189,258,703,727]" pageId="7" pageNumber="71">Male.</emphasis>
Forewing length
<quantity box="[471,556,703,727]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="7" pageNumber="71" unit="mm" value="18.0">18 mm</quantity>
. General body color as above. Ventral: aedeagal apex truncate with notched tip; aedeagus constricted subapically and bearing a pair of large membranous lobes covering most of apex and shoulders (
<figureCitation box="[566,652,847,871]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="7" pageNumber="71">Fig. 13</figureCitation>
); shoulders rounded and sclerotized; hooks typical in appearance, the space between hooks &quot;skeleton-key shaped&quot;; medial processes of hooks forming sharp corners; mediobasal area of hooks with bristles moderate in length (
<figureCitation box="[431,513,1027,1051]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="7" pageNumber="71">Fig. 13</figureCitation>
). Lateral: apex rounded frontally and with posterior projection with pointy tip; protuberance at median area and concave at base (
<figureCitation box="[263,351,1135,1159]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="7" pageNumber="71">Fig. 14</figureCitation>
). Dorsal: of aedeagus without keel; apex as discussed above; apex divergent (
<figureCitation box="[679,760,1171,1195]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="7" pageNumber="71">Fig. 15</figureCitation>
).
</paragraph>
<paragraph blockId="7.[189,797,199,1914]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[189,283,1206,1230]" pageId="7" pageNumber="71">Female.</emphasis>
Forewing length
<quantity box="[517,609,1206,1230]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="7" pageNumber="71" unit="mm" value="23.0">23 mm</quantity>
. General body color as above. Subgenital plate 4-lobed with slightly curved, setose posterior borders; lateral margins of lobes same in length as in their width; shallow U-shaped notch at middle of lobes. Sternum 9 without transverse sclerite; dark, mesal sclerite M-shaped with a deep and U- or V-shaped notch at apex, entire structure covered with long hairs; rest of sternum 9 mostly membranous with few hairs (
<figureCitation box="[313,394,1530,1554]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="7" pageNumber="71">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="71" type="diagnosis">
<paragraph blockId="7.[189,797,199,1914]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[189,319,1566,1590]" pageId="7" pageNumber="71">Diagnosis.</emphasis>
This species is similar in head and pronotal coloration to the recently described
<taxonomicName authority="Gutierrez-Fonseca &amp; Springer, 2015" authorityName="Gutierrez-Fonseca &amp; Springer" authorityYear="2015" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="71" phylum="Arthropoda" rank="species" species="quetzali">
<emphasis italics="true" pageId="7" pageNumber="71">A. quetzali</emphasis>
<bibRefCitation author="Gutierrez-Fonseca, P. E. &amp; M. Springer" box="[287,726,1638,1663]" pageId="7" pageNumber="71" pagination="595 - 600" refId="ref4260" refString="Gutierrez-Fonseca, P. E. &amp; M. Springer. 2015. A new species of Anacroneuria Klapalek 1909 (Plecoptera: Perlidae) and notes on the altitudinal distribution of the genus in Costa Rica. Zootaxa, 4058: 595 - 600." type="journal article" year="2015">Gutiérrez-Fonseca &amp; Springer, 2015</bibRefCitation>
</taxonomicName>
from
<collectingCountry box="[189,319,1675,1699]" name="Costa Rica" pageId="7" pageNumber="71">Costa Rica</collectingCountry>
. However,
<taxonomicName authorityName="Gutierrez-Fonseca &amp; Springer" authorityYear="2015" box="[457,577,1675,1699]" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="71" phylum="Arthropoda" rank="species" species="quetzali">
A.
<emphasis box="[493,577,1675,1699]" italics="true" pageId="7" pageNumber="71">quetzali</emphasis>
</taxonomicName>
is overall a larger species, its aedeagus subapex is not constricted, and its apical membranes are smaller than in
<emphasis italics="true" pageId="7" pageNumber="71">
<taxonomicName authorityName="Jewett. Moreover" authorityYear="1958" class="Insecta" family="Perlidae" genus="Anacroneuria" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="71" phylum="Arthropoda" rank="species" species="quadriloba">A. quadriloba</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="71" type="distribution">
<paragraph blockId="7.[189,797,199,1914]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[189,348,1818,1842]" pageId="7" pageNumber="71">Distribution.</emphasis>
<collectingCountry box="[356,487,1818,1842]" name="Guatemala" pageId="7" pageNumber="71">Guatemala</collectingCountry>
,
<collectingCountry box="[498,584,1818,1842]" name="Mexico" pageId="7" pageNumber="71">Mexico</collectingCountry>
(
<collectingRegion box="[600,699,1818,1842]" country="Mexico" name="Chiapas" pageId="7" pageNumber="71">Chiapas</collectingRegion>
,
<collectingRegion box="[710,797,1818,1842]" country="Mexico" name="Oaxaca" pageId="7" pageNumber="71">Oaxaca</collectingRegion>
and
<collectingRegion box="[241,352,1855,1879]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="7" pageNumber="71">Veracruz</collectingRegion>
),
<collectingCountry box="[370,465,1855,1878]" name="Panama" pageId="7" pageNumber="71">Panama</collectingCountry>
(
<bibRefCitation author="DeWalt, R. E. &amp; M. D. Maehr &amp; U. Neu-Becker &amp; G. Stueber" box="[480,698,1854,1878]" pageId="7" pageNumber="71" pagination="2016" refId="ref4080" refString="DeWalt, R. E., M. D. Maehr, U. Neu-Becker and G. Stueber. 2016. Plecoptera Species File Online. Version 5.0 / 5.0. 10 / 31 / 2016. http: // Plecoptera. SpeciesFile. org" type="book chapter" year="2016">DeWalt et al. 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="71" type="discussion">
<paragraph blockId="7.[189,797,199,1914]" lastBlockId="7.[835,1443,199,439]" pageId="7" pageNumber="71">
<emphasis bold="true" box="[189,328,1890,1914]" pageId="7" pageNumber="71">Comments.</emphasis>
This species was redescribed by
<bibRefCitation author="Stark, B. P. &amp; B. C. Kondratieff" box="[735,1084,199,1914]" pageId="7" pageNumber="71" pagination="1 - 64" refId="ref4432" refString="Stark, B. P. &amp; B. C. Kondratieff. 2004. Anacroneuria from Mexico and upper Mesoamerica (Plecoptera: Perlidae). Monographs of the Western North American Naturalist, 2: 1 - 64." type="journal article" year="2004">Stark &amp; Kondratieff (2004)</bibRefCitation>
, but because of the age of the material, only a diffuse pigment pattern was present on the head and pronotum. Examination of fresh specimens revealed a distinctive head and pronotal pattern (
<figureCitation box="[1087,1249,343,367]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="7" pageNumber="71">Figs. 11&amp;12</figureCitation>
). The female subgenital plate is illustrated from recently collected specimens (
<figureCitation box="[1083,1164,415,439]" captionStart="Figs" captionStartId="6.[189,241,1863,1887]" captionTargetBox="[362,1374,189,1816]" captionTargetId="figure-89@6.[362,1269,189,1096]" captionTargetPageId="6" captionText="Figs. 11-16. Anacroneuria quadriloba. 11. Male adult habitus. 12. Head and pronotum. 13. Aedeagus, ventral. 14. Aedeagus, dorsal. 15. Aedeagus, lateral. 16. Female subgenital plate." figureDoi="http://doi.org/10.5281/zenodo.4752826" httpUri="https://zenodo.org/record/4752826/files/figure.png" pageId="7" pageNumber="71">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>