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<mods:title id="D6B699F12A43C5471C0160F851BDF20B">A phylogenetic analysis and systematic revision of the cryptobranch dorids (Mollusca, Nudibranchia, Anthobranchia)</mods:title>
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<paragraph id="8B84CE18FFC36065FC09F9956A55D54E" blockId="44.[910,1318,1627,1651]" box="[910,1318,1627,1651]" pageId="44" pageNumber="579">
<heading id="D0CC7974FFC36065FC09F9956A55D54E" box="[910,1318,1627,1651]" centered="true" fontSize="9" level="2" pageId="44" pageNumber="579" reason="5">
GENUS
<taxonomicName id="4C3BB59BFFC36065FC61F9956A55D54E" ID-CoL="7NPCZ" ID-ENA="763087" authority="BERGH, 1879" authorityName="ANTILLENSIS BERGH" authorityYear="1879" box="[998,1318,1627,1651]" class="Gastropoda" family="Dorididae" genus="Aphelodoris" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="579" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFC36065FC61F9956BF4D54F" box="[998,1159,1627,1651]" italics="true" pageId="44" pageNumber="579">APHELODORIS</emphasis>
<bibRefCitation id="EFAAB3E9FFC36065FB08F9956A55D54E" author="Bergh R" box="[1167,1318,1627,1651]" pageId="44" pageNumber="579" pagination="87 - 116" refId="ref53956" refString="Bergh R. 1879. Neue Chromodoriden. Malakozoologische Blatter Neue Folge 1: 87 - 116, plate 4." type="journal article" year="1879">BERGH, 1879</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3219D93FFC36065FCA1F94A6A0BD585" pageId="44" pageNumber="579" type="reference_group">
<paragraph id="8B84CE18FFC36065FCA1F94A6A0BD585" blockId="44.[806,1423,1668,1720]" pageId="44" pageNumber="579">
<treatmentCitationGroup id="AB2BE936FFC36065FCA1F94A6A0BD585" pageId="44" pageNumber="579">
<treatmentCitation id="0A9AE809FFC36065FCA1F94A6BA0D5A4" author="Bergh R" box="[806,1235,1668,1690]" page="108" pageId="44" pageNumber="579" year="1879">
<taxonomicName id="4C3BB59BFFC36065FCA1F94A6BA0D5A4" ID-CoL="7NPCZ" authority="Bergh, 1879: 107 - 108" authorityName="ANTILLENSIS BERGH" authorityPageNumber="107 - 108" authorityYear="1879" box="[806,1235,1668,1690]" class="Gastropoda" family="Dorididae" genus="Aphelodoris" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="579" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFC36065FCA1F94A6CD9D5A4" box="[806,938,1668,1689]" italics="true" pageId="44" pageNumber="579">Aphelodoris</emphasis>
<bibRefCitation id="EFAAB3E9FFC36065FC39F94A6BA0D5A4" author="Bergh R" box="[958,1235,1668,1690]" pageId="44" pageNumber="579" pagination="87 - 116" refId="ref53956" refString="Bergh R. 1879. Neue Chromodoriden. Malakozoologische Blatter Neue Folge 1: 87 - 116, plate 4." type="journal article" year="1879">Bergh, 1879: 107108</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="548070BAFFC36065FB6DF94A6A51D5A4" box="[1258,1314,1668,1689]" pageId="44" pageNumber="579">Type</typeStatus>
species:
<taxonomicName id="4C3BB59BFFC36065FCB8F96C6BA0D585" authority="Bergh, 1879" authorityName="Bergh" authorityYear="1879" box="[831,1235,1698,1720]" class="Gastropoda" family="Dorididae" genus="Aphelodoris" kingdom="Animalia" order="Nudibranchia" pageId="44" pageNumber="579" phylum="Mollusca" rank="species" species="antillensis">
<emphasis id="B94F120AFFC36065FCB8F96C6B4DD58A" box="[831,1086,1698,1719]" italics="true" pageId="44" pageNumber="579">Aphelodoris antillensis</emphasis>
<bibRefCitation id="EFAAB3E9FFC36065FBC0F96C6BA0D585" author="Bergh R" box="[1095,1235,1698,1720]" pageId="44" pageNumber="579" pagination="87 - 116" refId="ref53956" refString="Bergh R. 1879. Neue Chromodoriden. Malakozoologische Blatter Neue Folge 1: 87 - 116, plate 4." type="journal article" year="1879">Bergh, 1879</bibRefCitation>
</taxonomicName>
, by monotypy.
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</paragraph>
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<subSubSection id="C3219D93FFC36066FCA1F9316DF1D041" lastPageId="47" lastPageNumber="582" pageId="44" pageNumber="579" type="diagnosis">
<paragraph id="8B84CE18FFC36065FCA1F9316CE0D429" blockId="44.[806,1422,1791,1904]" box="[806,915,1791,1812]" pageId="44" pageNumber="579">
<emphasis id="B94F120AFFC36065FCA1F9316CE0D429" box="[806,915,1791,1812]" italics="true" pageId="44" pageNumber="579">Diagnosis</emphasis>
</paragraph>
<paragraph id="8B84CE18FFC36066FCA1F8D36EC6D24E" blockId="44.[806,1422,1791,1904]" lastBlockId="47.[155,771,196,371]" lastPageId="47" lastPageNumber="582" pageId="44" pageNumber="579">Body soft, lacking integumentary spicules. Dorsum smooth, with no tubercles. Anterior border of the foot grooved but not notched. Head with two large and grooved lateral prolongations. Labial cuticle lacking rodlets. Radula composed of simple, hamate teeth. The innermost teeth may be simple or denticulate. Reproductive system with a tubular, granular and simple prostate. Penis and vagina unarmed. Vestibular or accessory glands absent.</paragraph>
<caption id="DF449E90FFC26064FF1CF8DE6A0CD47C" ID-DOI="http://doi.org/10.5281/zenodo.4634276" ID-Zenodo-Dep="4634276" httpUri="https://zenodo.org/record/4634276/files/figure.png" pageId="45" pageNumber="580" startId="45.[155,233,1808,1827]" targetBox="[155,1435,195,1776]" targetPageId="45">
<paragraph id="8B84CE18FFC26064FF1CF8DE6A0CD47C" blockId="45.[155,1435,1808,1857]" pageId="45" pageNumber="580">
<emphasis id="B94F120AFFC26064FF1CF8DE6E63D41E" bold="true" box="[155,272,1808,1827]" pageId="45" pageNumber="580">Figure 24.</emphasis>
<taxonomicName id="4C3BB59BFFC26064FE9BF8DE6D01D41E" authority="(SMNHI 1540)" baseAuthorityName="SMNHI" baseAuthorityYear="1540" box="[284,626,1808,1827]" class="Gastropoda" family="Cadlinidae" genus="Aldisa" kingdom="Animalia" order="Nudibranchia" pageId="45" pageNumber="580" phylum="Mollusca" rank="species" species="zetlandica">
<emphasis id="B94F120AFFC26064FE9BF8DE6EA3D41E" box="[284,464,1808,1827]" italics="true" pageId="45" pageNumber="580">Aldisa zetlandica</emphasis>
(SMNHI 1540)
</taxonomicName>
, SEM images of the radula and dorsal tubercles. A, general view of the lateral teeth; scale bar = 43 Mm. B, detail of the lateral tooth cusps; scale bar = 15 Mm. C, dorsal tubercles; scale bar = 250 Mm.
</paragraph>
</caption>
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<paragraph id="8B84CE18FFC16067FF09F8D66BAED45B" blockId="46.[142,1422,1816,1894]" pageId="46" pageNumber="581">
<emphasis id="B94F120AFFC16067FF09F8D66E77D416" bold="true" box="[142,260,1816,1835]" pageId="46" pageNumber="581">Figure 25.</emphasis>
<taxonomicName id="4C3BB59BFFC16067FE88F8D66D1ED416" authority="(SMNHI 1540)" baseAuthorityName="SMNHI" baseAuthorityYear="1540" box="[271,621,1816,1835]" class="Gastropoda" family="Cadlinidae" genus="Aldisa" kingdom="Animalia" order="Nudibranchia" pageId="46" pageNumber="581" phylum="Mollusca" rank="species" species="zetlandica">
<emphasis id="B94F120AFFC16067FE88F8D66EB5D416" box="[271,454,1816,1835]" italics="true" pageId="46" pageNumber="581">Aldisa zetlandica</emphasis>
(SMNHI 1540)
</taxonomicName>
. A, general view of the anatomy; scale bar = 1 mm. B, reproductive system; scale bar = 1 mm. C, detail of the ampulla and prostate; scale bar = 1 mm. D, central nervous system; scale bar = 1 mm. E, lateral view of the buccal bulb; scale bar = 1 mm. F, ventral view of the mouth area; scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B84CE18FFC06066FF1CFE776F8ED2F3" blockId="47.[155,771,441,892]" box="[155,253,441,462]" pageId="47" pageNumber="582">
<emphasis id="B94F120AFFC06066FF1CFE776F8ED2F3" box="[155,253,441,462]" italics="true" pageId="47" pageNumber="582">Remarks</emphasis>
</paragraph>
<paragraph id="8B84CE18FFC06066FF1CFE166DF1D041" blockId="47.[155,771,441,892]" pageId="47" pageNumber="582">
The genus
<taxonomicName id="4C3BB59BFFC06066FEA8FE166EC0D2D0" authorityName="ANTILLENSIS BERGH" authorityYear="1879" box="[303,435,472,493]" class="Gastropoda" family="Dorididae" genus="Aphelodoris" kingdom="Animalia" order="Nudibranchia" pageId="47" pageNumber="582" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFC06066FEA8FE166EC0D2D0" box="[303,435,472,493]" italics="true" pageId="47" pageNumber="582">Aphelodoris</emphasis>
</taxonomicName>
was introduced by
<bibRefCitation id="EFAAB3E9FFC06066FD3AFE166F90D130" author="Bergh R" pageId="47" pageNumber="582" pagination="87 - 116" refId="ref53956" refString="Bergh R. 1879. Neue Chromodoriden. Malakozoologische Blatter Neue Folge 1: 87 - 116, plate 4." type="journal article" year="1879">Bergh (1879)</bibRefCitation>
for
<taxonomicName id="4C3BB59BFFC06066FE8AFE386DEFD131" authority="Bergh, 1879" authorityName="Bergh" authorityYear="1879" box="[269,668,502,524]" class="Gastropoda" family="Dorididae" genus="Aphelodoris" kingdom="Animalia" order="Nudibranchia" pageId="47" pageNumber="582" phylum="Mollusca" rank="species" species="antillensis">
<emphasis id="B94F120AFFC06066FE8AFE386D79D136" box="[269,522,502,523]" italics="true" pageId="47" pageNumber="582">Aphelodoris antillensis</emphasis>
<bibRefCitation id="EFAAB3E9FFC06066FD96FE386DEFD131" author="Bergh R" box="[529,668,502,524]" pageId="47" pageNumber="582" pagination="87 - 116" refId="ref53956" refString="Bergh R. 1879. Neue Chromodoriden. Malakozoologische Blatter Neue Folge 1: 87 - 116, plate 4." type="journal article" year="1879">Bergh, 1879</bibRefCitation>
</taxonomicName>
, the
<typeStatus id="548070BAFFC06066FD56FE396C71D131" box="[721,770,503,524]" pageId="47" pageNumber="582">type</typeStatus>
species by monotypy. According to
<bibRefCitation id="EFAAB3E9FFC06066FDA9FDDB6DB6D116" author="Bergh R" box="[558,709,533,555]" pageId="47" pageNumber="582" pagination="87 - 116" refId="ref53956" refString="Bergh R. 1879. Neue Chromodoriden. Malakozoologische Blatter Neue Folge 1: 87 - 116, plate 4." type="journal article" year="1879">Bergh (1879)</bibRefCitation>
<taxonomicName id="4C3BB59BFFC06066FD49FDDB6F81D174" authorityName="ANTILLENSIS BERGH" authorityYear="1879" class="Gastropoda" family="Dorididae" genus="Aphelodoris" kingdom="Animalia" order="Nudibranchia" pageId="47" pageNumber="582" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFC06066FD49FDDB6F81D174" italics="true" pageId="47" pageNumber="582">Aphelodoris</emphasis>
</taxonomicName>
is very similar in body shape to the chromodorids, but it differs from this latter group in having a narrow mantle margin and a short posterior end of the foot. Other differences include the shape of the oral tentacles, which are grooved, and the presence of multipinnate branchial leaves. Internally the differences are even more obvious, with the absence of jaws and the presence of a single blood gland.
<bibRefCitation id="EFAAB3E9FFC06066FDEEFCC46C70D01C" author="Bergh R" box="[617,771,778,801]" pageId="47" pageNumber="582" pagination="87 - 116" refId="ref53956" refString="Bergh R. 1879. Neue Chromodoriden. Malakozoologische Blatter Neue Folge 1: 87 - 116, plate 4." type="journal article" year="1879">Bergh (1879)</bibRefCitation>
regarded
<taxonomicName id="4C3BB59BFFC06066FE95FCE76EE5D003" authorityName="ANTILLENSIS BERGH" authorityYear="1879" box="[274,406,809,830]" class="Gastropoda" family="Dorididae" genus="Aphelodoris" kingdom="Animalia" order="Nudibranchia" pageId="47" pageNumber="582" phylum="Mollusca" rank="genus">
<emphasis id="B94F120AFFC06066FE95FCE76EE5D003" box="[274,406,809,830]" italics="true" pageId="47" pageNumber="582">Aphelodoris</emphasis>
</taxonomicName>
as a member of the family
<taxonomicName id="4C3BB59BFFC06066FF1CFC866E10D063" authorityName="Bergh" authorityYear="1891" box="[155,355,840,862]" class="Gastropoda" family="Chromodorididae" kingdom="Animalia" order="Nudibranchia" pageId="47" pageNumber="582" phylum="Mollusca" rank="family">Chromodorididae</taxonomicName>
, but Odhner (see
<bibRefCitation id="EFAAB3E9FFC06066FDB4FC866DCFD063" author="Franc A" box="[563,700,840,862]" pageId="47" pageNumber="582" pagination="608 - 893" refId="ref55985" refString="Franc A. 1968. Mollusques, gasteropodes et scaphopodes. In: Grasse P, ed. Traite de Zoologie Anatomie, Systematique, Biologie, Vol. 5. Paris: Masson, 608 - 893." type="book chapter" year="1968">Franc, 1968</bibRefCitation>
) later transferred it to the family
<taxonomicName id="4C3BB59BFFC06066FE5CFCA86D0DD041" authorityName="Thiele" authorityYear="1931" box="[475,638,870,892]" class="Gastropoda" family="Asteronotidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nudibranchia" pageId="47" pageNumber="582" phylum="Mollusca" rank="family">Asteronotidae</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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