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<document id="B0845D278218583B7964CFCC3ABE4E73" ID-DOI="10.11646/zootaxa.5040.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5531247" ID-ZooBank="B10750F1-0CD9-4A61-B071-29989FB307B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632747454806" checkinUser="plazi" docAuthor="Alvarado, Mabel &amp; Palacio, Edgard" docDate="2021" docId="3A23BA1CFFF42E2B2697FD92FE6FFE1A" docLanguage="en" docName="zootaxa.5040.2.6.pdf" docOrigin="Zootaxa 5040 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Forrestopius nasutus Alvarado &amp; Palacio 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="273" masterDocId="C61AC264FFF32E232600FD05FFA8FFEF" masterDocTitle="Forrestopius Gauld &amp; Sithole, 2002 (Hymenoptera: Ichneumonidae: Metopiinae) in South America" masterLastPageNumber="282" masterPageNumber="265" pageNumber="272" updateTime="1699415403863" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="FCF19F52ACC18D8F62E7936789058EAE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AD93C54F19CD785C0A0EEA6CB11D1723">
<mods:title id="16BF780B4485491000D295599B884BC9">Forrestopius Gauld &amp; Sithole, 2002 (Hymenoptera: Ichneumonidae: Metopiinae) in South America</mods:title>
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<mods:name id="A4FA6396FB95A25C97911B5884D792FC" type="personal">
<mods:role id="3A6F38C066CCF07F958AA59DDE165723">
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<mods:namePart id="EE82BF3979B4F92F9CE65130716F5C17">Alvarado, Mabel</mods:namePart>
<mods:affiliation id="C7E3DD301E09AC33BC0C95B11A95434F">Departamento de Entomología, Museo de Historia Natural, Universidad Nacional Mayor de San Marcos. Av. Arenales 1256 Jesús María, Lima 14, Perú &amp; Bosque Llaqta, Av. Confraternidad Internacional Este N ° 364, Huaraz, Perú.</mods:affiliation>
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<mods:namePart id="5E710013876A28094C2CF92F78231B23">Palacio, Edgard</mods:namePart>
<mods:affiliation id="D1FF32141F10E58CF12DE1109B3F8EF1">Instituto Colombiano Agropecuario, Calle 18 Nº 50 - 32 Soledad, Barranquilla, Colombia &amp; Laboratorio de Entomología, Departamento de Biología, Pontificia Universidad Javeriana, Bogotá, Colombia.</mods:affiliation>
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<mods:title id="778EDE11CB427D96725760930F69E56A">Zootaxa</mods:title>
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<mods:date id="24EC44075F27BCB2E39F2A92793F1A90">2021</mods:date>
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<mods:number id="D0E0C95B11912748070B9E19A44187B9">2021-09-21</mods:number>
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<mods:detail id="70EBCB541974882AF915C307AA3FD6E4" type="volume">
<mods:number id="E78068E20960649DA4AAF442FEE84970">5040</mods:number>
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<mods:number id="0FD7E98E7B1F717F0A22EF346CDB5D46">2</mods:number>
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<mods:classification id="42B8ECBC2E10AB1B0EE6FB2F934E9F67">journal article</mods:classification>
<mods:identifier id="6FD011591ABC6BA849A4D433C95A953A" type="DOI">10.11646/zootaxa.5040.2.6</mods:identifier>
<mods:identifier id="8588452996BDF078A72FCB3BA3809A40" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="12FCECE2325D4DD7576CAD7DC89D907E" type="Zenodo-Dep">5531247</mods:identifier>
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<treatment id="3A23BA1CFFF42E2B2697FD92FE6FFE1A" ID-DOI="http://doi.org/10.5281/zenodo.5531301" ID-Zenodo-Dep="5531301" LSID="urn:lsid:plazi:treatment:3A23BA1CFFF42E2B2697FD92FE6FFE1A" httpUri="http://treatment.plazi.org/id/3A23BA1CFFF42E2B2697FD92FE6FFE1A" lastPageId="8" lastPageNumber="273" pageId="7" pageNumber="272">
<subSubSection id="FA905881FFF42E242697FD92FDF7FF5E" box="[151,607,151,177]" pageId="7" pageNumber="272" type="nomenclature">
<paragraph id="B2350B0AFFF42E242697FD92FDF7FF5E" blockId="7.[151,607,151,213]" box="[151,607,151,177]" pageId="7" pageNumber="272">
<heading id="E97DBC66FFF42E242697FD92FDF7FF5E" bold="true" box="[151,607,151,177]" fontSize="11" level="1" pageId="7" pageNumber="272" reason="1">
<emphasis id="80FED718FFF42E242697FD92FDF7FF5E" bold="true" box="[151,607,151,177]" pageId="7" pageNumber="272">
<taxonomicName id="758A7089FFF42E242697FD92FE52FF5E" authority="Palacio" authorityName="Alvarado &amp; Palacio" authorityYear="2021" box="[151,506,151,177]" class="Insecta" family="Ichneumonidae" genus="Forrestopius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="272" phylum="Arthropoda" rank="species" species="nasutus" status="sp. nov.">
<emphasis id="80FED718FFF42E242697FD92FE30FF5E" bold="true" box="[151,408,151,177]" italics="true" pageId="7" pageNumber="272">Forrestopius nasutus</emphasis>
Palacio
</taxonomicName>
<taxonomicNameLabel id="9BCD6A63FFF42E242402FD92FDF7FF5E" box="[514,607,151,177]" pageId="7" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FA905881FFF42E242697FDBEFE83FF3A" box="[151,299,187,213]" pageId="7" pageNumber="272" type="description">
<paragraph id="B2350B0AFFF42E242697FDBEFE83FF3A" blockId="7.[151,607,151,213]" box="[151,299,187,213]" pageId="7" pageNumber="272">
(
<figureCitation id="2AB1178FFFF42E24269FFDBEFF40FF3A" box="[159,232,187,213]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272">Figs 4</figureCitation>
,
<figureCitation id="2AB1178FFFF42E2426F4FDBEFE8CFF3A" box="[244,292,187,213]" captionStart="FIGURE 10" captionStartId="16.[151,250,881,905]" captionTargetBox="[151,1436,181,857]" captionTargetId="figure-23@16.[151,1436,181,858]" captionTargetPageId="16" captionText="FIGURE 10. Geographic distribution of the Forrestopius species" figureDoi="http://doi.org/10.5281/zenodo.5531269" httpUri="https://zenodo.org/record/5531269/files/figure.png" pageId="7" pageNumber="272">10C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FA905881FFF42E242697FC06FE66FEAE" pageId="7" pageNumber="272" type="materials_examined">
<paragraph id="B2350B0AFFF42E242697FC06FE66FEAE" blockId="7.[151,1437,259,1186]" pageId="7" pageNumber="272">
<materialsCitation id="02E20157FFF42E242697FC06FE66FEAE" collectingDate="2001-05-04" collectingDateMax="2001-05-06" collectingDateMin="2001-05-04" collectionCode="T, MPUJ, ENT" country="Colombia" elevation="2800" latitude="5.4166665" location="Boyaca" longLatPrecision="1305" longitude="-73.45" pageId="7" pageNumber="272" specimenCount="1" specimenCount-female="1" stateProvince="Boyaca" typeStatus="holotype">
<emphasis id="80FED718FFF42E242697FC06FEA3FEF2" bold="true" box="[151,267,259,285]" pageId="7" pageNumber="272">
<typeStatus id="6D31B5A8FFF42E242697FC06FEAEFEF2" box="[151,262,259,285]" pageId="7" pageNumber="272" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="A48CC083FFF42E242711FC01FE8EFEF1" box="[273,294,260,286]" pageId="7" pageNumber="272" type="female">♀</specimenCount>
, “
<collectingCountry id="CA9D4B9AFFF42E24273DFC06FE72FEF2" box="[317,474,259,285]" name="Colombia" pageId="7" pageNumber="272">COLOMBIA</collectingCountry>
:
<collectingRegion id="704EC5E8FFF42E2427E3FC06FD92FEF2" box="[483,570,259,285]" country="Colombia" name="Boyaca" pageId="7" pageNumber="272">Boyacá</collectingRegion>
, Arcabuco, finca El Edén,
<geoCoordinate id="D7BE6DCDFFF42E242561FC06FC1CFEF1" box="[865,948,259,286]" degrees="5" direction="north" minutes="25" orientation="latitude" pageId="7" pageNumber="272" precision="925" value="5.4166665">5°25’N</geoCoordinate>
<geoCoordinate id="D7BE6DCDFFF42E2425BAFC06FB89FEF2" box="[954,1057,259,285]" degrees="73" direction="west" minutes="27" orientation="longitude" pageId="7" pageNumber="272" precision="925" value="-73.45">73°27’W</geoCoordinate>
,
<date id="C6342DCAFFF42E24222BFC06FB0DFEF2" box="[1067,1189,259,285]" pageId="7" pageNumber="272" value="2001-05-04" valueMax="2001-05-06" valueMin="2001-05-04">
<collectingDate id="D670D422FFF42E24222BFC06FB0DFEF2" box="[1067,1189,259,285]" pageId="7" pageNumber="272" value="2001-05-04" valueMax="2001-05-06" valueMin="2001-05-04">4-6.v.2001</collectingDate>
</date>
,
<quantity id="7572A6EFFFF42E2422B0FC06FAA9FEF2" box="[1200,1281,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="272" unit="m" value="2800.0">
<elevation id="39A7EC39FFF42E2422B0FC06FAA9FEF2" box="[1200,1281,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="272" unit="m" value="2800.0">2800m</elevation>
</quantity>
,
<collectingMethod id="6BCB731DFFF42E24230BFC06FA30FEF1" box="[1291,1432,259,286]" pageId="7" pageNumber="272">Malaise trap</collectingMethod>
,
<collectionCode id="D49B93CFFFF42E242697FC2DFF01FEAD" box="[151,169,296,322]" name="Tavera, Department of Geology and Geophysics" pageId="7" pageNumber="272">T</collectionCode>
. Angarita” (
<collectionCode id="D49B93CFFFF42E242730FC22FED3FEAE" box="[304,379,295,321]" pageId="7" pageNumber="272">MPUJ</collectionCode>
_
<collectionCode id="D49B93CFFFF42E24278AFC22FE6BFEAE" box="[394,451,295,321]" country="Uganda" lsid="urn:lsid:biocol.org:col:13589" name="Ministry of Natural Resources" pageId="7" pageNumber="272" type="Herbarium">ENT</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="FA905881FFF42E2426C7FC4EFB95FDD2" pageId="7" pageNumber="272" type="diagnosis">
<paragraph id="B2350B0AFFF42E2426C7FC4EFB95FDD2" blockId="7.[151,1437,259,1186]" pageId="7" pageNumber="272">
<emphasis id="80FED718FFF42E2426C7FC4EFEEAFE8A" bold="true" box="[199,322,331,357]" pageId="7" pageNumber="272">Diagnosis.</emphasis>
This species can be recognized by the following combination of characters: mandibles with three teeth (
<figureCitation id="2AB1178FFFF42E2426DEFC6AFE9EFE66" box="[222,310,367,393]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272">Fig. 4E</figureCitation>
), labrum exposed when the mandibles are closed, clypeus strongly swollen with a nose-like prominence (
<figureCitation id="2AB1178FFFF42E2426EAFC96FEECFE42" box="[234,324,403,429]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272">Fig. 4B</figureCitation>
), with lower part concave (
<figureCitation id="2AB1178FFFF42E24247FFC96FD71FE42" box="[639,729,403,429]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272">Fig. 4C</figureCitation>
), face transversely striate (Fig, 4C), antenna with 30 flagellomeres, pronotum with a short wrinkle arising from pronotal pit reaching less than one third of way to anterior margin, body with intense blue to purple reflections. This species looks similar to
<taxonomicName id="758A7089FFF42E242206FCD9FBC5FE1A" authorityName="Gauld &amp; Sithole" authorityYear="2002" box="[1030,1133,475,501]" class="Insecta" family="Ichneumonidae" genus="Forrestopius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="272" phylum="Arthropoda" rank="species" species="judyae">
<emphasis id="80FED718FFF42E242206FCD9FBC5FE1A" box="[1030,1133,475,501]" italics="true" pageId="7" pageNumber="272">F. judyae</emphasis>
</taxonomicName>
, both have mandibles with three teeth, head subquadrate behind ocellar triangle, vertex long and almost flat, and occiput abruptly declivous.
<taxonomicName id="758A7089FFF42E242388FF05FF45FDD2" authorityName="Alvarado &amp; Palacio" authorityYear="2021" class="Insecta" family="Ichneumonidae" genus="Forrestopius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="272" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="80FED718FFF42E242388FF05FF45FDD2" italics="true" pageId="7" pageNumber="272">F. nasutus</emphasis>
</taxonomicName>
can be differentiated by having the clypeus nasute and the exposed labrum.
</paragraph>
</subSubSection>
<subSubSection id="FA905881FFF42E2B26C7FF42FE28FE66" lastPageId="8" lastPageNumber="273" pageId="7" pageNumber="272" type="description">
<paragraph id="B2350B0AFFF42E2426C7FF42FC4EFD8E" blockId="7.[151,1437,259,1186]" box="[199,998,583,610]" pageId="7" pageNumber="272">
<emphasis id="80FED718FFF42E2426C7FF42FE68FD8E" bold="true" box="[199,448,583,609]" pageId="7" pageNumber="272">Description. Female:</emphasis>
Body length
<quantity id="7572A6EFFFF42E242458FF42FD17FD8D" box="[600,703,583,610]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="7" pageNumber="272" unit="mm" value="13.9">13.9 mm</quantity>
. Fore wing length 9.0mm.
</paragraph>
<paragraph id="B2350B0AFFF42E2426C7FF6EFC2BFC6E" blockId="7.[151,1437,259,1186]" pageId="7" pageNumber="272">
<emphasis id="80FED718FFF42E2426C7FF6EFEA3FD6A" box="[199,267,619,645]" italics="true" pageId="7" pageNumber="272">Head.</emphasis>
Face and clypeus 0.7× as long as wide, face transversely striate (
<figureCitation id="2AB1178FFFF42E2425F6FF6EFBE7FD6A" box="[1014,1103,619,645]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272">Fig. 4C</figureCitation>
), centrally granulate, clypeus strongly swollen with a nose-like prominence, clearly differentiated from face (
<figureCitation id="2AB1178FFFF42E24220AFF8AFBCAFD46" box="[1034,1122,655,681]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272">Fig. 4B</figureCitation>
), sparsely punctate dorsally, ventrally concave and granulate with transverse wrinkles centrally; labrum exposed when mandibles are closed; mandibles stout, with three teeth (
<figureCitation id="2AB1178FFFF42E24240EFFD2FDCDFD1E" box="[526,613,727,753]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272">Fig. 4E</figureCitation>
); malar space 0.4× as long as basal mandibular width; lateral ocellus separated from compound eye by about 2.0× maximum ocellar diameter; distance between ocelli 1.0× maximum ocellar diameter; vertex behind ocellar triangle almost flat, occiput abruptly declivous far back; gena in lateral view about 2.2× as long as compound eye; antenna with 30 flagellomeres, ratio of length from second to fourth flagellomeres: 1.0:0.9:0.8, subapical flagellomere 0.8× as long as centrally broad.
</paragraph>
<paragraph id="B2350B0AFFF42E2426C7FE89FD4CFB4D" blockId="7.[151,1437,259,1186]" pageId="7" pageNumber="272">
<taxonomicName id="758A7089FFF42E2426C7FE89FE97FC4A" box="[199,319,908,933]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="272" phylum="Arthropoda" rank="genus">
<emphasis id="80FED718FFF42E2426C7FE89FE97FC4A" box="[199,319,908,933]" italics="true" pageId="7" pageNumber="272">Mesosoma</emphasis>
</taxonomicName>
. Pronotum with a short wrinkle arising from pronotal pit reaching less than one third of way to anterior margin (
<figureCitation id="2AB1178FFFF42E24273AFEAAFE3AFC26" box="[314,402,943,969]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272">Fig. 4B</figureCitation>
); metapleuron smooth and polished with sparse setae over its entire surface; submetapleural carina smooth, anteriorly expanded into a punctate triangular lobe. Propodeum with lateromedian longitudinal carina strong, slightly closer to each other anteriorly (
<figureCitation id="2AB1178FFFF42E2424D1FEF2FC83FBFE" box="[721,811,1015,1041]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272">Fig. 4D</figureCitation>
); area basalis+superomedia 3.6× as long as wide; lateral longitudinal carina complete; posterior transverse carina complete, enclosing area petiolaris and area coxalis. Fore wing with vein
<emphasis id="80FED718FFF42E242746F945FECFFBB5" box="[326,359,1088,1114]" italics="true" pageId="7" pageNumber="272">Cu</emphasis>
1
<emphasis id="80FED718FFF42E242775F945FE2BFBB6" box="[373,387,1088,1113]" italics="true" pageId="7" pageNumber="272">a</emphasis>
between
<emphasis id="80FED718FFF42E2427EEF945FDA7FBB5" box="[494,527,1088,1114]" italics="true" pageId="7" pageNumber="272">Cu</emphasis>
1
<emphasis id="80FED718FFF42E24241CF93AFD82FBB6" box="[540,554,1087,1113]" italics="true" pageId="7" pageNumber="272">b</emphasis>
and 2
<emphasis id="80FED718FFF42E24246FF945FD0FFBB6" box="[623,679,1088,1113]" italics="true" pageId="7" pageNumber="272">m-cu</emphasis>
1.0× as long as
<emphasis id="80FED718FFF42E24255CF945FCD5FBB5" box="[860,893,1088,1114]" italics="true" pageId="7" pageNumber="272">Cu</emphasis>
1 between
<emphasis id="80FED718FFF42E2425F6F945FBBAFBB6" box="[1014,1042,1088,1113]" italics="true" pageId="7" pageNumber="272">Rs</emphasis>
&amp;
<emphasis id="80FED718FFF42E242228F945FB97FBB6" box="[1064,1087,1088,1113]" italics="true" pageId="7" pageNumber="272">M</emphasis>
and 1
<emphasis id="80FED718FFF42E242284F945FB14FBB6" box="[1156,1212,1088,1113]" italics="true" pageId="7" pageNumber="272">m-cu</emphasis>
; 2
<emphasis id="80FED718FFF42E2422D8F945FAA3FBB6" box="[1240,1291,1088,1113]" italics="true" pageId="7" pageNumber="272">rs-m</emphasis>
0.8× as long as abscissa of
<emphasis id="80FED718FFF42E242736F961FEE5FB92" box="[310,333,1124,1149]" italics="true" pageId="7" pageNumber="272">M</emphasis>
between 2
<emphasis id="80FED718FFF42E2427C7F961FE52FB92" box="[455,506,1124,1149]" italics="true" pageId="7" pageNumber="272">rs-m</emphasis>
and 2
<emphasis id="80FED718FFF42E24243EF961FDDEFB92" box="[574,630,1124,1149]" italics="true" pageId="7" pageNumber="272">m-cu</emphasis>
. Hind wing with distal abscissae of
<emphasis id="80FED718FFF42E24220BF961FB8AFB92" box="[1035,1058,1124,1149]" italics="true" pageId="7" pageNumber="272">M</emphasis>
,
<emphasis id="80FED718FFF42E242231F961FBFAFB91" box="[1073,1106,1124,1150]" italics="true" pageId="7" pageNumber="272">Cu</emphasis>
1 and 1
<emphasis id="80FED718FFF42E2422A4F966FB1DFB92" box="[1188,1205,1123,1149]" italics="true" pageId="7" pageNumber="272">A</emphasis>
sclerotized throughout. Outer metatibial spur 0.7× as long as inner spur.
</paragraph>
<caption id="E6F55B82FFF42E242697FA9AFBCAF834" ID-DOI="http://doi.org/10.5281/zenodo.5531255" ID-Zenodo-Dep="5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="7" pageNumber="272" startId="7.[151,250,1951,1975]" targetBox="[151,1435,1209,1926]" targetPageId="7">
<paragraph id="B2350B0AFFF42E242697FA9AFBCAF834" blockId="7.[151,1437,1951,2011]" pageId="7" pageNumber="272">
<emphasis id="80FED718FFF42E242697FA9AFEBDF858" bold="true" box="[151,277,1951,1975]" pageId="7" pageNumber="272">FIGURE 4.</emphasis>
Details of
<taxonomicName id="758A7089FFF42E242789FAA5FDC8F858" authority="Alvarado &amp; Palacio, 2021" authorityName="Alvarado &amp; Palacio" authorityYear="2021" box="[393,608,1952,1975]" class="Insecta" family="Ichneumonidae" genus="Forrestopius" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="272" phylum="Arthropoda" rank="species" species="nasutus" status="sp. nov.">
<emphasis id="80FED718FFF42E242789FAA5FDC8F858" box="[393,608,1952,1975]" italics="true" pageId="7" pageNumber="272">Forrestopius nasutus</emphasis>
</taxonomicName>
,
<emphasis id="80FED718FFF42E24246FFA9AFD4AF858" bold="true" box="[623,738,1951,1975]" pageId="7" pageNumber="272">
<taxonomicNameLabel id="9BCD6A63FFF42E24246FFA9AFD69F858" box="[623,705,1951,1975]" pageId="7" pageNumber="272" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Habitus of female, in lateral view (scale bar = 2 mm)
<emphasis id="80FED718FFF42E24231BFA9AFA9AF858" bold="true" box="[1307,1330,1951,1975]" pageId="7" pageNumber="272">B.</emphasis>
Head and pronotum, in lateral view
<emphasis id="80FED718FFF42E24279EFAC6FE1EF834" bold="true" box="[414,438,1987,2011]" pageId="7" pageNumber="272">C.</emphasis>
Facial view
<emphasis id="80FED718FFF42E242438FAC6FDF9F834" bold="true" box="[568,593,1987,2011]" pageId="7" pageNumber="272">D.</emphasis>
Head and mesosoma, in dorsal view.
<emphasis id="80FED718FFF42E2425D2FAC6FC41F834" bold="true" box="[978,1001,1987,2011]" pageId="7" pageNumber="272">E.</emphasis>
Mandibles.
</paragraph>
</caption>
<paragraph id="B2350B0AFFFB2E2B26C7FD92FEDFFEF2" blockId="8.[151,1436,151,501]" pageId="8" pageNumber="273">
<taxonomicName id="758A7089FFFB2E2B26C7FD92FE97FF5F" box="[199,319,151,176]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="80FED718FFFB2E2B26C7FD92FE97FF5F" box="[199,319,151,176]" italics="true" pageId="8" pageNumber="273">Mesosoma</emphasis>
</taxonomicName>
. Tergite I 1.7× as long as posteriorly wide, lateromedian carina weak, extending about 0.3× length of tergite, smooth, with isolated punctures laterally on the apical half; tergite II 1.6× as long as posteriorly wide; laterotergite II very narrow, about 0.1× as wide as long; laterotergite III about 0.3× as wide as long, wedge-shaped, mesal edge straight.
</paragraph>
<paragraph id="B2350B0AFFFB2E2B26C7FC22FB15FE8A" blockId="8.[151,1436,151,501]" pageId="8" pageNumber="273">
<emphasis id="80FED718FFFB2E2B26C7FC22FEBEFEAE" box="[199,278,295,321]" italics="true" pageId="8" pageNumber="273">Colour</emphasis>
. Body entirely black with intense blue to purple reflections (
<figureCitation id="2AB1178FFFFB2E2B25B1FC22FB93FEAE" box="[945,1083,295,321]" captionStart="FIGURE 4" captionStartId="7.[151,250,1951,1975]" captionTargetBox="[151,1435,1209,1926]" captionTargetId="figure-457@7.[151,1436,1209,1927]" captionTargetPageId="7" captionText="FIGURE 4. Details of Forrestopius nasutus, sp. nov. A. Habitus of female, in lateral view (scale bar = 2 mm) B. Head and pronotum, in lateral view C. Facial view D. Head and mesosoma, in dorsal view. E. Mandibles." figureDoi="http://doi.org/10.5281/zenodo.5531255" httpUri="https://zenodo.org/record/5531255/files/figure.png" pageId="8" pageNumber="273">Figs 4A, 4D</figureCitation>
) except on clypeus, lower head, mandibles, palpi, antennae, tibiae, tarsi, laterotergites and sterna. Ovipositor valvae testaceous.
</paragraph>
<paragraph id="B2350B0AFFFB2E2B26C7FC6AFE28FE66" blockId="8.[151,1436,151,501]" box="[199,384,367,393]" pageId="8" pageNumber="273">
<emphasis id="80FED718FFFB2E2B26C7FC6AFEACFE66" bold="true" box="[199,260,367,393]" pageId="8" pageNumber="273">Male</emphasis>
. Unknown
</paragraph>
</subSubSection>
<subSubSection id="FA905881FFFB2E2B26C7FC96FBF9FE42" box="[199,1105,403,429]" pageId="8" pageNumber="273" type="discussion">
<paragraph id="B2350B0AFFFB2E2B26C7FC96FBF9FE42" blockId="8.[151,1436,151,501]" box="[199,1105,403,429]" pageId="8" pageNumber="273">
<emphasis id="80FED718FFFB2E2B26C7FC96FE93FE42" bold="true" box="[199,315,403,429]" pageId="8" pageNumber="273">Remarks.</emphasis>
This species occurs in Cloud forest of
<collectingCountry id="CA9D4B9AFFFB2E2B24F0FC96FCF6FE42" box="[752,862,403,429]" name="Colombia" pageId="8" pageNumber="273">Colombia</collectingCountry>
at
<quantity id="7572A6EFFFFB2E2B2580FC96FC66FE42" box="[896,974,403,429]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="273" unit="m" value="2800.0">2800m</quantity>
(
<figureCitation id="2AB1178FFFFB2E2B25DDFC96FBECFE42" box="[989,1092,403,429]" captionStart="FIGURE 10" captionStartId="16.[151,250,881,905]" captionTargetBox="[151,1436,181,857]" captionTargetId="figure-23@16.[151,1436,181,858]" captionTargetPageId="16" captionText="FIGURE 10. Geographic distribution of the Forrestopius species" figureDoi="http://doi.org/10.5281/zenodo.5531269" httpUri="https://zenodo.org/record/5531269/files/figure.png" pageId="8" pageNumber="273">Fig. 10C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="FA905881FFFB2E2B26C7FCB2FE6FFE1A" pageId="8" pageNumber="273" type="etymology">
<paragraph id="B2350B0AFFFB2E2B26C7FCB2FE6FFE1A" blockId="8.[151,1436,151,501]" pageId="8" pageNumber="273">
<emphasis id="80FED718FFFB2E2B26C7FCB2FEE0FE3E" bold="true" box="[199,328,439,465]" pageId="8" pageNumber="273">Etymology</emphasis>
. The specific epithet
<taxonomicName id="758A7089FFFB2E2B2437FCB2FD25FE3F" authorityName="Alvarado &amp; Palacio" authorityYear="2021" box="[567,653,439,464]" class="Insecta" family="Ichneumonidae" genus="Forrestopius" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="273" phylum="Arthropoda" rank="species" species="nasutus">
<emphasis id="80FED718FFFB2E2B2437FCB2FD25FE3F" box="[567,653,439,464]" italics="true" pageId="8" pageNumber="273">nasutus</emphasis>
</taxonomicName>
refers to the strongly prominent clypeus that distinguishes this species from all other in the genus.
</paragraph>
</subSubSection>
</treatment>
</document>
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