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<document ID-ISSN="0003-0090" approvalRequired="248" approvalRequired_for_originalDoi="1" approvalRequired_for_taxonomicNames="67" approvalRequired_for_textStreams="120" approvalRequired_for_treatments="60" checkinTime="1630255760165" checkinUser="felipe" docAuthor="Conrad, J. L." docDate="2008" docId="BF23879DD103FFE8FF33AB134F8BD616" docLanguage="en" docName="B310.pdf" docOrigin="Bulletin of the American Museum of Natural History 2008 (310)" docStyle="DocumentStyle:2B34016C1EBFE58B07368F9517618743.1:BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleId="2B34016C1EBFE58B07368F9517618743" docStyleName="BulAmeMusNatHis.2000-2010.journal_article.0cover.type1" docStyleVersion="1" docTitle="Hoplocercidae Frost and Etheridge 1989" docType="treatment" docVersion="2" lastPageNumber="83" masterDocId="431AFFE5D153FFBAFFA0AE674F56D039" masterDocTitle="Phylogeny And Systematics Of Squamata (Reptilia) Based On Morphology" masterLastPageNumber="182" masterPageNumber="1" pageNumber="81" updateTime="1630877461833" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>Phylogeny And Systematics Of Squamata (Reptilia) Based On Morphology</mods:title>
</mods:titleInfo>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Conrad, J. L.</mods:namePart>
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<mods:title>Bulletin of the American Museum of Natural History</mods:title>
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<mods:date>2008</mods:date>
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<mods:number>2008-06-03</mods:number>
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<mods:number>2008</mods:number>
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<mods:detail type="issue">
<mods:number>310</mods:number>
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<mods:start>1</mods:start>
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<mods:identifier type="ISSN">0003-0090</mods:identifier>
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<paragraph blockId="80.[147,634,1396,1448]" box="[147,634,1396,1418]" pageId="80" pageNumber="81">
<heading box="[147,634,1396,1418]" centered="true" fontSize="9" level="2" pageId="80" pageNumber="81" reason="2">
<taxonomicName authority="Frost and Etheridge, 1989" authorityName="Frost and Etheridge" authorityYear="1989" box="[147,634,1396,1418]" class="Reptilia" family="Hoplocercidae" kingdom="Animalia" order="Squamata" pageId="80" pageNumber="81" phylum="Chordata" rank="family">
Hoplocercidae
<bibRefCitation author="Frost, D. R. &amp; R. Etheridge" box="[325,634,1396,1418]" pageId="80" pageNumber="81" pagination="1 - 65" refId="ref89007" refString="Frost, D. R., and R. Etheridge. 1989. A phylogenetic analysis, and taxonomy of iguanian lizards (Reptilia: Squamata). University of Kansas Museum of Natural History Miscellaneous Publications 81: 1 - 65." type="journal article" year="1989">Frost and Etheridge, 1989</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="80.[147,634,1396,1448]" box="[264,517,1426,1448]" pageId="80" pageNumber="81">
<heading box="[264,517,1426,1448]" centered="true" fontSize="10" level="2" pageId="80" pageNumber="81" reason="4">(figs. 54A, 55A, 56A)</heading>
</paragraph>
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<subSubSection pageId="80" pageNumber="81" type="diagnosis">
<paragraph blockId="80.[128,654,1471,1733]" lastBlockId="80.[683,801,1160,1179]" pageId="80" pageNumber="81">DIAGNOSIS: The current analysis recovers five unambiguous synapomorphies for this clade. These are 44(1) enlarged lacrimal foramen (see character description above), 60(1) a dorsoventrally inflated frontal, 190(0) splenial extending anteriorly for more than two-thirds the dentary tooth row, 258(1) one or more pair(s) of postxiphisternal inscriptional ribs confluent at midline, and 285(2) Continued.</paragraph>
<paragraph blockId="80.[701,1228,1237,1735]" pageId="80" pageNumber="81">symphysial portion of the pubis more than half again as long as the tubercular portion.</paragraph>
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<subSubSection lastPageId="82" lastPageNumber="83" pageId="80" pageNumber="81" type="discussion">
<paragraph blockId="80.[701,1228,1237,1735]" pageId="80" pageNumber="81">
COMMENTS: This analysis includes
<taxonomicName class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="80" pageNumber="81" phylum="Chordata" rank="species" species="annularis">
<emphasis italics="true" pageId="80" pageNumber="81">Morunasaurus annularis</emphasis>
</taxonomicName>
,
<taxonomicName box="[966,1220,1326,1349]" class="Reptilia" family="Hoplocercidae" genus="Hoplocercus" kingdom="Animalia" order="Squamata" pageId="80" pageNumber="81" phylum="Chordata" rank="species" species="spinosus">
<emphasis box="[966,1220,1326,1349]" italics="true" pageId="80" pageNumber="81">Hoplocercus spinosus</emphasis>
</taxonomicName>
, and a composite
<taxonomicName authorityName="Boulenger" authorityYear="1885" box="[908,1039,1356,1379]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="80" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis box="[908,1039,1356,1379]" italics="true" pageId="80" pageNumber="81">Enyalioides</emphasis>
</taxonomicName>
scored from
<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1883" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="80" pageNumber="81" phylum="Chordata" rank="species" species="palpebralis">
<emphasis italics="true" pageId="80" pageNumber="81">E. palpebralis</emphasis>
</taxonomicName>
and
<taxonomicName box="[885,1003,1385,1408]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="80" pageNumber="81" phylum="Chordata" rank="species" species="laticeps">
<emphasis box="[885,1003,1385,1408]" italics="true" pageId="80" pageNumber="81">E. laticeps</emphasis>
</taxonomicName>
. A recent study by
<bibRefCitation author="Wiens, J. J. &amp; R. E. Etheridge" box="[701,1044,1416,1438]" pageId="80" pageNumber="81" pagination="375 - 398" refId="ref102174" refString="Wiens, J. J., and R. E. Etheridge. 2003. Phylogenetic relationships of hoplocercid lizards: coding and combining meristic, morphometric, and polymorphic data using step matrices. Herpetologica 59: 375 - 398." type="journal article" year="2003">Wiens and Etheridge (2003)</bibRefCitation>
with extensive sampling of hoplocercid taxa suggests the monophyly of
<taxonomicName authorityName="Boulenger" authorityYear="1885" box="[886,1016,1475,1498]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="80" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis box="[886,1016,1475,1498]" italics="true" pageId="80" pageNumber="81">Enyalioides</emphasis>
</taxonomicName>
. Even so, future versions of this analysis will eliminate the composite coding of
<taxonomicName authorityName="Boulenger" authorityYear="1885" box="[963,1094,1534,1557]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="80" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis box="[963,1094,1534,1557]" italics="true" pageId="80" pageNumber="81">Enyalioides</emphasis>
</taxonomicName>
and likely include more species of
<taxonomicName authorityName="Dunn" authorityYear="1933" box="[1001,1165,1565,1586]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="80" pageNumber="81" phylum="Chordata" rank="genus">
<emphasis box="[1001,1165,1565,1586]" italics="true" pageId="80" pageNumber="81">Morunasaurus</emphasis>
</taxonomicName>
. Importantly, the Adams consensus (figs. 55, 56) supports the topology of Wiens and Etheridge based on their analysis using mixed scaling of meristic characters (
<bibRefCitation author="Wiens, J. J. &amp; R. E. Etheridge" pageId="80" pageNumber="81" pagination="375 - 398" refId="ref102174" refString="Wiens, J. J., and R. E. Etheridge. 2003. Phylogenetic relationships of hoplocercid lizards: coding and combining meristic, morphometric, and polymorphic data using step matrices. Herpetologica 59: 375 - 398." type="journal article" year="2003">Wiens and Etheridge, 2003</bibRefCitation>
: fig. 4).
</paragraph>
<caption pageId="81" pageNumber="82" startId="81.[116,155,1656,1675]" targetBox="[118,1169,211,1626]" targetPageId="81">
<paragraph blockId="81.[93,1193,1656,1729]" pageId="81" pageNumber="82">
Fig. 55.
<emphasis bold="true" box="[224,285,1656,1675]" pageId="81" pageNumber="82">(AF)</emphasis>
Hypothesis of squamate interrelationships based on the current study; the Adams consensus with accompanying taxonomic scheme. Following (
<bibRefCitation author="Brochu, C. A." box="[751,894,1683,1702]" pageId="81" pageNumber="82" pagination="9 - 100" refId="ref84907" refString="Brochu, C. A. 1999. Phylogenetics, taxonomy, and historical biogeography of Alligatoroidea. Memoir: Society of Vertebrate Paleontology 6: 9 - 100." type="journal article" year="1999">Brochu, 1999</bibRefCitation>
), parentheses indicate stembased clade names, dots indicate node-based names.
</paragraph>
</caption>
<paragraph blockId="82.[578,659,1246,1265]" box="[578,659,1246,1265]" pageId="82" pageNumber="83">
<figureCitation box="[578,656,1246,1265]" captionStart="Fig" captionStartId="81.[116,155,1656,1675]" captionTargetBox="[118,1169,211,1626]" captionTargetId="figure-1@81.[118,1169,211,1626]" captionTargetPageId="81" captionText="Fig. 55. (AF) Hypothesis of squamate interrelationships based on the current study; the Adams consensus with accompanying taxonomic scheme. Following (Brochu, 1999), parentheses indicate stembased clade names, dots indicate node-based names." pageId="82" pageNumber="83">Fig. 55</figureCitation>
.
</paragraph>
<paragraph blockId="82.[128,654,1357,1583]" pageId="82" pageNumber="83">
Given the absence of a fossil record for this clade and the somewhat limited sampling for it, I refrain from attaching a definition to this taxon name. The term Hoplocercidae is usually used to refer to all species within
<taxonomicName authorityName="Boulenger" authorityYear="1885" box="[353,484,1502,1525]" class="Reptilia" family="Hoplocercidae" genus="Enyalioides" kingdom="Animalia" order="Squamata" pageId="82" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis box="[353,484,1502,1525]" italics="true" pageId="82" pageNumber="83">Enyalioides</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Fitzinger" authorityYear="1843" box="[505,646,1502,1525]" class="Reptilia" family="Hoplocercidae" genus="Hoplocercus" kingdom="Animalia" order="Squamata" pageId="82" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis box="[505,646,1502,1525]" italics="true" pageId="82" pageNumber="83">Hoplocercus</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Dunn" authorityYear="1933" box="[186,350,1533,1554]" class="Reptilia" family="Hoplocercidae" genus="Morunasaurus" kingdom="Animalia" order="Squamata" pageId="82" pageNumber="83" phylum="Chordata" rank="genus">
<emphasis box="[186,350,1533,1554]" italics="true" pageId="82" pageNumber="83">Morunasaurus</emphasis>
</taxonomicName>
; it is used here in that context.
</paragraph>
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</treatment>
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