148 lines
19 KiB
XML
148 lines
19 KiB
XML
<document id="6C1A3F28BD1073824218CD6F2D9EFD38" ID-CLB-Dataset="3199" ID-DOI="10.5252/geodiversitas2018v40a22" ID-GBIF-Dataset="e4383849-346d-4c1a-ab4a-d4f8bfd06d76" ID-ISSN="1638-9395" ID-Zenodo-Dep="5745669" ID-ZooBank="urn:lsid:zoobank.org:pub:61939862-091E-44F6-8811-4A1C4709ACBB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630291425266" checkinUser="marcus" docAuthor="Novo, Nelson M., Tejedor, Marcelo F. & González Ruiz, Laureano R." docDate="2018" docId="AB4887B0FFCAFFA3FF45FE7DFDD0FA9F" docLanguage="en" docName="Geodiversitas.40.22.529-535.pdf.imf" docOrigin="Geodiversitas 40 (22)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Homunculus patagonicus Ameghino 1891" docType="treatment" docVersion="5" lastPageNumber="531" masterDocId="5771FFC8FFCEFFA6FF9BFF8BFFDBFFB1" masterDocTitle="Previously unknown fossil platyrrhines (Primates) of Patagonia from the Tournouër collection at the Muséum national d’Histoire naturelle, Paris" masterLastPageNumber="535" masterPageNumber="529" pageNumber="531" updateTime="1699240401720" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="0334E15DCBA98CCBD283B8F60824D8C2">Previously unknown fossil platyrrhines (Primates) of Patagonia from the Tournouër collection at the Muséum national d’Histoire naturelle, Paris</mods:title>
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<mods:name id="24F4727F616E4B94BEFF47535AE60730" type="personal">
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<mods:roleTerm id="38B47C4539D39657B6E01870BE41A28B">Author</mods:roleTerm>
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<mods:namePart id="FA23BE8E6393C5EDEDFE7724E42AD60F">Novo, Nelson M.</mods:namePart>
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<mods:namePart id="AC1B169ACC131EA44DD9EB115CFB6333">Tejedor, Marcelo F.</mods:namePart>
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<mods:name id="DA5F55863621523AC79A7CF03132A214" type="personal">
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<mods:namePart id="602DD2F95DD3988242755BBDF39897A3">González Ruiz, Laureano R.</mods:namePart>
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<mods:title id="CD85D7DA03D6BB6017A875CC69E030ED">Geodiversitas</mods:title>
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<mods:part id="985EE2E8077D8F7A707438D1300B4BC5">
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<mods:date id="59D2686F49AEEC22550C7832501BBF5B">2018</mods:date>
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<mods:number id="6B2C45BF8609818C5EA3CCC0E43FD77E">2018-11-08</mods:number>
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<mods:number id="0A9F4E1BB24F00EBF84184611C11ADE2">40</mods:number>
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<treatment id="AB4887B0FFCAFFA3FF45FE7DFDD0FA9F" ID-DOI="http://doi.org/10.5281/zenodo.5745700" ID-GBIF-Taxon="190874952" ID-Zenodo-Dep="5745700" LSID="urn:lsid:plazi:treatment:AB4887B0FFCAFFA3FF45FE7DFDD0FA9F" httpUri="http://treatment.plazi.org/id/AB4887B0FFCAFFA3FF45FE7DFDD0FA9F" lastPageId="5" lastPageNumber="531" pageId="4" pageNumber="531">
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<subSubSection id="6BFB652DFFCAFFA2FF45FE7DFD78FD27" pageId="4" pageNumber="531" type="nomenclature">
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<paragraph id="235E36A6FFCAFFA2FF45FE7DFD76FDA1" blockId="4.[222,685,502,529]" box="[222,685,502,529]" pageId="4" pageNumber="531">
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<heading id="781681CAFFCAFFA2FF45FE7DFD76FDA1" box="[222,685,502,529]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="531" reason="2">
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<taxonomicName id="E4E14D25FFCAFFA2FF45FE7DFD76FDA1" authority="Ameghino, 1891" authorityName="Ameghino" authorityYear="1891" box="[222,685,502,529]" class="Mammalia" family="Atelidae" genus="Homunculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="4" pageNumber="531" phylum="Chordata" rank="species" species="patagonicus">
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<emphasis id="1195EAB4FFCAFFA2FF45FE7DFE34FDA1" bold="true" box="[222,495,502,529]" italics="true" pageId="4" pageNumber="531">Homunculus patagonicus</emphasis>
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<bibRefCitation id="47704B57FFCAFFA2FE6DFE7DFD76FDA1" author="AMEGHINO F." box="[502,685,502,528]" pageId="4" pageNumber="531" pagination="383 - 397" refId="ref4512" refString="AMEGHINO F. 1891. - Los monos fosiles del Eoceno de la Republica Argentina. Revista Argentina de Historia Natural 1: 383 - 397." type="journal article" year="1891">Ameghino, 1891</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="235E36A6FFCAFFA2FF1FFDBBFD78FD27" blockId="4.[132,776,559,663]" pageId="4" pageNumber="531">
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<typeStatus id="FC5A8804FFCAFFA2FF1FFDBBFF04FDF7" box="[132,223,560,583]" pageId="4" pageNumber="531" type="neotype">NEOTYPE</typeStatus>
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. — MACN-A 5757; partial mandible preserving a complete and worn dentition lacking crowns of the left canine and the incisors. This is a replacement (see
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<bibRefCitation id="47704B57FFCAFFA2FD82FDEFFF63FD26" author="TEJEDOR M. F. & ROSENBERGER A. L." pageId="4" pageNumber="531" pagination="68 - 82" refId="ref6497" refString="TEJEDOR M. F. & ROSENBERGER A. L. 2008. - A neotype for Homunculus patagonicus Ameghino, 1891, and a new interpretation of the taxon. Paleoanthropology 2008: 68 - 82." type="journal article" year="2008">Tejedor & Rosenberger 2008</bibRefCitation>
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) for the lost
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<typeStatus id="FC5A8804FFCAFFA2FEA6FDF5FE4EFD27" box="[317,405,638,662]" pageId="4" pageNumber="531" type="holotype">holotype</typeStatus>
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specimen, MACN-A 634.
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</paragraph>
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</subSubSection>
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<subSubSection id="6BFB652DFFCAFFA3FF1EFD3BFDD0FA9F" lastPageId="5" lastPageNumber="532" pageId="4" pageNumber="531" type="description">
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<paragraph id="235E36A6FFCAFFA2FF1EFD3BFE57FCFA" blockId="4.[131,775,686,843]" pageId="4" pageNumber="531">
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GEOGRAPHIC AND STRATIGRAPHIC PROVENANCE. — The
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<typeStatus id="FC5A8804FFCAFFA2FD2CFD24FCDDFD76" box="[695,774,687,711]" pageId="4" pageNumber="531" type="neotype">neotype</typeStatus>
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MACN-A 5757 comes from Corriguen Aike (= Puesto Estancia La Costa), on the Atlantic coast of
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<collectingRegion id="E125F844FFCAFFA2FE79FD68FD75FD4A" box="[482,686,739,763]" country="Argentina" name="Santa Cruz" pageId="4" pageNumber="531">Santa Cruz Province</collectingRegion>
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between the rivers Coyle and Gallegos,
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<collectingCountry id="5BF67636FFCAFFA2FE29FD76FDC2FCA4" box="[434,537,765,789]" name="Argentina" pageId="4" pageNumber="531">Argentina</collectingCountry>
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.
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<collectingRegion id="E125F844FFCAFFA2FDBAFD75FD4BFCA7" box="[545,656,766,790]" country="Argentina" name="Santa Cruz" pageId="4" pageNumber="531">Santa Cruz</collectingRegion>
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Formation, late-early Miocene (Burdigalian Age),
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<emphasis id="1195EAB4FFCAFFA2FE66FC92FDD7FC81" box="[509,524,793,816]" italics="true" pageId="4" pageNumber="531">c.</emphasis>
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17.5 to 16.7 Ma (
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<bibRefCitation id="47704B57FFCAFFA2FD5AFC93FF63FCFA" author="TAUBER A. A." pageId="4" pageNumber="531" pagination="413 - 426" refId="ref6345" refString="TAUBER A. A. 1997. - Bioestratigrafia de la Formacion Santa Cruz (Mioceno Inferior) en el extremo sudeste de la Patagonia. Ameghiniana 34 (4): 413 - 426." type="journal article" year="1997">Tauber 1997</bibRefCitation>
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;
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<bibRefCitation id="47704B57FFCAFFA2FF5DFCB9FEA5FCFA" author="FLEAGLE J. G. & PERKINS M. E. & HEIZLER M. T. & NASH B. & BOWN T. M. & TAUBER A. A. & DOZO M. T. & TEJEDOR M. F." box="[198,382,818,843]" pageId="4" pageNumber="531" pagination="41 - 58" refId="ref4810" refString="FLEAGLE J. G., PERKINS M. E., HEIZLER M. T., NASH B., BOWN T. M., TAUBER A. A., DOZO M. T. & TEJEDOR M. F. 2012. - Absolute and relative ages of fossil localities in the Santa Cruz and Pinturas Formations, in VIZCAINO S. F., KAY R. F. & BARGO M. S. (eds), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation. Cambridge University Press, Cambridge: 41 - 58. https: // doi. org / 10.1017 / CBO 9780511667381.004" type="book chapter" year="2012">
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Fleagle
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<emphasis id="1195EAB4FFCAFFA2FE88FCB8FE98FCFB" box="[275,323,818,842]" italics="true" pageId="4" pageNumber="531">et al.</emphasis>
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2012
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</bibRefCitation>
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).
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</paragraph>
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<paragraph id="235E36A6FFCAFFA2FF1FFCE8FEA5FC78" blockId="4.[132,776,866,969]" pageId="4" pageNumber="531">NEW REFERRED MATERIAL. — MNHN.F.SCZ215 (Fig. 2A-C), partial mandible preserving the left ramus with alveoli for left i1-2, c, p2, broken alveoli of p3-4, and complete m1; alveoli for right i1-2, and broken c.</paragraph>
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<paragraph id="235E36A6FFCAFFA2FF1EFC69FD70FBCC" blockId="4.[130,776,993,1149]" pageId="4" pageNumber="531">
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GEOGRAPHIC AND STRATIGRAPHIC PROVENANCE. —MNHN.F.SCZ215 is labeled as “
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<taxonomicName id="E4E14D25FFCAFFA2FF65FC70FEA9FBA2" authorityName="Ameghino" authorityYear="1891" box="[254,370,1019,1043]" class="Mammalia" family="Atelidae" genus="Homunculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="4" pageNumber="531" phylum="Chordata" rank="genus">
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<emphasis id="1195EAB4FFCAFFA2FF65FC70FEA9FBA2" box="[254,370,1019,1043]" italics="true" pageId="4" pageNumber="531">Homunculus</emphasis>
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</taxonomicName>
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// Coyle 1902-6” (Fig. 2E). Thus, the provenance is certainly nearby the Coyle river on the Atlantic coast of
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<collectingRegion id="E125F844FFCAFFA2FD4FFB9DFED0FBF9" country="Argentina" name="Santa Cruz" pageId="4" pageNumber="531">Santa Cruz Province</collectingRegion>
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,
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<collectingCountry id="5BF67636FFCAFFA2FE88FBBBFEAEFBF9" box="[275,373,1072,1096]" name="Argentina" pageId="4" pageNumber="531">Argentina</collectingCountry>
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.
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<collectingRegion id="E125F844FFCAFFA2FEE0FBBAFE3FFBF8" box="[379,484,1073,1097]" country="Argentina" name="Santa Cruz" pageId="4" pageNumber="531">Santa Cruz</collectingRegion>
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Formation, late-early Miocene (Burdigalian Age) (see
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<bibRefCitation id="47704B57FFCAFFA2FECFFBC0FE06FBD2" author="MARSHALL L. G." box="[340,477,1099,1123]" pageId="4" pageNumber="531" pagination="1129 - 1142" refId="ref5668" refString="MARSHALL L. G. 1976. - Fossil Localities for Santacrucian (Early Miocene) Mammals, Santa Cruz Province, Southern Patagonia, Argentina. Journal of Paleontology 50 (6): 1129 - 1142. https: // www. jstor. org / stable / 1303553" type="journal article" year="1976">Marshall 1976</bibRefCitation>
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;
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<bibRefCitation id="47704B57FFCAFFA2FE73FBC0FDBBFBD2" author="TAUBER A. A." box="[488,608,1099,1123]" pageId="4" pageNumber="531" pagination="413 - 426" refId="ref6345" refString="TAUBER A. A. 1997. - Bioestratigrafia de la Formacion Santa Cruz (Mioceno Inferior) en el extremo sudeste de la Patagonia. Ameghiniana 34 (4): 413 - 426." type="journal article" year="1997">Tauber 1997</bibRefCitation>
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;
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<bibRefCitation id="47704B57FFCAFFA2FDF0FBC0FD23FBD2" author="BUFFETAUT E." box="[619,760,1099,1123]" pageId="4" pageNumber="531" pagination="207 - 214" refId="ref4708" refString="BUFFETAUT E. 2014. - Tertiary ground birds from Patagonia (Argentina) in the Tournouer collection of the Museum national d'Histoire naturelle, Paris. Bulletin de la Societe geologique de France 185 (3): 207 - 214. https: // doi. org / 10.2113 / gssgfbull. 185.3.207" type="journal article" year="2014">Buffetaut 2014</bibRefCitation>
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), approximately 16.1-16.5 Ma (
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<bibRefCitation id="47704B57FFCAFFA2FE3EFBEEFD8AFBCC" author="FLEAGLE J. G. & PERKINS M. E. & HEIZLER M. T. & NASH B. & BOWN T. M. & TAUBER A. A. & DOZO M. T. & TEJEDOR M. F." box="[421,593,1125,1149]" pageId="4" pageNumber="531" pagination="41 - 58" refId="ref4810" refString="FLEAGLE J. G., PERKINS M. E., HEIZLER M. T., NASH B., BOWN T. M., TAUBER A. A., DOZO M. T. & TEJEDOR M. F. 2012. - Absolute and relative ages of fossil localities in the Santa Cruz and Pinturas Formations, in VIZCAINO S. F., KAY R. F. & BARGO M. S. (eds), Early Miocene Paleobiology in Patagonia: High-Latitude Paleocommunities of the Santa Cruz Formation. Cambridge University Press, Cambridge: 41 - 58. https: // doi. org / 10.1017 / CBO 9780511667381.004" type="book chapter" year="2012">
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Fleagle
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<emphasis id="1195EAB4FFCAFFA2FE76FBEDFDC0FBCC" box="[493,539,1125,1149]" italics="true" pageId="4" pageNumber="531">et al.</emphasis>
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2012
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</bibRefCitation>
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) (
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<figureCitation id="BBDA2A23FFCAFFA2FDFCFBEEFD46FBCC" box="[615,669,1125,1149]" captionStart="FIG" captionStartId="4.[813,824,1849,1867]" captionTargetBox="[828,1427,221,1754]" captionTargetId="figure-546@4.[813,1456,215,1776]" captionTargetPageId="4" captionText="FIG. 1. — MNHN.F.SCZ215, partial mandible of Homunculus patagonicus Ameghino, 1891 with left m1 in lateral left (A), lateral right (B) and dorsal (C) views; D, m2 of MNHN.F.SCZ215; E, MNHN.F.SCZ215 identification label from MNHN.Abbreviations:alv i1, first incisor alveolus;alv i2, second incisor alveolus; alv c, canine alveolus; alv pm2, second premolar alveolus; Man Sym, Mandibular symphysis; Hypd, Hypoconid; Prd, Protoconid; Trid, Trigonid basin; Med, Metaconid; Entd, Entoconido; Tald, Talonid basin. Orientation abbreviations: D, Distal; La, Labial; M, Mesial; Li, Lingual. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5745671" httpUri="https://zenodo.org/record/5745671/files/figure.png" pageId="4" pageNumber="531">Fig. 1</figureCitation>
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).
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</paragraph>
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<paragraph id="235E36A6FFCAFFA2FF1FFB3FFEC1FB7D" blockId="4.[131,776,1204,2028]" box="[132,282,1204,1230]" pageId="4" pageNumber="531">DESCRIPTION</paragraph>
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<paragraph id="235E36A6FFCAFFA2FF1FFB5FFE30F9FC" blockId="4.[131,776,1204,2028]" pageId="4" pageNumber="531">
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Based on comparisons, the mandible MNHN.F.SCZ215 is indistinguishable from the cast of the lost MACN-A 634, the type of
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<taxonomicName id="E4E14D25FFCAFFA2FEDAFA98FD89FA9C" authorityName="Ameghino" authorityYear="1891" box="[321,594,1299,1325]" class="Mammalia" family="Atelidae" genus="Homunculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="4" pageNumber="531" phylum="Chordata" rank="species" species="patagonicus">
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<emphasis id="1195EAB4FFCAFFA2FEDAFA98FD89FA9C" box="[321,594,1299,1325]" italics="true" pageId="4" pageNumber="531">Homunculus patagonicus</emphasis>
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</taxonomicName>
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. MACN-A 634 is strongly worn but shows several morphological similarities at the alveolar level as well as the crown outline and size of m1. The size and orientation of the alveoli indicate that the specimen had small roots and procumbent, staggered incisors, producing a V-shaped mandible. The small alveolus for the left canine, similar in size to the alveolus of p2, indicates that the canine was small, as seen in the
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<typeStatus id="FC5A8804FFCAFFA2FF1EF998FF39F99C" box="[133,226,1555,1581]" pageId="4" pageNumber="531" type="neotype">neotype</typeStatus>
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of
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<taxonomicName id="E4E14D25FFCAFFA2FE95F999FDF7F99D" authorityName="Ameghino" authorityYear="1891" box="[270,556,1554,1580]" class="Mammalia" family="Atelidae" genus="Homunculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="4" pageNumber="531" phylum="Chordata" rank="species" species="patagonicus">
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<emphasis id="1195EAB4FFCAFFA2FE95F999FDF7F99D" box="[270,556,1554,1580]" italics="true" pageId="4" pageNumber="531">Homunculus patagonicus</emphasis>
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</taxonomicName>
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(MACN-A 5757) (
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<bibRefCitation id="47704B57FFCAFFA2FF15F9B9FE01F9FC" author="TEJEDOR M. F. & ROSENBERGER A. L." box="[142,474,1586,1613]" pageId="4" pageNumber="531" pagination="68 - 82" refId="ref6497" refString="TEJEDOR M. F. & ROSENBERGER A. L. 2008. - A neotype for Homunculus patagonicus Ameghino, 1891, and a new interpretation of the taxon. Paleoanthropology 2008: 68 - 82." type="journal article" year="2008">Tejedor & Rosenberger 2008</bibRefCitation>
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).
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</paragraph>
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<paragraph id="235E36A6FFCAFFA3FF00F9D9FDD0FA9F" blockId="4.[131,776,1204,2028]" lastBlockId="5.[132,776,1140,1326]" lastPageId="5" lastPageNumber="532" pageId="4" pageNumber="531">
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Compared to MACN-A 635, this specimen has slightly larger alveoli for the canines and a somewhat more robust mandible. Remarkably, the m1 of this new specimen is one of the best preserved lower molars attributed to
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<taxonomicName id="E4E14D25FFCAFFA2FD31F938FF61F95D" authorityName="Ameghino" authorityYear="1891" class="Mammalia" family="Atelidae" genus="Homunculus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Primates" pageId="4" pageNumber="531" phylum="Chordata" rank="genus">
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<emphasis id="1195EAB4FFCAFFA2FD31F938FF61F95D" italics="true" pageId="4" pageNumber="531">Homunculus</emphasis>
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</taxonomicName>
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. Its morphology resembles MACN-A 5969b. It has two roots and the trigonid is higher than the talonid, with moderately high and sharp crests. The preprotocristid extends forward closing the trigonid mesially; the paraconid is absent, and there is an oblique cristid connecting the preparacristid. The metaconid is the highest cusp, and is situated relative to the protoconid in a more distal position. The hypoflexid appears relatively deep. It shows a tiny cuspule as seen in the m1 of MACN-A 5969b. The molar has a long cristid obliqua reaching the trigonid wall lingual to the protoconid, while the hypoconid is displaced labially. The entoconid is lower than the hypoconid, with a long entocristid. The talonid basin is relatively wide and deep, showing an incipient and centrally placed hypoconulid (Fig. 2).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |