311 lines
40 KiB
XML
311 lines
40 KiB
XML
<document id="1C662B24F28240906E2BFE0E2DE912CA" ID-DOI="10.11646/zootaxa.5260.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7794920" ID-ZooBank="7171477F-DB75-4CF6-9507-3329F5D7A9F7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1680527871645" checkinUser="plazi" docAuthor="Blake, James A. & Maciolek, Nancy J." docDate="2023" docId="03F1A22EFF93CB19FF7282C3FD50FD52" docLanguage="en" docName="zootaxa.5260.1.1.pdf" docOrigin="Zootaxa 5260 (1)" docSource="http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heterospio paulolanai Blake & Maciolek 2023, new species" docType="treatment" docUuid="9D2C1C8E-4088-4EA7-B564-EAFB7B64AA2E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="34" masterDocId="FFC8DA56FF8CCB38FFE58038FFEAFFA3" masterDocTitle="New species and records of Heterospio (Annelida, Longosomatidae) from continental shelf, slope and abyssal depths of the Atlantic Ocean, Pacific Ocean, Indian Ocean and adjacent seas" masterLastPageNumber="74" masterPageNumber="1" pageNumber="32" updateTime="1699481072705" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="EF06AA4E5D4EF49475F1881A96A56294">New species and records of Heterospio (Annelida, Longosomatidae) from continental shelf, slope and abyssal depths of the Atlantic Ocean, Pacific Ocean, Indian Ocean and adjacent seas</mods:title>
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<mods:roleTerm id="83FBE92039553A13170C7EF57D395FC4">Author</mods:roleTerm>
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<mods:namePart id="289297E55947D3B561C9E492468CAF7A">Blake, James A.</mods:namePart>
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<mods:nameIdentifier id="8C22BBF773106E2C80E8871D89F5C846" type="email">jablake9@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="5A04E1517E8F2D18DEA2EF905E30D363">Author</mods:roleTerm>
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<mods:namePart id="114DB1B45AE7E08BB032A3B384DDC627">Maciolek, Nancy J.</mods:namePart>
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<mods:nameIdentifier id="5097E67A09CE6E932EEB72957F9C7B5D" type="email">njmaciolek@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="E138155FE0CBC4B6947FEE270FD75700">text</mods:typeOfResource>
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<mods:title id="D6C9D6E9CDE62868E1A92A4E794062CA">Zootaxa</mods:title>
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<mods:part id="D97DB62408D5AC9FAEB388A6D120FEA0">
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<mods:date id="341E8FF3507FE0995DF64B1F5B48CA21">2023</mods:date>
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<mods:number id="09D982E0A66204DF96231E449D6AF4C2">2023-04-03</mods:number>
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<mods:number id="1285E7E993DBE8D77FC29016F0ED7FF6">5260</mods:number>
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<mods:detail id="C17FBF5BDAB2A8490F188BF380241750" type="issue">
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<mods:number id="EFB16A321F19CBB4E1E11BD9CA15CF96">1</mods:number>
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<mods:url id="24CF1AB68801CA094C3CA9709409890F">http://dx.doi.org/10.1094/PDIS-04-22-0755-PDN</mods:url>
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<mods:classification id="B7A3C3BD31F58728BABC65E51373CDFF">journal article</mods:classification>
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<mods:identifier id="3D0FA37B301941FA7DD7430BE7DFEB08" type="DOI">10.11646/zootaxa.5260.1.1</mods:identifier>
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<mods:identifier id="0D013A6C2274D28261B3DA4561502D27" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="0025B9F5E166B42F1EA64A941CE52870" type="ZooBank">7171477F-DB75-4CF6-9507-3329F5D7A9F7</mods:identifier>
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<treatment id="03F1A22EFF93CB19FF7282C3FD50FD52" ID-DOI="http://doi.org/10.5281/zenodo.7798811" ID-Zenodo-Dep="7798811" LSID="urn:lsid:zoobank.org:act:9D2C1C8E-4088-4EA7-B564-EAFB7B64AA2E" httpUri="http://treatment.plazi.org/id/03F1A22EFF93CB19FF7282C3FD50FD52" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
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<subSubSection id="C34240B3FF93CB27FF7282C3FDD6FCB7" box="[151,572,762,789]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8BE71338FF93CB27FF7282C3FDD6FCB7" blockId="31.[151,985,762,861]" box="[151,572,762,789]" pageId="31" pageNumber="32">
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<heading id="D0AFA454FF93CB27FF7282C3FDD6FCB7" bold="true" box="[151,572,762,789]" fontSize="11" level="2" pageId="31" pageNumber="32" reason="4">
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<emphasis id="B92CCF2AFF93CB27FF7282C3FDD6FCB7" bold="true" box="[151,572,762,789]" pageId="31" pageNumber="32">
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<taxonomicName id="4C5868BBFF93CB27FF7282C3FE4FFCB6" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[151,421,762,789]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="31" pageNumber="32" phylum="Annelida" rank="species" species="paulolanai" status="sp. nov.">
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<emphasis id="B92CCF2AFF93CB27FF7282C3FE4FFCB6" bold="true" box="[151,421,762,789]" italics="true" pageId="31" pageNumber="32">Heterospio paulolanai</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A21F7251FF93CB27FE4982C3FDD6FCB7" box="[428,572,762,789]" pageId="31" pageNumber="32" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34240B3FF93CB27FF728327FC33FCFE" pageId="31" pageNumber="32" type="description">
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<paragraph id="8BE71338FF93CB27FF728327FEE9FC9A" blockId="31.[151,985,762,861]" box="[151,259,799,825]" pageId="31" pageNumber="32">
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<figureCitation id="13630FBDFF93CB27FF728327FEE9FC9A" box="[151,259,799,825]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Figure 13</figureCitation>
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</paragraph>
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<paragraph id="8BE71338FF93CB27FF72837AFC33FCFE" blockId="31.[151,985,762,861]" box="[151,985,834,861]" pageId="31" pageNumber="32">
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<uri id="FFC91F3AFF93CB27FF72837AFC33FCFE" box="[151,985,834,861]" pageId="31" pageNumber="32">
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urn:lsid:zoobank.org:act:
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<uuid id="FFFE29EDFF93CB27FE49837AFC33FCFE" box="[428,985,834,861]" pageId="31" pageNumber="32">9D2C1C8E-4088-4EA7-B564-EAFB7B64AA2E</uuid>
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</uri>
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</subSubSection>
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<subSubSection id="C34240B3FF93CB27FF7283B2FC07FC6A" pageId="31" pageNumber="32" type="materials_examined">
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<paragraph id="8BE71338FF93CB27FF7283B2FC07FC6A" blockId="31.[151,1437,906,1977]" pageId="31" pageNumber="32">
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<emphasis id="B92CCF2AFF93CB27FF7283B2FE93FC07" bold="true" box="[151,377,906,932]" pageId="31" pageNumber="32">Material examined</emphasis>
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.
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<materialsCitation id="3B301965FF93CB27FE6283B2FCA1FC07" box="[391,843,906,932]" country="Brazil" location="South Atlantic Ocean" pageId="31" pageNumber="32" specimenCount="1" stateProvince="South Atlantic Ocean">
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<emphasis id="B92CCF2AFF93CB27FE6283B2FCA1FC07" bold="true" box="[391,843,906,932]" pageId="31" pageNumber="32">
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<collectingRegion id="499CDDDAFF93CB27FE6283B2FD6DFC07" box="[391,647,906,932]" name="South Atlantic Ocean" pageId="31" pageNumber="32">South Atlantic Ocean</collectingRegion>
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,
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<collectingCountry id="F34F53A8FF93CB27FD7683B2FD34FC07" box="[659,734,906,932]" name="Brazil" pageId="31" pageNumber="32">Brazil</collectingCountry>
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, offshore
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</emphasis>
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</materialsCitation>
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<materialsCitation id="3B301965FF93CB27FCB783B2FC03FC6A" collectingDate="1985-11-14" collectionCode="MZUSP" collectorName="Sueste IV & Sta." country="Brazil" elevation="69" latitude="-24.735" location="Sao Paulo" longLatPrecision="74" longitude="-45.917" pageId="31" pageNumber="32" specimenCode="MZUSP 5588" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
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<emphasis id="B92CCF2AFF93CB27FCB783B2FB54FC07" bold="true" box="[850,1214,906,933]" pageId="31" pageNumber="32">
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<collectingRegion id="499CDDDAFF93CB27FCB783B2FC21FC06" box="[850,971,906,933]" country="Brazil" name="Sao Paulo" pageId="31" pageNumber="32">São Paulo</collectingRegion>
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,
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<collectorName id="26AD76EEFF93CB27FC3283B2FBAFFC06" box="[983,1093,906,933]" pageId="31" pageNumber="32">
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<emphasis id="B92CCF2AFF93CB27FC3283B2FBC8FC07" bold="true" box="[983,1058,906,932]" italics="true" pageId="31" pageNumber="32">Súeste</emphasis>
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IV
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</collectorName>
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,
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<collectorName id="26AD76EEFF93CB27FBB483B2FB95FC07" box="[1105,1151,906,932]" pageId="31" pageNumber="32">Sta.</collectorName>
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6672
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</emphasis>
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, coll.
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<date id="FFE635F8FF93CB27FAE183B3FA72FC06" box="[1284,1432,906,933]" pageId="31" pageNumber="32" value="1985-11-14">
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<collectingDate id="EFA2CC10FF93CB27FAE183B3FA72FC06" box="[1284,1432,906,933]" pageId="31" pageNumber="32" value="1985-11-14">14 Nov 1985</collectingDate>
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||
</date>
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||
,
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||
<geoCoordinate id="EE6C75FFFF93CB27FF728396FEE8FC6A" box="[151,258,942,969]" degrees="24.735" direction="south" orientation="latitude" pageId="31" pageNumber="32" precision="55" value="-24.735">24.735°S</geoCoordinate>
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,
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||
<geoCoordinate id="EE6C75FFFF93CB27FEE88396FE6AFC6A" box="[269,384,942,969]" degrees="45.917" direction="west" orientation="longitude" pageId="31" pageNumber="32" precision="55" value="-45.917">45.917°W</geoCoordinate>
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,
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<quantity id="4CA0BEDDFF93CB27FE6E8397FE2CFC6A" box="[395,454,943,969]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="31" pageNumber="32" unit="m" value="69.0">
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<elevation id="0075F40BFF93CB27FE6E8397FE2CFC6A" box="[395,454,943,969]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="31" pageNumber="32" unit="m" value="69.0">69 m</elevation>
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</quantity>
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, mud and shell gravel,
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<emphasis id="B92CCF2AFF93CB27FD2F8397FCDBFC6A" bold="true" box="[714,817,943,969]" pageId="31" pageNumber="32">
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||
<typeStatus id="54E3AD9AFF93CB27FD2F8397FCDBFC6A" box="[714,817,943,969]" pageId="31" pageNumber="32" type="holotype">holotype</typeStatus>
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</emphasis>
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||
(
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<specimenCode id="DBFEBB43FF93CB27FCDA8397FC0BFC6A" box="[831,993,942,969]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="31" pageNumber="32">MZUSP 5588</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34240B3FF93CB19FF2283EAFE4AFEBE" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="description">
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<paragraph id="8BE71338FF93CB27FF2283EAFB7CFB66" blockId="31.[151,1437,906,1977]" pageId="31" pageNumber="32">
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<emphasis id="B92CCF2AFF93CB27FF2283EAFEBBFC4F" bold="true" box="[199,337,978,1004]" pageId="31" pageNumber="32">Description</emphasis>
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.
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<typeStatus id="54E3AD9AFF93CB27FEBA83EBFE2DFC4E" box="[351,455,979,1005]" pageId="31" pageNumber="32" type="holotype">Holotype</typeStatus>
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(MZUSP 5588) only specimen, incomplete with 12 setigers, long, narrow,
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<quantity id="4CA0BEDDFF93CB27FAEA83EBFAB7FC4E" box="[1295,1373,979,1005]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="31" pageNumber="32" unit="mm" value="27.0">27 mm</quantity>
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long, 1.0 mm wide across thoracic setigers,
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<quantity id="4CA0BEDDFF93CB27FDB783CFFD41FBB2" box="[594,683,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="32" unit="mm" value="0.8">0.8 mm</quantity>
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wide across abdominal segments. Specimen with some segments (setigers 1, 9, and 11) damaged, partially dissected by a previous worker. Thoracic region with nine setigers, 1–8 short, 9 transitional, about 2.5 times as long as setiger 8 (
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<figureCitation id="13630FBDFF93CB27FCC58407FC60FBFA" box="[800,906,1087,1113]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13A</figureCitation>
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). Thoracic segments with defined dorso-lateral setal fascicles transitioning to abdominal segments with setae arising from broad cincture-like fascicles with rows of setae mostly surrounding anterior margin of each segment, but with wide dorsal and ventral gaps. Abdominal segments 10–12 very long, each longer than entire thoracic region. Color in alcohol brown.
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</paragraph>
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<paragraph id="8BE71338FF93CB27FF2284F7FA82FA46" blockId="31.[151,1437,906,1977]" pageId="31" pageNumber="32">
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Pre-setiger region triangular in shape, about as long as first two thoracic setigers (
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<figureCitation id="13630FBDFF93CB27FB9684F7FB14FB4A" box="[1139,1278,1231,1257]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13A–B</figureCitation>
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). Prostomium rhomboid-shaped, tapering to rounded tip (
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<figureCitation id="13630FBDFF93CB27FD8984CBFD38FAAE" box="[620,722,1267,1293]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13A</figureCitation>
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); eyespots absent, nuchal organs narrow slits laterally at posterior junction of prostomium and peristomium (
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<figureCitation id="13630FBDFF93CB27FD8B852FFD3CFA92" box="[622,726,1303,1329]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13A</figureCitation>
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). Peristomium with two dorsolateral rings apparent dorsally and laterally, separated by groove from which dorsal tentacles arise medially; no tentacles present, but scars evident in grooves. Peristomium incomplete dorsally, interrupted by posterior extension of prostomium merging with dorsal surface, dorsal crest lacking. Ventral surface of peristomium broad a single broad surface, anteriorly encompassing mouth, merging posteriorly with broad ventral surface of thoracic segments (
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||
<figureCitation id="13630FBDFF93CB27FBF6859FFB96FA62" box="[1043,1148,1447,1473]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13B</figureCitation>
|
||
). Mouth a transverse slit consisting of simple opening between two large lateral lobes and simple anterior and posterior lips (
|
||
<figureCitation id="13630FBDFF93CB27FB1085F3FAB1FA46" box="[1269,1371,1483,1509]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE71338FF93CB27FF2285D7FB9FF9F2" blockId="31.[151,1437,906,1977]" pageId="31" pageNumber="32">
|
||
Branchiae or their stubs present on setigers 2–8 (
|
||
<figureCitation id="13630FBDFF93CB27FD1585D7FCB2F9AA" box="[752,856,1519,1545]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13A</figureCitation>
|
||
), not observed on setiger 9 possibly due to damage from an earlier dissection; most branchiae when present long, thin, rounded in cross section, tapering to rounded tip; branchiae with narrow ciliated groove; internal blood vessel extends along entire length.
|
||
</paragraph>
|
||
<paragraph id="8BE71338FF93CB27FF228663FCBAF836" blockId="31.[151,1437,906,1977]" pageId="31" pageNumber="32">
|
||
All parapodia biramous with setae emerging from near anterior edge of each segment (
|
||
<figureCitation id="13630FBDFF93CB27FB7E8663FACFF9D6" box="[1179,1317,1627,1653]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13A–B</figureCitation>
|
||
). Thoracic setigers with notches from which dense setal fascicles in dorsolateral locations arise on each segment; postsetal lamellae not observed, but short elevated ridge present on posterior margin of notch from which thoracic neurosetae arise; thoracic notosetae with numerous capillaries (25+) in dense fascicle with distinct rows not defined; neurosetae similarly arranged with capillaries even more numerous (~60+). Abdominal setigers from setiger 10 with setae arranged in long double rows nearly surrounding body, with relatively wide dorsal and ventral gaps. Setiger 10 with all capillaries arranged in two rows; setiger 11 with anterior row of aristate spines (
|
||
<figureCitation id="13630FBDFF93CB27FB75870BFB1FF8EE" box="[1168,1269,1843,1869]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13C</figureCitation>
|
||
) and posterior row consisting of thickened capillaries (
|
||
<figureCitation id="13630FBDFF93CB27FDBA876FFD23F8D2" box="[607,713,1879,1905]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13D</figureCitation>
|
||
); setiger 12 with anterior row of acicular spines (
|
||
<figureCitation id="13630FBDFF93CB27FB1B876FFA88F8D2" box="[1278,1378,1879,1905]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13E</figureCitation>
|
||
) and posterior row of thick, curved, pointed subuluncini (
|
||
<figureCitation id="13630FBDFF93CB27FD058743FCAEF836" box="[736,836,1915,1941]" captionStart="FIGURE 13" captionStartId="32.[151,250,1820,1844]" captionTargetBox="[162,1419,196,1783]" captionTargetId="figure-18@32.[151,1436,181,1796]" captionTargetPageId="32" captionText="FIGURE 13. Heterospio paulolanai n. sp. Holotype (MZUSP 5588):A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12." figureDoi="http://doi.org/10.5281/zenodo.7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="31" pageNumber="32">Fig. 13F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE71338FF93CB27FF2287A7FDEFF81A" blockId="31.[151,1437,906,1977]" box="[199,517,1951,1977]" pageId="31" pageNumber="32">Posterior region not present.</paragraph>
|
||
<caption id="DF2743B0FFACCB18FF728724FCDFF8DE" ID-DOI="http://doi.org/10.5281/zenodo.7794946" ID-Zenodo-Dep="7794946" httpUri="https://zenodo.org/record/7794946/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,250,1820,1844]" targetBox="[162,1419,196,1783]" targetPageId="32">
|
||
<paragraph id="8BE71338FFACCB18FF728724FCDFF8DE" blockId="32.[151,1437,1820,1917]" pageId="32" pageNumber="33">
|
||
<emphasis id="B92CCF2AFFACCB18FF728724FEF2F897" bold="true" box="[151,280,1820,1844]" pageId="32" pageNumber="33">FIGURE 13</emphasis>
|
||
.
|
||
<taxonomicName id="4C5868BBFFACCB18FEC18725FDEDF897" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[292,519,1820,1844]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="32" pageNumber="33" phylum="Annelida" rank="species" species="paulolanai" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFACCB18FEC18725FDEDF897" box="[292,519,1820,1844]" italics="true" pageId="32" pageNumber="33">Heterospio paulolanai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFACCB18FDE98725FDA9F897" box="[524,579,1821,1845]" pageId="32" pageNumber="33" rank="species">
|
||
<emphasis id="B92CCF2AFFACCB18FDE98725FDD7F896" bold="true" box="[524,573,1821,1845]" pageId="32" pageNumber="33">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Holotype (MZUSP 5588): A, anterior end (setiger 1–10), dorsal view; B, anterior end (setiger 1–5), ventral view; C, neuropodial aristate spines, setiger 11; D, neuropodial capillary seta, setiger 11; E, neuropodial acicular spine, setiger 12; F, neuropodial subuluncinus, setiger 12.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE71338FFADCB19FF2280AFFE4AFEBE" blockId="33.[151,1437,150,753]" pageId="33" pageNumber="34">
|
||
<emphasis id="B92CCF2AFFADCB19FF2280AFFE0AFF13" bold="true" box="[199,480,150,177]" pageId="33" pageNumber="34">Methyl Green staining</emphasis>
|
||
. Methyl Green producing prominent pattern on pre-setiger and anterior thoracic segments. Stain concentrates on posterior and lateral margins of prostomium and both peristomial rings on dorsal and lateral surfaces. Stain also concentrates laterally on setigers 1–5 producing dark-green parapodia. Rest of body lacking distinct pattern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34240B3FFADCB19FF22811FFA68FDC2" pageId="33" pageNumber="34" type="discussion">
|
||
<paragraph id="8BE71338FFADCB19FF22811FFA68FDC2" blockId="33.[151,1437,150,753]" pageId="33" pageNumber="34">
|
||
<emphasis id="B92CCF2AFFADCB19FF22811FFEDEFEE2" bold="true" box="[199,308,295,321]" pageId="33" pageNumber="34">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C5868BBFFADCB19FEA4811FFDD6FEE2" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[321,572,295,321]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="paulolanai" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FEA4811FFDD6FEE2" box="[321,572,295,321]" italics="true" pageId="33" pageNumber="34">Heterospio paulolanai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FDA6811FFD6BFEE2" box="[579,641,295,321]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FDA6811FFD90FEE2" bold="true" box="[579,634,295,321]" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
belongs to a group of
|
||
<taxonomicName id="4C5868BBFFADCB19FC9F811FFC1FFEE3" authorityName="Ehlers" authorityYear="1874" box="[890,1013,295,320]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="genus">
|
||
<emphasis id="B92CCF2AFFADCB19FC9F811FFC1FFEE3" box="[890,1013,295,320]" italics="true" pageId="33" pageNumber="34">Heterospio</emphasis>
|
||
</taxonomicName>
|
||
having a narrow conical prostomium, two peristomial rings, 7–8 pairs of branchiae, and setiger 9 as the first elongate segment about 2–3 times longer than setiger 8. Of these,
|
||
<taxonomicName id="4C5868BBFFADCB19FE958157FDE1FE2A" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[368,523,367,393]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="paulolanai" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FE958157FDE1FE2A" box="[368,523,367,393]" italics="true" pageId="33" pageNumber="34">H. paulolanai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FDF78157FDBAFE2A" box="[530,592,367,393]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FDF78157FDA3FE2A" bold="true" box="[530,585,367,393]" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is most similar to
|
||
<taxonomicName id="4C5868BBFFADCB19FCFB8157FC69FE2A" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[798,899,367,393]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="aruba" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FCFB8157FC69FE2A" box="[798,899,367,393]" italics="true" pageId="33" pageNumber="34">H. aruba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FC6F8157FC22FE2A" box="[906,968,367,393]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FC6F8157FC2BFE2A" bold="true" box="[906,961,367,393]" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C5868BBFFADCB19FC338157FBACFE2B" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[982,1094,367,392]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="guiana" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FC338157FBACFE2B" box="[982,1094,367,392]" italics="true" pageId="33" pageNumber="34">H. guiana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FBA88157FB60FE2A" box="[1101,1162,367,393]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FBA88157FB6EFE2A" bold="true" box="[1101,1156,367,393]" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="4C5868BBFFADCB19FB2D8157FAC7FE2B" authorityName="Wu & Chen" authorityYear="1960" box="[1224,1325,367,392]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="sinica">
|
||
<emphasis id="B92CCF2AFFADCB19FB2D8157FAC7FE2B" box="[1224,1325,367,392]" italics="true" pageId="33" pageNumber="34">H. sinica</emphasis>
|
||
</taxonomicName>
|
||
in having all capillaries on setiger 10 and following setigers with at least a few aristate spines. However,
|
||
<taxonomicName id="4C5868BBFFADCB19FB2481ABFACCFE0E" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[1217,1318,403,429]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="aruba" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FB2481ABFACCFE0E" box="[1217,1318,403,429]" italics="true" pageId="33" pageNumber="34">H. aruba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FAC881ABFA87FE0E" box="[1325,1389,403,429]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FAC881ABFA8FFE0E" bold="true" box="[1325,1381,403,429]" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C5868BBFFADCB19FF72818FFED8FE72" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[151,306,439,465]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="paulolanai" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FF72818FFED8FE72" box="[151,306,439,465]" italics="true" pageId="33" pageNumber="34">H. paulolanai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FEDD818FFE9CFE72" box="[312,374,439,465]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FEDD818FFE85FE72" bold="true" box="[312,367,439,465]" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
also have acicular spines that are lacking in
|
||
<taxonomicName id="4C5868BBFFADCB19FC87818FFC3BFE73" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[866,977,439,464]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="guiana" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FC87818FFC3BFE73" box="[866,977,439,464]" italics="true" pageId="33" pageNumber="34">H. guiana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FC32818FFBFFFE72" box="[983,1045,439,465]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FC32818FFBE4FE72" bold="true" box="[983,1038,439,465]" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C5868BBFFADCB19FBAF818FFB44FE73" authorityName="Wu & Chen" authorityYear="1960" box="[1098,1198,439,464]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="sinica">
|
||
<emphasis id="B92CCF2AFFADCB19FBAF818FFB44FE73" box="[1098,1198,439,464]" italics="true" pageId="33" pageNumber="34">H. sinica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C5868BBFFADCB19FB5E818FFA6AFE72" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[1211,1408,439,465]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="aruba" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FB5E818FFA6AFE72" box="[1211,1408,439,465]" italics="true" pageId="33" pageNumber="34">Heterospio aruba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FA63818FFF53FE56" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FA63818FFF5BFE56" bold="true" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C5868BBFFADCB19FEAF81E3FE0EFE56" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[330,484,475,501]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="paulolanai" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FEAF81E3FE0EFE56" box="[330,484,475,501]" italics="true" pageId="33" pageNumber="34">H. paulolanai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FE0F81E3FDC3FE56" box="[490,553,475,501]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FE0F81E3FDCBFE56" bold="true" box="[490,545,475,501]" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
in having prominent postsetal lamellae on thoracic neuropodia, an oral opening that is a narrow transverse slit with no obvious lobes or surrounding papillae, and a distinct ventral ridge that extends from the mouth posteriorly over the peristomium and first seven setigers. In contrast,
|
||
<taxonomicName id="4C5868BBFFADCB19FB80821BFAE8FD9E" authority="Blake & Maciolek, 2023" authorityName="Blake & Maciolek" authorityYear="2023" box="[1125,1282,547,573]" class="Polychaeta" family="Longosomatidae" genus="Heterospio" kingdom="Animalia" order="Spionida" pageId="33" pageNumber="34" phylum="Annelida" rank="species" species="paulolanai" status="sp. nov.">
|
||
<emphasis id="B92CCF2AFFADCB19FB80821BFAE8FD9E" box="[1125,1282,547,573]" italics="true" pageId="33" pageNumber="34">H. paulolanai</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21F7251FFADCB19FAEF821BFAA1FD9E" box="[1290,1355,547,573]" pageId="33" pageNumber="34" rank="species">
|
||
<emphasis id="B92CCF2AFFADCB19FAEF821BFAA9FD9E" bold="true" box="[1290,1347,547,573]" pageId="33" pageNumber="34">n. sp</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
has no obvious thoracic neuropodial lamellae, an oral opening bordered by two large lateral lobes, and no ventral ridge.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34240B3FFADCB19FF228253FAC6FD6E" pageId="33" pageNumber="34" type="etymology">
|
||
<paragraph id="8BE71338FFADCB19FF228253FAC6FD6E" blockId="33.[151,1437,150,753]" pageId="33" pageNumber="34">
|
||
<emphasis id="B92CCF2AFFADCB19FF228253FEA2FD26" bold="true" box="[199,328,619,645]" pageId="33" pageNumber="34">Etymology</emphasis>
|
||
. This species is named for the late Professor Paulo da
|
||
<collectingCountry id="F34F53A8FFADCB19FC538253FC15FD26" box="[950,1023,619,645]" name="Saint Helena, Ascension and Tristan da Cunha" pageId="33" pageNumber="34">Cunha</collectingCountry>
|
||
Lana, prominent Brazilian authority on annelids. Prof. Lana was host of the sixth International Polychaete Conference held in Curitiba,
|
||
<collectingCountry id="F34F53A8FFADCB19FB0482B7FACFFD0A" box="[1249,1317,655,681]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
|
||
(
|
||
<date id="FFE635F8FFADCB19FAD682B7FF3AFD6E" pageId="33" pageNumber="34" value="1998-08-02" valueMax="1998-08-07" valueMin="1998-08-02">Aug 2–7, 1998</date>
|
||
) and later elected the ninth President of the International Polychaetology Association (2010–2013).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34240B3FFADCB19FF2282EEFD50FD52" box="[199,698,726,753]" pageId="33" pageNumber="34" type="distribution">
|
||
<paragraph id="8BE71338FFADCB19FF2282EEFD50FD52" blockId="33.[151,1437,150,753]" box="[199,698,726,753]" pageId="33" pageNumber="34">
|
||
<emphasis id="B92CCF2AFFADCB19FF2282EEFEB3FD53" bold="true" box="[199,345,726,752]" pageId="33" pageNumber="34">Distribution</emphasis>
|
||
. Off southeastern
|
||
<collectingCountry id="F34F53A8FFADCB19FDCD82EFFD85FD52" box="[552,623,727,753]" name="Brazil" pageId="33" pageNumber="34">Brazil</collectingCountry>
|
||
,
|
||
<specimenCount id="9D5ED8B1FFADCB19FD9F82EFFD50FD52" box="[634,698,727,753]" pageId="33" pageNumber="34" type="generic">
|
||
<quantity id="4CA0BEDDFFADCB19FD9F82EFFD5CFD52" box="[634,694,727,753]" metricMagnitude="1" metricUnit="m" metricValue="6.9" pageId="33" pageNumber="34" unit="m" value="69.0">69 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |