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<document id="71615C72B03FDDFEFDE435DA5FCC9065" ID-CLB-Dataset="101998" ID-DOI="10.11646/zootaxa.5168.1.6" ID-GBIF-Dataset="4e44572a-bd6a-4f63-89fd-02699f087ab1" ID-ISSN="1175-5326" ID-Zenodo-Dep="6876467" ID-ZooBank="21BFA0D5-3F5C-440A-9291-A2D96A31242A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658418833899" checkinUser="plazi" docAuthor="Liu, Liyuan, Wang, Yan, Guo, Qiuhong, Chen, Xiaosheng &amp; Wang, Xingmin" docDate="2022" docId="2A00879F6B27583AFF49FA52FACAEA1A" docLanguage="en" docName="zootaxa.5168.1.6.pdf" docOrigin="Zootaxa 5168 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sulcolotis threadis Liu &amp; Wang &amp; Guo &amp; Chen &amp; Wang 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="78" masterDocId="D639FFE76B255839FFDEFF8DFFA8E803" masterDocTitle="Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines" masterLastPageNumber="82" masterPageNumber="75" pageNumber="77" updateTime="1699361246700" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="BE3C61C7037E1827C340EBF464011A57">Three new species of the genus Sulcolotis Miyatake (Coleoptera, Coccinellidae Sticholotidini) from the Philippines</mods:title>
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<mods:namePart id="7E7CBAA5562FC5DD41C5232BD0938428">Liu, Liyuan</mods:namePart>
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<mods:affiliation id="58298537DF182E7B290EB02B2D919CFD">Key Laboratory of Bio-Pesticide Innovation and Application, Engineering Technology Research Center of Agricultural Pest Biocontrol, Guangdong Province; Department of Entomology, South China Agricultural University, Guangzhou, 510640, China. Department of Forest Protection, College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou, 510640, China. xshchen @ scau. edu. cn; https: // orcid. org / 0000 - 0001 - 8253 - 4943 &amp; 332132824 @ qq. com; https: // orcid. org / 0000 - 0003 - 1160 - 8209</mods:affiliation>
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<mods:namePart id="CB5A95A1009DFB2CEE9B946EF33B2A21">Wang, Yan</mods:namePart>
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<mods:namePart id="0FAF17CAC94C399C7BE5BCC4CFC2E8BC">Guo, Qiuhong</mods:namePart>
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<mods:affiliation id="31EF0E091A486D9D963AFD75541E6790">Key Laboratory of Bio-Pesticide Innovation and Application, Engineering Technology Research Center of Agricultural Pest Biocontrol, Guangdong Province; Department of Entomology, South China Agricultural University, Guangzhou, 510640, China. Department of Forest Protection, College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou, 510640, China. xshchen @ scau. edu. cn; https: // orcid. org / 0000 - 0001 - 8253 - 4943 &amp; qiuhongdd @ 163. com; https: // orcid. org / 0000 - 0003 - 1586 - 893 X</mods:affiliation>
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<mods:namePart id="E04CD09D6F73E54D806A057565B56511">Chen, Xiaosheng</mods:namePart>
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<mods:affiliation id="D01AEE7682E618D79FC2B488B6DB2275">Key Laboratory of Bio-Pesticide Innovation and Application, Engineering Technology Research Center of Agricultural Pest Biocontrol, Guangdong Province; Department of Entomology, South China Agricultural University, Guangzhou, 510640, China. Department of Forest Protection, College of Forestry and Landscape Architecture, South China Agricultural University, Guangzhou, 510640, China. xshchen @ scau. edu. cn; https: // orcid. org / 0000 - 0001 - 8253 - 4943</mods:affiliation>
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<mods:namePart id="2A0D0B5E2AAF38B262492038ACF791D1">Wang, Xingmin</mods:namePart>
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<treatment id="2A00879F6B27583AFF49FA52FACAEA1A" ID-DOI="http://doi.org/10.5281/zenodo.6876457" ID-GBIF-Taxon="197836909" ID-Zenodo-Dep="6876457" LSID="urn:lsid:plazi:treatment:2A00879F6B27583AFF49FA52FACAEA1A" httpUri="http://treatment.plazi.org/id/2A00879F6B27583AFF49FA52FACAEA1A" lastPageId="3" lastPageNumber="78" pageId="2" pageNumber="77">
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<paragraph id="A21636896B27583BFF49FA52FD28EDFA" blockId="2.[151,640,1503,1565]" box="[151,640,1503,1530]" pageId="2" pageNumber="77">
<heading id="F95E81E56B27583BFF49FA52FD28EDFA" bold="true" box="[151,640,1503,1530]" fontSize="11" level="1" pageId="2" pageNumber="77" reason="1">
<emphasis id="90DDEA9B6B27583BFF49FA52FD28EDFA" bold="true" box="[151,640,1503,1530]" pageId="2" pageNumber="77">
<taxonomicName id="65A94D0A6B27583BFF49FA52FDB2EDFA" authority="Liu &amp; Wang" authorityName="Liu &amp; Wang &amp; Guo &amp; Chen &amp; Wang" authorityYear="2022" box="[151,538,1503,1530]" class="Insecta" family="Coccinellidae" genus="Sulcolotis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="threadis" status="sp. nov.">
<emphasis id="90DDEA9B6B27583BFF49FA52FED0EDF9" bold="true" box="[151,376,1503,1530]" italics="true" pageId="2" pageNumber="77">Sulcolotis threadis</emphasis>
Liu &amp; Wang
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<taxonomicNameLabel id="8BEE57E06B27583BFDFCFA52FD28EDFA" box="[546,640,1503,1529]" pageId="2" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="EAB365026B27583BFF49F98EFEBAEE1E" box="[151,274,1539,1565]" pageId="2" pageNumber="77" type="description">
<paragraph id="A21636896B27583BFF49F98EFEBAEE1E" blockId="2.[151,640,1503,1565]" box="[151,274,1539,1565]" pageId="2" pageNumber="77">
(
<figureCitation id="3A922A0C6B27583BFF41F98EFEA0EE1E" box="[159,264,1539,1565]" captionStart="FIGURE 1" captionStartId="2.[151,250,1361,1385]" captionTargetBox="[152,1430,316,1325]" captionTargetId="figure-66@2.[151,1437,306,1334]" captionTargetPageId="2" captionText="FIGURE 1. Sulcolotis threadis Liu et Wang sp. n.: a) head, ventral; b. head, clypeus; c. antenna; d. labrum; e. left mandible; f. right mandible; g. maxilla; h. labium; i. prothorax, ventral; j. meso- and metaventrite; k. spermatheca; l. female genitalia." figureDoi="http://doi.org/10.5281/zenodo.6876469" httpUri="https://zenodo.org/record/6876469/files/figure.png" pageId="2" pageNumber="77">Figs 1al</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="EAB365026B27583BFF49F9C6FEBAEEF5" pageId="2" pageNumber="77" type="diagnosis">
<paragraph id="A21636896B27583BFF49F9C6FEBAEEF5" blockId="2.[151,1437,1611,2033]" pageId="2" pageNumber="77">
<emphasis id="90DDEA9B6B27583BFF49F9C6FEBAEE66" bold="true" box="[151,274,1611,1637]" pageId="2" pageNumber="77">Diagnosis.</emphasis>
This species can be easily separated from
<taxonomicName id="65A94D0A6B27583BFD32F9C1FCDFEE66" authorityName="Miyatake" authorityYear="1994" box="[748,887,1612,1638]" class="Insecta" family="Coccinellidae" genus="Sulcolotis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="seminigra">
<emphasis id="90DDEA9B6B27583BFD32F9C1FCDFEE66" box="[748,887,1612,1638]" italics="true" pageId="2" pageNumber="77">S. seminigra</emphasis>
</taxonomicName>
by the following characters: elytron yellow, with a large black spot situated at 1/3 elytral length to base. In
<taxonomicName id="65A94D0A6B27583BFCFCF9FDFC06EE8A" authorityName="Miyatake" authorityYear="1994" box="[802,942,1648,1674]" class="Insecta" family="Coccinellidae" genus="Sulcolotis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="seminigra">
<emphasis id="90DDEA9B6B27583BFCFCF9FDFC06EE8A" box="[802,942,1648,1674]" italics="true" pageId="2" pageNumber="77">S. seminigra</emphasis>
</taxonomicName>
, elytra are black in basal half and yellow in apical half, without dark spots. It is also similar to
<taxonomicName id="65A94D0A6B27583BFD17F919FC8FEEAE" authorityName="Liu &amp; Wang &amp; Guo &amp; Chen &amp; Wang" authorityYear="2022" box="[713,807,1683,1710]" class="Insecta" family="Coccinellidae" genus="Sulcolotis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="90DDEA9B6B27583BFD17F919FC8FEEAE" box="[713,807,1683,1710]" italics="true" pageId="2" pageNumber="77">S. ovalis</emphasis>
</taxonomicName>
but can be separated from the latter by the bright yellow head and penis guide nearly as long as parameres. In
<taxonomicName id="65A94D0A6B27583BFD2FF935FCF8EED2" authorityName="Liu &amp; Wang &amp; Guo &amp; Chen &amp; Wang" authorityYear="2022" box="[753,848,1719,1746]" class="Insecta" family="Coccinellidae" genus="Sulcolotis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="77" phylum="Arthropoda" rank="species" species="ovalis">
<emphasis id="90DDEA9B6B27583BFD2FF935FCF8EED2" box="[753,848,1719,1746]" italics="true" pageId="2" pageNumber="77">S. ovalis</emphasis>
</taxonomicName>
, head is reddish brown and penis guide shorter than parameres.
</paragraph>
</subSubSection>
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<paragraph id="A21636896B27583BFF19F972FAECEF3D" blockId="2.[151,1437,1611,2033]" pageId="2" pageNumber="77">
<emphasis id="90DDEA9B6B27583BFF19F972FEF1EF1A" bold="true" box="[199,345,1791,1817]" pageId="2" pageNumber="77">Description.</emphasis>
TL:
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, TW:
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, TH:
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, TL/TW: 1.191.22, PL/PW: 0.260.29, EL/EW: 0.880.89, HW/PW: 0.560.63, PW/TW: 0.630.66, HW/TW: 0.380.40, Eye W/HW: 0.540.59.
</paragraph>
<paragraph id="A21636896B27583BFF19F8CAFB25EF86" blockId="2.[151,1437,1611,2033]" pageId="2" pageNumber="77">Head yellow. Pronotum, scutellar shield, and elytra bright yellow, with small dense punctures. Underside yellowish brown, except metaventrite and the middle part of the first ventrite blackish brown.</paragraph>
<paragraph id="A21636896B27583BFF19F802FEDDEFF2" blockId="2.[151,1437,1611,2033]" pageId="2" pageNumber="77">
Body rounded, weakly convex, dosally shiny and glabrous (
<figureCitation id="3A922A0C6B27583BFCB8F802FC70EFAA" box="[870,984,1935,1961]" captionStart="FIGURE 2" captionStartId="3.[151,250,1560,1584]" captionTargetBox="[160,1431,570,1528]" captionTargetId="figure-174@3.[151,1435,561,1537]" captionTargetPageId="3" captionText="FIGURE 2. Sulcolotis threadis Liu et Wang sp. n.: a) dorsal habitus; b. lateral habitus; c. frontal habitus; d. tegmen, ventral view; e. tegmen, lateral view; f. penis; g. abdomen, male." figureDoi="http://doi.org/10.5281/zenodo.6876473" httpUri="https://zenodo.org/record/6876473/files/figure.png" pageId="2" pageNumber="77">Figs 2ac</figureCitation>
). Head 0.39 times of elytral width (HW/ TW=1:2.6). Eyes finely faceted interocular distance 0.57 times of head width. Frons broad with punctures uniform and dense (
<figureCitation id="3A922A0C6B27583BFEC8F85AFEC0EFF2" box="[278,360,2007,2033]" captionStart="FIGURE 2" captionStartId="3.[151,250,1560,1584]" captionTargetBox="[160,1431,570,1528]" captionTargetId="figure-174@3.[151,1435,561,1537]" captionTargetPageId="3" captionText="FIGURE 2. Sulcolotis threadis Liu et Wang sp. n.: a) dorsal habitus; b. lateral habitus; c. frontal habitus; d. tegmen, ventral view; e. tegmen, lateral view; f. penis; g. abdomen, male." figureDoi="http://doi.org/10.5281/zenodo.6876473" httpUri="https://zenodo.org/record/6876473/files/figure.png" pageId="2" pageNumber="77">Fig. 2c</figureCitation>
).
</paragraph>
<paragraph id="A21636896B26583AFF19FF1AFD45E8D6" blockId="3.[151,1437,150,537]" pageId="3" pageNumber="78">
Pronotum 0.65 times of elytral width (PW/TW=1:1.5), moderately transverse, punctures uniform (
<figureCitation id="3A922A0C6B26583AFAC3FF1AFA26E8B2" box="[1309,1422,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1560,1584]" captionTargetBox="[160,1431,570,1528]" captionTargetId="figure-174@3.[151,1435,561,1537]" captionTargetPageId="3" captionText="FIGURE 2. Sulcolotis threadis Liu et Wang sp. n.: a) dorsal habitus; b. lateral habitus; c. frontal habitus; d. tegmen, ventral view; e. tegmen, lateral view; f. penis; g. abdomen, male." figureDoi="http://doi.org/10.5281/zenodo.6876473" httpUri="https://zenodo.org/record/6876473/files/figure.png" pageId="3" pageNumber="78">Figs 2a, c</figureCitation>
). Elytra with uniform and dense punctures (
<figureCitation id="3A922A0C6B26583AFDB0FF36FD49E8D6" box="[622,737,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1560,1584]" captionTargetBox="[160,1431,570,1528]" captionTargetId="figure-174@3.[151,1435,561,1537]" captionTargetPageId="3" captionText="FIGURE 2. Sulcolotis threadis Liu et Wang sp. n.: a) dorsal habitus; b. lateral habitus; c. frontal habitus; d. tegmen, ventral view; e. tegmen, lateral view; f. penis; g. abdomen, male." figureDoi="http://doi.org/10.5281/zenodo.6876473" httpUri="https://zenodo.org/record/6876473/files/figure.png" pageId="3" pageNumber="78">Figs 2ac</figureCitation>
).
</paragraph>
<paragraph id="A21636896B26583AFF19FF52FB77E8FA" blockId="3.[151,1437,150,537]" box="[199,1247,223,249]" pageId="3" pageNumber="78">
Ventrite 6 of posterior margin at middle slightly concave and ventrite 7 subtrianglar (
<figureCitation id="3A922A0C6B26583AFBA0FF52FB7AE8FA" box="[1150,1234,223,249]" captionStart="FIGURE 2" captionStartId="3.[151,250,1560,1584]" captionTargetBox="[160,1431,570,1528]" captionTargetId="figure-174@3.[151,1435,561,1537]" captionTargetPageId="3" captionText="FIGURE 2. Sulcolotis threadis Liu et Wang sp. n.: a) dorsal habitus; b. lateral habitus; c. frontal habitus; d. tegmen, ventral view; e. tegmen, lateral view; f. penis; g. abdomen, male." figureDoi="http://doi.org/10.5281/zenodo.6876473" httpUri="https://zenodo.org/record/6876473/files/figure.png" pageId="3" pageNumber="78">
Fig.
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).
</paragraph>
<paragraph id="A21636896B26583AFF19FE8EFE71E966" blockId="3.[151,1437,150,537]" pageId="3" pageNumber="78">
Male genitalia (
<figureCitation id="3A922A0C6B26583AFEAAFE8EFE4DE91E" box="[372,485,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1560,1584]" captionTargetBox="[160,1431,570,1528]" captionTargetId="figure-174@3.[151,1435,561,1537]" captionTargetPageId="3" captionText="FIGURE 2. Sulcolotis threadis Liu et Wang sp. n.: a) dorsal habitus; b. lateral habitus; c. frontal habitus; d. tegmen, ventral view; e. tegmen, lateral view; f. penis; g. abdomen, male." figureDoi="http://doi.org/10.5281/zenodo.6876473" httpUri="https://zenodo.org/record/6876473/files/figure.png" pageId="3" pageNumber="78">Figs 2df</figureCitation>
): penis long and slender, apex pointed, penis capsule with inner arm reduced and outer arm well developed; tegmen with penis guide as long as parameres, in lateral view stout, widest at base, gradually narrowing to a pointed apex.
</paragraph>
</subSubSection>
<subSubSection id="EAB365026B26583AFF19FEE2FB75E9D2" pageId="3" pageNumber="78" type="materials_examined">
<paragraph id="A21636896B26583AFF19FEE2FB75E9D2" blockId="3.[151,1437,150,537]" pageId="3" pageNumber="78">
<emphasis id="90DDEA9B6B26583AFF19FEE2FEDDE98B" bold="true" box="[199,373,366,393]" pageId="3" pageNumber="78">Type material.</emphasis>
<materialsCitation id="12C13CD46B26583AFEA5FEE2FB88E98A" ID-GBIF-Occurrence="3864664304" box="[379,1056,366,393]" collectionCode="USNM" country="Philippines" location="Baker" pageId="3" pageNumber="78" specimenCount="1" specimenCount-male="1" stateProvince="Baguio" typeStatus="holotype">
<emphasis id="90DDEA9B6B26583AFEA5FEE2FE40E98A" bold="true" box="[379,488,367,393]" pageId="3" pageNumber="78">
<typeStatus id="7D12882B6B26583AFEA5FEE2FE40E98A" box="[379,488,367,393]" pageId="3" pageNumber="78" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="B4AFFD006B26583AFE29FEE2FDA6E98A" box="[503,526,367,393]" pageId="3" pageNumber="78" type="male"></specimenCount>
,
<collectingCountry id="DABE76196B26583AFDC4FEE2FD33E98A" box="[538,667,367,393]" name="Philippines" pageId="3" pageNumber="78">Philippines</collectingCountry>
,
<collectingRegion id="606DF86B6B26583AFD78FEE2FD52E98A" box="[678,762,367,393]" country="Philippines" name="Baguio" pageId="3" pageNumber="78">Baguio</collectingRegion>
,
<collectingRegion id="606DF86B6B26583AFCDAFEE2FCCCE98A" box="[772,868,367,393]" country="Philippines" name="Benguet" pageId="3" pageNumber="78">Benguet</collectingRegion>
,
<location id="A77660526B26583AFCB1FEE2FC1AE98A" LSID="urn:lsid:plazi:treatment:2A00879F6B27583AFF49FA52FACAEA1A:A77660526B26583AFCB1FEE2FC1AE98A" box="[879,946,367,393]" country="Philippines" name="Baker" pageId="3" pageNumber="78" stateProvince="Baguio">Baker</location>
(
<emphasis id="90DDEA9B6B26583AFC1CFEE3FBBCE98B" bold="true" box="[962,1044,366,392]" pageId="3" pageNumber="78">
<collectionCode id="C4B8AE4C6B26583AFC1CFEE3FBBCE98B" box="[962,1044,366,392]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="78" type="Museum">USNM</collectionCode>
</emphasis>
)
</materialsCitation>
.
<materialsCitation id="12C13CD46B26583AFBF2FEE2FE37E9AE" ID-GBIF-Occurrence="3864664303" collectionCode="USNM, ANIC" country="Philippines" location="Baker" pageId="3" pageNumber="78" specimenCount="6" specimenCount-female="6" stateProvince="Baguio" typeStatus="paratype">
<emphasis id="90DDEA9B6B26583AFBF2FEE2FB0CE98A" bold="true" box="[1068,1188,367,393]" pageId="3" pageNumber="78">
<typeStatus id="7D12882B6B26583AFBF2FEE2FB0CE98A" box="[1068,1188,367,393]" pageId="3" pageNumber="78" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="B4AFFD006B26583AFB6CFEE2FB7EE98A" box="[1202,1238,367,393]" pageId="3" pageNumber="78" type="female">6♀</specimenCount>
, same data as holotype (
<emphasis id="90DDEA9B6B26583AFF30FE1FFE3DE9AF" bold="true" box="[238,405,402,428]" pageId="3" pageNumber="78">
<collectionCode id="C4B8AE4C6B26583AFF30FE1FFEEAE9AF" box="[238,322,402,428]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="3" pageNumber="78" type="Museum">USNM</collectionCode>
,
<collectionCode id="C4B8AE4C6B26583AFE93FE1FFE3DE9AF" box="[333,405,402,428]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="3" pageNumber="78">ANIC</collectionCode>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="12C13CD46B26583AFE72FE1EFACDE9AE" ID-GBIF-Occurrence="3864664307" box="[428,1381,402,429]" collectionCode="MNHB" collectorName="G. Boellcher" country="Philippines" location="Benguet" pageId="3" pageNumber="78" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Benguet" typeStatus="paratype">
<specimenCount id="B4AFFD006B26583AFE72FE1EFE79E9AE" box="[428,465,403,429]" pageId="3" pageNumber="78" type="male">1♂</specimenCount>
<specimenCount id="B4AFFD006B26583AFE0EFE1EFE5CE9AE" box="[464,500,403,429]" pageId="3" pageNumber="78" type="female">4♀</specimenCount>
,
<collectingCountry id="DABE76196B26583AFDDFFE1EFD2AE9AE" box="[513,642,403,429]" name="Philippines" pageId="3" pageNumber="78">Philippines</collectingCountry>
, Luzon
<collectingRegion id="606DF86B6B26583AFD3CFE1EFC97E9AE" box="[738,831,403,429]" country="Philippines" name="Benguet" pageId="3" pageNumber="78">Benguet</collectingRegion>
LaTrinidadv. 1914,
<collectorName id="0F5C535F6B26583AFBFAFE1EFB04E9AE" box="[1060,1196,403,429]" pageId="3" pageNumber="78">G.Boellcher</collectorName>
6500 (
<emphasis id="90DDEA9B6B26583AFADDFE1EFAF2E9AE" bold="true" box="[1283,1370,403,429]" pageId="3" pageNumber="78">
<collectionCode id="C4B8AE4C6B26583AFADDFE1EFAF2E9AE" box="[1283,1370,403,429]" pageId="3" pageNumber="78">MNHB</collectionCode>
</emphasis>
)
</materialsCitation>
;
<materialsCitation id="12C13CD46B26583AFAACFE1EFB70E9D2" ID-GBIF-Occurrence="3864664305" collectionCode="BNHM" collectorName="Mindanao &amp; Dec. Bolm" country="Philippines" elevation="1600" location="Philippines" pageId="3" pageNumber="78" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="B4AFFD006B26583AFAACFE1EFA3DE9AE" box="[1394,1429,403,429]" pageId="3" pageNumber="78" type="female">1♀</specimenCount>
,
<collectingCountry id="DABE76196B26583AFF49FE3AFEBDE9D2" box="[151,277,439,465]" name="Philippines" pageId="3" pageNumber="78">Philippines</collectingCountry>
<quantity id="65519B6C6B26583AFEC2FE3AFEC4E9D2" box="[284,364,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="78" unit="m" value="1600.0">
<elevation id="2984D1BA6B26583AFEC2FE3AFEC4E9D2" box="[284,364,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="78" unit="m" value="1600.0">1600m</elevation>
</quantity>
,
<collectorName id="0F5C535F6B26583AFEA6FE3AFE44E9D2" box="[376,492,439,465]" pageId="3" pageNumber="78">Mindanao</collectorName>
,
<quantity id="65519B6C6B26583AFE26FE3AFD97E9D2" box="[504,575,439,465]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="78" unit="km" value="30.0">30 km</quantity>
W of MARAMAG, 2830.
<collectorName id="0F5C535F6B26583AFCA5FE3AFC5CE9D2" box="[891,1012,439,465]" pageId="3" pageNumber="78">Dec. Bolm</collectorName>
1gt. 1990 (
<emphasis id="90DDEA9B6B26583AFBA8FE3AFB65E9D2" bold="true" box="[1142,1229,439,465]" pageId="3" pageNumber="78">
<collectionCode id="C4B8AE4C6B26583AFBA8FE3AFB65E9D2" box="[1142,1229,439,465]" country="China" httpUri="http://grbio.org/cool/p1gh-na36" name="Beijing Natural History Museum" pageId="3" pageNumber="78">BNHM</collectionCode>
</emphasis>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EAB365026B26583AFF19FE57FD7CE9F6" box="[199,724,474,501]" pageId="3" pageNumber="78" type="distribution">
<paragraph id="A21636896B26583AFF19FE57FD7CE9F6" blockId="3.[151,1437,150,537]" box="[199,724,474,501]" pageId="3" pageNumber="78">
<emphasis id="90DDEA9B6B26583AFF19FE57FEC9E9F7" bold="true" box="[199,353,474,500]" pageId="3" pageNumber="78">Distribution.</emphasis>
<collectingCountry id="DABE76196B26583AFEB9FE56FE4DE9F6" box="[359,485,475,501]" name="Philippines" pageId="3" pageNumber="78">Philippines</collectingCountry>
(Mindanao, Luzon).
</paragraph>
</subSubSection>
<subSubSection id="EAB365026B26583AFF19FE72FACAEA1A" box="[199,1378,511,537]" pageId="3" pageNumber="78" type="etymology">
<paragraph id="A21636896B26583AFF19FE72FACAEA1A" blockId="3.[151,1437,150,537]" box="[199,1378,511,537]" pageId="3" pageNumber="78">
<emphasis id="90DDEA9B6B26583AFF19FE72FEE6EA1A" bold="true" box="[199,334,511,537]" pageId="3" pageNumber="78">Etymology.</emphasis>
The specific name
<taxonomicName id="65A94D0A6B26583AFDF7FE72FD2CEA1A" authorityName="Liu &amp; Wang &amp; Guo &amp; Chen &amp; Wang" authorityYear="2022" box="[553,644,511,537]" class="Insecta" family="Coccinellidae" genus="Sulcolotis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="threadis">
<emphasis id="90DDEA9B6B26583AFDF7FE72FD2CEA1A" box="[553,644,511,537]" italics="true" pageId="3" pageNumber="78">threadis</emphasis>
</taxonomicName>
is adopted as meaning thread, referring to its thin and long penis.
</paragraph>
</subSubSection>
</treatment>
</document>