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<document id="3230D37BD3970DE456B5714E308F7214" ID-DOI="10.25674/so95iss2id315" ID-ISSN="2509-9523" ID-Zenodo-Dep="13193924" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722631509206" checkinUser="felipe" docAuthor="Seifert, Bernhard" docDate="2023" docId="03FAE936FFCBFA7E375EFF29FB98EE7F" docLanguage="en" docName="SoilOrg.95.2.129-142.pdf" docOrigin="Soil Organisms 95 (2)" docSource="http://dx.doi.org/10.25674/so95iss2id315" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Formicoxenus gebaueri Seifert, 2023, n.sp." docType="treatment" docVersion="3" lastPageNumber="137" masterDocId="FFC3914EFFC3FA763477FFD8FFBAE831" masterDocTitle="Two new species of Formicoxenus Mayr 1855 and Leptothorax Mayr 1855 from Tibet (Hymenoptera: Formicidae)" masterLastPageNumber="142" masterPageNumber="129" pageNumber="137" updateTime="1722679370904" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="93BB70804D9527E761B841BF21D1CBC6">Two new species of Formicoxenus Mayr 1855 and Leptothorax Mayr 1855 from Tibet (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="2A406B334ED08D6B15AE14935F7BC406">Seifert, Bernhard</mods:namePart>
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<mods:title id="99A8E03E1621C878CED2666E953F0AB9">Soil Organisms</mods:title>
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<mods:date id="6363800F2E3A9B1F4F2F9CA6FEE2300D">2023</mods:date>
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<mods:number id="87A17311A29C26ABB5D73A5C6F1636A8">2023-08-01</mods:number>
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<mods:identifier id="279E4C71E7E7B4DE7480489A89D5AA43" type="DOI">10.25674/so95iss2id315</mods:identifier>
<mods:identifier id="98724318EFCD15CDCEACB5BF1CF297CD" type="ISSN">2509-9523</mods:identifier>
<mods:identifier id="4440FA6CB8E4ED90CC57BE200C446AA7" type="Zenodo-Dep">13193924</mods:identifier>
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<treatment id="03FAE936FFCBFA7E375EFF29FB98EE7F" ID-DOI="http://doi.org/10.5281/zenodo.13193106" ID-Zenodo-Dep="13193106" LSID="urn:lsid:plazi:treatment:03FAE936FFCBFA7E375EFF29FB98EE7F" httpUri="http://treatment.plazi.org/id/03FAE936FFCBFA7E375EFF29FB98EE7F" lastPageNumber="137" pageId="8" pageNumber="137">
<subSubSection id="C3490BABFFCBFA7E375EFF29FBCCE93B" box="[809,1142,241,266]" pageId="8" pageNumber="137" type="nomenclature">
<paragraph id="8BEC5820FFCBFA7E375EFF29FBCCE93B" blockId="8.[809,1142,241,266]" box="[809,1142,241,266]" pageId="8" pageNumber="137">
<heading id="D0A4EF4CFFCBFA7E375EFF29FBCCE93B" box="[809,1142,241,266]" fontSize="10" level="2" pageId="8" pageNumber="137" reason="4">
<emphasis id="B9278432FFCBFA7E375EFF29FBCCE93B" bold="true" box="[809,1142,241,266]" pageId="8" pageNumber="137">
<taxonomicName id="4C5323A3FFCBFA7E375EFF29FB8DE938" authorityName="Seifert" authorityYear="2023" box="[809,1079,241,265]" class="Insecta" family="Formicidae" genus="Formicoxenus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="gebaueri" status="sp. nov.">
<emphasis id="B9278432FFCBFA7E375EFF29FB8DE938" bold="true" box="[809,1079,241,265]" italics="true" pageId="8" pageNumber="137">Formicoxenus gebaueri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2143949FFCBFA7E3048FF2AFBCCE93B" box="[1087,1142,242,266]" pageId="8" pageNumber="137" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3490BABFFCBFA7E375EFEEBFAD4E97A" box="[809,1390,307,331]" pageId="8" pageNumber="137" type="etymology">
<paragraph id="8BEC5820FFCBFA7E375EFEEBFAD4E97A" blockId="8.[786,1399,307,1614]" box="[809,1390,307,331]" pageId="8" pageNumber="137">Etymology: dedicated to the collector Axel Gebauer.</paragraph>
</subSubSection>
<subSubSection id="C3490BABFFCBFA7E375EFE8CFC4CE9DE" pageId="8" pageNumber="137" type="materials_examined">
<paragraph id="8BEC5820FFCBFA7E375EFE8CFC6CE95D" blockId="8.[786,1399,307,1614]" box="[809,982,340,364]" pageId="8" pageNumber="137">
<emphasis id="B9278432FFCBFA7E375EFE8CFC6CE95D" bold="true" box="[809,982,340,364]" pageId="8" pageNumber="137">Type material:</emphasis>
</paragraph>
<paragraph id="8BEC5820FFCBFA7E375EFEADFC4CE9DE" blockId="8.[786,1399,307,1614]" pageId="8" pageNumber="137">
<materialsCitation id="3B3B527DFFCBFA7E375EFEADFC4CE9DE" collectingDate="1996-05-30" collectorName="Gerollhang. A. Gebauer" country="China" elevation="2600" latitude="36.82" location="Beishan Nat. Park" longLatPrecision="711" longitude="102.53" pageId="8" pageNumber="137" specimenCount="7" specimenCount-worker="7" typeStatus="holotype">
<typeStatus id="54E8E682FFCBFA7E375EFEADFC35E9BC" box="[809,911,373,397]" pageId="8" pageNumber="137" type="holotype">Holotype</typeStatus>
and
<specimenCount id="9D5593A9FFCBFA7E37A2FEAEFB00E9BC" box="[981,1210,373,398]" count="7" pageId="8" pageNumber="137" type="worker" typeStatus="paratype">7 paratype workers</specimenCount>
from the same nest labelled
<collectingCountry id="F34418B0FFCBFA7E37CBFE4EFBA1E99F" box="[956,1051,406,430]" name="China" pageId="8" pageNumber="137">CHINA</collectingCountry>
:
<geoCoordinate id="EE673EE7FFCBFA7E3047FE4EFB36E99F" box="[1072,1164,406,430]" degrees="36.82" direction="north" orientation="latitude" pageId="8" pageNumber="137" precision="555" value="36.82">36.82°N</geoCoordinate>
,
<geoCoordinate id="EE673EE7FFCBFA7E30D5FE4EFAB2E99F" box="[1186,1288,406,430]" degrees="102.53" direction="east" orientation="longitude" pageId="8" pageNumber="137" precision="555" value="102.53">102.53°E</geoCoordinate>
,
<location id="8E8C0EFBFFCBFA7E3169FE4EFC3AE9FE" LSID="urn:lsid:plazi:treatment:03FAE936FFCBFA7E375EFF29FB98EE7F:8E8C0EFBFFCBFA7E3169FE4EFC3AE9FE" country="China" latitude="36.82" longLatPrecision="711" longitude="102.53" name="Beishan Nat. Park" pageId="8" pageNumber="137">Beishan Nat. Park</location>
,
<quantity id="4CABF5C5FFCBFA7E37F8FE6FFC53E9FF" box="[911,1001,439,463]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="137" unit="m" value="2600.0">
<elevation id="007EBF13FFCBFA7E37F8FE6FFC53E9FF" box="[911,1001,439,463]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="137" unit="m" value="2600.0">2600 m</elevation>
</quantity>
, Picea-Pinus-Wald,
<collectorName id="26A63DF6FFCBFA7E30B9FE6FFCD5E9C1" pageId="8" pageNumber="137">Geröllhang. A. Gebauer</collectorName>
<date id="FFED7EE0FFCBFA7E3701FE0FFC48E9DE" box="[886,1010,471,495]" pageId="8" pageNumber="137" value="1996-05-30">
<collectingDate id="EFA98708FFCBFA7E3701FE0FFC48E9DE" box="[886,1010,471,495]" pageId="8" pageNumber="137" value="1996-05-30">1996.05.30</collectingDate>
</date>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3490BABFFCBFA7E375EFE20FB0DEDFB" pageId="8" pageNumber="137" type="diagnosis">
<paragraph id="8BEC5820FFCBFA7E375EFE20FB7AED17" blockId="8.[786,1399,307,1614]" pageId="8" pageNumber="137">
<emphasis id="B9278432FFCBFA7E375EFE20FC20EA21" bold="true" box="[809,922,504,528]" pageId="8" pageNumber="137">Diagnosis</emphasis>
: --Worker (
<figureCitation id="136844A5FFCBFA7E306BFE20FB2CEA20" box="[1052,1174,504,529]" captionStart="Figure 1417" captionStartId="9.[190,254,1204,1226]" captionTargetBox="[190,1437,227,1177]" captionTargetPageId="9" captionText="Figure 1417: Formicoxenus gebaueri n.sp., holotype; (14) head in dorsal view; (15) lateral view (flipped horizontally); (16) dorsal view; (17) dorsolateral view on eye. China: Beishan National Park, 1995.05.30" figureDoi="http://doi.org/10.5281/zenodo.13193936" httpUri="https://zenodo.org/record/13193936/files/figure.png" pageId="8" pageNumber="137">Figs 1417</figureCitation>
, key). Numeric data given are arithmetic means based on measurement of
<specimenCount id="9D5593A9FFCBFA7E3765FDE2FC18EA63" box="[786,930,570,594]" count="5" pageId="8" pageNumber="137" type="generic">5 specimens</specimenCount>
(for standard deviation, minimum and maximum values see
<tableCitation id="C6D16D9BFFCBFA7E3079FD83FBEFEA42" box="[1038,1109,603,627]" captionStart="Table 1" captionStartId="5.[189,242,246,267]" captionText="Table 1. Measurements of workers of the genera Leptothorax and Formicoxenus. Data are given in the sequence arithmetic mean ± standard deviation [minimum, maximum]." pageId="8" pageNumber="137">Tab. 1</tableCitation>
). Medium-sized, CS 738 µm.Head elongated, CL/CW 1.253. Genae in dorsal view nearly parallel. Median third of anterior margin of clypeus and hind margin of vertex in dorsal view straight. Frons narrower than in
<taxonomicName id="4C5323A3FFCBFA7E30FEFD07FAB3EAC6" baseAuthorityName="Forel" baseAuthorityYear="1899" box="[1161,1289,735,759]" class="Insecta" family="Formicidae" genus="Formicoxenus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="sibiricus">
<emphasis id="B9278432FFCBFA7E30FEFD07FAB3EAC6" box="[1161,1289,735,759]" italics="true" pageId="8" pageNumber="137">F. sibiricus</emphasis>
</taxonomicName>
(FRS/CS 0.432), scape slightly shorter (SL/CS 0.712. Eye very small, EYE/CS 0.164. Dorsal profile of promesonotum and propodeum slightly convex, metanotal depression moderately deep (MGr/CS 2.95 %). Spines rather short and acute (SP/CS 0.153), their bases moderately distant (SPBA/CS 0.289). Petiole with strong anterolateral corners, in profile without peduncle and with a less broad subpetiolar lobe. Postpetiolar sternite with a well-developed spine. All surfaces of head, mesosoma, waist and appendages matt due to a reticulate microsculpture. Dorsal vertex strongly longitudinally carinulate-rugulose. Gaster smooth and shiny but in contrast to
<taxonomicName id="4C5323A3FFCBFA7E3765FB53FC22EC93" baseAuthorityName="Forel" baseAuthorityYear="1899" box="[786,920,1162,1186]" class="Insecta" family="Formicidae" genus="Formicoxenus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="sibiricus">
<emphasis id="B9278432FFCBFA7E3765FB53FC22EC93" box="[786,920,1162,1186]" italics="true" pageId="8" pageNumber="137">F. sibiricus</emphasis>
</taxonomicName>
with subdecumbent to semierect setae. Head, mesosoma, waist, and gaster with long (PnHL/ CS 0.128) setae, tapering apicad (
<figureCitation id="136844A5FFCBFA7E30EBFB14FB54ECD5" box="[1180,1262,1228,1252]" captionStart="Figure 1417" captionStartId="9.[190,254,1204,1226]" captionTargetBox="[190,1437,227,1177]" captionTargetPageId="9" captionText="Figure 1417: Formicoxenus gebaueri n.sp., holotype; (14) head in dorsal view; (15) lateral view (flipped horizontally); (16) dorsal view; (17) dorsolateral view on eye. China: Beishan National Park, 1995.05.30" figureDoi="http://doi.org/10.5281/zenodo.13193936" httpUri="https://zenodo.org/record/13193936/files/figure.png" pageId="8" pageNumber="137">Fig. 17</figureCitation>
) and round in cross-section. Color of head, mesosoma, waist and gaster homogenously yellowish brown.
</paragraph>
<paragraph id="8BEC5820FFCBFA7E375EFAF6FB0DEDFB" blockId="8.[786,1399,307,1614]" pageId="8" pageNumber="137">
<emphasis id="B9278432FFCBFA7E375EFAF6FB96ED76" bold="true" box="[809,1068,1326,1351]" pageId="8" pageNumber="137">Taxonomic comments</emphasis>
. As a combination of long, tapering setae, well-developed sculpture on head, mesosoma, waist and appendages and narrower frons,
<taxonomicName id="4C5323A3FFCBFA7E3765FA4AFBA4ED98" authorityName="Seifert" authorityYear="2023" box="[786,1054,1425,1449]" class="Insecta" family="Formicidae" genus="Formicoxenus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="gebaueri" status="sp. nov.">
<emphasis id="B9278432FFCBFA7E3765FA4AFBA4ED98" box="[786,1054,1425,1449]" italics="true" pageId="8" pageNumber="137">Formicoxenus gebaueri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2143949FFCBFA7E305CFA4AFBE5ED98" box="[1067,1119,1426,1449]" pageId="8" pageNumber="137" rank="species">n.sp.</taxonomicNameLabel>
is unmistakable among the Palaearctic
<taxonomicName id="4C5323A3FFCBFA7E37CCFA6BFB0DEDFB" box="[955,1207,1458,1482]" class="Insecta" family="Formicidae" genus="Formicoxenus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="137" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9278432FFCBFA7E37CCFA6BFBE2EDFB" box="[955,1112,1459,1482]" italics="true" pageId="8" pageNumber="137">Formicoxenus</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3490BABFFCBFA7E375EFA0BFB98EE7F" pageId="8" pageNumber="137" type="biology_ecology">
<paragraph id="8BEC5820FFCBFA7E375EFA0BFB98EE7F" blockId="8.[786,1399,307,1614]" pageId="8" pageNumber="137">
<emphasis id="B9278432FFCBFA7E375EFA0BFC3BEDDA" bold="true" box="[809,897,1491,1515]" pageId="8" pageNumber="137">Biology</emphasis>
. The
<typeStatus id="54E8E682FFCBFA7E37C9FA0BFC55EDDA" box="[958,1007,1491,1515]" pageId="8" pageNumber="137">type</typeStatus>
sample was collected below a stone within a
<emphasis id="B9278432FFCBFA7E3703FA2DFC41EE3D" box="[884,1019,1525,1548]" italics="true" pageId="8" pageNumber="137">Picea-Pinus</emphasis>
forest on a slope with rubble. The host species was an undescribed species of the subgenus
<taxonomicName id="4C5323A3FFCBFA7E3765F9EEFBA7EE7F" authority="Forel 1913" authorityName="Forel" authorityYear="1913" box="[786,1053,1590,1614]" pageId="8" pageNumber="137" rank="subGenus" subGenus="Serviformica">
<emphasis id="B9278432FFCBFA7E3765F9EEFC18EE7F" box="[786,930,1590,1614]" italics="true" pageId="8" pageNumber="137">Serviformica</emphasis>
Forel 1913
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>