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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.541.6058" ID-GBIF-Dataset="fa40454d-8f0e-4387-86ee-8dfc421d0e05" ID-PMC="PMC4714381" ID-Pensoft-Pub="1313-2970-541-109" ID-PubMed="26798284" ID-ZBK="C336A3C4DBCB49C5898C8FA38BDFF0C0" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-541-109" ModsDocOrigin="ZooKeys 541" ModsDocTitle="Systematics of Nothopsini (Serpentes, Dipsadidae), with a new species of Synophis from the Pacific Andean slopes of southwestern Ecuador" checkinTime="1451243656936" checkinUser="pensoft" docAuthor="Pyron, R. Alexander, Guayasamin, Juan M., Penafiel, Nicolas, Bustamante, Lucas &amp; Arteaga, Alejandro" docDate="2015" docId="31983D5A90E401E9D297A1C2429F3C22" docLanguage="en" docName="ZooKeys 541: 109-147" docOrigin="ZooKeys 541" docSource="http://dx.doi.org/10.3897/zookeys.541.6058" docTitle="Nothopsini Cope 1871" docType="treatment" docVersion="4" lastPageNumber="129" masterDocId="FF8E84462424FFD3A52779666640FF82" masterDocTitle="Systematics of Nothopsini (Serpentes, Dipsadidae), with a new species of Synophis from the Pacific Andean slopes of southwestern Ecuador" masterLastPageNumber="147" masterPageNumber="109" pageNumber="129" updateTime="1668162264059" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Systematics of Nothopsini (Serpentes, Dipsadidae), with a new species of Synophis from the Pacific Andean slopes of southwestern Ecuador</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pyron, R. Alexander</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guayasamin, Juan M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Penafiel, Nicolas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bustamante, Lucas</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Arteaga, Alejandro</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>541</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>109</mods:start>
<mods:end>147</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.541.6058</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.541.6058</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-541-109</mods:identifier>
<mods:identifier type="ZBK">C336A3C4DBCB49C5898C8FA38BDFF0C0</mods:identifier>
<mods:identifier type="ZooBank">C336A3C4DBCB49C5898C8FA38BDFF0C0</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="127891637" LSID="urn:lsid:plazi:treatment:31983D5A90E401E9D297A1C2429F3C22" httpUri="http://treatment.plazi.org/id/31983D5A90E401E9D297A1C2429F3C22" lastPageId="21" lastPageNumber="129" pageId="20" pageNumber="129">
<subSubSection pageId="20" pageNumber="129" type="multiple">
<paragraph pageId="20" pageNumber="129">Taxon classification Animalia Squamata Colubridae</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="129" type="nomenclature">
<paragraph pageId="20" pageNumber="129">
Tribe
<taxonomicName authority="Cope, 1871" authorityName="Cope" authorityYear="1871" lsidName="" pageId="20" pageNumber="129" rank="tribe" tribe="Nothopsini">Nothopsini Cope, 1871</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="129" type="reference_group">
<paragraph pageId="20" pageNumber="129">
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Nothopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothopsis" order="Squamata" pageId="20" pageNumber="129" phylum="Chordata" rank="genus">Nothopsis</taxonomicName>
Cope, 1871 (type genus by monotypy)
</paragraph>
<paragraph pageId="20" pageNumber="129">
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Nothopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothopsis rugosus" order="Squamata" pageId="20" pageNumber="129" phylum="Chordata" rank="species" species="rugosus">Nothopsis rugosus</taxonomicName>
Cope, 1871
</paragraph>
<paragraph pageId="20" pageNumber="129">
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Nothopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothopsis affinis" order="Squamata" pageId="20" pageNumber="129" phylum="Chordata" rank="species" species="affinis">Nothopsis affinis</taxonomicName>
Boulenger, 1895 (Holotype BMNH 1946.1.15.62, &quot;Salidero, NW Ecuador, 350ft&quot;) [subjective junior synonym of
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Nothopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothopsis rugosus" order="Squamata" pageId="20" pageNumber="129" phylum="Chordata" rank="species" species="rugosus">Nothopsis rugosus</taxonomicName>
fide Dunn &amp; Dowling 1957]
</paragraph>
<paragraph pageId="20" pageNumber="129">
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Nothopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothopsis torresi" order="Squamata" pageId="20" pageNumber="129" phylum="Chordata" rank="species" species="torresi">Nothopsis torresi</taxonomicName>
Taylor, 1951 (Holotype KU 28710,
<normalizedToken originalValue="“Morehead">&quot;'Morehead'</normalizedToken>
Finca, 5 miles southwest of Turrialba, Costa Rica&quot;) [subjective junior synonym of
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Nothopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothopsis rugosus" order="Squamata" pageId="20" pageNumber="129" phylum="Chordata" rank="species" species="rugosus">Nothopsis rugosus</taxonomicName>
fide Dunn &amp; Dowling 1957]
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="129" type="holotype">
<paragraph pageId="20" pageNumber="129">Holotype.</paragraph>
<paragraph pageId="20" pageNumber="129">USNM 12427, type locality &quot;Isthmus of Darien [Panama]&quot;</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="129" type="etymology">
<paragraph pageId="20" pageNumber="129">Etymology.</paragraph>
<paragraph pageId="20" pageNumber="129">
From the Greek nothos for
<normalizedToken originalValue="“bastard”">&quot;bastard&quot;</normalizedToken>
and opsis for
<normalizedToken originalValue="“appearance,”">&quot;appearance,&quot;</normalizedToken>
with
<bibRefCitation author="Cope, ED" journalOrPublisher="Proceedings of the Academy of Natural Sciences Philadelphia" pageId="24" pageNumber="133" pagination="200 - 224" title="Ninth contribution to the herpetology of Tropical America." volume="23" year="1871">Cope (1871)</bibRefCitation>
apparently referring to putative mimicry of
<taxonomicName class="Reptilia" family="Viperidae" genus="Bothrops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bothrops atrox" order="Squamata" pageId="20" pageNumber="129" phylum="Chordata" rank="species" species="atrox">Bothrops atrox</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="20" pageNumber="129" type="description">
<paragraph pageId="20" pageNumber="129">Description.</paragraph>
<paragraph pageId="20" pageNumber="129">
A relatively small-sized (&lt;350mm SVL) dipsadine snake, ranging in Central and South America from Honduras to Colombia and Ecuador, in lowland and middle-elevation rainforests, 250-900m, distinguishable from nearly all other similar or related snakes in the area by the rugose, granular nature of the dorsal scales, in particular lacking differentiation of the cephalic scales with the exception of well-defined internasals and poorly defined frontal and parietals, which are separated by rows of irregular, undifferentiated scales. Color pattern consists of irregular and poorly defined blotches of blackish or light, dark, and yellowish brown. With respect to the characters described here for diaphorolepidine species,
<taxonomicName class="Reptilia" family="Dipsadidae" genus="Nothopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothopsis rugosus" order="Squamata" pageId="20" pageNumber="129" phylum="Chordata" rank="species" species="rugosus">Nothopsis rugosus</taxonomicName>
typically exhibits 19-21 maxillary teeth, 9-13 supralabials, 11-16 infralabials, 149-162 ventrals, 81-112 subcaudals, dorsal scales in (24
<normalizedToken originalValue="30)-(2630)-(22">-30)-(26-30)-(22-</normalizedToken>
26) rows, SVL of 151-320mm, and tail length of 61-133mm (see
<bibRefCitation author="Dunn, ER" journalOrPublisher="Copeia" pageId="24" pageNumber="133" pagination="255 - 261" title="The neotropical snake genus Nothopsis Cope." url="10.2307/1439148" volume="1957" year="1957">Dunn and Dowling 1957</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="21" lastPageNumber="130" pageId="20" pageNumber="129" type="notes">
<paragraph pageId="20" pageNumber="129">Notes.</paragraph>
<paragraph lastPageId="21" lastPageNumber="130" pageId="20" pageNumber="129">
This taxon has historically been divided up into as many as three species (see
<bibRefCitation author="Dunn, ER" journalOrPublisher="Copeia" pageId="24" pageNumber="133" pagination="255 - 261" title="The neotropical snake genus Nothopsis Cope." url="10.2307/1439148" volume="1957" year="1957">Dunn and Dowling 1957</bibRefCitation>
), though only a single species is currently recognized. There may be cryptic variation or undiscovered diversity within this group. Note that
<pageBreakToken pageId="21" pageNumber="130" start="start">the</pageBreakToken>
family name was originally spelled
<taxonomicName above-genus="Nothopidae" family="Nothopidae" lsidName="" pageId="21" pageNumber="130" rank="above-genus">Nothopidae</taxonomicName>
by
<bibRefCitation author="Cope, ED" journalOrPublisher="Proceedings of the Academy of Natural Sciences Philadelphia" pageId="24" pageNumber="133" pagination="200 - 224" title="Ninth contribution to the herpetology of Tropical America." volume="23" year="1871">Cope (1871)</bibRefCitation>
, but
<normalizedToken originalValue="ops">-ops-</normalizedToken>
is the correct stem from -opsis, and
<taxonomicName above-genus="Nothopsidae" family="Nothopsidae" lsidName="" pageId="21" pageNumber="130" rank="above-genus">Nothopsidae</taxonomicName>
(and
<taxonomicName lsidName="" pageId="21" pageNumber="130" rank="tribe" tribe="Nothopsini">Nothopsini</taxonomicName>
) is thus the correct spelling, as adopted by later authors.
</paragraph>
</subSubSection>
</treatment>
</document>