732 lines
92 KiB
XML
732 lines
92 KiB
XML
<document id="847897C88D802BBE04E7F4BE77DD6C08" ID-CLB-Dataset="25304" ID-DOI="10.5852/ejt.2019.561" ID-GBIF-Dataset="f0779c03-8e65-424d-b680-ff8f219fd345" ID-Zenodo-Dep="3474359" ID-ZooBank="6C81FC6F-2CDD-46B8-9825-6D09DF9475F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570396803884" checkinUser="plazi" docAuthor="Prokofiev, Artem M." docDate="2019" docId="03E72F184000FFF9FD9857AFFAECFD43" docLanguage="en" docName="ejt-561_prokofiev.pdf.imf" docOrigin="European Journal of Taxonomy 561" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Philacelota leucothea Prokofiev 2019, sp. nov." docType="treatment" docUuid="BF182F51-5D7C-4F3E-937D-ED8458AADF21" docUuidSource="ZooBank" docVersion="8" lastPageNumber="12" masterDocId="FFDE57604005FFF2FFFE5345FF9AFFE5" masterDocTitle="Two new species of Engertia Dalla Torre, 1913 and Philacelota Heller, 1900 from Indonesia and the Philippines with a revised identification key (Coleoptera: Scarabaeidae: Melolonthinae)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="6" updateTime="1698757900835" updateUser="ExternalLinkService">
|
||
<mods:mods id="DA2BAB1C9312DD8EDF66C6F306053F34" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3946E18E63B2561840479D0E7FED4441">
|
||
<mods:title id="C4FB0D7CDF7708AB50AC6D5CE9F88CB4">Two new species of Engertia Dalla Torre, 1913 and Philacelota Heller, 1900 from Indonesia and the Philippines with a revised identification key (Coleoptera: Scarabaeidae: Melolonthinae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="DA361D839CE47D5874F0058D79042C5B" type="personal">
|
||
<mods:role id="7709EBB1F4FA28E9FAE984A90C7C5E7D">
|
||
<mods:roleTerm id="0DC68E92E9B0AE3D4136C8D2F340E0D8">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2FD58F1B4179DF5F2B62130C16A69EDC">Prokofiev, Artem M.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="25B5F6C71DC37F01B3C0777A061DD600">text</mods:typeOfResource>
|
||
<mods:relatedItem id="1AAC818465937545D14C4CC86543721F" type="host">
|
||
<mods:titleInfo id="FC115F3098B7E4E2B501B1A3ED727F26">
|
||
<mods:title id="8D42D8C7031D501A97646B908B0C3398">European Journal of Taxonomy</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="7AA0DD88D35EC717EC39D66C1E1F45A1">
|
||
<mods:date id="672AF4B34A83EB9DD79E586C0D11F2C2">2019</mods:date>
|
||
<mods:detail id="099B509C168370222D0BD2772777605C" type="pubDate">
|
||
<mods:number id="6F14C83DEC249ABD15E4D1184E55C188">2019-10-03</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="62BC4C80A01A3E8ABAAAD673F651FD17" type="volume">
|
||
<mods:number id="CAE906ED529F905FEEF0F1F4DFA464AE">561</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="85A4AF7BBBE735FAED9CFEA74ED96C3F" unit="page">
|
||
<mods:start id="EFB7CF964B14F10EA8314D6DA8973767">1</mods:start>
|
||
<mods:end id="CA8B98C2949F51C10908347E920F4073">13</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="A674AD4A8058EE90CDCDF9EF9ECD5305">journal article</mods:classification>
|
||
<mods:identifier id="5052B9F1C270B9B34CA2D32123FB92A9" type="CLB-Dataset">25304</mods:identifier>
|
||
<mods:identifier id="028E079A6E222E5B8BB23F5F005E4BD0" type="DOI">10.5852/ejt.2019.561</mods:identifier>
|
||
<mods:identifier id="42DC3C4BDC093A7A9025D80A0797DBF9" type="GBIF-Dataset">f0779c03-8e65-424d-b680-ff8f219fd345</mods:identifier>
|
||
<mods:identifier id="67575B16B124FBF0AEB8CD0F46771408" type="Zenodo-Dep">3474359</mods:identifier>
|
||
<mods:identifier id="B6C8B2B98F12A23CF565A89E8568160C" type="ZooBank">6C81FC6F-2CDD-46B8-9825-6D09DF9475F3</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03E72F184000FFF9FD9857AFFAECFD43" ID-DOI="http://doi.org/10.5281/zenodo.3475273" ID-GBIF-Taxon="159647023" ID-Zenodo-Dep="3475273" LSID="urn:lsid:zoobank.org:act:BF182F51-5D7C-4F3E-937D-ED8458AADF21" httpUri="http://treatment.plazi.org/id/03E72F184000FFF9FD9857AFFAECFD43" lastPageId="11" lastPageNumber="12" pageId="5" pageNumber="6">
|
||
<subSubSection id="C354CD854000FFF7FD9857AFFB50FACF" pageId="5" pageNumber="6" type="nomenclature">
|
||
<paragraph id="8BF19E0E4000FFF7FD9857AFFC54FAE0" blockId="5.[362,1226,1258,1359]" box="[614,974,1258,1285]" pageId="5" pageNumber="6">
|
||
<heading id="D0B929624000FFF7FD9857AFFC54FAE0" box="[614,974,1258,1285]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="6" reason="2">
|
||
<taxonomicName id="4C4EE58D4000FFF7FD9857AFFCF1FAE0" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[614,875,1258,1285]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="leucothea" status="sp. nov.">
|
||
<emphasis id="B93A421C4000FFF7FD9857AFFCF1FAE0" bold="true" box="[614,875,1258,1285]" italics="true" pageId="5" pageNumber="6">Philacelota leucothea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF674000FFF7FC8D57AEFC54FAE0" box="[883,974,1259,1285]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FE94564AFB50FACF" blockId="5.[362,1226,1258,1359]" box="[362,1226,1295,1322]" pageId="5" pageNumber="6">
|
||
<uri id="FFDF920C4000FFF7FE94564AFB50FACF" box="[362,1226,1295,1322]" pageId="5" pageNumber="6">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FFE8A4DB4000FFF7FD76564AFB50FACF" box="[648,1226,1295,1322]" pageId="5" pageNumber="6">BF182F51-5D7C-4F3E-937D-ED8458AADF21</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354CD854000FFF7FD425671FCEDFAAA" box="[700,887,1332,1359]" pageId="5" pageNumber="6" type="description">
|
||
<paragraph id="8BF19E0E4000FFF7FD425671FCEDFAAA" blockId="5.[362,1226,1258,1359]" box="[700,887,1332,1359]" pageId="5" pageNumber="6">
|
||
<figureCitation id="1375828B4000FFF7FD425671FCBFFAAB" box="[700,805,1332,1358]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[189,232,1965,1991]" captionStartId-1="7.[189,232,1894,1920]" captionTargetBox-0="[277,1302,936,1922]" captionTargetBox-1="[276,1320,722,1829]" captionTargetId-0="figure@6.[260,1322,924,1928]" captionTargetId-1="figure@7.[265,1333,722,1841]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 3. Philacelota leucothea sp. nov., habitus. A. ♂, holotype (MNHN). B. ♀, paratype (MNHN). Scale bars: 2.5 mm." captionText-1="Fig. 4. Philacelota leucothea sp. nov., holotype, ♂ (MNHN) (A–C) and paratype, ♀ (MNHN) (D– G). A. Antenna, lateral view. B. Propygidium. C. Pygidium. D. Antenna, lateral view. E. Antenna, dorsolateral view showing relative size of club antennomeres. F. Propygidium. G. Pygidium. Scale bars: A, C–E = 1.5 mm; B = 2 mm; F–G = 2.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.3474365" figureDoi-1="http://doi.org/10.5281/zenodo.3474367" httpUri-0="https://zenodo.org/record/3474365/files/figure.png" httpUri-1="https://zenodo.org/record/3474367/files/figure.png" pageId="5" pageNumber="6">Figs 3–4</figureCitation>
|
||
,
|
||
<figureCitation id="1375828B4000FFF7FCCF5671FCA4FAAA" box="[817,830,1332,1359]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[202,1387,1426,1882]" captionTargetId="figure@8.[189,1396,1422,1886]" captionTargetPageId="8" captionText="Fig. 5. A–B. Philacelota leucothea sp. nov., holotype, ♂ (MNHN). A. Aedeagus, lateral view. B. Parameres, dorsal view. — C–D. P. submaculata Heller, 1900, from Kema, Sulawesi, ♂ (MNHN). C. Aedeagus, lateral view. D. Parameres, dorsal view. Scale bars: A–B = 1.5 mm; C–D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3474369" httpUri="https://zenodo.org/record/3474369/files/figure.png" pageId="5" pageNumber="6">5</figureCitation>
|
||
A–B
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354CD854000FFF7FF435632FAEDF9B1" pageId="5" pageNumber="6" type="diagnosis">
|
||
<paragraph id="8BF19E0E4000FFF7FF435632FEACFA74" blockId="5.[189,310,1399,1425]" box="[189,310,1399,1425]" pageId="5" pageNumber="6">
|
||
<heading id="D0B929624000FFF7FF435632FEACFA74" bold="true" box="[189,310,1399,1425]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="B93A421C4000FFF7FF435632FEACFA74" bold="true" box="[189,310,1399,1425]" pageId="5" pageNumber="6">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FF4356E2FAEDF9B1" blockId="5.[189,1399,1447,1620]" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C4EE58D4000FFF7FF4356E2FEDEFA24" authorityName="Heller" authorityYear="1900" box="[189,324,1447,1473]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93A421C4000FFF7FF4356E2FEDEFA24" box="[189,324,1447,1473]" italics="true" pageId="5" pageNumber="6">Philacelota</emphasis>
|
||
</taxonomicName>
|
||
with squamose vestiture. Disc of pronotum with narrow, longitudinal callose stripe. Elytra evenly scaled in males, while in females the scaled areas are intermixed with glabrous ones to form a variegated pattern. Propygidium of males with short acicular setae and sparse long setae, of females with short setae only; pygidium fully scaled in males, but with glabrous medial longitudinal stripe in females. Scales along midline of abdominal ventrites 2 to 4 evenly distributed. Aedeagus of characteristic shape.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354CD854000FFF7FF435538FB69F90E" pageId="5" pageNumber="6" type="etymology">
|
||
<paragraph id="8BF19E0E4000FFF7FF435538FEDEF972" blockId="5.[189,324,1661,1687]" box="[189,324,1661,1687]" pageId="5" pageNumber="6">
|
||
<heading id="D0B929624000FFF7FF435538FEDEF972" bold="true" box="[189,324,1661,1687]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="B93A421C4000FFF7FF435538FEDEF972" bold="true" box="[189,324,1661,1687]" pageId="5" pageNumber="6">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FF4355E8FB69F90E" blockId="5.[189,1398,1709,1772]" pageId="5" pageNumber="6">
|
||
The name of this new species reflects the white scale covering of the beetles.
|
||
<emphasis id="B93A421C4000FFF7FBD555E8FB3FF922" box="[1067,1189,1709,1735]" italics="true" pageId="5" pageNumber="6">Leucothea</emphasis>
|
||
(‘white goddess’) is a poetic epithet of a Greek goddess. The species epithet is treated as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354CD854000FFF7FF435450FBBDF80E" pageId="5" pageNumber="6" type="materials_examined">
|
||
<paragraph id="8BF19E0E4000FFF7FF435450FEF6F8CB" blockId="5.[189,364,1812,1839]" box="[189,364,1812,1839]" pageId="5" pageNumber="6">
|
||
<heading id="D0B929624000FFF7FF435450FEF6F8CB" bold="true" box="[189,364,1812,1839]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="B93A421C4000FFF7FF435450FEF6F8CB" bold="true" box="[189,364,1812,1839]" pageId="5" pageNumber="6">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FF145400FEC6F8BA" blockId="5.[189,996,1861,1924]" box="[234,348,1861,1887]" pageId="5" pageNumber="6">
|
||
<emphasis id="B93A421C4000FFF7FF145400FEC6F8BA" bold="true" box="[234,348,1861,1887]" pageId="5" pageNumber="6">
|
||
<typeStatus id="54F520AC4000FFF7FF145400FEC6F8BA" box="[234,348,1861,1887]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FF43542CFC7EF861" blockId="5.[189,996,1861,1924]" box="[189,996,1897,1924]" pageId="5" pageNumber="6">
|
||
<materialsCitation id="3B2694534000FFF7FF43542CFC7EF861" ID-GBIF-Occurrence="2424121668" box="[189,996,1897,1924]" collectionCode="MNHN" collectorName="Philippines Ch. Semper" country="Philippines" location="PHILIPPINES" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
|
||
<collectingCountry id="F359DE9E4000FFF7FF43542CFEF0F866" box="[189,362,1897,1923]" name="Philippines" pageId="5" pageNumber="6">PHILIPPINES</collectingCountry>
|
||
•
|
||
<specimenCount id="9D4855874000FFF7FE7D542FFE06F861" box="[387,412,1898,1924]" pageId="5" pageNumber="6" type="male">♂</specimenCount>
|
||
; “N O Luzon /
|
||
<collectorName id="26BBFBD84000FFF7FDAE542CFCF6F866" box="[592,876,1897,1923]" pageId="5" pageNumber="6">Philippines Ch. Semper</collectorName>
|
||
”;
|
||
<collectionCode id="ED5F06CB4000FFF7FC7C542FFC7AF861" box="[898,992,1898,1924]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="6">MNHN</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FF1454E9FEFDF823" blockId="5.[189,1063,1964,2027]" box="[234,359,1964,1990]" pageId="5" pageNumber="6">
|
||
<emphasis id="B93A421C4000FFF7FF1454E9FEFDF823" bold="true" box="[234,359,1964,1990]" pageId="5" pageNumber="6">
|
||
<typeStatus id="54F520AC4000FFF7FF1454E9FEFDF823" box="[234,359,1964,1990]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4000FFF7FF435494FBBDF80E" blockId="5.[189,1063,1964,2027]" box="[189,1063,2001,2027]" pageId="5" pageNumber="6">
|
||
<materialsCitation id="3B2694534000FFF7FF435494FBB9F80E" ID-GBIF-Occurrence="2424121666" box="[189,1059,2001,2027]" collectionCode="MNHN" collectorName="Philippines Ch. Semper" country="Philippines" location="PHILIPPINES" pageId="5" pageNumber="6" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
|
||
<collectingCountry id="F359DE9E4000FFF7FF435494FEF0F80E" box="[189,362,2001,2027]" name="Philippines" pageId="5" pageNumber="6">PHILIPPINES</collectingCountry>
|
||
•
|
||
<specimenCount id="9D4855874000FFF7FE7D5494FE2BF80E" box="[387,433,2001,2027]" pageId="5" pageNumber="6" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D4855874000FFF7FE435494FD9AF80E" box="[445,512,2001,2027]" pageId="5" pageNumber="6" type="female">2 ♀♀</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED5F06CB4000FFF7FC3B5494FBB9F80E" box="[965,1059,2001,2027]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="6">MNHN</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354CD854003FFFBFF43524EFB94FCED" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7" type="description">
|
||
<paragraph id="8BF19E0E4003FFF4FF43524EFE4AFEC3" blockId="6.[189,464,267,294]" box="[189,464,267,294]" pageId="6" pageNumber="7">
|
||
<emphasis id="B93A421C4003FFF4FF43524EFED4FEC0" bold="true" box="[189,334,267,293]" pageId="6" pageNumber="7">Description</emphasis>
|
||
(
|
||
<typeStatus id="54F520AC4003FFF4FEA3524EFE5DFEC3" box="[349,455,267,294]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4003FFF4FF14527FFEB0FEB1" blockId="6.[189,1398,314,410]" box="[234,298,314,340]" pageId="6" pageNumber="7">
|
||
<emphasis id="B93A421C4003FFF4FF14527FFEB0FEB1" bold="true" box="[234,298,314,340]" pageId="6" pageNumber="7">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4003FFF4FF43521BFE81FE7F" blockId="6.[189,1398,314,410]" pageId="6" pageNumber="7">
|
||
BODY (
|
||
<figureCitation id="1375828B4003FFF4FEF35218FEF3FE92" box="[269,361,349,376]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[277,1302,936,1922]" captionTargetId="figure@6.[260,1322,924,1928]" captionTargetPageId="6" captionText="Fig. 3. Philacelota leucothea sp. nov., habitus. A. ♂, holotype (MNHN). B. ♀, paratype (MNHN). Scale bars: 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474365" httpUri="https://zenodo.org/record/3474365/files/figure.png" pageId="6" pageNumber="7">Fig. 3A</figureCitation>
|
||
). Length 15.0 mm, greatest width
|
||
<quantity id="4CB633EB4003FFF4FD085218FCC8FE9D" box="[758,850,349,376]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="6" pageNumber="7" unit="mm" value="6.5">6.5 mm</quantity>
|
||
. Integument reddish-brown; scales and pilosity whitish.
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E4003FFF4FF435287FD8FFCD9" blockId="6.[189,1399,450,828]" pageId="6" pageNumber="7">
|
||
HEAD. Clypeus nearly crescent-shaped, with anterior margin arcuate in frontal view and weakly reflexed; surface roughly rugo-punctate, with long spiniform (slightly broadened at base, pointed at tip) scales. Fronto-clypeal suture conspicuous, almost straight. Frons and vertex roughly rugo-punctate; punctures denser on anterior half of frons; posterior half of frons and vertex with a narrow longitudinal callosity along midlength. Frons and vertex covered by elongate spiniform scales, denser toward inner margins of eyes, intermixed with sparse, moderately long setae. Antennae consist of 10 antennomeres, with seven antennomeres in club (
|
||
<figureCitation id="1375828B4003FFF4FE3851D0FDBEFD4A" box="[454,548,661,687]" captionStart="Fig" captionStartId="7.[189,232,1894,1920]" captionTargetBox="[276,1320,722,1829]" captionTargetId="figure@7.[265,1333,722,1841]" captionTargetPageId="7" captionText="Fig. 4. Philacelota leucothea sp. nov., holotype, ♂ (MNHN) (A–C) and paratype, ♀ (MNHN) (D– G). A. Antenna, lateral view. B. Propygidium. C. Pygidium. D. Antenna, lateral view. E. Antenna, dorsolateral view showing relative size of club antennomeres. F. Propygidium. G. Pygidium. Scale bars: A, C–E = 1.5 mm; B = 2 mm; F–G = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474367" httpUri="https://zenodo.org/record/3474367/files/figure.png" pageId="6" pageNumber="7">Fig. 4A</figureCitation>
|
||
); all antennomeres of club of equal length; 3
|
||
<superScript id="7C3B33464003FFF4FBD551D6FBA3FD44" attach="left" box="[1067,1081,659,673]" fontSize="6" pageId="6" pageNumber="7">rd</superScript>
|
||
antennomere with a strong anteromedial process; antennomeres of club strongly shagreened, antennomeres of funicle glabrous. Last maxillary palpomere elongately fusiform, with outer side impressed. Labrum weakly bilobed, covered by spiniform scales and sparse long setae; mentum deeply concave, bearing long setae along its lateral and posterior margins.
|
||
</paragraph>
|
||
<caption id="DF31CE864003FFF4FF4354E8FEC5F80E" ID-DOI="http://doi.org/10.5281/zenodo.3474365" ID-Zenodo-Dep="3474365" httpUri="https://zenodo.org/record/3474365/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1965,1991]" targetBox="[277,1302,936,1922]" targetPageId="6">
|
||
<paragraph id="8BF19E0E4003FFF4FF4354E8FEC5F80E" blockId="6.[189,1399,1965,2027]" pageId="6" pageNumber="7">
|
||
<emphasis id="B93A421C4003FFF4FF4354E8FE93F822" bold="true" box="[189,265,1965,1991]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
|
||
<taxonomicName id="4C4EE58D4003FFF4FEEE54E8FD95F822" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[272,527,1965,1991]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="leucothea" status="sp. nov.">
|
||
<emphasis id="B93A421C4003FFF4FEEE54E8FD95F822" box="[272,527,1965,1991]" italics="true" pageId="6" pageNumber="7">Philacelota leucothea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF674003FFF4FDEB54EBFDEBF82D" box="[533,625,1966,1992]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, habitus.
|
||
<emphasis id="B93A421C4003FFF4FD2054E8FD69F822" bold="true" box="[734,755,1965,1991]" pageId="6" pageNumber="7">A</emphasis>
|
||
. ♂, holotype (MNHN).
|
||
<emphasis id="B93A421C4003FFF4FBF254E8FBBAF822" bold="true" box="[1036,1056,1965,1991]" pageId="6" pageNumber="7">B</emphasis>
|
||
. ♀, paratype (MNHN). Scale bars: 2.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF19E0E4002FFFAFF435249FCE0FD0A" blockId="7.[189,1399,267,645]" lastBlockId="8.[189,1399,267,752]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
THORAX AND ABDOMEN. Pronotum narrower at base than base of elytra, 1.5 times as broad as long, broadest in middle, distinctly more convergent anteriad than posteriad; sides straight anteriad, but very weakly concave posteriad; anterior and posterior angles obtuse; anterior margin strongly concave, slightly bisinuate in middle; posterior margin strongly convex before scutellar shield; all sides of pronotum unbeaded. Pronotum not roughly rugo-punctate, with a narrow longitudinal medial callosity along its anterior half, covered by acicular to narrowly lanceolate scales being much smaller in size on sides than on disc, with setae along its anterior and lateral margins only. Propleura setose, with few small acicular scales along lateral margins only. Scutellar shield broadly rounded apically, not roughly rugo-punctate, covered with narrowly lanceolate scales becoming acicular and much denser along lateral margins. Elytra with three indistinct costae bearing scales; base of elytron with a swollen margin; humeral and apical umbones somewhat callose, glabrous; surface of elytra somewhat impressed behind humeral umbones. Interstices shallowly, unevenly and irregularly punctured; punctures somewhat rugose; scales small, rather evenly covering entire surface of elytra, lanceolate to narrowly lanceolate, becoming acicular close to margins. Propygidium (
|
||
<figureCitation id="1375828B400DFFFAFDCD5217FD14FE89" box="[563,654,338,364]" captionStart="Fig" captionStartId="7.[189,232,1894,1920]" captionTargetBox="[276,1320,722,1829]" captionTargetId="figure@7.[265,1333,722,1841]" captionTargetPageId="7" captionText="Fig. 4. Philacelota leucothea sp. nov., holotype, ♂ (MNHN) (A–C) and paratype, ♀ (MNHN) (D– G). A. Antenna, lateral view. B. Propygidium. C. Pygidium. D. Antenna, lateral view. E. Antenna, dorsolateral view showing relative size of club antennomeres. F. Propygidium. G. Pygidium. Scale bars: A, C–E = 1.5 mm; B = 2 mm; F–G = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474367" httpUri="https://zenodo.org/record/3474367/files/figure.png" pageId="8" pageNumber="9">Fig. 4B</figureCitation>
|
||
) finely and densely punctured; its basal half covered with small acicular setae intermixed with sparse long setae along basal margin; setae replaced on distal half by small ovoid scales. Pygidium (
|
||
<figureCitation id="1375828B400DFFFAFDDE52DDFDE3FE57" box="[544,633,408,434]" captionStart="Fig" captionStartId="7.[189,232,1894,1920]" captionTargetBox="[276,1320,722,1829]" captionTargetId="figure@7.[265,1333,722,1841]" captionTargetPageId="7" captionText="Fig. 4. Philacelota leucothea sp. nov., holotype, ♂ (MNHN) (A–C) and paratype, ♀ (MNHN) (D– G). A. Antenna, lateral view. B. Propygidium. C. Pygidium. D. Antenna, lateral view. E. Antenna, dorsolateral view showing relative size of club antennomeres. F. Propygidium. G. Pygidium. Scale bars: A, C–E = 1.5 mm; B = 2 mm; F–G = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474367" httpUri="https://zenodo.org/record/3474367/files/figure.png" pageId="8" pageNumber="9">Fig. 4C</figureCitation>
|
||
) fully covered by small ovoid scales, setose at apical margin only. Prosternum, mesoventrite and metaventrite densely and finely rugo-punctate, densely covered with long adpressed setae; metepimeron and metepisternum possessing scales; metaventrite lacking scales. First abdominal ventrite carinate in middle, covered with short, adpressed, hair-like setae replaced by densely setting, scale-like setae along posterior margin; 2
|
||
<superScript id="7C3B3346400DFFFAFCE65166FCB3FDD4" attach="left" box="[792,809,547,561]" fontSize="6" pageId="8" pageNumber="9">nd</superScript>
|
||
abdominal ventrite covered by small lanceolate scales intermixed with adpressed setae on its anterior half; remaining abdominal ventrites covered by small lanceolate and ovoid scales evenly distributed, but concentrated into dense patches on uppermost parts of ventrites; 3
|
||
<superScript id="7C3B3346400DFFFAFE5A51C8FE28FD7E" attach="left" box="[420,434,653,667]" fontSize="6" pageId="8" pageNumber="9">rd</superScript>
|
||
to 5
|
||
<superScript id="7C3B3346400DFFFAFE1751C8FE6CFD7E" attach="left" box="[489,502,653,667]" fontSize="6" pageId="8" pageNumber="9">th</superScript>
|
||
abdominal ventrites with few isolated long setae on lateral thirds only; last visible ventrite with several rows of moderately short, adpressed setae along its apical margin; sutures between 2
|
||
<superScript id="7C3B3346400DFFFAFECB5196FEDCFD04" attach="left" box="[309,326,723,737]" fontSize="6" pageId="8" pageNumber="9">nd</superScript>
|
||
to 5
|
||
<superScript id="7C3B3346400DFFFAFE845196FE1DFD04" attach="left" box="[378,391,723,737]" fontSize="6" pageId="8" pageNumber="9">th</superScript>
|
||
abdominal ventrites obliterated in middle.
|
||
</paragraph>
|
||
<caption id="DF31CE864002FFF5FF435423FC8DF80E" ID-DOI="http://doi.org/10.5281/zenodo.3474367" ID-Zenodo-Dep="3474367" httpUri="https://zenodo.org/record/3474367/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1894,1920]" targetBox="[276,1320,722,1829]" targetPageId="7">
|
||
<paragraph id="8BF19E0E4002FFF5FF435423FC8DF80E" blockId="7.[189,1399,1894,2027]" pageId="7" pageNumber="8">
|
||
<emphasis id="B93A421C4002FFF5FF435423FE94F865" bold="true" box="[189,270,1894,1920]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
|
||
<taxonomicName id="4C4EE58D4002FFF5FEE75423FD84F865" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[281,542,1894,1920]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="leucothea" status="sp. nov.">
|
||
<emphasis id="B93A421C4002FFF5FEE75423FD84F865" box="[281,542,1894,1920]" italics="true" pageId="7" pageNumber="8">Philacelota leucothea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF674002FFF5FDD75422FD10F864" box="[553,650,1895,1921]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (MNHN) (A–C) and paratype, ♀ (MNHN) (D– G).
|
||
<emphasis id="B93A421C4002FFF5FF1154CCFE9EF846" bold="true" box="[239,260,1929,1955]" pageId="7" pageNumber="8">A</emphasis>
|
||
. Antenna, lateral view.
|
||
<emphasis id="B93A421C4002FFF5FDD554CFFDA5F841" bold="true" box="[555,575,1930,1956]" pageId="7" pageNumber="8">B</emphasis>
|
||
. Propygidium.
|
||
<emphasis id="B93A421C4002FFF5FD0054CCFC89F846" bold="true" box="[766,787,1929,1955]" pageId="7" pageNumber="8">C</emphasis>
|
||
. Pygidium.
|
||
<emphasis id="B93A421C4002FFF5FC5554CFFC5AF841" bold="true" box="[939,960,1930,1956]" pageId="7" pageNumber="8">D</emphasis>
|
||
. Antenna, lateral view.
|
||
<emphasis id="B93A421C4002FFF5FB1954CFFB61F841" bold="true" box="[1255,1275,1930,1956]" pageId="7" pageNumber="8">E</emphasis>
|
||
. Antenna, dorsolateral view showing relative size of club antennomeres.
|
||
<emphasis id="B93A421C4002FFF5FC4754E8FC51F822" bold="true" box="[953,971,1965,1991]" pageId="7" pageNumber="8">F</emphasis>
|
||
. Propygidium.
|
||
<emphasis id="B93A421C4002FFF5FB7754E8FB3AF822" bold="true" box="[1161,1184,1965,1991]" pageId="7" pageNumber="8">G</emphasis>
|
||
. Pygidium. Scale bars: A, C–E = 1.5 mm; B = 2 mm; F–G = 2.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF19E0E400DFFFAFF435052FE7BFC05" blockId="8.[189,1398,790,993]" pageId="8" pageNumber="9">
|
||
LEGS. Protibiae unidentate, with spur short and thick; middle and hind tibiae gracile; apex of hind tibiae with 13 separated bristles. Femora setose and moderately densely covered with lanceolate scales; tibiae with sparse lanceolate scales and spiniform setae; tarsi with setae only. Tarsomeres 1
|
||
<emphasis id="B93A421C400DFFFAFB2A5018FB79FC92" bold="true" box="[1236,1251,861,887]" pageId="8" pageNumber="9">–</emphasis>
|
||
4 shortened, especially of the fore tarsi; 1
|
||
<superScript id="7C3B3346400DFFFAFDF7503AFD8EFC68" attach="left" box="[521,532,895,909]" fontSize="6" pageId="8" pageNumber="9">st</superScript>
|
||
tarsomere of hind tarsi only slightly longer than 2
|
||
<superScript id="7C3B3346400DFFFAFBA3503BFBF4FC69" attach="none" box="[1117,1134,894,908]" fontSize="6" pageId="8" pageNumber="9">nd</superScript>
|
||
. Claws bifurcate, with lower lobe slightly closer to upper one than to base and almost parallel to upper one, distinctly shorter, but as broad as the latter.
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400DFFFAFF43574DFDBAFB8D" blockId="8.[189,1398,1032,1128]" pageId="8" pageNumber="9">
|
||
GENITALIA. Aedeagus (
|
||
<figureCitation id="1375828B400DFFFAFE31574DFD80FBC6" box="[463,538,1032,1059]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[202,1387,1426,1882]" captionTargetId="figure@8.[189,1396,1422,1886]" captionTargetPageId="8" captionText="Fig. 5. A–B. Philacelota leucothea sp. nov., holotype, ♂ (MNHN). A. Aedeagus, lateral view. B. Parameres, dorsal view. — C–D. P. submaculata Heller, 1900, from Kema, Sulawesi, ♂ (MNHN). C. Aedeagus, lateral view. D. Parameres, dorsal view. Scale bars: A–B = 1.5 mm; C–D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3474369" httpUri="https://zenodo.org/record/3474369/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
||
A–B) strongly arcuate; phallobase conspicuously humped dorsally at its distal extremity; parameres almost uniformly deep along their length in lateral view, with apical extensions large, boot-shaped.
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400DFFFAFF4357D5FC94FAF6" blockId="8.[189,1399,1167,1300]" pageId="8" pageNumber="9">
|
||
VARIATION. Length 14.0 mm, greatest width
|
||
<quantity id="4CB633EB400DFFFAFD3B57CAFCB9FB4F" box="[709,803,1167,1194]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="8" pageNumber="9" unit="mm" value="5.5">5.5 mm</quantity>
|
||
. Margins of clypeus less angular than in
|
||
<typeStatus id="54F520AC400DFFFAFAF657D5FAE9FB4F" box="[1288,1395,1168,1194]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
. Fronto-clypeal suture somewhat carinate in middle. Scales on dorsal surface of head, pronotum and elytra smaller and sparser than in
|
||
<typeStatus id="54F520AC400DFFFAFDB35793FD22FB15" box="[589,696,1238,1264]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
. Elytra with four costae more pronounced than in
|
||
<typeStatus id="54F520AC400DFFFAFAF65793FAE9FB15" box="[1288,1395,1238,1264]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
. Abdominal ventrites more setose than in
|
||
<typeStatus id="54F520AC400DFFFAFD6157BCFC90FAF6" box="[671,778,1273,1299]" pageId="8" pageNumber="9" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF31CE86400DFFFAFF4354CFFABBF80E" ID-DOI="http://doi.org/10.5281/zenodo.3474369" ID-Zenodo-Dep="3474369" httpUri="https://zenodo.org/record/3474369/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1930,1956]" targetBox="[202,1387,1426,1882]" targetPageId="8">
|
||
<paragraph id="8BF19E0E400DFFFAFF4354CFFABBF80E" blockId="8.[189,1399,1930,2027]" pageId="8" pageNumber="9">
|
||
<emphasis id="B93A421C400DFFFAFF4354CFFEC3F841" bold="true" box="[189,345,1930,1956]" pageId="8" pageNumber="9">Fig. 5. A–B</emphasis>
|
||
.
|
||
<taxonomicName id="4C4EE58D400DFFFAFE8F54CFFDE3F841" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[369,633,1930,1956]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="leucothea" status="sp. nov.">
|
||
<emphasis id="B93A421C400DFFFAFE8F54CFFDE3F841" box="[369,633,1930,1956]" italics="true" pageId="8" pageNumber="9">Philacelota leucothea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF67400DFFFAFD7754CFFD75F841" box="[649,751,1930,1956]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype, ♂ (MNHN).
|
||
<emphasis id="B93A421C400DFFFAFBD454CFFBA5F841" bold="true" box="[1066,1087,1930,1956]" pageId="8" pageNumber="9">A</emphasis>
|
||
. Aedeagus, lateral view.
|
||
<emphasis id="B93A421C400DFFFAFF4354E8FF4BF822" bold="true" box="[189,209,1965,1991]" pageId="8" pageNumber="9">B</emphasis>
|
||
. Parameres, dorsal view. —
|
||
<emphasis id="B93A421C400DFFFAFDD654E8FDFBF822" bold="true" box="[552,609,1965,1991]" pageId="8" pageNumber="9">C–D</emphasis>
|
||
.
|
||
<taxonomicName id="4C4EE58D400DFFFAFD8C54EBFC5EF822" authority="Heller, 1900" authorityName="Heller" authorityYear="1900" box="[626,964,1965,1991]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="submaculata">
|
||
<emphasis id="B93A421C400DFFFAFD8C54EBFCBCF822" box="[626,806,1965,1991]" italics="true" pageId="8" pageNumber="9">P. submaculata</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400DFFFAFCD054E8FC5EF822" author="Heller" box="[814,964,1965,1991]" firstAuthor="Heller" pageId="8" pageNumber="9" pagination="1 - 46" refId="ref5896" refString="Heller K. M. 1900. Neue Kafer von Celebes IV. Abhandlungen und Berichte des koniglichen zoologischen und anthropologisch-ethnographischen Museums zu Dresden 9: 1 - 46. Available from https: // biodiversitylibrary. org / page / 46189178 [accessed 10 Sep. 2019]." type="journal article" year="1900">Heller, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
, from Kema, Sulawesi, ♂ (MNHN).
|
||
<emphasis id="B93A421C400DFFFAFF435495FF48F80F" bold="true" box="[189,210,2000,2026]" pageId="8" pageNumber="9">C</emphasis>
|
||
. Aedeagus, lateral view.
|
||
<emphasis id="B93A421C400DFFFAFE065495FD97F80F" bold="true" box="[504,525,2000,2026]" pageId="8" pageNumber="9">D</emphasis>
|
||
. Parameres, dorsal view. Scale bars: A–B = 1.5 mm; C–D = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF19E0E400CFFFBFF14524EFEDFFEC0" blockId="9.[189,1399,267,776]" box="[234,325,267,293]" pageId="9" pageNumber="10">
|
||
<emphasis id="B93A421C400CFFFBFF14524EFEDFFEC0" bold="true" box="[234,325,267,293]" pageId="9" pageNumber="10">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400CFFFBFF435275FB94FCED" blockId="9.[189,1399,267,776]" pageId="9" pageNumber="10">
|
||
Habitus: see
|
||
<figureCitation id="1375828B400CFFFBFEAA5275FE35FEAF" box="[340,431,304,330]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[277,1302,936,1922]" captionTargetId="figure@6.[260,1322,924,1928]" captionTargetPageId="6" captionText="Fig. 3. Philacelota leucothea sp. nov., habitus. A. ♂, holotype (MNHN). B. ♀, paratype (MNHN). Scale bars: 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474365" httpUri="https://zenodo.org/record/3474365/files/figure.png" pageId="9" pageNumber="10">Fig. 3B</figureCitation>
|
||
. Length 16.0
|
||
<emphasis id="B93A421C400CFFFBFDB65275FDCDFEAF" bold="true" box="[584,599,304,330]" pageId="9" pageNumber="10">–</emphasis>
|
||
17.0 mm, greatest width 6.5
|
||
<emphasis id="B93A421C400CFFFBFC635275FC36FEAF" bold="true" box="[925,940,304,330]" pageId="9" pageNumber="10">–</emphasis>
|
||
7.0 mm. Fronto-clypeal suture concave in middle. Antennal club (
|
||
<figureCitation id="1375828B400CFFFBFE0D5210FDA2FE8A" box="[499,568,341,367]" captionStart="Fig" captionStartId="7.[189,232,1894,1920]" captionTargetBox="[276,1320,722,1829]" captionTargetId="figure@7.[265,1333,722,1841]" captionTargetPageId="7" captionText="Fig. 4. Philacelota leucothea sp. nov., holotype, ♂ (MNHN) (A–C) and paratype, ♀ (MNHN) (D– G). A. Antenna, lateral view. B. Propygidium. C. Pygidium. D. Antenna, lateral view. E. Antenna, dorsolateral view showing relative size of club antennomeres. F. Propygidium. G. Pygidium. Scale bars: A, C–E = 1.5 mm; B = 2 mm; F–G = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474367" httpUri="https://zenodo.org/record/3474367/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
|
||
D–E) consists of five antennomeres: 2
|
||
<superScript id="7C3B3346400CFFFBFC035216FB94FE84" attach="left" box="[1021,1038,339,353]" fontSize="6" pageId="9" pageNumber="10">nd</superScript>
|
||
antennomere 1.4–1.7 times as long as 1
|
||
<superScript id="7C3B3346400CFFFBFEDA523CFEB5FE62" attach="left" box="[292,303,377,391]" fontSize="6" pageId="9" pageNumber="10">st</superScript>
|
||
(1
|
||
<superScript id="7C3B3346400CFFFBFEB0523CFEC3FE62" attach="left" box="[334,345,377,391]" fontSize="6" pageId="9" pageNumber="10">st</superScript>
|
||
antennomere sometimes only slightly exceeding anteromedial process of last antennomere of funicle in length), 2
|
||
<superScript id="7C3B3346400CFFFBFE3352DBFE44FE49" attach="left" box="[461,478,414,428]" fontSize="6" pageId="9" pageNumber="10">nd</superScript>
|
||
antennomere one-third to two-thirds as long as 3
|
||
<superScript id="7C3B3346400CFFFBFBCE52DBFBA4FE49" attach="none" box="[1072,1086,414,428]" fontSize="6" pageId="9" pageNumber="10">rd</superScript>
|
||
, 3
|
||
<superScript id="7C3B3346400CFFFBFBA052DBFBF6FE49" attach="left" box="[1118,1132,414,428]" fontSize="6" pageId="9" pageNumber="10">rd</superScript>
|
||
antennomere equal to or slightly shorter than 4
|
||
<superScript id="7C3B3346400CFFFBFE1A5286FE6BFE34" attach="none" box="[484,497,451,465]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
|
||
; 4
|
||
<superScript id="7C3B3346400CFFFBFDEE5286FD87FE34" attach="left" box="[528,541,451,465]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
|
||
and 5
|
||
<superScript id="7C3B3346400CFFFBFD975286FDECFE34" attach="left" box="[617,630,451,465]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
|
||
antennomeres of equal length. Funicle: 4
|
||
<superScript id="7C3B3346400CFFFBFB9B5286FBE8FE34" attach="left" box="[1125,1138,451,465]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
|
||
and 5
|
||
<superScript id="7C3B3346400CFFFBFB435286FB50FE34" attach="left" box="[1213,1226,451,465]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
|
||
antennomeres with conspicuous anteromedial process, 3
|
||
<superScript id="7C3B3346400CFFFBFD5B52ADFD29FE13" attach="left" box="[677,691,488,502]" fontSize="6" pageId="9" pageNumber="10">rd</superScript>
|
||
antennomere as long as 4
|
||
<superScript id="7C3B3346400CFFFBFC1C52ADFC75FE13" attach="left" box="[994,1007,488,502]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
|
||
and 5
|
||
<superScript id="7C3B3346400CFFFBFBC852ADFBD9FE13" attach="left" box="[1078,1091,488,502]" fontSize="6" pageId="9" pageNumber="10">th</superScript>
|
||
combined. Last maxillary palpomere more attenuated than in male. Scales on pronotum and especially on elytra unevenly distributed, intermixed with glabrous spaces forming a variegated pattern. Elytra with three–four more or less developed costae; humeral umbones conspicuously produced backward, with surface of elytra distinctly impressed behind them. Propygidium lacking long setae (
|
||
<figureCitation id="1375828B400CFFFBFC18513BFBD8FD7D" box="[998,1090,638,664]" captionStart="Fig" captionStartId="7.[189,232,1894,1920]" captionTargetBox="[276,1320,722,1829]" captionTargetId="figure@7.[265,1333,722,1841]" captionTargetPageId="7" captionText="Fig. 4. Philacelota leucothea sp. nov., holotype, ♂ (MNHN) (A–C) and paratype, ♀ (MNHN) (D– G). A. Antenna, lateral view. B. Propygidium. C. Pygidium. D. Antenna, lateral view. E. Antenna, dorsolateral view showing relative size of club antennomeres. F. Propygidium. G. Pygidium. Scale bars: A, C–E = 1.5 mm; B = 2 mm; F–G = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474367" httpUri="https://zenodo.org/record/3474367/files/figure.png" pageId="9" pageNumber="10">Fig. 4F</figureCitation>
|
||
). Pygidium with glabrous medial stripe, sometimes callose on its basal half (
|
||
<figureCitation id="1375828B400CFFFBFCDA51E6FC1DFD5B" box="[804,903,675,702]" captionStart="Fig" captionStartId="7.[189,232,1894,1920]" captionTargetBox="[276,1320,722,1829]" captionTargetId="figure@7.[265,1333,722,1841]" captionTargetPageId="7" captionText="Fig. 4. Philacelota leucothea sp. nov., holotype, ♂ (MNHN) (A–C) and paratype, ♀ (MNHN) (D– G). A. Antenna, lateral view. B. Propygidium. C. Pygidium. D. Antenna, lateral view. E. Antenna, dorsolateral view showing relative size of club antennomeres. F. Propygidium. G. Pygidium. Scale bars: A, C–E = 1.5 mm; B = 2 mm; F–G = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474367" httpUri="https://zenodo.org/record/3474367/files/figure.png" pageId="9" pageNumber="10">Fig. 4G</figureCitation>
|
||
). Abdominal ventrites more inflated than in male. Protibial spur longer than in male. Apex of hind tibiae with 16 short, separated bristles. Tarsi shorter than in males; 1
|
||
<superScript id="7C3B3346400CFFFBFE2E51A9FE41FD1F" attach="left" box="[464,475,748,762]" fontSize="6" pageId="9" pageNumber="10">st</superScript>
|
||
and 2
|
||
<superScript id="7C3B3346400CFFFBFDDD51A9FDAEFD1F" attach="left" box="[547,564,748,762]" fontSize="6" pageId="9" pageNumber="10">nd</superScript>
|
||
tarsomeres of hind tarsi equal in length.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354CD85400CFFFBFF435074FBD2FB65" pageId="9" pageNumber="10" type="diagnosis">
|
||
<paragraph id="8BF19E0E400CFFFBFF435074FE53FCAE" blockId="9.[189,457,817,843]" box="[189,457,817,843]" pageId="9" pageNumber="10">
|
||
<heading id="D0B92962400CFFFBFF435074FE53FCAE" bold="true" box="[189,457,817,843]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="B93A421C400CFFFBFF435074FE53FCAE" bold="true" box="[189,457,817,843]" pageId="9" pageNumber="10">Differential diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400CFFFBFF435024FBD2FB65" blockId="9.[189,1399,865,1152]" pageId="9" pageNumber="10">
|
||
This new species is most similar to
|
||
<taxonomicName id="4C4EE58D400CFFFBFDAC5027FC02FC99" authority="Heller, 1900" authorityName="Heller" authorityYear="1900" box="[594,920,865,892]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="submaculata">
|
||
<emphasis id="B93A421C400CFFFBFDAC5027FC99FC99" box="[594,771,866,892]" italics="true" pageId="9" pageNumber="10">P. submaculata</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400CFFFBFCF65024FC02FC99" author="Heller" box="[776,920,865,892]" firstAuthor="Heller" pageId="9" pageNumber="10" pagination="1 - 46" refId="ref5896" refString="Heller K. M. 1900. Neue Kafer von Celebes IV. Abhandlungen und Berichte des koniglichen zoologischen und anthropologisch-ethnographischen Museums zu Dresden 9: 1 - 46. Available from https: // biodiversitylibrary. org / page / 46189178 [accessed 10 Sep. 2019]." type="journal article" year="1900">Heller, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
from northern
|
||
<collectingRegion id="498A50EC400CFFFBFBBA5024FB34FC9E" box="[1092,1198,865,891]" country="Indonesia" name="Sulawesi Utara" pageId="9" pageNumber="10">Sulawesi</collectingRegion>
|
||
in the possession of squamose vestiture (vs setose in
|
||
<taxonomicName id="4C4EE58D400CFFFBFDAE50CDFCAAFC44" authority="Zidek, 2018" authorityName="Zidek" authorityYear="2018" box="[592,816,903,929]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="jakli">
|
||
<emphasis id="B93A421C400CFFFBFDAE50CDFD04FC44" box="[592,670,903,929]" italics="true" pageId="9" pageNumber="10">P. jakli</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400CFFFBFD5A50C2FCAAFC44" author="Zidek" box="[676,816,903,929]" firstAuthor="Zidek" pageId="9" pageNumber="10" pagination="1 - 6" refId="ref6147" refString="Zidek J. 2018. A new species of Philacelota Heller (Scarabaeidae: Melolonthinae: Melolonthini) from Flores Island, Indonesia. Insecta Mundi 0679: 1 - 6. Available from http: // digitalcommons. unl. edu / insectamundi / 1176 [accessed 19 Sep. 2019]." type="book chapter" year="2018">Zidek, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EE58D400CFFFBFC9A50CDFBFCFC44" authority="Heller, 1900" authorityName="Heller" authorityYear="1900" box="[868,1126,903,929]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sulana">
|
||
<emphasis id="B93A421C400CFFFBFC9A50CDFC57FC44" box="[868,973,903,929]" italics="true" pageId="9" pageNumber="10">P. sulana</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400CFFFBFC2C50C2FBFCFC44" author="Heller" box="[978,1126,903,929]" firstAuthor="Heller" pageId="9" pageNumber="10" pagination="1 - 46" refId="ref5896" refString="Heller K. M. 1900. Neue Kafer von Celebes IV. Abhandlungen und Berichte des koniglichen zoologischen und anthropologisch-ethnographischen Museums zu Dresden 9: 1 - 46. Available from https: // biodiversitylibrary. org / page / 46189178 [accessed 10 Sep. 2019]." type="journal article" year="1900">Heller, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C4EE58D400CFFFBFB8650C2FAECFC44" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[1144,1398,903,929]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="leucothea" status="sp. nov.">
|
||
<emphasis id="B93A421C400CFFFBFB8650C2FAECFC44" box="[1144,1398,903,929]" italics="true" pageId="9" pageNumber="10">Philacelota leucothea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF67400CFFFBFF4350E9FE83FC23" box="[189,281,940,966]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="4C4EE58D400CFFFBFE4850E8FDF0FC23" authorityName="Heller" authorityYear="1900" box="[438,618,940,966]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="submaculata">
|
||
<emphasis id="B93A421C400CFFFBFE4850E8FDF0FC23" box="[438,618,940,966]" italics="true" pageId="9" pageNumber="10">P. submaculata</emphasis>
|
||
</taxonomicName>
|
||
in the following respects: long hair-like setae on the basal half of the propygidium very sparse in males and absent in females (vs dense in both sexes), much less callose pronotum and upper surface of the head, less costate elytra, and finer and sparser puncturation of their interstices. Furthermore, the males of
|
||
<taxonomicName id="4C4EE58D400CFFFBFD8A5759FC9BFBD0" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[628,769,1051,1077]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="leucothea" status="sp. nov.">
|
||
<emphasis id="B93A421C400CFFFBFD8A5759FC9BFBD0" box="[628,769,1051,1077]" italics="true" pageId="9" pageNumber="10">P. leucothea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF67400CFFFBFCF95759FCFBFBD3" box="[775,865,1052,1078]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differ from the males of
|
||
<taxonomicName id="4C4EE58D400CFFFBFB805759FAAAFBD0" authorityName="Heller" authorityYear="1900" box="[1150,1328,1051,1077]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="submaculata">
|
||
<emphasis id="B93A421C400CFFFBFB805759FAAAFBD0" box="[1150,1328,1051,1077]" italics="true" pageId="9" pageNumber="10">P. submaculata</emphasis>
|
||
</taxonomicName>
|
||
in the elytra uniformly covered with scales (vs elytral costae glabrous), in the absence of the bare longitudinal stripe on the pygidium (vs present), and in the shape of the aedeagus (
|
||
<figureCitation id="1375828B400CFFFBFC0F5720FBA3FB65" box="[1009,1081,1125,1152]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[202,1387,1426,1882]" captionTargetId="figure@8.[189,1396,1422,1886]" captionTargetPageId="8" captionText="Fig. 5. A–B. Philacelota leucothea sp. nov., holotype, ♂ (MNHN). A. Aedeagus, lateral view. B. Parameres, dorsal view. — C–D. P. submaculata Heller, 1900, from Kema, Sulawesi, ♂ (MNHN). C. Aedeagus, lateral view. D. Parameres, dorsal view. Scale bars: A–B = 1.5 mm; C–D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3474369" httpUri="https://zenodo.org/record/3474369/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C354CD85400CFFF9FF4357ECFAECFD43" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="key">
|
||
<paragraph id="8BF19E0E400CFFFBFF4357ECFC33FB21" blockId="9.[189,937,1193,1220]" box="[189,937,1193,1220]" pageId="9" pageNumber="10">
|
||
<heading id="D0B92962400CFFFBFF4357ECFC33FB21" bold="true" box="[189,937,1193,1220]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
|
||
<emphasis id="B93A421C400CFFFBFF4357ECFC33FB21" bold="true" box="[189,937,1193,1220]" pageId="9" pageNumber="10">
|
||
Identification key for the species of
|
||
<taxonomicName id="4C4EE58D400CFFFBFD8657ECFD78FB26" authorityName="Dalla Torre" authorityYear="1913" box="[632,738,1193,1219]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93A421C400CFFFBFD8657ECFD78FB26" bold="true" box="[632,738,1193,1219]" italics="true" pageId="9" pageNumber="10">Engertia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EE58D400CFFFBFCDE57ECFC33FB21" authorityName="Heller" authorityYear="1900" box="[800,937,1193,1220]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93A421C400CFFFBFCDE57ECFC33FB21" bold="true" box="[800,937,1193,1220]" italics="true" pageId="9" pageNumber="10">Philacelota</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<key id="FFDFC93F400CFFF9FF43579FFAECFD43" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
|
||
<keyStep id="30BA86AB400CFFFBFF43579FFAECFA6C" pageId="9" pageNumber="10">
|
||
<paragraph id="8BF19E0E400CFFFBFF43579FFAEDFAFC" blockId="9.[189,1399,1241,1417]" pageId="9" pageNumber="10">
|
||
<keyLead id="30BF3D3B400CFFFBFF43579FFAEDFAFC" pageId="9" pageNumber="10">1. Antennal club with 5–7 lamellae in males and 4 or 5 lamellae in females, those of males conspicuously elongated; scales on elytral interstices aggregated into spots or stripes .............................. 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400CFFFBFF3F5661FAECFA6C" blockId="9.[189,1399,1241,1417]" pageId="9" pageNumber="10">
|
||
<keyLead id="30BF3D3B400CFFFBFF3F5661FAECFA6C" pageId="9" pageNumber="10">
|
||
– Antennal club with 3–5 lamellae in both sexes (rarely more than 3); when more than
|
||
<quantity id="4CB633EB400CFFFBFB1E5661FA94FADB" box="[1248,1294,1316,1342]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="9" pageNumber="10" unit="in" value="3.0">3 in</quantity>
|
||
number, club lamellae not conspicuously elongated in males, and scales of elytral interstices never aggregated into spots or stripes ..........................................................other Indo-Australian leucopholine genera
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA86AB400CFFFBFF4356F4FAEDF9DE" pageId="9" pageNumber="10">
|
||
<paragraph id="8BF19E0E400CFFFBFF4356F4FAEDFA14" blockId="9.[189,1399,1456,1595]" pageId="9" pageNumber="10">
|
||
<keyLead id="30BF3D3B400CFFFBFF4356F4FAEDFA14" pageId="9" pageNumber="10">
|
||
2. Antennal club of males with 7 lamellae; 3
|
||
<superScript id="7C3B3346400CFFFBFD0956F5FC9FFA5B" attach="left" box="[759,773,1456,1470]" fontSize="6" pageId="9" pageNumber="10">rd</superScript>
|
||
antennomere elongated in both sexes (
|
||
<figureCitation id="1375828B400CFFFBFB1556F4FAAFFA29" box="[1259,1333,1457,1484]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[202,1386,1381,1876]" captionTargetId="figure@10.[189,1396,1381,1882]" captionTargetPageId="10" captionText="Fig. 6. Shape of 3rd antennomere (arrowed) in males (A–B) and females (C–D) of the type species of the genera Engertia Dalla Torre, 1913 and Philacelota Heller, 1900. A, C. Engertia amboinae (Brenske, 1897). B, D. Philacelota submaculata Heller, 1900. Scale bars = 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474371" httpUri="https://zenodo.org/record/3474371/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
|
||
B–D) (
|
||
<taxonomicName id="4C4EE58D400CFFFBFF0A5693FEE1FA15" authorityName="Heller" authorityYear="1900" box="[244,379,1494,1520]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93A421C400CFFFBFF0A5693FEE1FA15" box="[244,379,1494,1520]" italics="true" pageId="9" pageNumber="10">Philacelota</emphasis>
|
||
</taxonomicName>
|
||
) ...................................................................................................................................... 3
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400CFFFBFF3F56B9FAEDF9DE" blockId="9.[189,1399,1456,1595]" pageId="9" pageNumber="10">
|
||
<keyLead id="30BF3D3B400CFFFBFF3F56B9FAEDF9DE" pageId="9" pageNumber="10">
|
||
– Antennal club of males with 5 or 6 lamellae; 3
|
||
<superScript id="7C3B3346400CFFFBFCCC56BFFCDAF9ED" attach="left" box="[818,832,1530,1544]" fontSize="6" pageId="9" pageNumber="10">rd</superScript>
|
||
antennomere short in both sexes (
|
||
<figureCitation id="1375828B400CFFFBFB1556B9FAAFF9F2" box="[1259,1333,1532,1559]" captionStart="Fig" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[202,1386,1381,1876]" captionTargetId="figure@10.[189,1396,1381,1882]" captionTargetPageId="10" captionText="Fig. 6. Shape of 3rd antennomere (arrowed) in males (A–B) and females (C–D) of the type species of the genera Engertia Dalla Torre, 1913 and Philacelota Heller, 1900. A, C. Engertia amboinae (Brenske, 1897). B, D. Philacelota submaculata Heller, 1900. Scale bars = 1.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474371" httpUri="https://zenodo.org/record/3474371/files/figure.png" pageId="9" pageNumber="10">Fig. 6</figureCitation>
|
||
A–C) (
|
||
<taxonomicName id="4C4EE58D400CFFFBFF0A5567FEC1F9DE" authorityName="Dalla Torre" authorityYear="1913" box="[244,347,1570,1595]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93A421C400CFFFBFF0A5567FEC1F9DE" box="[244,347,1570,1595]" italics="true" pageId="9" pageNumber="10">Engertia</emphasis>
|
||
</taxonomicName>
|
||
)........................................................................................................................................... 6
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph id="8BF19E0E400CFFFBFF435521FAEDF941" blockId="9.[189,1399,1636,1700]" pageId="9" pageNumber="10">
|
||
<keyLead id="30BF3D3B400CFFFBFF435521FAEDF941" pageId="9" pageNumber="10">
|
||
<keyStep id="30BA86AB400CFFFBFF435521FAEDF941" pageId="9" pageNumber="10">
|
||
3. Body vestiture squamose; parameres compressed laterally, with irregular outlines ........................ 4 – Body vestiture setose; parameres tube-like with expanded tips (unknown in
|
||
<taxonomicName id="4C4EE58D400CFFFBFB9855CFFB4AF946" authorityName="Heller" authorityYear="1900" box="[1126,1232,1673,1699]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="sulana">
|
||
<emphasis id="B93A421C400CFFFBFB9855CFFB4AF946" box="[1126,1232,1673,1699]" italics="true" pageId="9" pageNumber="10">P. sulana</emphasis>
|
||
</taxonomicName>
|
||
) ................. 5
|
||
</keyStep>
|
||
</keyLead>
|
||
</paragraph>
|
||
<keyStep id="30BA86AB400CFFFBFF435588FAEDF80E" pageId="9" pageNumber="10">
|
||
<paragraph id="8BF19E0E400CFFFBFF435588FAEDF8B3" blockId="9.[189,1399,1741,2027]" pageId="9" pageNumber="10">
|
||
<keyLead id="30BF3D3B400CFFFBFF435588FAEDF8B3" pageId="9" pageNumber="10">
|
||
4. Upper surface of head and disc of pronotum with irregular callose areas of variable expression in both sexes. Males: basal half of propygidium with dense long setae; elytral costae glabrous; aedeagus as on
|
||
<figureCitation id="1375828B400CFFFBFE5C5452FE7FF8D7" box="[418,485,1815,1842]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[202,1387,1426,1882]" captionTargetId="figure@8.[189,1396,1422,1886]" captionTargetPageId="8" captionText="Fig. 5. A–B. Philacelota leucothea sp. nov., holotype, ♂ (MNHN). A. Aedeagus, lateral view. B. Parameres, dorsal view. — C–D. P. submaculata Heller, 1900, from Kema, Sulawesi, ♂ (MNHN). C. Aedeagus, lateral view. D. Parameres, dorsal view. Scale bars: A–B = 1.5 mm; C–D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3474369" httpUri="https://zenodo.org/record/3474369/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
|
||
C–D. Females: basal half of propygidium with long setae; puncturation of the elytral interstices dense and coarse (
|
||
<collectingCountry id="F359DE9E400CFFFBFD6F5479FC93F8B3" box="[657,777,1852,1878]" name="Indonesia" pageId="9" pageNumber="10">Indonesia</collectingCountry>
|
||
:
|
||
<collectingRegion id="498A50EC400CFFFBFCEA5479FC1AF8B2" box="[788,896,1852,1879]" country="Indonesia" name="Sulawesi Utara" pageId="9" pageNumber="10">Sulawesi</collectingRegion>
|
||
) .....................
|
||
<taxonomicName id="4C4EE58D400CFFFBFBD85478FAEDF8B3" authority="Heller, 1900" authorityName="Heller" authorityYear="1900" box="[1062,1399,1852,1879]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="submaculata">
|
||
<emphasis id="B93A421C400CFFFBFBD85478FB44F8B2" bold="true" box="[1062,1246,1852,1879]" italics="true" pageId="9" pageNumber="10">P. submaculata</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400CFFFBFB1B5479FAEDF8B3" author="Heller" box="[1253,1399,1852,1878]" firstAuthor="Heller" pageId="9" pageNumber="10" pagination="1 - 46" refId="ref5896" refString="Heller K. M. 1900. Neue Kafer von Celebes IV. Abhandlungen und Berichte des koniglichen zoologischen und anthropologisch-ethnographischen Museums zu Dresden 9: 1 - 46. Available from https: // biodiversitylibrary. org / page / 46189178 [accessed 10 Sep. 2019]." type="journal article" year="1900">Heller, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400CFFFBFF3F5427FAEDF80E" blockId="9.[189,1399,1741,2027]" pageId="9" pageNumber="10">
|
||
<keyLead id="30BF3D3B400CFFFBFF3F5427FAEDF80E" pageId="9" pageNumber="10">
|
||
– Upper surface of head and disc of pronotum with longitudinal callose stripe in both sexes. Males: basal half of propygidium with minute acicular and sparse long setae; elytral costae scaled; aedeagus as on
|
||
<figureCitation id="1375828B400CFFFBFED354EEFEEAF823" box="[301,368,1963,1990]" captionStart="Fig" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[202,1387,1426,1882]" captionTargetId="figure@8.[189,1396,1422,1886]" captionTargetPageId="8" captionText="Fig. 5. A–B. Philacelota leucothea sp. nov., holotype, ♂ (MNHN). A. Aedeagus, lateral view. B. Parameres, dorsal view. — C–D. P. submaculata Heller, 1900, from Kema, Sulawesi, ♂ (MNHN). C. Aedeagus, lateral view. D. Parameres, dorsal view. Scale bars: A–B = 1.5 mm; C–D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3474369" httpUri="https://zenodo.org/record/3474369/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
|
||
A–B. Females: basal half of propygidium with minute acicular setae; puncturation of the elytral interstices sparse and fine (
|
||
<collectingCountry id="F359DE9E400CFFFBFD875494FC99F80E" box="[633,771,2001,2027]" name="Philippines" pageId="9" pageNumber="10">Philippines</collectingCountry>
|
||
: Luzon) ......................................
|
||
<taxonomicName id="4C4EE58D400CFFFBFB7D5494FA8FF80E" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[1155,1301,2000,2027]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="leucothea" status="sp. nov.">
|
||
<emphasis id="B93A421C400CFFFBFB7D5494FA8FF80E" bold="true" box="[1155,1301,2000,2027]" italics="true" pageId="9" pageNumber="10">P. leucothea</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF67400CFFFBFAE25494FAEDF80E" box="[1308,1399,2001,2027]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA86AB400FFFF8FF43524EFAEDFE6A" pageId="10" pageNumber="11">
|
||
<paragraph id="8BF19E0E400FFFF8FF43524EFAEDFEAC" blockId="10.[189,1399,267,1320]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF43524EFAEDFEAC" pageId="10" pageNumber="11">
|
||
5. Protibiae unidentate in male; pronotum widest at base; setae of body vestiture very short and sparse (
|
||
<collectingCountry id="F359DE9E400FFFF8FF0C526BFEF0FEAD" box="[242,362,302,328]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
|
||
: Sula Mangoli) ................................................................................
|
||
<taxonomicName id="4C4EE58D400FFFF8FB91526AFAEDFEAC" authority="Heller, 1900" authorityName="Heller" authorityYear="1900" box="[1135,1399,302,329]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="sulana">
|
||
<emphasis id="B93A421C400FFFF8FB91526AFB44FEAC" bold="true" box="[1135,1246,302,329]" italics="true" pageId="10" pageNumber="11">P. sulana</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FB1B526BFAEDFEAC" author="Heller" box="[1253,1399,302,329]" firstAuthor="Heller" pageId="10" pageNumber="11" pagination="1 - 46" refId="ref5896" refString="Heller K. M. 1900. Neue Kafer von Celebes IV. Abhandlungen und Berichte des koniglichen zoologischen und anthropologisch-ethnographischen Museums zu Dresden 9: 1 - 46. Available from https: // biodiversitylibrary. org / page / 46189178 [accessed 10 Sep. 2019]." type="journal article" year="1900">Heller, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400FFFF8FF3F5217FAEDFE6A" blockId="10.[189,1399,267,1320]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF3F5217FAEDFE6A" pageId="10" pageNumber="11">
|
||
– Protibiae weakly bidentate in male; pronotum widest at midlength; setae of body vestiture long and dense (
|
||
<collectingCountry id="F359DE9E400FFFF8FEC35230FE2FFE6A" box="[317,437,373,399]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
|
||
:
|
||
<collectingCountry id="F359DE9E400FFFF8FE3E5230FD91FE75" box="[448,523,373,400]" name="Indonesia" pageId="10" pageNumber="11">Flores</collectingCountry>
|
||
)......................................................................................
|
||
<taxonomicName id="4C4EE58D400FFFF8FB715230FAEDFE6A" authority="Zidek, 2018" authorityName="Zidek" authorityYear="2018" box="[1167,1399,372,399]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="jakli">
|
||
<emphasis id="B93A421C400FFFF8FB715230FB78FE6A" bold="true" box="[1167,1250,372,399]" italics="true" pageId="10" pageNumber="11">P. jakli</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FB175230FAEDFE6A" author="Zidek" box="[1257,1399,373,399]" firstAuthor="Zidek" pageId="10" pageNumber="11" pagination="1 - 6" refId="ref6147" refString="Zidek J. 2018. A new species of Philacelota Heller (Scarabaeidae: Melolonthinae: Melolonthini) from Flores Island, Indonesia. Insecta Mundi 0679: 1 - 6. Available from http: // digitalcommons. unl. edu / insectamundi / 1176 [accessed 19 Sep. 2019]." type="book chapter" year="2018">Zidek, 2018</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA86AB400FFFF8FF4352F3FAECFDF2" pageId="10" pageNumber="11">
|
||
<paragraph id="8BF19E0E400FFFF8FF4352F3FAEDFE34" blockId="10.[189,1399,267,1320]" box="[189,1399,438,465]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF4352F3FAEDFE34" box="[189,1399,438,465]" pageId="10" pageNumber="11">
|
||
6. Antennal club with conspicuously elongated antennomeres. Males (unknown for
|
||
<taxonomicName id="4C4EE58D400FFFF8FB5D52F2FAAEFE35" baseAuthorityName="Brenske" baseAuthorityYear="1896" box="[1187,1332,438,464]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="maculosa">
|
||
<emphasis id="B93A421C400FFFF8FB5D52F2FAAEFE35" box="[1187,1332,438,464]" italics="true" pageId="10" pageNumber="11">E. maculosa</emphasis>
|
||
</taxonomicName>
|
||
) .... 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400FFFF8FF3F529FFAECFDF2" blockId="10.[189,1399,267,1320]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF3F529FFAECFDF2" pageId="10" pageNumber="11">
|
||
– Antennal club with short antennomeres. Females (unknown for
|
||
<taxonomicName id="4C4EE58D400FFFF8FC20529FFBFFFE11" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[990,1125,474,500]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="allolepis" status="sp. nov.">
|
||
<emphasis id="B93A421C400FFFF8FC20529FFBFFFE11" box="[990,1125,474,500]" italics="true" pageId="10" pageNumber="11">E. allolepis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF67400FFFF8FB93529FFB51FE11" box="[1133,1227,474,500]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C4EE58D400FFFF8FB29529FFAECFE16" authorityName="Prokofiev" authorityYear="2016" box="[1239,1398,474,499]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="germanica">
|
||
<emphasis id="B93A421C400FFFF8FB29529FFAECFE16" box="[1239,1398,474,499]" italics="true" pageId="10" pageNumber="11">E. germanica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EE58D400FFFF8FEE252BBFE3EFDF2" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[284,420,510,535]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis id="B93A421C400FFFF8FEE252BBFE3EFDF2" box="[284,420,510,535]" italics="true" pageId="10" pageNumber="11">E. papuana</emphasis>
|
||
</taxonomicName>
|
||
)............................................................................................................................... 12
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyLead id="30BF3D3B400FFFF8FF43517AFAEDFD99" pageId="10" pageNumber="11">
|
||
<keyStep id="30BA86AB400FFFF8FF43517AFAEDFD99" pageId="10" pageNumber="11">
|
||
<paragraph id="8BF19E0E400FFFF8FF43517AFAEDFDBD" blockId="10.[189,1399,267,1320]" box="[189,1399,574,601]" pageId="10" pageNumber="11">
|
||
7. Antennal club with 6 lamellae (
|
||
<collectingCountry id="F359DE9E400FFFF8FDA5517BFD7FFDBD" box="[603,741,574,600]" name="Philippines" pageId="10" pageNumber="11">Philippines</collectingCountry>
|
||
:
|
||
<collectingRegion id="498A50EC400FFFF8FD0F517AFCADFDBC" box="[753,823,575,601]" country="Philippines" name="Southern Leyte" pageId="10" pageNumber="11">Leyte</collectingRegion>
|
||
,
|
||
<collectingRegion id="498A50EC400FFFF8FCBC517BFC3DFDBC" box="[834,935,574,601]" country="Philippines" name="Palawan" pageId="10" pageNumber="11">Palawan</collectingRegion>
|
||
)..................................
|
||
<taxonomicName id="4C4EE58D400FFFF8FB54517AFAEDFDBD" authority="Keith, 2006" authorityName="Keith" authorityYear="2006" box="[1194,1399,574,601]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lii">
|
||
<emphasis id="B93A421C400FFFF8FB54517AFB7EFDBC" bold="true" box="[1194,1252,574,601]" italics="true" pageId="10" pageNumber="11">E. lii</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FB12517BFAEDFDBD" author="Keith" box="[1260,1399,574,600]" firstAuthor="Keith" pageId="10" pageNumber="11" pagination="90 - 94" refId="ref5946" refString="Keith D. 2006. Reflexions sur le genre Engertia et description d'une nouvelle espece des Philippines. Lambillionea 106: 90 - 94." type="journal article" year="2006">Keith, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400FFFF8FF3F5127FAEDFD99" blockId="10.[189,1399,267,1320]" box="[193,1399,609,636]" pageId="10" pageNumber="11">– Antennal club with 5 lamellae ........................................................................................................... 8</paragraph>
|
||
</keyStep>
|
||
</keyLead>
|
||
<keyStep id="30BA86AB400FFFF8FF4351E6FAEDFCAF" pageId="10" pageNumber="11">
|
||
<paragraph id="8BF19E0E400FFFF8FF4351E6FAEDFCE1" blockId="10.[189,1399,267,1320]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF4351E6FAEDFCE1" pageId="10" pageNumber="11">
|
||
8. Scales heterogeneous in shape in two outermost interstices and along apical margin of elytra. Aedeagus more arcuate in profile and more robust in shape (as on
|
||
<figureCitation id="1375828B400FFFF8FC0D5183FBD4FD05" box="[1011,1102,710,736]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="10" pageNumber="11">Fig. 2G</figureCitation>
|
||
) (
|
||
<collectingCountry id="F359DE9E400FFFF8FB985183FB44FD05" box="[1126,1246,710,736]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
|
||
:
|
||
<collectingCountry id="F359DE9E400FFFF8FB165183FAD8FD04" box="[1256,1346,710,737]" name="Indonesia" pageId="10" pageNumber="11">Ambon</collectingCountry>
|
||
)..... ..............................................................................................................................
|
||
<taxonomicName id="4C4EE58D400FFFF8FB7051AFFA8FFCE1" authority="Prokofiev, 2019" authorityName="Prokofiev" authorityYear="2019" box="[1166,1301,745,772]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="allolepis" status="sp. nov.">
|
||
<emphasis id="B93A421C400FFFF8FB7051AFFA8FFCE1" bold="true" box="[1166,1301,745,772]" italics="true" pageId="10" pageNumber="11">E. allolepis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A209FF67400FFFF8FAE251AFFAEDFCE1" box="[1308,1399,746,772]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400FFFF8FF3F5048FAEDFCAF" blockId="10.[189,1399,267,1320]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF3F5048FAEDFCAF" pageId="10" pageNumber="11">
|
||
– Scales homogeneous in shape in all interstices and along apical margin of elytra. Aedeagus less arcuate in profile (except
|
||
<taxonomicName id="4C4EE58D400FFFF8FDE85075FD2EFCAC" authorityName="Prokofiev" authorityYear="2016" box="[534,692,816,841]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="germanica">
|
||
<emphasis id="B93A421C400FFFF8FDE85075FD2EFCAC" box="[534,692,816,841]" italics="true" pageId="10" pageNumber="11">E. germanica</emphasis>
|
||
</taxonomicName>
|
||
) and always more gracile in shape (as on
|
||
<figureCitation id="1375828B400FFFF8FB5F5075FB68FCAF" box="[1185,1266,816,842]" captionStart="Fig" captionStartId="4.[189,232,1662,1688]" captionTargetBox="[201,1389,302,1606]" captionTargetId="figure@4.[189,1398,300,1614]" captionTargetPageId="4" captionText="Fig. 2. A–G. Engertia allolepis sp. nov., holotype, ♂ (MNHN). A. Antenna. B. Lateral part of the elytron. C. Apical margin of elytra and propygidium. D. Pygidium. E. Abdominal ventrites 2–4. F. Parameres, dorsal view. G. Aedeagus, lateral view. — H–J. E. papuana (Moser, 1913), ♂, from Sentani (cAP). H. Lateral part of elytron. I. Apical margin of elytra and propygidium. J. Aedeagus, lateral view. — K. E. amboinae (Brenske, 1897), ♂, from Ambon (MNHN), lateral part of elytron. — L. E. setifera (Moser, 1913), ♂, from Seram, pygidium (cAP). Scale bars are in common for B, H and K, and for F–G and J. Scale bars: A–B, H, K = 1.5 mm, C–G, I–J, L = 2.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3474363" httpUri="https://zenodo.org/record/3474363/files/figure.png" pageId="10" pageNumber="11">Fig. 2J</figureCitation>
|
||
).............. 9
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyLead id="30BF3D3B400FFFF8FF435034FAEFFC4B" pageId="10" pageNumber="11">
|
||
<keyStep id="30BA86AB400FFFF8FF435034FAEFFC4B" pageId="10" pageNumber="11">
|
||
<paragraph id="8BF19E0E400FFFF8FF435034FAECFC6E" blockId="10.[189,1399,267,1320]" box="[189,1398,881,908]" pageId="10" pageNumber="11">9. Elytra covered with ovoid to lanceolate scales; scales present on labrum and metaventrite........... 10</paragraph>
|
||
<paragraph id="8BF19E0E400FFFF8FF3F50D0FAEFFC4B" blockId="10.[189,1399,267,1320]" box="[193,1397,916,943]" pageId="10" pageNumber="11">– Elytra covered with setiform scales; labrum and metaventrite with setae only................................11</paragraph>
|
||
</keyStep>
|
||
</keyLead>
|
||
<keyStep id="30BA86AB400FFFF8FF435093FAECFBBC" pageId="10" pageNumber="11">
|
||
<paragraph id="8BF19E0E400FFFF8FF435093FAECFBF6" blockId="10.[189,1399,267,1320]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF435093FAECFBF6" pageId="10" pageNumber="11">
|
||
10. Elytra covered with ovoid scales; scales along midline of abdominal ventrites not aggregated into spots (
|
||
<collectingCountry id="F359DE9E400FFFF8FEC850BCFE34FBF6" box="[310,430,1017,1043]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
|
||
: islands of
|
||
<collectingCountry id="F359DE9E400FFFF8FDCE50BCFD11FBF6" box="[560,651,1017,1043]" name="Indonesia" pageId="10" pageNumber="11">Ambon</collectingCountry>
|
||
, Yapen and Miosidi) ....................
|
||
<taxonomicName id="4C4EE58D400FFFF8FBE750BCFAECFBF6" authority="(Brenske, 1897)" authorityName="Brenske" authorityYear="1897" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[1049,1398,1016,1043]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="amboinae">
|
||
<emphasis id="B93A421C400FFFF8FBE750BCFB28FBF6" bold="true" box="[1049,1202,1016,1043]" italics="true" pageId="10" pageNumber="11">E. amboinae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FB3C50BCFAF4FBF6" author="Brenske" box="[1218,1390,1017,1043]" firstAuthor="Brenske" pageId="10" pageNumber="11" pagination="96 - 120" refId="ref5871" refString="Brenske E. 1897. Neue Gattungen und Arten der Melolonthiden aus Afrika und Asien. Stettiner entomologische Zeitung 58: 96 - 120." type="journal article" year="1897">Brenske, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400FFFF8FF3F5758FAECFBBC" blockId="10.[189,1399,267,1320]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF3F5758FAECFBBC" pageId="10" pageNumber="11">
|
||
– Elytra covered with lanceolate scales; scales along midline of abdominal ventrites aggregated into small spots (New
|
||
<collectingCountry id="F359DE9E400FFFF8FE43577AFD8EFBBF" box="[445,532,1087,1114]" name="Guinea-Bissau" pageId="10" pageNumber="11">Guinea</collectingCountry>
|
||
)..........................................................................
|
||
<taxonomicName id="4C4EE58D400FFFF8FBC25705FAECFBBC" authority="(Moser, 1913)" authorityName="Moser" authorityYear="1913" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[1084,1398,1087,1114]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="papuana">
|
||
<emphasis id="B93A421C400FFFF8FBC25705FB52FBBF" bold="true" box="[1084,1224,1088,1114]" italics="true" pageId="10" pageNumber="11">E. papuana</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FB26577AFAF4FBBC" author="Moser" box="[1240,1390,1087,1113]" firstAuthor="Moser" pageId="10" pageNumber="11" pagination="271 - 297" refId="ref5971" refString="Moser J. 1913. Beitrag zur Kenntnis der Melolonthiden I. Deutsche entomologische Zeitschrift 3: 271 - 297. Available from https: // archive. org / details / biostor- 101429 [accessed 10 Sep. 2019]." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA86AB400FFFF8FF4357C4FAEDFAC2" pageId="10" pageNumber="11">
|
||
<paragraph id="8BF19E0E400FFFF8FF4357C4FAECFB04" blockId="10.[189,1399,267,1320]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF4357C4FAECFB04" pageId="10" pageNumber="11">
|
||
11. Setiform scales on elytra thick; pygidium mainly setigerous; scales along midline of abdominal ventrites not aggregated into spots (
|
||
<collectingCountry id="F359DE9E400FFFF8FD7257E1FC9EFB5B" box="[652,772,1188,1214]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
|
||
: islands of Seram,
|
||
<collectingCountry id="F359DE9E400FFFF8FC1E57E1FBA2FB5B" box="[992,1080,1188,1214]" name="Indonesia" pageId="10" pageNumber="11">Ambon</collectingCountry>
|
||
and Saparua) .................... .....................................................................................................................
|
||
<taxonomicName id="4C4EE58D400FFFF8FBB3578DFAECFB04" authority="(Moser, 1913)" authorityName="Moser" authorityYear="1913" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[1101,1398,1223,1250]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="setifera">
|
||
<emphasis id="B93A421C400FFFF8FBB3578DFB53FB04" bold="true" box="[1101,1225,1223,1250]" italics="true" pageId="10" pageNumber="11">E. setifera</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FB265782FAF4FB04" author="Moser" box="[1240,1390,1223,1249]" firstAuthor="Moser" pageId="10" pageNumber="11" pagination="271 - 297" refId="ref5971" refString="Moser J. 1913. Beitrag zur Kenntnis der Melolonthiden I. Deutsche entomologische Zeitschrift 3: 271 - 297. Available from https: // archive. org / details / biostor- 101429 [accessed 10 Sep. 2019]." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400FFFF8FF3F57AEFAEDFAC2" blockId="10.[189,1399,267,1320]" pageId="10" pageNumber="11">
|
||
<keyLead id="30BF3D3B400FFFF8FF3F57AEFAEDFAC2" pageId="10" pageNumber="11">
|
||
– Setiform scales on elytra thin; pygidium mainly scaly; scales along midline of abdominal ventrites aggregated into spots (New
|
||
<collectingCountry id="F359DE9E400FFFF8FDCD5648FD10FACD" box="[563,650,1293,1320]" name="Guinea-Bissau" pageId="10" pageNumber="11">Guinea</collectingCountry>
|
||
)....................................................
|
||
<taxonomicName id="4C4EE58D400FFFF8FBED564BFAEDFAC2" authority="Prokofiev, 2016" authorityName="Prokofiev" authorityYear="2016" box="[1043,1399,1293,1320]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="germanica">
|
||
<emphasis id="B93A421C400FFFF8FBED564BFB2CFACD" bold="true" box="[1043,1206,1294,1320]" italics="true" pageId="10" pageNumber="11">E. germanica</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FB435648FAEDFAC2" author="Prokofiev A. M." box="[1213,1399,1293,1319]" journalOrPublisher="Euroasian Entomological Journal" pageId="10" pageNumber="11" pagination="68 - 74" part="15" refId="ref6048" refString="Prokofiev A. M. 2016 a. New genera and species of the Papuan Leucopholini and Melolonthini (Coleoptera, Scarabaeidae: Melolonthinae). Euroasian Entomological Journal 15: 68 - 74. [In Russian.]" title="New genera and species of the Papuan Leucopholini and Melolonthini (Coleoptera, Scarabaeidae: Melolonthinae)" type="journal article" year="2016">Prokofiev, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<caption id="DF31CE86400FFFF8FF4354CDFB80F80C" ID-DOI="http://doi.org/10.5281/zenodo.3474371" ID-Zenodo-Dep="3474371" httpUri="https://zenodo.org/record/3474371/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1928,1954]" targetBox="[202,1386,1381,1876]" targetPageId="10">
|
||
<paragraph id="8BF19E0E400FFFF8FF4354CDFB80F80C" blockId="10.[189,1398,1927,2026]" pageId="10" pageNumber="11">
|
||
<emphasis id="B93A421C400FFFF8FF4354CDFE96F847" bold="true" box="[189,268,1928,1954]" pageId="10" pageNumber="11">Fig. 6.</emphasis>
|
||
Shape of 3
|
||
<superScript id="7C3B3346400FFFF8FE6854C2FE3EF870" attach="left" box="[406,420,1927,1941]" fontSize="6" pageId="10" pageNumber="11">rd</superScript>
|
||
antennomere (arrowed) in males (A–B) and females (C–D) of the type species of the genera
|
||
<taxonomicName id="4C4EE58D400FFFF8FEC254E9FDECF823" authority="Dalla Torre, 1913" authorityName="Dalla Torre" authorityYear="1913" box="[316,630,1964,1990]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93A421C400FFFF8FEC254E9FE39F820" box="[316,419,1964,1989]" italics="true" pageId="10" pageNumber="11">Engertia</emphasis>
|
||
Dalla Torre, 1913
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4EE58D400FFFF8FD5354E9FC54F823" authority="Heller, 1900" authorityName="Heller" authorityYear="1900" box="[685,974,1964,1990]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93A421C400FFFF8FD5354E9FCAEF823" box="[685,820,1964,1990]" italics="true" pageId="10" pageNumber="11">Philacelota</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FCC454E9FC54F823" author="Heller" box="[826,974,1964,1990]" firstAuthor="Heller" pageId="10" pageNumber="11" pagination="1 - 46" refId="ref5896" refString="Heller K. M. 1900. Neue Kafer von Celebes IV. Abhandlungen und Berichte des koniglichen zoologischen und anthropologisch-ethnographischen Museums zu Dresden 9: 1 - 46. Available from https: // biodiversitylibrary. org / page / 46189178 [accessed 10 Sep. 2019]." type="journal article" year="1900">Heller, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B93A421C400FFFF8FC2654EEFC77F820" bold="true" box="[984,1005,1963,1989]" pageId="10" pageNumber="11">A</emphasis>
|
||
,
|
||
<emphasis id="B93A421C400FFFF8FC0554EEFB8AF820" bold="true" box="[1019,1040,1963,1989]" pageId="10" pageNumber="11">C</emphasis>
|
||
.
|
||
<taxonomicName id="4C4EE58D400FFFF8FBE054E9FE9FF80C" authority="(Brenske, 1897)" authorityName="Brenske" authorityYear="1897" baseAuthorityName="Brenske" baseAuthorityYear="1897" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="amboinae">
|
||
<emphasis id="B93A421C400FFFF8FBE054E9FB65F823" box="[1054,1279,1964,1990]" italics="true" pageId="10" pageNumber="11">Engertia amboinae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FAF354E9FF66F80C" author="Brenske" firstAuthor="Brenske" pageId="10" pageNumber="11" pagination="96 - 120" refId="ref5871" refString="Brenske E. 1897. Neue Gattungen und Arten der Melolonthiden aus Afrika und Asien. Stettiner entomologische Zeitung 58: 96 - 120." type="journal article" year="1897">Brenske, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B93A421C400FFFF8FEEE548AFEBEF80C" bold="true" box="[272,292,1999,2025]" pageId="10" pageNumber="11">B</emphasis>
|
||
,
|
||
<emphasis id="B93A421C400FFFF8FECC548AFEDDF80C" bold="true" box="[306,327,1999,2025]" pageId="10" pageNumber="11">D</emphasis>
|
||
.
|
||
<taxonomicName id="4C4EE58D400FFFF8FEA8548AFC8DF80C" authority="Heller, 1900" authorityName="Heller" authorityYear="1900" box="[342,791,1999,2025]" class="Insecta" family="Melolonthidae" genus="Philacelota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="submaculata">
|
||
<emphasis id="B93A421C400FFFF8FEA8548AFDE1F80C" box="[342,635,1999,2025]" italics="true" pageId="10" pageNumber="11">Philacelota submaculata</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400FFFF8FD7C548AFC8DF80C" author="Heller" box="[642,791,1999,2025]" firstAuthor="Heller" pageId="10" pageNumber="11" pagination="1 - 46" refId="ref5896" refString="Heller K. M. 1900. Neue Kafer von Celebes IV. Abhandlungen und Berichte des koniglichen zoologischen und anthropologisch-ethnographischen Museums zu Dresden 9: 1 - 46. Available from https: // biodiversitylibrary. org / page / 46189178 [accessed 10 Sep. 2019]." type="journal article" year="1900">Heller, 1900</bibRefCitation>
|
||
</taxonomicName>
|
||
. Scale bars = 1.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<keyStep id="30BA86AB400EFFF9FF43524EFAECFE8B" pageId="11" pageNumber="12">
|
||
<paragraph id="8BF19E0E400EFFF9FF43524EFAEDFEAC" blockId="11.[189,1399,267,679]" pageId="11" pageNumber="12">
|
||
<keyLead id="30BF3D3B400EFFF9FF43524EFAEDFEAC" pageId="11" pageNumber="12">
|
||
12. Antennal club with 5 lamellae, first two abbreviated (
|
||
<collectingCountry id="F359DE9E400EFFF9FCA5524EFC7FFEC0" box="[859,997,267,293]" name="Philippines" pageId="11" pageNumber="12">Philippines</collectingCountry>
|
||
:
|
||
<collectingRegion id="498A50EC400EFFF9FC0E5249FBACFEC3" box="[1008,1078,268,294]" country="Philippines" name="Southern Leyte" pageId="11" pageNumber="12">Leyte</collectingRegion>
|
||
,
|
||
<collectingRegion id="498A50EC400EFFF9FBBC524EFB3DFEC3" box="[1090,1191,267,294]" country="Philippines" name="Palawan" pageId="11" pageNumber="12">Palawan</collectingRegion>
|
||
) ......................... ..................................................................................................................................
|
||
<taxonomicName id="4C4EE58D400EFFF9FB545275FAEDFEAC" authority="Keith, 2006" authorityName="Keith" authorityYear="2006" box="[1194,1399,303,330]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="lii">
|
||
<emphasis id="B93A421C400EFFF9FB545275FB7EFEAF" bold="true" box="[1194,1252,303,330]" italics="true" pageId="11" pageNumber="12">E. lii</emphasis>
|
||
<bibRefCitation id="EFDFE3FF400EFFF9FB12526AFAEDFEAC" author="Keith" box="[1260,1399,303,329]" firstAuthor="Keith" pageId="11" pageNumber="12" pagination="90 - 94" refId="ref5946" refString="Keith D. 2006. Reflexions sur le genre Engertia et description d'une nouvelle espece des Philippines. Lambillionea 106: 90 - 94." type="journal article" year="2006">Keith, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400EFFF9FF3F5211FAECFE8B" blockId="11.[189,1399,267,679]" box="[193,1398,339,366]" pageId="11" pageNumber="12">
|
||
<keyLead id="30BF3D3B400EFFF9FF3F5211FAECFE8B" box="[193,1398,339,366]" pageId="11" pageNumber="12">– Antennal club with 4 lamellae, only first one abbreviated .............................................................. 13</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA86AB400EFFF9FF4352D3FAECFE1D" pageId="11" pageNumber="12">
|
||
<paragraph id="8BF19E0E400EFFF9FF4352D3FAECFE30" blockId="11.[189,1399,267,679]" pageId="11" pageNumber="12">
|
||
<keyLead id="30BF3D3B400EFFF9FF4352D3FAECFE30" pageId="11" pageNumber="12">
|
||
13. Scales on elytra aggregated into spots; clypeus semicircular (
|
||
<collectingCountry id="F359DE9E400EFFF9FC4852D3FBB6FE54" box="[950,1068,406,433]" name="Sri Lanka" pageId="11" pageNumber="12">Sri Lanka</collectingCountry>
|
||
)........................................... ..............................................................................................................
|
||
<taxonomicName id="4C4EE58D400EFFF9FBE252FFFAECFE30" authority="(Brenske, 1896)" authorityName="Brenske" authorityYear="1896" baseAuthorityName="Brenske" baseAuthorityYear="1896" box="[1052,1398,441,469]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="maculosa">
|
||
<emphasis id="B93A421C400EFFF9FBE252FFFB28FE31" bold="true" box="[1052,1202,441,468]" italics="true" pageId="11" pageNumber="12">E. maculosa</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF400EFFF9FB3C52FFFAF4FE30" author="Brenske" box="[1218,1390,442,469]" firstAuthor="Brenske" pageId="11" pageNumber="12" pagination="178 - 205" refId="ref5850" refString="Brenske E. 1896. Neue Melolonthiden aus Africa und Asien. Stettiner entomologische Zeitschrift 57: 178 - 205." type="journal article" year="1896">Brenske, 1896</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400EFFF9FF3F529BFAECFE1D" blockId="11.[189,1399,267,679]" box="[193,1398,478,504]" pageId="11" pageNumber="12">
|
||
<keyLead id="30BF3D3B400EFFF9FF3F529BFAECFE1D" box="[193,1398,478,504]" pageId="11" pageNumber="12">– Scales on elytra aggregated into longitudinal stripes; clypeus polygonal ....................................... 14</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30BA86AB400EFFF9FF435165FAECFD43" pageId="11" pageNumber="12">
|
||
<paragraph id="8BF19E0E400EFFF9FF435165FAECFDBB" blockId="11.[189,1399,267,679]" pageId="11" pageNumber="12">
|
||
<keyLead id="30BF3D3B400EFFF9FF435165FAECFDBB" pageId="11" pageNumber="12">
|
||
14. Scales on elytra ovoid (
|
||
<collectingCountry id="F359DE9E400EFFF9FE005165FDECFDDF" box="[510,630,544,570]" name="Indonesia" pageId="11" pageNumber="12">Indonesia</collectingCountry>
|
||
: islands of
|
||
<collectingCountry id="F359DE9E400EFFF9FD065165FCC9FDDF" box="[760,851,544,570]" name="Indonesia" pageId="11" pageNumber="12">Ambon</collectingCountry>
|
||
, Yapen and Miosidi) ....................................... ..............................................................................................................
|
||
<taxonomicName id="4C4EE58D400EFFF9FBE75100FAECFDBB" authority="(Brenske, 1897)" authorityName="Brenske" authorityYear="1897" baseAuthorityName="Brenske" baseAuthorityYear="1897" box="[1049,1398,580,607]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="amboinae">
|
||
<emphasis id="B93A421C400EFFF9FBE75100FB28FDBA" bold="true" box="[1049,1202,580,607]" italics="true" pageId="11" pageNumber="12">E. amboinae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF400EFFF9FB3C5101FAF4FDBB" author="Brenske" box="[1218,1390,580,606]" firstAuthor="Brenske" pageId="11" pageNumber="12" pagination="96 - 120" refId="ref5871" refString="Brenske E. 1897. Neue Gattungen und Arten der Melolonthiden aus Afrika und Asien. Stettiner entomologische Zeitung 58: 96 - 120." type="journal article" year="1897">Brenske, 1897</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BF19E0E400EFFF9FF3F512CFAECFD43" blockId="11.[189,1399,267,679]" pageId="11" pageNumber="12">
|
||
<keyLead id="30BF3D3B400EFFF9FF3F512CFAECFD43" pageId="11" pageNumber="12">
|
||
– Scales on elytra setiform (
|
||
<collectingCountry id="F359DE9E400EFFF9FDE3512DFD0FFD67" box="[541,661,616,642]" name="Indonesia" pageId="11" pageNumber="12">Indonesia</collectingCountry>
|
||
: islands of Seram,
|
||
<collectingCountry id="F359DE9E400EFFF9FC8E512DFC52FD67" box="[880,968,616,642]" name="Indonesia" pageId="11" pageNumber="12">Ambon</collectingCountry>
|
||
and Saparua) ................................... .....................................................................................................................
|
||
<taxonomicName id="4C4EE58D400EFFF9FBB351C8FAECFD43" authority="(Moser, 1913)" authorityName="Moser" authorityYear="1913" baseAuthorityName="Moser" baseAuthorityYear="1913" box="[1101,1398,652,679]" class="Insecta" family="Melolonthidae" genus="Engertia" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="setifera">
|
||
<emphasis id="B93A421C400EFFF9FBB351C8FB53FD43" bold="true" box="[1101,1225,652,679]" italics="true" pageId="11" pageNumber="12">E. setifera</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDFE3FF400EFFF9FB2651C9FAF4FD43" author="Moser" box="[1240,1390,652,678]" firstAuthor="Moser" pageId="11" pageNumber="12" pagination="271 - 297" refId="ref5971" refString="Moser J. 1913. Beitrag zur Kenntnis der Melolonthiden I. Deutsche entomologische Zeitschrift 3: 271 - 297. Available from https: // archive. org / details / biostor- 101429 [accessed 10 Sep. 2019]." type="journal article" year="1913">Moser, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |